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Leptosia Hübner, 1818
Leptosia Hübner, 1818; Zuträge Samml. exot. Schmett. 1 : 13, TS: Leptosia chlorographa Hübner
Nina Horsfield, [1829]; Descr. Cat. lep. Ins. Mus. East India Coy (2): 140, TS: Papilio nina Fabricius
Nychitona Butler, 1870; Cistula ent. 1 (3) : 34, 41, TS: Papilio dorothea Fabricius
Leptosia, NSG Voucher Specimen [Wahlberg]
Leptosia ; Winhard, 2000, Butterflies of the world, 10: 31
Nychitona (Pierinae) ; Moore, [1881], Lepid. Ceylon 1 (3): 117
Leptosia (Pierinae) ; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 103
Leptosia (Anthocharidini) ; [AfrL]

Leptosia alcesta (Stoll, [1782])
Natal, Swaziland, Moçambique, Rhodesia, Tropical Africa. See [About maps] African Wood White Flip Flop
Papilio alcesta Stoll, [1782]; in Cramer, Uitl. Kapellen 4 (32-32): 175, pl. 379, f. A, B
Papilio dorothea Fabricius, 1793; Ent. Syst. 3 (1): 194
?uganda Neustetter, 1927²; Int. ent. Z. 21 : 7
?Leptosia uganda ; [EBW]
Leptosia alcesta ; [EBW]; [AfrL]

Larva on Richea [EOB], Capparis fascicularis var. zeyheri [PBSA]

W.Africa Leptosia alcesta alcesta ; [BAFR, 103 (text)]; [AfrL]
L. a. inalcesta Bernardi, 1959
Ethiopia, Sudan, Uganda, Zaire, Kenya, Tanzania - Natal Leptosia alcesta inalcesta ; [BAFR, 103]; [BK, 147]; Winhard, 2000, Butterflies of the world, 10: 31, pl. 48, f. 14; [AfrL]
L. a. sylvicola (Boisduval, 1833)
Madagascar Leucophasia sylvicola Boisduval, 1833; , TL: Madagascar
Leptosia alcesta sylvicola ; [BAFR, 103 (text)]; [MadL]; [AfrL]
L. a. pseudonuptilla Bernardi, 1959
Ethiopia Leptosia nupta pseudonuptilla ; [BAFR, 103 (text)]
Leptosia pseudonuptilla ; Winhard, 2000, Butterflies of the world, 10: 31, pl. 48, f. 14
Leptosia alcesta pseudonuptilla ; [AfrL]

Leptosia bastini Hecq, 1997
Leptosia bastini ; [AfrL]

Leptosia hybrida Bernardi, 1952
-. See [About maps] Hybrid Wood White
Leptosia hybrida ; [AfrL]
Guinea - Zaire Leptosia hybrida hybrida ; [BAFR, 103]; [AfrL]
L. h. somereni Bernardi, 1959
E.Zaire, Uganda, W.Kenya Leptosia hybrida somereni ; [BAFR, 103 (text)]; [BK, 147]; Winhard, 2000, Butterflies of the world, 10: 31, pl. 48, f. 17; [AfrL]

Leptosia marginea (Mabille, 1890)
Ivory Coast - Cameroun, W.Zaire. See [About maps] Leptosia marginea ; [BAFR, 103]; Winhard, 2000, Butterflies of the world, 10: 31, pl. 48, f. 16; [AfrL]

Leptosia medusa (Cramer, [1777])
W.Africa, Sierra Leone - Ghana. See [About maps] Papilio medusa Cramer, [1777]; Uitl. Kapellen 2 (9-16): 86, pl. 150, f. F
Pontia medusa ; Boisduval, 1836, Hist. nat. Ins., Spec. gén. Lépid. 1 : 433
Leptosia medusa ; [EBW]; [BAFR, 103]; [AfrL]

Leptosia nina (Fabricius, 1793)
India, Ceylon, Burma, AU. See [About maps] Psyche
Papilio nina Fabricius, 1793; Ent. Syst. 3 (1): 194, no. 604, TL: India
Leptosia chlorographa Hübner, 1818; Zuträge Samml. exot. Schmett. 1 : 13, : pl. [9], f. 47-48, TL: Java
Papilio xiphia Fabricius, 1781; Spec. Ins. 2 : 43 (preocc.)
Pontia crokera MacLeay, [1826]; in King, Narr. Surv. Austr. 2: 458
Leptosia xiphia micropunctata van Eecke, 1918; Zool. Meded. 4 (7) : 73, TL: Pulu Lasia
= Leptosia nina micropunctata ; [NHM]
Leptosia nina nina, Butterflies in Indo-China [Yutaka Inayoshi]
Pontia crokera ; Boisduval, 1836, Hist. nat. Ins., Spec. gén. Lépid. 1 : 431
Nychitona xiphia ; Moore, [1881], Lepid. Ceylon 1 (3): 118
Leptosia nina ; [EBW]; [BOR, 122]; [MRS, 262]; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 104
Leptosia xiphia ; [BOW: pl. 160, f. 29]
[nina-1v.jpg] 769x513 (~65Kb) underside Thailand, Krabi 24.12.2003-7.1.2004, Photo © Olli Vesikko

Larva on Capparidaceae [BIR], Capparis heyneana, Crateva religiosa [MRS], Capparis, Cleome, Crateva, Polanisia Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 104


India, Ceylon - Indo-China, Thailand, Langkawi Is, Andamans Leptosia nina nina ; [BOR, 122]; [BMP: 85, pl. 8, f. 1]; Winhard, 2000, Butterflies of the world, 10: 31, pl. 48, f. 12
[nina_nina-1.jpg] 930x852 (~74Kb) Thailand, Chon Buri Province, Pattaya, a small grassy swamp with cattail and reed, surrounded by trees, at the Jomtien beach, between B.O. Guesthouse and Jomtien Metro Condotel. 26th January 2005, Photo © Oleg Kosterin

Java Leptosia nina chlorographa ; [BOR, 122]
L. n. niobe (Wallace, 1866)
Taiwan Pontia niobe Wallace, 1866; Proc. zool. Soc. Lond. 1866 (2) : 357, TL: Formosa
Leptosia nina niobe ; [MRS, 262]; [BOR, 122]

Larva on Capparis acutifolia, C. floribunda, C. lanceolaris, C. micracantha, C. sikkimensis, Cleome gynandra, Crateva adansonii, Polanisia viscosa [MRS]

L. n. dione (Wallace, 1867)
S.Sulawesi Pontia dione Wallace, 1867; Trans. ent. Soc. Lond. (3) 4 (3) : 319, TL: Macassar
Leptosia nina dione ; [BOR, 122]; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 104
L. n. nicobarica (Doherty, 1886)
Nicobars Nychitona xiphia var. nicobarica Doherty, 1886; J. asiat. Soc. Bengal 55 Pt.II (3) : 262, TL: Nicobars
Leptosia nina nicobarica ; [BOR, 122]
L. n. fumigata Fruhstorfer, 1903
Lombok, Sumbawa, Flores, Solor Leptosia xiphia fumigata Fruhstorfer, 1903; Dt. ent. Z. Iris 15 (2) : 269, TL: Lombok
Leptosia xiphia fumigata Fruhstorfer, 1903; Soc. Ent. 18 (3): 18, TL: Lombok
?aurisparsa Fruhstorfer
Leptosia nina fumigata ; [BOR, 122]
L. n. terentia Fruhstorfer²
S.Philippines Leptosia nina terentia ; [BOR, 122]
L. n. georgi Fruhstorfer²
N.Philippines Leptosia nina georgi ; [BOR, 122]
L. n. comma Fruhstorfer, 1903
Timor - Tanimbar Leptosia xiphia comma Fruhstorfer, 1903; Dt. ent. Z. Iris 15 (2) : 269, TL: Timor; Wetter; Kalao; Tanah; Djampea; Selaru; Key
Leptosia xiphia comma Fruhstorfer, 1903; Soc. Ent. 18 (3): 18, TL: Timor; Wetter; Kalao; Tanah; Djampea; Selaru; Key
Leptosia nina comma ; [BAUR, 124]
L. n. malayana Fruhstorfer, 1910
Peninsular Malaya, Singapore, Sumatra, Borneo, Bangka, Biliton Leptosia nina malayana Fruhstorfer, 1910; in Seitz, Gross-Schmett. Erde 9 : 121, (44) , TL: Malaysia
Leptosia nina malayana, Butterflies in Indo-China [Yutaka Inayoshi]
Leptosia nina malayana ; [BOR, 122]

Larva on Capparis heyneana, Crateva religiosa [BMP]

L. n. aebutia Fruhstorfer, 1910²
Tanahdjampea, Kalao Leptosia nina aebutia ; [BOR, 122]; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 104

Leptosia lignea (Vollenhoven, 1865)
Sulawesi. See [About maps] Pontia lignea Vollenhoven, 1865; Faune ent. 2 (Monogr. Pierid.) : 4, pl. 2, f. 1a-b, TL: Celebes
Leptosia nina lignea ; [BOR, 122]; Winhard, 2000, Butterflies of the world, 10: 31, pl. 48, f. 13
Leptosia lignea ; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 104

Larva on Capparis Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 104


Leptosia nupta (Butler, 1873)
-. See [About maps] Immaculate Wood White
Nychitona nupta Butler, 1873; Cist. Ent. 1 (7) : 175, TL: Angola, Bembe Mines
Leptosia nupta ; [AfrL]

Larva on Capparidaceae? [BK]

Nigeria - Angola Leptosia nupta nupta ; [BK, 147]; [BAFR, 103]; [AfrL]
L. n. pseudonupta Bernardi, 1959
Zaire, Uganda - Tanzania Leptosia nupta pseudonupta ; [BAFR, 103 (text)]; [AfrL]
L. n. viettei Bernardi, 1959
Madagascar Leptosia nupta viettei Bernardi, 1959; , TL: Madagascar, Massif de l'Ankaratra, Forêt d'Ambahona, Manjakatompo
Leptosia nupta viettei ; [BAFR, 103 (text)]; [MadL]; [AfrL]

Leptosia wigginsi (Dixey, 1915)
-. See [About maps] Opaque Wood White
Nychitona wigginsi Dixey, 1915; Trans. Ent. Soc. 1915 (1) : 7, pl. 1, f. 9-12
Leptosia wigginsi ; [BK, 148]; [AfrL]
Uganda, SW.Kenya, Tanzania, E.Zaire Leptosia wigginsi wigginsi ; [BAFR, 103]; [AfrL]
L. w. pseudalcesta Bernardi, 1965
Senegal - Ghana Leptosia wigginsi pseudalcesta; [BAFR, 103 (text)#]
Leptosia wigginsi pseudalcesta ; [AfrL]


30.6.2012 (9)

References:
[About maps]
Warning! The maps are automatically generated from the textual information, and the process does not always produce acceptable result; See about maps for more info.
[AfrL]
Mark C. Williams; Checklist of Afrotropical Papilionoidea and Hesperoidea; Compiled by Mark C. Williams, 7th ed. (2008) (April 2007); Afrototropical Butterflies; e-mail: Mark C. Williams
[BAFR]
D'Abrera, 1980; Butterflies of the Afrotropical Region
[BAUR]
D'Abrera, 1977; Butterflies of the Australian Region (2nd edn)
[BIR]
Wynter-Blyth, 1957; Butterflies of the Indian Region; (1982 Reprint)
[BK]
Larsen, 1996; The Butterflies of Kenya and their natural history
[BMP]
Eliot (Ed.), Corbet & Pendlebury, 1992; The Butterflies of The Malay Peninsula; 4th edition; ISBN 983-9681-05-2
[BOR]
D'Abrera, 1982, 1985, 1986; Butterflies of the Oriental Region, Part I-III
[BOW]
Lewis, 1974; Butterflies of the World; ISBN 0 245 52097 X
[EBW]
Smart, Paul, 1976; The Illustrated Encyclopedia of the Butterfly World; ISBN 0 600 31381 6
[EOB]
Feltwell; The Encyclopedia of Butterflies; Prentice Hall, 1993; ISBN 0-671-86828-4
[HBN]
Francis Hemming, 1937; HÜBNER, A bibliographical and systematic account of the entomological works of Jacob Hübner and of the supplements thereto by Carl Geyer, Gottfried Franz von Frölich and Gottlieb August Wilhelm Herrich-Schäffer, Vol 1 & 2
[LAMAS-2005]
Gerardo Lamas; Entomofauna 26 (6): 57-100; A Bibliography of the Zoological Publications of Hans Fruhstorfer (1886* - 1922+)
[LAMAS-NEO]
Gerardo Lamas; Bibliography of butterflies; An Annocated Bibliography of the Neotropical Butterflies and Skipper (Lepididoptera: Papilionoidea and Hesperioidea); Revised Electronic Edition (2007)
[MRS]
Chou Io (Ed.); Monographia Rhopalocerum Sinensium, 1-2
[MadL]
Lees, D.C., Kremen, C. and Raharitsimba, H. 2003; in S.M. Goodman & J. P. Benstead (eds.), The Natural History of Madagascar; Classification, Diversity and Endemism of the Butterflies (Papilionoidea and Hesperioidea): A Revised Species Checklist: : pp. 762-793; University of Chicago Press. 1709 pp.
[NBC]
Netherlands Centre for Biodiversity Naturalis; NBC naturalis
[NHM]
Natural History Museum; The Global Lepidoptera Names Index
[Oleg Kosterin]
Institue of Cytology & Genetics; Siberian Division of Russian Academy of Sciences; Novosibirsk; e-mail: Oleg Kosterin; http://pisum.bionet.nsc.ru/kosterin/
[Olli Vesikko]
http://koti.welho.com/ovesikko/
[PBSA]
Dickson, C.G.C (ed.), Kroon, D.M.; Pennington's Butterflies of Southern Africa; AD. DONKER 1978
[Wahlberg]
Niklas Wahlberg; The NSG's voucher specimen database; Nymphalidae Systematics Group
[Yutaka Inayoshi]
A Check List of Butterflies in Indo-China; Chiefly from THAILAND, LAOS & VIETNAM
[²]
This may require parentheses or not. I don't have the necessary information for this taxon.


Some related literature:
Boisduval, 1836
Histoire Naturelle des Insectes. Species Général des Lépidoptéres. Tome Premier Hist. nat. Ins., Spec. gén. Lépid. 1 : 1-690, pl. 1-24
Butler, 1870
A revision of the Genera of the Sub-family Pierinae Cistula ent. 1 (3) : 33-58, pl. 1-4
Butler, 1873
Descriptions of new species of Lepidoptera Cist. Ent. 1 (7) : 151-177
Cramer, [1777]
Uitlandsche Kapellen (Papillons exotiques) Uitl. Kapellen 2 (9-16): 1-152, pl. 97-192 (1777)
Dixey, 1915
New species and subspecies of Pierinae Trans. Ent. Soc. 1915 (1) : 1-15, pl. 1-2
Doherty, 1886
Additional Notes on new or rare Indian Butterflies J. asiat. Soc. Bengal 55 Pt.II (3) : 256-265
van Eecke, 1918
Studies on Indo-Australian Lepidoptera--III. Some Rhopalocera and Netrocera[sic] from Simalur, Pulu Lasia, Pulu Babi and Sumatra Zool. Meded. 4 (7) : 70-101, 2pls
Fabricius, 1781
Species Insectorum Exhibentes Eorum Differentia Specifica, Synonyma Auctorum, Loca Natalia, Metamorphosin Adiectis, Observationibus, Descriptionibus. Tom II Spec. Ins. 2 : 1-494 (1781), : 495-514 (appendix), 515-517 (index) ([1782])
Fruhstorfer, 1903
Verzeichniss der in Tonkin, Annam und Siam gesammelten Pieriden und Besprechung verwandter Formen Dt. ent. Z. Iris 15 (2) : 269-305 (1 May)
Fruhstorfer, 1903
Verzeichniss der in Hinter-Indien gesammelten Pieriden Soc. Ent. 18 (3): 18-19 (1 May), (4): 25-26 (15 May)
Fruhstorfer, 1910
2. Familie: Pieridae, Weisslinge in Seitz, Gross-Schmett. Erde 9 : 119-120 (1909), : 121-144 (14 January, 1910), : 145-152 (11 February, 1910), : 153-168 (2 April, 1910), : 169-176 (4 April, 1910), : 177-190 (20 May, 1910)
Horsfield, [1829]
Descriptive Catalogue of the Lepidopterous Insects contained in the Museum of the Horourable East-India Company, illustrated by coloured figures of new species Descr. Cat. lep. Ins. Mus. East India Coy (1): 1-80, pl. 1-4 (1828), (2): 81-144, pl. 5-8, (expl.) [1-4] (1829)
Hübner, 1818
Zuträge zur Sammlung exotischer Schmettlinge, Vol. 1 [1808-] 1818 Zuträge Samml. exot. Schmett. 1 : [1-3], 4-6, [7], 8-32, [33-40] (post 22nd December 1818), : pl. [1-2], f. 1-12 (1808 - 1809), : pl. [3-25], f. 13-146 (1809 - 1813), : pl. [26-35], f. 147-200 (1814 - 1818)
MacLeay, [1826]
in King, Narrative of a Survey of the Intertropical and Western Coasts of Australia. 2 vols. in King, Narr. Surv. Austr. 2: -
Moore, [1881]
The Lepidoptera of Ceylon Lepid. Ceylon 1 (1): 1-40, pl. 1-18 (Dec 1880), (2): 41-80, pl. 19-36 (Jan 1881), (3): 81-136, pl. 37-54 (Jun 1881), (4): 137-190, pl. 55-71 (Dec 1881)
Neustetter, 1927
Neue afrikanische Tagfalter Int. ent. Z. 21 : 7-8,14-16,23-24,32
Stoll, [1782]
Uitlandsche Kapellen (Papillons exotiques) in Cramer, Uitl. Kapellen 4 (26b-28): 29-90, pl. 305-336 (1780), (29-31): 91-164, pl. 337-372 (1781), (32-32): 165-252, 1-29, pl. 373-400 (1782)
Vane-Wright & de Jong, 2003
The butterflies of Sulawesi: annotated checklist for a critical island faunda Zool. Verh. Leiden 343: 3-268, pl. 1-16
Vollenhoven, 1865
Essai d'une Faune Entomologique de l'Archipel Indo-Néerlandais, Seconde monographie: Famille des Piérides Faune ent. 2 (Monogr. Pierid.) : 1-70, pl. 1-7
Wallace, 1866
List of Lepidopterous insects collected at Takow, Formosa, by Mr. Robert Swinhoe Proc. zool. Soc. Lond. 1866 (2) : 355-365
Wallace, 1867
On the Pieridae of the Indian and Australian Regions Trans. ent. Soc. Lond. (3) 4 (3) : 301-416, pl. 6-9
Winhard, 2000
Pieridae I Butterflies of the world, 10: (1) 40pp, (2) 48pls


If you have corrections, comments or information to add into these pages, just send mail to Markku Savela
Keep in mind that the taxonomic information is copied from various sources, and may include many inaccuracies. Expert help is welcome.