Maculinea

Maculinea van Eecke, 1915

Maculinea van Eecke, 1915; Zool. Meded. 1 (3): 28; TS: Papilio alcon Denis & Schiffermüller
- = Phengaris; [FE]
Argus Boisduval, 1832; in d'Urville, Voy. Astrolabe (Faune ent. Pacif.) 1: 49; TS: Papilio alcon Denis & Schiffermüller

Subgenus Maculinea (Phengaris); Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 81
Argus; [NHM-BMW] ; [NHM-BMW]
Maculinea; [CoL] ; [NHM-BMW]
Maculinea (Section Glaucopsyche); Eliot, 1973, Bull. Br. Mus. nat. Hist. (Ent.) 28 (6): 449; Ugelvig, Vila, Pierce & Nash, 2011, Molec. Phyl. Evol. 61: 238
Maculinea (Lycaenidae); [BRU2], 154
Maculinea (Scolitantidina); Stradomsky, 2016, Caucasian Ent. Bull. 12 (1): 151

Maculinea arion (Linnaeus, 1758)

. See [maps] Schwarzflecken Blauling Svartfläckig blåvinge Large Blue Modraszek arion Fläckblåvinge Hormiguera de lunares Muurahaissinisiipi

Papilio arion Linnaeus, 1758; Syst. Nat. (Edn 10) 1:; TL: Germany, Nurnberg
Papilio telegone Bergsträsser, [1779]; Nomen. Ins. 3: 8, pl. 52, f. 5-6
Papilio varietas Bergsträsser, [1779]
Lycaena arthurus Melvill, 1873; Ent. mon. Mag. 9: 263; TL: Chamounix, Switzerland
Lycaena arion var. obscura Christ, 1880; Lep. Schweiz.: 22; TL: Zermatt
- = Phengaris arion; [FE]
Lycaena arion ab. supra-impunctata Oberthür, 1896; Étud. d'Ent. 20: 15, pl. 3, f. 19
Lycaena arion ab. subtus-impunctata Oberthür, 1896; Étud. d'Ent. 20: 15, pl. 3, f. 20
Lycaena arion ab. subtus-maclulis-extensis Oberthür, 1896; Étud. d'Ent. 20: pl. 3, f. 21
immaculata (Morsley, 1896); Nat. J. Suppl.: 10, pl. 4, f. 13
Lycaena arion f. tripuncta Courvoisier, 1903; Mitt. schweiz. ent. Ges. 11: 22
Lycaena arion f. impuncta Courvoisier, 1903; Mitt. schweiz. ent. Ges. 11: 24
Lycaena arion f. unipuncta Courvoisier, 1903; Mitt. schweiz. ent. Ges. 11: 24
Lycaena arion ab. coalescens Gillmer, 1904; Soc. ent. 18 (23): 180
Lycaena arion ab. fasciata Gillmer, 1904; Soc. ent. 18 (23): 180
Lycaena arion f. unipuncta Courvoisier, 1907; Zs. wiss. Insektenbiol. 3 (3): 74
Lycaena arion ab. parvimacula Aigner, 1907; Rovart. Lapok 14 (9-10): 214
Lycaena arion ab. ♂ pseudo-alcon le Chamberlain, 1908; Entomologist 41: 202; TL: Cotswold
Lycaena arion ab. ♀ imperialis le Chamberlain, 1908; Entomologist 41: 202; TL: Cotswold
Lycaena arion ab. multo-maculata le Chamberlain, 1908; Entomologist 41: 202; TL: Cotswold
Lycaena arion ab. marginata le Chamberlain, 1908; Entomologist 41: 202; TL: Cotswold
Lycaena arion ab. cotswoldensis le Chamberlain, 1908; Entomologist 41: 202; TL: Cotswold
Lycaena arion ab. pallida le Chamberlain, 1908; Entomologist 41: 202; TL: Cotswold
Lycaena arion ab. occidentalis le Chamberlain, 1908; Entomologist 41: 202; TL: Cotswold
Lycaena arion ab. oolitica le Chamberlain, 1908; Entomologist 41: 202; TL: Cotswold
Lycaena arion ab. punctifera Grund, 1908; Int. ent. Zs. 2 (14): 88
Lycaena arion var. (ab.) ofenia Tutt, 1909; Ent. Rec. J. Var. 21: 198
Lycaena arion ab. bipuncta Rebel, [1909]; in Berge, Schmetterlingsbuch (edn 9): 75
Lycaena arion ab. magnifica Heydemann, 1910; Ent. Zs. 23 (40): 177
Lycaena arion f. retro-juncta Courvoisier, 1911; Ent. Zs. 25 (13): 83
Lycaena arion ab. fasciata Skala, 1912; Ver. naturf. Ver. Brünn 50: 135
Lycaena arion ab. ♂ grisea Courvoisier, 1913; Ent. Mitt. 2 (10): 294
Lycaena arion ab. caeruleo-marginata Tutt, 1914; Nat. Hist. Br. Lepid. 4: 309, 314
Lycaena arion var. amurensis Tutt, 1914; Nat. Hist. Br. Lepid. 4: 312
Lycaena arion ab. obscura-major Tutt, 1914; Nat. Hist. Br. Lepid. 4: 317
Lycaena arion ab. punctatissima Tutt, 1914; Nat. Hist. Br. Lepid. 4: 319
Lycaena arion ab. basipuncta Tutt, 1914; Nat. Hist. Br. Lepid. 4: 322
Lycaena arion ab. conjuncta Tutt, 1914; Nat. Hist. Br. Lepid. 4: 322
Lycaena arion ab. postero-immaculata Tutt, 1914; Nat. Hist. Br. Lepid. 4: 324
Lycaena arion ab. addenda Tutt, 1914; Nat. Hist. Br. Lepid. 4: 325
Lycaena arion ab. striata Tutt, 1914; Nat. Hist. Br. Lepid. 4: 325
Lycaena arion ab. antico-obsoleta Tutt, 1914; Nat. Hist. Br. Lepid. 4: 326
obsoleta (Frohawk, 1914) (Nomiades); Brit. Butts.: pl. 54, f. 23
Lycaena arion aglaophon Fruhstorfer, 1915; Soc. ent. 30 (12): 68; TL: Pyrenees
?Lycaena arion tainaron Fruhstorfer, 1915; Soc. ent. 30 (12): 68; TL: Wallis, Val Entremonts
Lycaena arion sosinomus Fruhstorfer, 1915; Soc. ent. 30 (12): 68; TL: Tian-Shan
vernetensis (Oberthür, 1916) (Lycaena); Ètud. Lépid. Comp. 12 (1): 486
Lycaena arion australpina Verity, 1924; Ent. Rec. J. Var. 36: 109
sub-unicolor (Pionneau, 1928) (Lycaena); Echange 44: 4
schmidti (Kardakoff, 1928) (Lycaena); Ent. Mitt. 17 (4): 272
Lycaena arion sabinorum Dannehl, 1933; Ent. Zs. 46 (23): 246
Glaucopsyche arion ab. suprapuncta Obraztsov, 1936; Z. Öst. EntVer. 21: 48
semiobscura Frohawk, 1938; Var. Brit. Butterfl. : pl. 32, f. 3
naryoniensis (Beuret, 1938) (Lycaeides); Lambillionea 38: 50
Lycaena arion ab. argentoazurea Silbernagel, 1946; Acta Soc. ent. bohem. 37: 118, 119
microarion Beuret, 1964; Die Lycaeniden der Schweiz (2): 250
subtusbrunea Beuret, 1964; Die Lycaeniden der Schweiz (2): 257
tainaronobscura Beuret, 1964; Die Lycaeniden der Schweiz (2): 257

Glaucopsyche arion;
Lycaena arion obscura; Fruhstorfer, 1917, Dt. ent. Z. Iris 31 (1/2): 30
Lycaena arthurus; [CoL]
Maculinea aglaophon; [iNaturalist]
Maculinea amurensis; [CoL]
Maculinea arion aglaophon; [CoL]
Maculinea arion australpina; [CoL]
Maculinea arion lutshniki; [CoL]
Maculinea arion microarion; [CoL]
Maculinea arion naryoniensis; [CoL]
Maculinea arion obscura; [CoL]
Maculinea arion ofenia; [CoL]
Maculinea arion sabinorum; [CoL]
Maculinea arion sosinomos; [CoL]
Maculinea arion subtusbrunea; [CoL]
Maculinea arion tainaron; [CoL]
Maculinea arion vernetensis; [CoL]
Maculinea arion; [H&R]; [SPRK]; [BARU], #375; Munguira, García-Barros & Martín, 1997, Boletín de la Asociación española de Entomología, 21; [BOW]: pl. 10, f. 4; [MRS], 683; Sibatani, Saigusa & Hirowatari, 1994, Tyô to Ga 44 (4): 179; [BRU2]: 155, pl. 65, f. 28-36; [Otakar Kudrna]; [CoL] ; [Heiner Ziegler] ; [Mario Maier]
Phengaris (Maculinea) arion; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 82; [RFEL]
Phengaris arion; [FE]; [iNaturalist] ; [LEPIFORUM] ; [MBE] ; [SVF]

500x360 (~38Kb) underside Axat, Pyrenees Orientales, France 24/07/96, Photo © Simon Coombes

830x570 (~78Kb) underside Finland, Ruokolahti 680:57, 05.07.1985, Photo © Pertti Pakkanen

M. a. arion EU, W.Siberia, Altai, NW.Kazakhstan, Ukraine

alconides Aurivillius, 1888; Nordens Fjärilar. Handbok I: 15
Lycaena arion var. rühli [= ruhli] Krulikovsky, 1892; Soc. Ent. 7 (1): 1
Lycaena unicolor Hormuzaki, 1892; Ent. Nachr. 18 (1): 1
baschkiria Krulikovsky, 1897
Lycaena jasilkowskii Hormuzaki, 1897; Verh. zool.-bot. Ges. Wien. 47: 138
Lycaena uralensis Elwes, 1899; Trans. ent. Soc. Lond. 1899 (3): 329
alpina Favre, 1903
caucasica Jachontov, 1914; (nom.nud.)
Lycaena arion antesion Fruhstorfer, 1917; Dt. ent. Z. Iris 31 (1/2): 27; TL: Bosnia, Koricna and Maklenpass
lutschniki (Krulikovsky, 1926) (Lycaena); Trav. Mus. Zool. Kieff 1: 29
Glaucopsyche arion sergeji Obraztsov, 1936; Z. Öst. EntVer. 21: 47
animula Szabó, 1956; Folia ent. hung. (n.s.) 9: 266, 340
vagula Szabó, 1956; Folia ent. hung. (n.s.) 9: 266, 340

Lacaena arion arion; Fruhstorfer, 1917, Dt. ent. Z. Iris 31 (1/2): 29 (note)
Lycaena arion unicolor; Fruhstorfer, 1917, Dt. ent. Z. Iris 31 (1/2): 29 (note)
Maculinea arion animula; [CoL]
Maculinea arion antesion; [CoL]
Maculinea arion arion; [BRU2], 155
Maculinea arion jasilkowskii; [CoL]
Maculinea arion ruehli[sic, ruhli]; [Oleg Kosterin]
Maculinea arion ruhli; [CoL]
Maculinea arion sergeji; [CoL]
Maculinea arion unicolor; [CoL]
Maculinea arion uralensis; [CoL]
Maculinea arion vagula; [CoL]
Phengaris arion arion; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 82
Phengaris arion rueli[sic, ruhli]; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 82
antesion; Abadjiev, 2002, Neue Ent. Nachr. 53: 21

533x400 (~48Kb) ♂ ♀ underside A meadow in the Opalikha Rivulet valley just upstream of its fall into the Koen River, Iskitim District, Novosibirsk Province, West Siberia, Russia. 11th July 1992, Photo © Oleg Kosterin

M. a. ligurica Wagner, 1904

Maculinea arion var. ligurica Wagner, 1904; Soc. Ent. 19 (1): 1; TL: Riviera
- = Phengaris arion; [FE]

Lycaena arion ligurica; Fruhstorfer, 1915, Soc. ent. 30 (12): 68
Maculinea arion ligurica; [CoL]

800x595 (~87Kb) HUNGARY, Borsod-Abaúj-Zemplén county, Baskó, 8.8.1998, Photo © Tamás Baranyi

M. a. delphinatus (Fruhstorfer, 1910)

Lycaena arion delphinatus Fruhstorfer, 1910; Ent. Zs. 24 (26): 144; TL: "Dauphine"

Lycaena arion delphinatus; Fruhstorfer, 1915, Soc. ent. 30 (12): 68; Fruhstorfer, 1917, Dt. ent. Z. Iris 31 (1/2): 30
Maculinea arion delphinatus; [CoL]

M. a. laranda (Fruhstorfer, 1910) S.Tirol

Lycaena arion laranda Fruhstorfer, 1910; Int. ent. Zs. 4 (10): 55; TL: S.Tirol, nr. Klausen and Atzwang

Lycaena arion laranda; Fruhstorfer, 1915, Soc. ent. 30 (12): 68; Fruhstorfer, 1917, Dt. ent. Z. Iris 31 (1/2): 29
Maculinea arion laranda; [CoL]

M. a. arcina (Fruhstorfer, 1910)

Lycaena arion arcina Fruhstorfer, 1910; Int. ent. Zs. 4 (10): 55; TL: Arcina

Lycaena arion arcina; Fruhstorfer, 1915, Soc. ent. 30 (12): 67; Fruhstorfer, 1917, Dt. ent. Z. Iris 31 (1/2): 30
Maculinea arion arcina; [CoL]

M. a. naruena (Courvoisier, 1911)

Lycaena arion f. naruena Courvoisier, 1911; Ent. Zs. 24 (37): 202

Lycaena arion naruena; Fruhstorfer, 1915, Soc. ent. 30 (12): 68
Maculinea arion naruena; [CoL]

M. a. insubrica (Vorbrodt, 1912)

Lycaena arion insubrica Vorbrodt, 1912; Die Schmett. Schweiz 1: 156

Lycaena arion insubrica; Fruhstorfer, 1917, Dt. ent. Z. Iris 31 (1/2): 30
Maculinea arion insubrica; [CoL]

M. a. eutyphron (Fruhstorfer, 1915) England

Lycaena arion eutyphron Fruhstorfer, 1915; Soc. ent. 30 (12): 67; TL: England, Cornwall

Maculinea arion eutyphron; [CoL]

M. a. nepete (Fruhstorfer, 1915) Kansu

Lycaena arion nepete Fruhstorfer, 1915; Soc. ent. 30 (12): 68; TL: Kansu, Lanchowfu

Maculinea arion nepete; [CoL]

M. a. major (Oberthür, 1916)

Lycaena arion major Oberthür, 1916; Ètud. Lépid. Comp. 12 (1): 486

Maculinea arion major; [CoL]

M. a. vesubia (Fruhstorfer, 1917)

Lycaena arion vesubia Fruhstorfer, 1917; Dt. ent. Z. Iris 31 (1/2): 26; TL: St. Martin
Lycaena arion vesubia f. alpina Fruhstorfer, 1917; Dt. ent. Z. Iris 31 (1/2): 26; TL: St. Martin

Maculinea arion vesubia; [CoL]

M. a. ussuriensis (Sheljuzhko, 1928) Amur, Ussuri

Lycaena arion ussuriensis Sheljuzhko, 1928; Lep. Rdsch. 2 (6): 62; TL: S.Ussuri
Lycaena arion ussuriensis ab. hironobui Sugitani, 1934; Zephyrus 5: 168

Maculinea arion ussuriensis; [BARU](note) , ; Sibatani, Saigusa & Hirowatari, 1994, Tyô to Ga 44 (4): 179; [CoL]
Maculinea cyanecula ussuriensis; [BRU2], 156
Phengaris arion ussuriensis; [RFEL]
Phengaris cyanecula ussuriensis; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 82

M. a. zara Jachontov, 1935 Caucasus, Armenia

arion zara Jachontov, 1935; Our Butterflies: [?]
- = Phengaris arion obscura; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 82
?obscura (Christoph, 1878) (Lycaena)

Maculinea arion zara; [BRU2], 155
Phengaris arion obscura; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 82

M. a. buholzeri Rezbanyai, 1978 Switzerland

Maculinea arion buholzeri Rezbanyai, 1978; Mitt. schweiz. Ent. Ges. 51 (4): 399

Maculinea arion buholzeri; [CoL]

M. a. xiaheana (Murayama, 1991) China (Gansu)

Glaucopsyche xiaheana Murayama, 1991; Nature & Ins. 26 (3): 21, f. 5; TL: Gansu, Prov. Xiahe, 3200-3500m

Maculinea arion xiaheana; Sibatani, Saigusa & Hirowatari, 1994, Tyô to Ga 44 (4): 186

M. a. inferna Sibatani, Saigusa & Hirowatari, 1994 Sichuan

Maculinea arion inferna Sibatani, Saigusa & Hirowatari, 1994; Tyô to Ga 44 (4): 188 (repl. Lycaena arion tatsienluica Oberthür, 1910)
Lycaena arion tatsienluica Oberthür, 1912; Étud. Lépid. Comp. 6: 328, pl. 126, f. 1121-1122

Lycaena arion tatsienluica; Fruhstorfer, 1915, Soc. ent. 30 (12): 68
Maculinea arion inferna; Huang & Wu, 2003, Neue Ent. Nachr. 55: 128
Maculinea arion tatsienlouica; [CoL]

Maculinea cyanecula (Eversmann, 1848)

Mongolia. See [maps]

Lycaena cyanecula Eversmann, 1848; Bull. Soc. Imp. Nat. Moscou 21 (3): 207; TL: Kiachta im östlichen Sibirien [Kyakhta, Buryatia]

Lycaena arion cyanecula; Fruhstorfer, 1915, Soc. ent. 30 (12): 68
Maculinea arion cyanecula; [BARU](note) , ; [MRS], 683; [CoL]
Maculinea cyanecula; [BRU2]: 155, pl. 66, f. 1-15
Phengaris (Maculinea) cyanecula; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 82
Phengaris cyanecula; [iNaturalist]

900x838 (~140Kb) ♂ upperside China, Qinghai prov., Caka nuo vill (3500m), 20.7.2010, Photo © A. Timchenko leg.	1300x839 (~186Kb) ♂ underside China, Qinghai prov., Caka nuo vill (3500m), 20.7.2010, Photo © A. Timchenko leg.	900x1250 (~245Kb) ♂ upperside China, Qinghai prov., Caka nuo vill (3500m), 20.7.2010, Photo © A. Timchenko leg.
600x789 (~111Kb) ♂ underside Russia, West Siberia, NW Altai Mts., Ust'-Kan District, Bashchelakskii Range, meadow steppe 1 km SSW of Keley village. 3th July 2005 (on Onobrychis arenaria), Photo © Oleg Kosterin

M. c. cyanecula Transbaikalia, S.Siberia (Tuva, Khakassia)

Lycaena arion philidor Fruhstorfer, 1915; Soc. ent. 30 (12): 68; TL: Mongolia

Maculinea arion philidor; Sibatani, Saigusa & Hirowatari, 1994, Tyô to Ga 44 (4): 180
Maculinea cyanecula cyanecula; [BRU2], 156
Phengaris cyanecula cyanecula; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 82

?M. c. obscurior (Staudinger, 1901) Tian-Shan

cyanecula obscurior Staudinger, 1901; Cat. Lep. palaearct. Faunengeb. 1: 90 [ ?? lapsus from desc.?]

Maculinea cyanecula obscurior; [BRU2], 156

M. c. taras (Fruhstorfer, 1915) Alai

Lycaena arion taras Fruhstorfer, 1915; Soc. ent. 30 (12): 68; TL: Mt. Aurunei, Petralla Valley, 1200m

Lycaena arion taras; Fruhstorfer, 1917, Dt. ent. Z. Iris 31 (1/2): 30
Maculinea arion taras; [CoL]
Maculinea cyanecula taras; [BRU2], 156

M. c. sauron Korb, 2003 Kirghizia

cyanecula sauron Korb, 2003; Bull. Soc. Nat. Mulhouse 59 (3): 46

Phengaris cyanecula sauron; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 82

Maculinea arionides (Staudinger, 1887)

Amur, Ussur, Korea, Japa, NE.China. See [maps]

Lycaena arionides Staudinger, 1887; in Romanoff, Mém. Lépid. 3: 141, pl. 7, f. 1a-c; TL: Vladivostok

Lycaena arionides; [CoL]
Maculinea arionides; [BARU], #372; [BOW]: pl. 206, f. 14; [MRS], 684; Sibatani, Saigusa & Hirowatari, 1994, Tyô to Ga 44 (4): 192; [BRU2]: 156, pl. 65, f. 25-27
Phengaris (Maculinea) arionides; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 82
Phengaris (Phengaris) arionides; [RFEL]
Phengaris arionides; [iNaturalist]

M. a. arionides Amur, Ussuri

Lycaena selzeri Warnecke, 1924; Dt. ent. Z. Iris 38: 149
Lycaena arionides sugitanii Matsumura, 1927; Ins. Matsumurana 2 (2): 118, pl. 3, f. 10; TL: Corea
- = ; Sibatani, Saigusa & Hirowatari, 1994, Tyô to Ga 44 (4): 194
Lycaena arionides ab. arionidula Kardakov, 1928; Ent. Mitt. 17 (4): 272, pl. 7, f. 19

Lycaena arionides selzeri; [CoL]
Maculinea arionides arionides; Sibatani, Saigusa & Hirowatari, 1994, Tyô to Ga 44 (4): 192; [BRU2], 156

M. a. takamukui (Matsumura, 1919) Japan

Lycaena arionides var. takamukui Matsumura, 1919; Thous. Ins. Japan. Addit. 3: 653, pl. 49, f. 32
Lycaena arion shiranensis Scriba, 1921; Ent. Rundschau. 38 (6): 25; TL: Shirane
- = ; Sibatani, Saigusa & Hirowatari, 1994, Tyô to Ga 44 (4): 194

Lycaena arionides takamukui; [CoL]
Maculinea arion shiranensis; [CoL]
Maculinea arionides takamukui; Kudrna, 1974, Atalanta 5: 114; Sibatani, Saigusa & Hirowatari, 1994, Tyô to Ga 44 (4): 194

Maculinea alcon (Denis & Schiffermüller, 1775)

-. See [maps] Alcon Blue Alkonblåvinge Hormiguera Modraszek alkon

Papilio alcon Denis & Schiffermüller, 1775; Ank. syst. Schmett. Wienergegend: 182
Papilio mamers Bergsträsser, [1779]; Nomen. Ins. 3: pl. 59, f. 1-2
Lycaena alcon ab. ♀ nigra Wheeler, 1903; Butts. Switzerland: 21
Lycaena alcon ab. marginepunctata Gillmer, 1904; Soc. ent. 18 (23): 179
Lycaena alcon ab. pallidior Schultz, 1906; Ent. Zs. 19 (36): 215
spormanni (Pfau, 1928) (Lycaena); Int. ent. Z. 22: 193
Lycaena alcon senilis Dannehl, 1933; Ent. Zs. 46 (23): 246; TL: Beuerberg; Fletzen
Lycaena alcon ab. crassipuncta Lempke, 1936; Tijdschr. Ent. 79: 307
Lycaena alcon ab. disco-elongata Lempke, 1936; Tijdschr. Ent. 79: 307
lunulata (Warnecke, 1942) (Lycaena); Dt. ent. Z. Iris 56: 103
paucipuncta (Lempke, 1946) (Lycaena); Natuurh. Maandbl. 31: 107
caerulescens (Lempke, 1946) (Lycaena); Natuurh. Maandbl. 31: 107
Maculinea rebeli macroconia Beuret, 1949; Bull. Soc. ent Mulhouse 5 (1949): 34; TL: Appenzell, Narwie
?tenuiconia Beuret, 1949; Bull. Soc. ent Mulhouse 5 (1949): 34
Maculinea alcon pseudoroboris Beuret, 1949; Bull. Soc. ent Mulhouse 5 (1949): 41; TL: Tessin, Ascona
hospitali Vilarrubia, 1950; Trab. Mus. Cienc. nat. Barcelona (n.s.) 1 (2): 6
Maculinea alcon f. ♂ supralunulata Lempke, 1955; Tijdschr. Ent. 98: 293; TL: Hilversum
Maculinea alcon f. ♀ supracaeca Lempke, 1955; Tijdschr. Ent. 98: 293; TL: Hezingen
Maculinea alcon f. ♂ marginemaculata Lempke, 1955; Tijdschr. Ent. 98: 293; TL: Wassenaar
Maculinea alcon f. marginedistincta Lempke, 1955; Tijdschr. Ent. 98: 294; TL: Slangenburg

Maculinea alcon caerulescens; [CoL]
Maculinea alcon hospitali; [CoL]
Maculinea alcon lunulata; [CoL]
Maculinea alcon macroconia; [CoL]
Maculinea alcon marginedistincta; [CoL]
Maculinea alcon marginemaculata; [CoL]
Maculinea alcon marginepunctata; [CoL]
Maculinea alcon pseudoroboris; [CoL]
Maculinea alcon spormanni; [CoL]
Maculinea alcon supracaeca; [CoL]
Maculinea alcon supralunulata; [CoL]
Maculinea alcon tenuiconia; [CoL]
Maculinea alcon; [H&R]; [BARU], #373; Munguira, García-Barros & Martín, 1997, Boletín de la Asociación española de Entomología, 21; [EBW], (name); [BOW]: pl. 10, f. 4 (text only); [BRU2]: 154, pl. 66, f. 19-24, 34; Sibatani, Saigusa & Hirowatari, 1994, Tyô to Ga 44 (4): 173; [Otakar Kudrna]; [CoL] ; [Heiner Ziegler] ; [Mario Maier]
Maculinea macroconia; [iNaturalist]
Phengaris (Maculinea) alcon; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 81; [RFEL]
Phengaris alcon; [FE]; [iNaturalist] ; [LEPIFORUM] ; [MBE] ; [SVF]

620x444 (~39Kb) ♂ underside A bushy western (right) slope of the Anui River valley with limestone rocks at Denisova Cave stationary, Soloneshino District, Altaiskii Krai Province, West Siberia, Russia. 2nd July 2001, Photo © Oleg Kosterin

M. a. alcon C.Europe

Lycaena alcon alcon; Fruhstorfer, 1917, Dt. ent. Z. Iris 31 (1/2): 25
Maculinea alcon alcon; [BRU2], 154
Phengaris alcon alcon; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 81

M. a. haurii (Fruhstorfer, 1917) Switzerland

Lycaena alcon haurii Fruhstorfer, 1917; Soc. ent. 32 (6): 26; TL: "Graubünden, Filisur", 1000-1100m
Lycaena alcon haurii Fruhstorfer, 1917; Dt. ent. Z. Iris 31 (1/2): 24; TL: "Graubünden, Filisur", 1000-1100m

Maculinea alcon haurii; [CoL]

M. a. tolistus (Fruhstorfer, 1917) Bosnia

Lycaena alcon tolistus Fruhstorfer, 1917; Dt. ent. Z. Iris 31 (1/2): 24; TL: Bosnia, Koricna

Maculinea alcon tolistus; [CoL]

M. a. turatiana (Verity, 1923)

Lycaena alcon turatiana Verity, 1923; Ent. Rec. (suppl.) 35: 7
Lycaena alcon f. italica Verity, 1919; Atti Soc. ital. Sci. nat. 58: 168

Maculinea alcon turatiana; [CoL]

M. a. jenissejensis (Sheljuzhko, 1928) S.Siberia

Lycaena alcon jenissejensis Sheljuzhko, 1928; Lep. Rdsch. 2 (4): 46

Maculinea alcon jeniseiensis[sic, jenissejensis]; [BARU](note) ,
Maculinea alcon jenissejensis; [BRU2], 154; [SZMN]
Phengaris alcon jenissejensis; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 81

M. a. nestae (Higgins, 1930)

alcon nestae (Higgins, 1930) (Lycaena); Entomologist 63: 100; TL: Piedmont

Maculinea alcon nestae; [CoL]

M. a. sevastos (Rebel & Zerny, 1931) Carpathians

Lycaena alcon sevastos Rebel & Zerny, 1931; Denkschr. Akad. Wiss. Wien 103: 82; TL: Macedonia
Maculinea alcon limitanea Bálint, 1985; Galathea 1: 66
- = Glaucopsyche alcon sevastos; Bálint, 1990, Janus Pannonius Múz. 34: 51

Maculinea alcon limitanea; [CoL]
Maculinea alcon sevastos; [BRU2], 154; [CoL]
Phengaris alcon sevastos; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 81

M. a. xerophila Berger, 1946 S.Europe

alcon xerophila Berger, 1946; Lambillionea 46: 106
arcasoides Berger, 1946; Lambillionea 46: 106
paucipuncta Berger, 1946; Lambillionea 46: 106
curiosa Szabó, 1956; Folia ent. hung. (n.s.) 9: 264, 339

Maculinea alcon arcasoides; [CoL]
Maculinea alcon curiosa; [CoL]
Maculinea alcon xerophila; [BRU2], 154; [CoL]
?Phengaris alcon xerophyla[sic, xerophila]; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 81

M. a. arirang Sibatani, Saigusa & Hirowatari, 1994 Korea

Maculinea alcon arirang Sibatani, Saigusa & Hirowatari, 1994; Tyô to Ga 44 (4): 175; TL: Ryanggang Do

M. a. aulendil Korb, 2009 Buryatia

alcon aulendil Korb, 2009; Alexanor 23 (3): 133; TL: Buryatia, Khamar-Daban, 1500m

Phengaris alcon aulendil; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 81

M. a. shaposhnikovi Korb, 2010 Kirgistan

Maculinea alcon shaposhnikovi Korb, 2010; Eversmannia (23-24): 15; TL: Kyrghyzstan, Sary-Chelek Nature Reserve, Sary-Chelek lake, 2000m

Phengaris alcon shaposhnikovi; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 81

Maculinea rebeli (Hirschke, 1905)

-. See [maps]

Lycaena alcon var. rebeli Hirschke, 1905; Jber. Wien. ent. Ver. 15: 109, pl. 2, f. 1-2; TL: Hochschwab [Austria]
- = Phengaris alcon; [FE]
Lycaena alcon ab. ♂ latimargo Courvoisier, 1913; Ent. Mitt. 2 (10): 294
caerulea Vorbrodt, 1912
Maculinea rebeli alpicola Beuret, 1949; Bull. Soc. ent Mulhouse 5 (1949): 41; TL: Wallis, Zermatt, 200m
Maculinea rebeli magnalpicola Beuret, 1949; Bull. Soc. ent Mulhouse 5 (1949): 42; TL: Tessin, Altanca, Fussio
Maculinea rebeli gadmensis Beuret, 1949; Bull. Soc. ent Mulhouse 5 (1949): 43; TL: Bern, Gadmen
Maculinea rebeli cruciata Beuret, 1949; Bull. Soc. ent Mulhouse 5 (1949): 50; TL: Jura

Maculinea alcon alpicola; [CoL]
Maculinea alcon cruciata; [CoL]
Maculinea alcon gadmensis; [CoL]
Maculinea alcon magnalpicola; [CoL]
Maculinea alcon rebeli; [CoL]
Maculinea rebeli; Munguira, García-Barros & Martín, 1997, Boletín de la Asociación española de Entomología, 21; [BRU2]: 154, pl. 66, f. 16-18, 25-33, 35-36; [Heiner Ziegler]

M. r. rebeli CEU, SEU

Lycaena alcon rebeli; Fruhstorfer, 1917, Dt. ent. Z. Iris 31 (1/2): 25
Maculinea rebeli rebeli; [BRU2], 154

M. r. monticola (Staudinger, 1901) Caucasus Minor, Armenia

Lycaena alcon v. monticola Staudinger, 1901; Cat. Lep. palaearct. Faunengeb. 1: 90

Lycaena alcon monticola; Fruhstorfer, 1917, Dt. ent. Z. Iris 31 (1/2): 25
Maculinea alcon monticola; [CoL]
Maculinea rebeli monticola; [BRU2], 154
Phengaris alcon monticola; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 81

M. r. sordidula (Jachontov, [1909]) Caucasus Major

Lycaena alcon var. sordidula Jachontov, [1909]; Revue Russe Ent. 8: 292

Maculinea alcon sordidula; [CoL]
Maculinea rebeli cordidula[sic, sordidula]; [BRU2], 154
Phengaris alcon sordidula; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 81

M. r. kondakovi Kurentzov, 1970 Transbaikalia, Amur, Ussuri, NE.China

Lycaena kondakovi Kurentzov, 1970; 142 (part)

Maculinea alcon kondakovi; Sibatani, Saigusa & Hirowatari, 1994, Tyô to Ga 44 (4): 173; [BARU](note) ,
Maculinea rebeli kondakovi; [BRU2], 154
Phengaris alcon kondakovi; [RFEL]
?Phengaris rebeli kondakovi; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 81

M. r. imitator Tuzov, 2000 N.Tian-Shan, Dzhungarsky Alatau

Maculinea rebeli imitator Tuzov, 2000; [BRU2]: 155 (repl. Maculinea alcon alconides Korb, 1997)
Maculinea alcon alconides Korb, 1997; Alexanor 20 (1): (61-64) (preocc. Lycaena arion alconides Aurivillius, 1888, Lycaena alcon alconides Dahlstrom, 1899)

Phengaris alcon imitator; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 81; [iNaturalist]

ssp. Altai, Sayan

Maculinea rebeli ssp.; [BRU2], 154

Maculinea teleius (Bergsträsser, [1779])

-. See [maps] Limbada Scarce Large Blue Modraszek telejus

Papilio teleius Bergsträsser, [1779]; Nomen. Ins. 2: 71, pl. 43, f. 4; TL: Germany, Hanau-Munzberg
Papilio diomedes Rottemburg, 1775; Der Naturforscher 6: 26, n. 14 (preocc. Linnaeus)
Papilio telegonus Bergsträsser, [1779]; Nomen. Ins. 2: 72, pl. 4, f. 1-2
Papilio arctophylas Bergsträsser, [1779]; Nomen. Ins. 3: pl. 51, f. 1-2 (arctophylax?)
Papilio arctophonus Bergsträsser, [1779]; Nomen. Ins. 3: pl. 53, f. 7-8
Papilio argiades Fabricius, 1787; Mantissa Insectorum 2: 76
Papilio euphemus Hübner, [1799-1800]; Samml. eur. Schmett. [1]: pl. 54, f. 257-259
- = Phengaris teleius; [FE]
Lycaena euphemus ab. striata Gillmer, 1904; Soc. ent. 18 (23): 179
Lycaena euphemus ab. obsoleta Gillmer, 1904; Soc. ent. 18 (23): 179
Lycaena euphemus ab. paula Schultz, 1906; Ent. Zs. 19 (36): 215
Lycaena euphemus ab. sanguisorbae Oberthür, 1910; Ètud. Lépid. Comp. 4: 333, pl. 55, f. 472
Lycaena euphemus ab. albocuneata Spuler, 1908; Schmett. Eur. 1: 68
Lycaena euphemus unicolor Vorbrodt & Müller-Rutz, 1911; Die Schmett. Schweiz 1: 155
Lycaena euphemus f. caeca Courvoisier, 1914; Dt. ent. Z. Iris 28 (2) (3): 189
subunicolor (Pionneau, 1928) (Lycaena); Echange 44: 51
Lycaena euphemus ab. grisescens Lempke, 1936; Tijdschr. Ent. 79: 308
Lycaena euphemus ab. centro-elongata Lempke, 1936; Tijdschr. Ent. 79: 308
Maculinea teleius f. infracrassipuncta Lempke, 1955; Tijdschr. Ent. 98: 289; TL: Roermond
Maculinea teleius f. infrapaucipuncta Lempke, 1955; Tijdschr. Ent. 98: 290; TL: HerkenBosch
Maculinea teleius f. marginereducta Lempke, 1955; Tijdschr. Ent. 98: 290; TL: Swalmen

Cupido diomedes; [CoL]
Maculinea teleia; [MRS], 684
Maculinea teleius infracrassipuncta; [CoL]
Maculinea teleius infrapaucipuncta; [CoL]
Maculinea teleius marginereducta; [CoL]
Maculinea teleius; [H&R]; [BARU], #374; Sibatani, Saigusa & Hirowatari, 1994, Tyô to Ga 44 (4): 202; [EBW]; [BOW]: pl. 10, f. 5; [BRU2]: 156, pl. 67, f. 1-15; [Otakar Kudrna]; [CoL] ; [Heiner Ziegler] ; [Mario Maier]
Papilio telegonus; [iNaturalist]
Phengaris (Maculinea) teleius; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 81; [RFEL]
Phengaris teleius; [FE]; [iNaturalist] ; [LEPIFORUM] ; [MBE]

900x338 (~75Kb) HUNGARY, Borsod-Abaúj-Zemplén county, Baskó, 4.7.1999 ♂, 8.8.1998 ♀, Photo © Tamás Baranyi

M. t. teleius CEU, W.Siberia, Caucasus

Lycaena euphemus bajuvaricus Fruhstorfer, 1917; Dt. ent. Z. Iris 31 (1/2): 25; TL: "Oberbayern, Kochelsee"
Lycaena euphemus thersandrus Fruhstorfer, 1917; Dt. ent. Z. Iris 31 (1/2): 26; TL: "Canton Vaud, Eclepens"
Lycaena euphemus peninsulae Verity & Querci, 1923; Ent. Rec. (suppl.) 35: 7; TL: Sibillini Mts

Lycaena euphemia bajuvaricus; [CoL]
Lycaena euphemia peninsulae; [CoL]
Lycaena euphemia thersandrus; [CoL]
Maculinea teleius teleius; [BRU2], 156
Phengaris teleius teleius; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 81

M. t. kazamoto (Druce, 1875) Japan (Honshu)

Lycaena kazamoto Druce, 1875; 361; TL: Yokohama
Lycaena euphemus kazamoto ab. oiwakeana Matsumura, 1926; Ins. Matsumurana 1 (1): 27, pl. 2, f. 16 ♂; TL: Honshu, Oiwake

Lycaena kazamoto; [CoL]
Maculinea teleius kazamoto; Kudrna, 1974, Atalanta 5: 114; Sibatani, Saigusa & Hirowatari, 1994, Tyô to Ga 44 (4): 211

M. t. euphemia (Staudinger, 1887) Amur, Ussuri

Lycaena euphemia Staudinger, 1887; in Romanoff, Mém. Lépid. 3: 142; TL: Amurland
Lycaena euphemus insignis Sheljuzhko, 1928; Lep. Rdsch. 2 (5): 51; TL: Ussuri
- = ; Sibatani, Saigusa & Hirowatari, 1994, Tyô to Ga 44 (4): 204

Lycaena euphemia; [CoL]
Maculinea teleia euphemia; [MRS], 684
Maculinea teleius euphemia; Sibatani, Saigusa & Hirowatari, 1994, Tyô to Ga 44 (4): 202
Maculinea telejus euphemia; [BARU](note) ,
Phengaris teleius euphemia; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 82 (?); [RFEL]

M. t. obscurata (Staudinger, 1892) Transbaikalia

Lycaena obscurata Staudinger, 1892; Dt. Ent. Z. Iris 5 (2): 320; TL: Kentei

Maculinea teleius obscurata; [BRU2], 156; [CoL] ; [SZMN]
Maculinea telejus obscurata; [BARU](note) ,
Phengaris teleius obscurata; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 81

M. t. ogumae (Matsumura, 1910) Sakhalin, Kuriles, Japan (Hokkaido, Honshu)

Licaena[sic] euphemus var. ogumae Matsumura, 1910; Ent. Zs. 23 (50): 221; TL: Sakhalin
Lycaena euphemus doii Matsumura, 1928; Insecta Matsumurana 2 (4): 199; TL: Kurile Is.
- = Phengaris teleius ogumae; [RFEL]
Lycaena euphemus naganumai Kanda, 1934; Ageha 3: 8

Lycaena euphemia doii; [CoL]
Lycaena euphemia ogumae; [CoL]
Maculinea ogumae doii; [BRU2], 157
Maculinea ogumae ogumae; [BRU2], 157
Maculinea ogumae; [BRU2]: 157, pl. 67, f. 16-20
Maculinea teleius ogumae; Sibatani, Saigusa & Hirowatari, 1994, Tyô to Ga 44 (4): 211
Maculinea telejus doii; [BARU](note) ,
Maculinea telejus ogumae; [BARU](note) ,
Phengaris teleius doii; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 82 (?)
Phengaris teleius ogumae; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 82 (?); [RFEL]

M. t. splendens (Kozhantshikov, 1924) Altais, Sayan

Lycaena splendens Kozhantshikov, 1924; Jb. Martjanov Staatmus. Minussinsk 2 (1): 67
pseudobajuvaricus (Kozhanchikov, 1936) (Lycaena); Folia zool. hydrobiol. 9 (1): 25

Lycaena euphemia splendens; [CoL]
Maculinea teleius splendens; [BRU2], 156
Maculinea telejus ab. splendens; [BARU](note) ,

M. t. daisensis (Matsumura, 1926) Japan (Honshu)

Lycaena euphemus daisensis Matsumura, 1926; Ins. Matsumurana 1 (1): 27, pl. 2, f. 8 ♀; TL: Tottori-ken, Mt. Daisen
Glaucopsyche teleius daisensis ab. yoshisakai Murayama, 1963; Tyô to Ga 14 (2): 48, f. 39-40, 42-43

Lycaena euphemia daisensis; [CoL]
Maculinea teleius daisensis; Sibatani, Saigusa & Hirowatari, 1994, Tyô to Ga 44 (4): 209

M. t. shiriyensis (Matsumura, 1927)

Lycaena euphemus shiriyensis Matsumura, 1927; Ins. Matsumurana 2 (2): 118, pl. 3, f. 9; TL: Honshu (Aomori)

Lycaena euphemia shiriyensis; [CoL]

M. t. chosensis (Matsumura, 1927) S.Ussuri

Lycaena euphemus coreana Matsumura, 1926; Ins. Matsumurana 1 (1): 28, pl. 2, f. 5 ♂ (preocc.); TL: Korea
- = Maculinea teleius euphemia; Sibatani, Saigusa & Hirowatari, 1994, Tyô to Ga 44 (4): 204
Lycaena euphemia chosensis Matsumura, 1927; Insecta Matsumurana 1 (4): 161 (repl. Lycaena euphemus coreana Matsumura, 1926); TL: Korea
- = Maculinea teleius euphemia; Sibatani, Saigusa & Hirowatari, 1994, Tyô to Ga 44 (4): 204
Lycaena hozanensis Matsumura, 1927; Insecta Matsumurana 1 (4): 168, pl. 5, f. 10; TL: Korea
- = Maculinea teleius euphemia; Sibatani, Saigusa & Hirowatari, 1994, Tyô to Ga 44 (4): 204
Maculinea teleius melancholica Bryk, 1946; Ark. Zool. 38 A (3): 58; TL: Korea
- = Maculinea teleius euphemia; Sibatani, Saigusa & Hirowatari, 1994, Tyô to Ga 44 (4): 204

Lycaena euphemia coreana; [CoL]
Lycaena hozanensis; [CoL]
Maculinea teleius choesenses[sic, chosensis]; [BRU2], 156 (missp.)
Maculinea teleius chosensis; [CoL]
Maculinea teleius melancholica; [CoL]

M. t. macromelanica Verity, 1946 Italy

teleius macromelanica Verity, 1946; Le Farfalle diurn. d'Italia 2: 147

Maculinea teleius macromelanica; [CoL]

M. t. matsumurai Shirôzu, 1953 Japan

teleius matsumurai Shirôzu, 1953; Sieboldia 1 (1952): 155 (repl. jezoensis Matsumura, 1936)
Lycaena euphemus jezoensis Matsumura, 1926; Ins. Matsumurana 1 (1): 28, pl. 2, f. 14 ♂ (preocc.)
Lycaena euphemus jezoensis ab. teshionis Matsumura, 1926; Ins. Matsumurana 1 (1): 28, pl. 2, f. 18 ♀; TL: Hokkaido, Teshio and Hikada
Lycaena euphemus jezoensis ab. muratae Matsumura, 1926; Ins. Matsumurana 1 (1): 28, pl. 2, f. 11 ♀; TL: Hokkaido

Maculinea teleius matsumurai; [CoL]

M. t. discoobsoleta Beuret, 1958 Germany

Maculinea teleius discoobsoleta Beuret, 1958; Mitt. ent. Ges. Basel (n.f.) 8 (1): 4

Maculinea teleius discoobsoleta; [CoL]

M. t. hosonoi Takahashi, 1973 Japan (Honshu)

Maculinea teleius hosonoi Takahashi, 1973; Tyô to Ga 23 (3-4): 80; TL: Nagano-ken, Mt Karamatu, Niigata-ken
Maculinea teleis hakusanensis Fujioka, 1975; 143, pl. 73, f. 53-63
- = ; Sibatani, Saigusa & Hirowatari, 1994, Tyô to Ga 44 (4): 211

Maculinea teleius hosonoi; Sibatani, Saigusa & Hirowatari, 1994, Tyô to Ga 44 (4): 211; [CoL]

M. t. sinalcon Murayama, 1992 China?

Maculinea sinalcon Murayama, 1992; Nature & Ins. 27 (5): 37

Maculinea teleia sinalcon; Chou, 1994, Mon. Rhop. Sinesium: 684
Maculinea teleius sinalcon; Sibatani, Saigusa & Hirowatari, 1994, Tyô to Ga 44 (4): 206

Maculinea kurentzovi Sibatani, Saigusa & Hirowatari, 1994

N.China, N.Korea. See [maps]

Maculinea kurentzovi Sibatani, Saigusa & Hirowatari, 1994; Tyô to Ga 44 (4): 196, f. 9-10, 18-24, 40; TL: N. Korea, Yangdo, Mandaeri

Maculinea kurentzovi; [BARU], -; [BRU2]: 157, pl. 67, f. 22-24; [SZMN] ; [SZMN]
Phengaris (Maculinea) kurentzovi; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 82; [RFEL]
Phengaris kurentzovi; [iNaturalist]

M. k. kurentzovi Amur, Ussuri

Maculinea kurentzovi kurentzovi; [BRU2], 157; [SZMN]
Phengaris kurentzovi kurentzovi; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 82

M. k. daurica Dubatolov, 1999 Transbaikalia

Maculinea kurentzovi daurica Dubatolov, 1999; Proc. Dahur. State Bios. Nat. Res. 2: 180; TL: Mongolia, East Aimak, 25-28km S of station Erentsav

Maculinea kurentzovi daurica; [BRU2], 157
Phengaris kurentzovi daurica; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 82

Maculinea nausithous (Bergsträsser, [1779])

SEU, CEU, S.Urals - SW.Siberia, Caucasus, Turkey. See [maps] Dusky Large Blue Modraszek nausitous

Papilio nausithous Bergsträsser, [1779]; Nomen. Ins. 2: 70, pl. 43, f. 1; TL: Germnay, Hanau-Munzberg
Papilio arcas Rottemburg, 1775; Der Naturforscher 6: 25 (preocc. Papilio arcas Drury, [1773])
- = Phengaris nausithous; [FE]
Papilio nausithoe Bergsträsser, [1779]
Papilio erebus Knoch, 1782; Beitr. Insektengesch. 2: 93, pl. 6, f. 6-7
Lycaena arcas ab. minor Frey, 1882; Mitt. schweiz. ent. Ges. 6 (7): 352
Lycaena arcas ab. inocellata Sohn, 1893; Soc. Ent. 8 (10): 77
Lycaena arcas ab. ab. impunctata Hauder, 1901; Jahresb. ver. naturk. Linz 30: 15
Lycaena arcas ab. ♂ lucida Geest, 1903; Allg. Zs. Ent. 8: 311
Lycaena arcas ♂ ab. lycaonius Schultz, 1904; Ent. Zs. 18 (24): 94
Lycaena arcas ab. ♂ emutata Marschner, 1909; Zs. wiss. InsektBiol. 5: 166
paucipuncta (Courvoisier, 1912) (Lycaena); Dt. ent. Z. Iris 26 (1): 63
Lycaena arcas kijevensis Sheljuzhko, 1928; Lep. Rdsch. 2 (6): 64; TL: Kijev, Ukraine
supraddenda Verity, 1943; Le Farfalle diurn. d'Italia 2: 152
agenjoi Vilarrubia, 1950; Trab. Mus. Cienc. nat. Barcelona (n.s.) 1 (2): 8
Maculinea nausithous f. ♂ clara Lempke, 1955; Tijdschr. Ent. 98: 287; TL: Holland
Maculinea nausithous f. supraelongata Lempke, 1955; Tijdschr. Ent. 98: 287; TL: Holland
maculata Beuret, 1964; Die Lycaeniden der Schweiz (2): 237
argoviensis Beuret, 1964; Die Lycaeniden der Schweiz (2): 238

Lycaena arcus; [CoL]
Maculinea arion supraddenda; [CoL]
Maculinea nausithous agenjoi; [CoL]
Maculinea nausithous argoviensis; [CoL]
Maculinea nausithous clara; [CoL]
Maculinea nausithous kijevensis; [CoL]
Maculinea nausithous supraelongata; [CoL]
Maculinea nausithous; [H&R]; [BARU], #371; Munguira, García-Barros & Martín, 1997, Boletín de la Asociación española de Entomología, 21; [EBW]; [BOW]: pl. 10, f. 5 (text only); Sibatani, Saigusa & Hirowatari, 1994, Tyô to Ga 44 (4): 213; [BRU2]: 157, pl. 65, f. 37-39; [Otakar Kudrna]; [CoL] ; [Heiner Ziegler] ; [Mario Maier]
Phengaris (Maculinea) nausithous; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 81
Phengaris nausithous; [FE]; [iNaturalist] ; [LEPIFORUM] ; [MBE]

Misidentified/misapplied:

Papilio arcas; Esper, 1777, Die Schmett. Th. I, Bd. 1 (9): 338, (6): pl. 34, f. 4-5

533x400 (~38Kb) ♀ underside A forest-steppe landscape in the vicinity of Akademgorodok [Academy Town], Novosibirsk, West Siberia, Russia. 7th August 1988 (on the larval foodplant Sanguisorba officinalis L.), Photo © Oleg Kosterin

nigricans (Kitt.) (Lycaena)

Lycaena arion nigricans; Fruhstorfer, 1917, Dt. ent. Z. Iris 31 (1/2): 28-29

taimaron (Fruhstorfer) (Lycaena)

Lycaena arion taimaron; Fruhstorfer, 1917, Dt. ent. Z. Iris 31 (1/2): 29

Polyommatus alcon Stephens, 1828; Ill. Br. Ent. (Haustellata) 1 (2): 88 (Fabricius); TL: Anglia

Maculinea arion alcon; [CoL]

obscura (Kaucki, 1924) (Lycaena); Polskie Pismo 3: 91

aihona (Matsumura, 1939) (Lycaena); Bull. biogeogr. Soc. Japan 9 (20): 357

Lycaena euphemia aihona; [CoL]

burdigalensis (Stempffer, 1932) (Lycaena); Amat. Pap. 6: 108

Lycaena euphemia burdigalensis; [CoL]

Lycaena euphemus tsugaruensis Watari, 1937; Zephyrus 7: 26

Lycaena euphemia tsugaruensis; [CoL]

Maculinea anresion; [CoL]

References:

[A. Timchenko]
Andrew Timchenko; Beijing, China (ex-Moscow, Russia)
[CEB]
Caucasian Entomological Bulletin; issues (PDF)
[CiNii]
http://ci.nii.ac.jp/
[CoL] Catalogue of Life
Bánki, O., Roskov, Y., Döring, M., Ower, G., Hernández Robles, D. R., Plata Corredor, C. A., Stjernegaard Jeppesen, T., Örn, A., Vandepitte, L., Hobern, D., Schalk, P., DeWalt, R. E., Ma, K., Miller, J., Orrell, T., Aalbu, R., Abbott, J., Adlard, R., Aedo, C., et al. (2024). Catalogue of Life Checklist (Version 2024-03-26); https://doi.org/10.48580/dfz8d
[ESP] :ESP#c Heppner, 1981
[EVERSMANNIA]
Eversmannia contents
[FE] Fauna Europaea
Fauna Europaea
[HBN] Francis Hemming, 1937
HÜBNER, A bibliographical and systematic account of the entomological works of Jacob Hübner and of the supplements thereto by Carl Geyer, Gottfried Franz von Frölich and Gottlieb August Wilhelm Herrich-Schäffer, Vol 1 & 2
[Heiner Ziegler]
Switzerland / Schweiz; www.pieris.ch
[LAMAS-2005] Gerardo Lamas
Entomofauna 26 (6): 57-100; A Bibliography of the Zoological Publications of Hans Fruhstorfer (1886* - 1922+)
[LAMAS-NEO] Gerardo Lamas
Bibliography of butterflies; An Annocated Bibliography of the Neotropical Butterflies and Skipper (Lepididoptera: Papilionoidea and Hesperioidea); Revised Electronic Edition (2007)
[LEPIFORUM]
Lepiforum; Bestimmung von Schmetterlingen (Lepidoptera) und ihren Präimaginalstadien
[MBE] Moths and Butteflies of Europe
Paolo Mazzei, Diego Reggianti and Ilaria Pimpinelli; http://www.leps.it/
[Mario Maier]
Europäische Schmetterlinge; e-mail: Mario Maier
[NBC] Netherlands Centre for Biodiversity Naturalis
NBC naturalis
[NHM-BMW] Natural History Museum
Butterflies and Moths of the World; Generic Names and their Type-species
[Oleg Kosterin] Institue of Cytology & Genetics
Siberian Division of Russian Academy of Sciences; Novosibirsk; e-mail: Oleg Kosterin; http://pisum.bionet.nsc.ru/kosterin/
[Pertti Pakkanen]
e-mail: Pertti Pakkanen; Butterfly and moth observations from Finland
[SEG]
Schweizerische Entomologische Gesellschaft; Mitteilungen der Schweizerischen Entomologischen Gesellschaft; = Bulletin de la Société Entomologique Suisse = Journal of the Swiss Entomological Society
[SVF]
Svenska fjärilar; Naturhistoriska riksmuseet; (Sweden)
[SZMN] Lepidoptera
Siberian Zoological Museum
[Simon Coombes]
1 Park Street, Plymouth, PL3 4BL, UK; 01752 607854; e-mail: Simon Coombes; http://www.butterfly-guide.co.uk
[Tamás Baranyi]
Tamás Baranyi
[ZOBODAT] Oberösterreichische Landesmuseum
OÖ. Landesmuseum; ZOBODAT - Literatur
[iNaturalist]
iNaturalist
[maps]
Warning! The maps are automatically generated from the textual information, and the process does not always produce acceptable result; See about maps for more info.

Some related literature:

Abadjiev, 2002
Types of Balkan butterflies in the collection of The Natural History Museum, London Neue Ent. Nachr. 53 : 3-53
Aigner, 1907
A magyar lepke-fauna gyarapodása 1906-ban Rovart. Lapok 14 (9-10) : 212-215
Aurivillius, 1888
Sveriges, Norges, Danmarks och Finlands. Macrolepidoptera Nordens Fjärilar. Handbok I: 277pp, Lii, 49pls (1888-1891) : i-lii, 1-277, pl. 1-50
Bálint, 1985
Maculinea alcon limitanea nov. ssp. (Lepidoptera: Lycaenidae) from Transsylvania, Rumania Galathea 1 : 62-74
Bálint, 1990
A Kárpát-medence boglárkalepkéinek reviziója (Lepidoptera, Lycaenida) Janus Pannonius Múz. 34 : 47-62
Bergsträsser, [1779]
Nomenclatur und Beschreibung der Insecten in der Graftschaft Hanau-Münzenburg Nomen. Ins. 2: 79pp, pl. 15-48
Bergsträsser, [1779]
Nomenclatur und Beschreibung der Insecten in der Graftschaft Hanau-Münzenburg Nomen. Ins. 3 : 1-48, pl. 49-72
Beuret, 1958
Maculinea teleius Bergsträsser discoobsoleta m. ssp. nova (Lycaenidae, Lep.) Mitt. ent. Ges. Basel (n.f.) 8 (1) : 4-5
Boisduval, 1832
Voyage de découvertes de l'Astrolabe exécuté par ordre du Roi, pendant les années 1826-1827-1828-1829, sous le commandément de M. J. Dumont d'Urville. Faune entomologique de l'Océan Pacifique, avec l'illustration des insectes nouveaux recueillis pendant le voyage. Lépidoptères in d'Urville, Voy. Astrolabe (Faune ent. Pacif.) 1 : 1-267, pl. 1-5
Bryk, 1946
Zur Kenntnis der Grossschmetterlinge von Korea. I Ark. Zool. 38 A (3) : 1-74, pl. 1-5
le Chamberlain, 1908
Lycaena arion in the Cotswolds Entomologist 41 : 201-202
[MRS]; Chou, 1994
Monografia Rhopalocerorum Sinensium 1-2 Mon. Rhop. Sinesium : 1-854
Courvoisier, 1903
Ueber Aberrationen der Lycaeniden Mitt. schweiz. ent. Ges. 11 : 18-25, pl. 2
Courvoisier, 1907
Ueber Zeichnungs-Aberrationen bei Lycaeniden Zs. wiss. Insektenbiol. 3 (1): 8-11, (2): 33-39, (3): 73-79
Courvoisier, 1911
Entdeckungsreisen und kritische Spaziergänge ins Gebiet der Lycaeniden Ent. Zs. 24 (12): 59-60, (13): 65-67, (14): 70-71, (15): 77-78, (16): 81-82, (17): 88-89, (18): 92-94, (19): 99-101, (20): 106-108, (21): 112, (23): 125-127, (24): 131-132, (26): 141-142, (27): 147-149, (28): 156, (31): 167-170, (32): 175-177, (33): 181-182, (34): 185-188, (35): 190-192, (36): 196-200, (37): 201-204, (38): 205-209, (39): 210-214, (40): 227-228, (41): 230-231, (42): 233-236, (43): 237-238, (44): 241-242, (45): 245-247, (46): 249-250, (47): 253-255, (48): 257-259, (49): 261-263, (50): 265-266, (51/52): 269-270
Courvoisier, 1911
Entdeckungsreisen und kritische Spaziergänge ins Gebiet der Lycaeniden Ent. Zs. 25 (1): 2-7, (2): 9-11, (3): 13-17, (4): 22-24, (5): 25-27, (6): 33-34, (7): 37-39, (8): 45-46, (9): 49-50, (11): 61-63, (12): 69-70, (13): 82-84
Courvoisier, 1912
Ueber Zeichnungs-Aberrationen bei Lycaeniden Dt. ent. Z. Iris 26 (1) : 38-65
Courvoisier, 1913
Einige neue oder wenig bekannte Lycaeniden-Formen Ent. Mitt. 2 (10) : 289-297
Courvoisier, 1914
Zur Synonymie des genus Lycaena Dt. ent. Z. Iris 28 (2) (2): 143-176, (3): 177-229
Dannehl, 1933
Neues aus meiner Sammlung. (Macrolepidoptera) Ent. Zs. 46 (22): 220-232, (23): 244-247, (24): 259-260
Denis & Schiffermüller, 1775
Ankündung eines systematischen Werkes von den Schmetterlingen der Wienergegend Ank. syst. Schmett. Wienergegend : 1-322, pl. 1a-b
Eliot, 1973
The higher classification of the Lycaenidae (Lepidoptera): a tentative Arrangement Bull. Br. Mus. nat. Hist. (Ent.) 28 (6) : 373-505 (162fgs), pl. 1-6
van Eecke, 1915
Bijdrage tot de kennis der Nederlandsche Lycaena-soorten Zool. Meded. 1 (3): 22-31
Elwes, 1899
On the Lepidoptera of the Altai Mountains Trans. ent. Soc. Lond. 1899 (3) : 295-367, pl. 11-14
Esper, 1777
Die Schmetterlinge in Abbildungen nach der Natur mit Beschreibungen. Theil I. Die Tagschmetterlinge. Band 1 Die Schmett. Th. I, Bd. 1 (1): 1-40, pl. 1-6 (1776), (2): 41-72, pl. 7-12 (1776), (3): 73-104, pl. 13-18 (1777), (4): 105-120, pl. 19-24 (1777), (5): 121-144, pl. 25-30 (1777), (6): 145-176, pl. 31-36 (1777), (7): 177-200, pl. 37-46 (1778), (8): 201-216, pl. 43-48 (1778), (9): 217-388, pl. 49-50 (1779)
Eversmann, 1848
Beschreibung einigen neuen falter Russlands Bull. Soc. Imp. Nat. Moscou 21 (3) : 205-232
Fabricius, 1787
Mantissa insectorum sistens species nuper detectas adiectis synonymis, observationibus, descriptionibus, emendationibus Mantissa Insectorum 2 : 1-382
Christ in Frey, 1880
Die Lepidopteren der Schweiz Lep. Schweiz. : 1-454
Frey, 1882
Zweiter Nachtrag zur Lepidopteren-Fauna der Schweiz Mitt. schweiz. ent. Ges. 6 (7) : 349-375
Fruhstorfer, 1910
Neue Lycaeniden des palaearktischen Gebietes Int. ent. Zs. 4 (10): 55 (4 June), (12): 62-63 (18 June)
Fruhstorfer, 1910
Neue palaearktische Rhopaloceren Ent. Zs. 24 (26) : 144 (1 October)
Fruhstorfer, 1915
Neue palaearktische Lycaeniden Soc. ent. 30 (11): 63 (15 October), (12): 67-68 (12 November), (13): 73 (10 December)
Fruhstorfer, 1917
Eine neue Lycaena aus der Schweiz Soc. ent. 32 (6) : 26 (25 May)
Fruhstorfer, 1917
Neue palaearktische Lycaeniden Dt. ent. Z. Iris 31 (1/2) : 24-43 (1 July)
Geest, 1903
Neue Schmetterlings-Aberrationen Allg. Zs. Ent. 8 : 308-313
Gillmer, 1904
Referat über: The Butterflies of Switzerland and the Alps of Central Europe Soc. ent. 18 (20): 154-157, (21): 161-163, (22): 170-172, (23): 178-181, (24): 186
Grund, 1908
Die Lycaeniden der Umgebung von Agram (Zagreb, Kroatien) Int. ent. Zs. 2 (11): 66-67, (12): 70-71, (13): 78-79, (14): 87-88
Hauder, 1901
Beitrag zur Macrolepidoptera-Fauna von Österreich ob der Enns Jahresb. ver. naturk. Linz 30 : 1-120
Heydemann, 1910
Einige neue paläarktische Lepidopteren-Formen Ent. Zs. 23 (40) : 177-178
[H&R]; Higgins & Riley, 1970
A Field Guide to the Butterflies of Britain and Europe
Hirschke, 1905
Eine neue hochalpine Form der Lycaena Alcon F. aus den steirischen Alpen Jber. Wien. ent. Ver. 15 : 109-111
Hormuzaki, 1892
Beschreibung einiger neuer Tagfaltervarietäten aus der Bucovina und den Nachbargebieten Ent. Nachr. 18 (1) : 1-3
Hormuzaki, 1897
Die Schmetterlinge (Lepidoptera) der Bukowina Verh. zool.-bot. Ges. Wien. 47 : 70-103, : 120-168, : 233-246, : 312-341
Huang & Wu, 2003
New and little known Chinese butteflies in the collection of the Institue of Zoology, Academia Sinica, Beijing - 1 Neue Ent. Nachr. 55 : 115-143
Hübner, [1799-1800]
Sammlung europäischer Schmetterlinge. I. Papiliones - Falter ("Erste Band") Samml. eur. Schmett. [1] : 3-74, : pl. 1-88, f. 1-446 (1799-1800), : pl. 89-96, f. 447-490 (1800-1803), : pl. 97-114, f. 491-588 (1803-1804), : pl. 115-119, f. 589-612 (1805), : pl. 120-124, f. 613-636 (1805-1806), : pl. 125-128, f. 637-652 (1806-1808), : pl. 129-144, f. 653-727 (1808-1813), : pl. 145-147, f. 728-741 (1814-1816), : pl. 148-150, f. 742-756 (1816-1817), : pl. 151, f. 757-762 (1818-1819), : pl. 152-154, f. 757-774 (1819-1822), : pl. 155, f. 775-778 (1822-1823), : pl. 156-161, f. 779-806 (1823), : pl. 162-175, f. 807-874 (1823-1824), : pl. 176, f. 869-874 (1824-1825), : pl. 177-181, f. 875-895 (1825-1826), : pl. 182-187, f. 896-981 (1827-1828), : pl. 188-195, f. 982-968 (1828-1832), : pl. 196, f. 969-974 (1832-1833), : pl. 197-201, f. 975-997 (1834-1836), : pl. 202-203, f. 998-1009 (1836-1838), : pl. 204-207, f. 1010-1029 (1837-1838)
Jachontov, [1909]
[Notice sur les Lèpidoptères Rhopalocères du Caucase] Revue Russe Ent. 8 : 282-292
Jachontov, 1935
[Our Butterflies. A guide] [in Russian] Our Butterflies: 1-160
Kanda, 1934
A new subspecies of Lycaenidae from Kuriles Ageha 3 : 8-9
Kardakoff, 1928; Kardakov, 1928
Zur Kenntnis der Lepidopteren des Ussuri-Gebietes Ent. Mitt. 17 (4): 261-273, (8): 414-425
Knoch, 1782
Beiträge zur Insektengeschichte Beitr. Insektengesch. 2 : 1-102, pl. 1-6
Korb, 1997
Description d'un Lycene nouveau d'Asie Moyenne: Maculinea alcon alconides ssp.n. (Lepidoptera, Lycaenidae) Alexanor 20 (1): 61-64
Korb, 2003
Notes sur la systématique et la répartition des Lycènes paléarctiques (III) (Lepidoptera, Lycaenidae) Bull. Soc. Nat. Mulhouse 59 (3): 46-....
Korb, 2010
[New taxa of satyrids an dblues (Lepidoptera: Satyridae, Lycaenidae) from Kirghizia] (in Russian) Eversmannia (23-24) : 10-16
Korb & Bolshakov, 2011
[A catalogue of butterflies (Lepidoptera: Papilioformes) of the former USSR. Second edition, reformatted and updated] (in Russian) Eversmannia Suppl. 2 : 1-123
[BARU]; Korshunov & Gorbunov, 1995
Dnevnye babochki Aziatskoi chasti Rossii. Spravochnik [Butterflies of Asian part of Russia. Guide.] [in Russian] Butts. Asian Russia : 1-202, (English): English translation by Oleg Kosterin
Kozhantshikov, 1924
Materialen zur Macrolepidopterenfauna des Minussinsk Bezirks II Jb. Martjanov Staatmus. Minussinsk 2 (1) : 66-75 [?]
Kozhanchikov, 1936
Lepidopterologisches aus Zentral-Sibirien Folia zool. hydrobiol. 9 (1) : 22-29
Krulikovsky, 1892
Lycaena Arion L., var. Rühli, nova Soc. Ent. 7 (1) : 1
Kudrna, 1974
An annotated List of Japanese Butterflies Atalanta 5 : 92-120
[RFEL]; Leley, 2016
Annotated catalogue of the insects of Russian Far East. Volume II. Lepidoptera Cat. ins. Russian Far East. 2 : 1-812
Lempke, 1936
Catalogus der Nederlandsche Macrolepidoptera Tijdschr. Ent. 79 : 238-315
Lempke, 1955
Catalogus der Nederlandse Macrolepidoptera (Derde Supplement) Tijdschr. Ent. 98 : 283-345
[BOW]; Lewis, 1974
Butterflies of the World
Linnaeus, 1758
Systema Naturae per Regna Tria Naturae, Secundum Clases, Ordines, Genera, Species, cum Characteribus, Differentiis, Symonymis, Locis. Tomis I. 10th Edition Syst. Nat. (Edn 10) 1 : 1-338, : 339-824
Marschner, 1909
Lycaena arcas Rott. ab. ♂ emutata Marschner und monographische Behandlung benannter paläarktischer Arcas-Formen Zs. wiss. InsektBiol. 5 : 166-168
Matsumura, 1910
Lycaeniden Japans Ent. Zs. 23 (49): 217-218, (50): 221-222
Matsumura, 1919
Thousand Insects of Japan. Additamenta 3 [Shin Nihon senchu zukai] Thous. Ins. Japan. Addit. 3 : 475-742, pl. 26-53
Matsumura, 1926
Some new and unrecorded Lycaenids-species from Japan, Corea and Formosa Ins. Matsumurana 1 (1) : 23-31
Matsumura, 1927
A list of the butterflies of Corea, with description of new species, subspecies and aberrations Insecta Matsumurana 1 (4) : 159-170
Matsumura, 1927
Some new Butterflies Ins. Matsumurana 2 (2) : 114-118
Matsumura, 1928
New Butterflies especially from the Kuriles Insecta Matsumurana 2 (4) : 191-201
Matsumura, 1939
The butterflies from Jehol (Nekka) Manchoukuo, collected by Marqis Y. Yamashina Bull. biogeogr. Soc. Japan 9 (20) : 343-359, pl. 13-14
Melvill, 1873
Description of Lycaena arthurus, a new European butterfly Ent. mon. Mag. 9 : 263
Munguira, García-Barros & Martín, 1997
Plantas nutricias de los licénidos y satirinos españoles (Lepidoptera: Lycaenidae y Nymphalidae) Boletín de la Asociación española de Entomología, 21 : 29-53
Murayama, 1963
Remarks on some butterflies from Japan and Korea, with descriptions of 2 races, 1 form, 4 aberrant forms Tyô to Ga 14 (2): 43-50
Murayama, 1991
Remarks on new taxa of Satyridae & Lycaenidae from North China and Chejudo Island Nature & Ins. 26 (3): 20-21
Murayama, 1992
Some new Lycaenid species of Chinese Rhopalocera Nature & Ins. 27 (5) : 37-41
Oberthür, 1910
Notes pour servir à établir la Faune Française et Algérienne des Lépidoptères Ètud. Lépid. Comp. 4 : 15-691, pl. 35-58
Oberthür, 1916
Considérations sur plusiuers Espèces de Lycaena Ètud. Lépid. Comp. 12 (1): 453-520 (1916), 13: 44-45, pl. 433-434 (1917)
Obraztsov, 1936
Neue palaearktische Rhopaloceren-Formen Z. Öst. EntVer. 21 : 46-48
Rebel, [1909]
Fr. Berge's Schmetterlingsbuch nach dem gegenwärtigen Stande der Lepidopterologie in Berge, Schmetterlingsbuch (edn 9) : 1-509
Rebel & Zerny, 1931
Die Lepidopterenfauna Albaniens (mit Berücksichtigung der Nacharbegiete) Denkschr. Akad. Wiss. Wien 103 : 37-161, pl. 1
Rezbanyai, 1978
Maculinea arion buholzeri n. ssp. und das arion-Problem in der Zertalschweiz Mitt. schweiz. Ent. Ges. 51 (4) : 399-406
Rottemburg, 1775
Unmertungen zu den Hufnagelifchen Tabellen der Schmetterlinge Der Naturforscher 6 : 1-34
Schultz, 1904
Ueber einige Aberrationen aus der Gruppe der Lycaeniden Ent. Zs. 18 (24) : 93-94
Schultz, 1906
Ueber einige Aberrationen aus der Gruppe der Lycaeniden II Ent. Zs. 19 (36) : 214-215
Scriba, 1921
Drei neue Schmetterlinge aus Central-Japan Ent. Rundschau. 38 (6): 25, f. 1-4, (7): 29
[SPRK]; Seppänen, 1970
Suomen suurperhostoukkien ravintokasvit Animalia Fennica 14
Sheljuzhko, 1928
Neue palaearktische Lycaeniden Lep. Rdsch. 2 (4): 44-46, (5): 51-52, (6): 62-64
Shirôzu, 1953
New or little know butterflies from the North-Eastern Asia, with some synonymic notes. (1) & (2) Sieboldia 1 (1952) : 11-37 (11pls), : 149-159
Sibatani, Saigusa & Hirowatari, 1994
The genus Maculinea van Eecke, 1915 (Lepidoptera, Lycaenidae) from the East Palaearctic Region Tyô to Ga 44 (4) : 157-220
Silbernagel, 1946
[Neue aberrationen der gattung Lycaena (Macrolep.)] Acta Soc. ent. bohem. 37 : 116-119
Skala, 1912
Die Lepidopterenfauna Mährens. I Teil Ver. naturf. Ver. Brünn 50 : 63-241
[EBW]; Smart, 1976
The Illustrated Encyclopedia of the Butterfly World Illust. Encyp. Butt. World: -
Sohn, 1893
Lycaena Arcas ab. inocellata n. ab. Soc. Ent. 8 (10) : 77
Spuler, 1908
Die Schmetterlinge Europas. Dritte Auflage. I. Band Schmett. Eur. 1 : 1-385
Staudinger, 1887
Neue Arten und Varietäten von Lepidopteren aus dem Amur-Gebiete in Romanoff, Mém. Lépid. 3 : 126-232, pl. 6-12, 15-17
Staudinger, 1892
Lepidopteren des Kentei-Gebirges Dt. Ent. Z. Iris 5 (2) : 300-393
Staudinger in Staudinger & Rebel, 1901
Catalog der Lepidopteren des palaearktischen Faunengebiets. I. Famil. Papilionidae - Hepialidae Cat. Lep. palaearct. Faunengeb. 1 : 1-411
Stephens, 1828
Illustrations of British Entomology; or, a synopsis of indigenous insects: containing their generic and specific distinctions; with an account of their metamorphoses, times of appearance, localities, food, and economy, as far as practicable Ill. Br. Ent. (Haustellata) 1 (1): 1-56, pl. 1-9 (1827), (2): 57-150, pl. 10-12 (1828)
Stradomsky, 2016
A molecular phylogeny of the subfamily Polyommatinae (Lepidoptera: Lycaenidae) Caucasian Ent. Bull. 12 (1) : 145-156
Sugitani, 1934
Corean butterflies (3) Zephyrus 5 : 165-176
Szabó, 1956
Magyarország Lycaenidái Folia ent. hung. (n.s.) 9 : 235-362
Takahashi, 1973
Maculinea teleius Bergstraesser at high elevation in the Central Honshû, Japan, with description of a new subspecies Tyô to Ga 23 (3-4) : 75-85
Verity, 1919
A 1000 metri sull'Appennino Modenese - note di lepidotterologia e descrizione di tre nuove specie di micri Atti Soc. ital. Sci. nat. 58 : 147-187
Tutt, 1909
Lepidoptera of the Grisons-Zernetz and Ofen Pass Ent. Rec. J. Var. 21 : 197-201
Tutt, 1914
A natural history of the British Butterflies. Their world-wide variation and geographical distribution. A text-book for students and collectors Nat. Hist. Br. Lepid. 4 : 1-373, pl. 1-45 (pl. 3, 12, 20 not publ.) (1910-1914)
[BRU2]; Tuzov, Bogdanov, Churkin, Devyatkin, Dantchenko, Murzin, Samodurov, Zhdanko, 2000
Guide to the Butterflies of Russia and adjacent territories: Libytheidae, Danaidae, Nymphalidae, Riodinidae, Lycaenidae Butts. Russia adj. terr. 2 : 1-580
Ugelvig, Vila, Pierce & Nash, 2011
A phylogenetic revision of the Glaucopsyche section (Lepidoptera: Lycaenidae), with special focus on the Phengaris-Maculinea clade Molec. Phyl. Evol. 61 : 237-243
Verity, 1924
Additions and Corrections to "List of Grypocera and Rhopalocera of Peninsular Italy" Ent. Rec. J. Var. 36 : 106-111
Verity & Querci, 1923; Verity in Verity & Querci, 1923
Races and Seasonal Polymorphism of the Grypocera and of the Rhopalocera of Peninsular Italy Ent. Rec. (suppl.) 35: 1-20 (1923), 36: 21-44,45-46 [?] (1924)
Vorbrodt & Müller-Rutz, 1911; Vorbrodt, 1912
Die Schmetterlinge der Schweiz. I. Band Die Schmett. Schweiz 1 : 1-489
Wagner, 1904
Lycaena arion L. nov. var. Soc. Ent. 19 (1) : 1
Warnecke, 1924
Doerriesia (Bemerkungen über ostasiatische und sibirische Großschmetterlinge) Dt. ent. Z. Iris 38 : 149-154
Watari, 1937
On a new subspecies of Lycaena euphemus Zephyrus 7 : 26-28
Wheeler, 1903
The Butterflies of Switzerland and the Alps of Central Europe Butts. Switzerland : 1-162

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