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Ypthima Hübner, 1818
Rings Ringlets
Ypthima Hübner, 1818; Zuträge Samml. exot. Schmett. 1 : 17 [missid. TS Papilio philomela Linnaeus], TS: Ypthima huebneri Kirby
Xois Hewitson, 1865; Trans. ent. Soc. Lond. (3) 2 (4) : 282, TS: Xois sesara Hewitson
Kolasa Moore, 1891; (nomen nudum)
Thymipa Moore, 1891; (nomen nudum)
Kolasa Moore, 1893; Lepidoptera Indica 2 (14): 57, (15): 82, TS: Satyrus chenui Guérin-Méneville
Thymipa Moore, 1893; Lepidoptera Indica 2 (14): 57, 58, TS: Papilio baldus Fabricius
Nadiria Moore, 1893; Lepidoptera Indica 2 (14): 57, (15): 85, TS: Ypthima bolanica Marshall
Pandima Moore, 1893; Lepidoptera Indica 2 (14): 58, (15): 86, TS: Satyrus nareda Kollar
Lohana Moore, 1893; Lepidoptera Indica 2 (14): 58, (16): 92, TS: Yphthima[sic] inica Hewitson
Dallacha Moore, 1893; Lepidoptera Indica 2 (14): 58, (16): 94, TS: Yphthima[sic] hyagriva Moore
Shania Evans, 1912; J. Bombay nat. Hist. Soc. 21 (2): 564, TS: Ypthima megalia de Nicéville
Sundaypthima Uemura, 1996
Ypthima, NSG Voucher Specimen [Wahlberg]
Iphthima ; Westwood, [1851], Gen. diurn. Lep. (2): pl. 67 (missp.)
Yphthima ; Scudder, 1875, Proc. Amer. Acad. Arts Sci. 10 (2) : 289 (missp.)
Thympia ; Hemming, 1967, Bull. Br. Mus. (Ent.) Suppl. 9 : 442 (missp.)
Ypthima (Satyrinae) ; Moore, [1880], Lepid. Ceylon 1 (1): 24
Ypthima (Satyrina) ; [Wahlberg]
Ypthima (Ypthimina) ; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 183; [AfrL]


[sp-1u.jpg] 816x612 (~79Kb) upperside Thailand, Chantaburi Province, Khao-Khitchakut National Park, a floating bog on a bank of the pond at headquarters. 30th January 2005, Photo © Oleg Kosterin [sp-1v.jpg] 743x786 (~67Kb) underside Thailand, Chantaburi Province, Khao-Khitchakut National Park, a floating bog on a bank of the pond at headquarters. 30th January 2005, Photo © Oleg Kosterin [sp-2v.jpg] 600x400 (~50Kb) underside India: Melghat, 9.10.2008, Photo © Kedar Tokekar
[sp-3v.jpg] 928x820 (~101Kb) underside Thailand, Chantaburi Province, Khao-Khitchakut National Park, a bank of the pond at headquarters. 7th January 2006, Photo © Oleg Kosterin [sp-4u.jpg] 512x341 (~35Kb) upperside India: Guwahati, 17.11.2005, Photo © Kedar Tokekar [sp-5.jpg] 512x341 (~36Kb) India: Ziro, 9.11.2005, Photo © Kedar Tokekar
[sp-6mfv.jpg] 1137x989 (~250Kb) underside male female Thailand, Phuket, a garden in the Nai Yang beach area. 16th February 2009, Photo © Oleg Kosterin [sp-7v.jpg] 810x827 (~203Kb) underside Thailand, Phuket, a bank of a pond at a road 1 km NE of Khao Sai Tan Kliang Mt., 18th February 2009, Photo © Oleg Kosterin

The exact identification of these species is still unknown, but tentatively assumed to belong into this group.


Subgenus Thymipa Moore, 1893
Thymipa Moore, 1893; Lepidoptera Indica 2 (14): 57, 58, TS: Papilio baldus Fabricius
Thymipa ; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80

The sakra species-group
sakra ; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80

Ypthima similis Elwes & Edwards, 1893
Assam, Manipur, Burma, Thailand, Laos. See [About maps] Eastern Five-Ring
Ypthima similis Elwes & Edwards, 1893; Trans. Ent. Soc. Lond. 1893 (1) : 29, pl. 1, f. 19, TL: Karen Hills
yoma Evans, 1924²; J. Bombay nat. Hist. Soc. 29 (3) (3): 787, TL: Pegu
= ; [NHM]
Ypthima similis, Butterflies in Indo-China [Yutaka Inayoshi]
Ypthima similis ; [BOR, 465]
Ypthima (Thymipa) similis ; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80
[similis-1v.jpg] 861x909 (~89Kb) underside Thailand, Chon Buri Province, Pattaya, a vasting land with open tree stand behind the Jomtien beach, 1st February, 2005, Photo © Oleg Kosterin [similis-2u.jpg] 684x754 (~82Kb) upperside Thailand, Chon Buri Province, Pattaya, Ko Khram Island, Saikaew beach, a bank of a pond in a wood near the coast, 3rd February, 2006, Photo © Oleg Kosterin

S.Yunnan, ... Ypthima similis similis ; Huang & Xue, 2004, Neue Ent. Nachr. 57: 141 (name), pl. 11, f. 7

Ypthima iarba de Nicéville, 1895
-. See [About maps] Ypthima iarba de Nicéville, 1895; J. Bombay nat. Hist. Soc. 10 (1): 18, pl. R, f. 7-8, TL: Sumatra
Ypthima iarba, Butterflies in Indo-China [Yutaka Inayoshi]
Ypthima jarbas[sic] ; [NHM]
Ypthima iarba ; [BOR, 472]
Ypthima (Thymipa) iarba ; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80
Burma, Sumatra, ?Peninsular Malaya Ypthima iarba iarba ; [BOR, 472]
Y. i. gaugamela Fruhstorfer, 1911²
E.Java iarba gaugamela Fruhstorfer, 1911²; in Seitz, Gross-Schmett. Erde 9: pl. 99 d
Ypthima iarba gaugamela ; [BOR, 472]
Ypthima (Thymipa) gaugamela ; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80
Y. i. eupeithes Fruhstorfer, 1911²
W.Java iarba eupeithes Fruhstorfer, 1911²; in Seitz, Gross-Schmett. Erde 9 : 287, pl. 99 f
Ypthima iarba eupeithes ; [BOR, 472]
Ypthima (Thymipa) eupeithes ; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80

Ypthima affectata Elwes & Edwards, 1893
Assam, Manipur. See [About maps] Ypthima affectata Elwes & Edwards, 1893; Trans. Ent. Soc. Lond. 1893 (1) : 30, pl. 1, f. 30, TL: Terria Ghat; Mamloo, Khasias
Ypthima affectata, Butterflies in Indo-China [Yutaka Inayoshi]
Ypthima affectata ; [BOR, 465]
Ypthima (Thymipa) affectata ; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80

Ypthima sobrina Elwes & Edwards, 1893
Assam, Burma, Thailand, Laos, S.Yunnan. See [About maps] Ypthima sobrina Elwes & Edwards, 1893; Trans. Ent. Soc. Lond. 1893 (1) : 29, pl. 1, f. 17-18, TL: Karen Hills
Ypthima sobrina, Butterflies in Indo-China [Yutaka Inayoshi]
Ypthima sobrina ; [BOR, 465]
Ypthima (Thymipa) sobrina ; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80
Ypthima sobrina ; Huang & Xue, 2004, Neue Ent. Nachr. 57: 141 (name)

Ypthima savara Grose-Smith, 1887
Manipur - Burma - Indo China. See [About maps] Pallid Five-Ring
Yphthima[sic] savara Grose-Smith, 1887; Ann. Mag. nat. Hist. (5) 20 : 267, TL: Burma-Siam frontier
Ypthima savara savara, Butterflies in Indo-China [Yutaka Inayoshi]
Ypthima savara ; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1) : 37, pl. 2, f. 30; [BOW: pl. 173, f. 8]; [BOR, 471]
Ypthima (Thymipa) savara ; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80
Burma, Thailand, Laos, S.Yunnan Ypthima savara savara ; Huang & Xue, 2004, Neue Ent. Nachr. 57: 141 (name), pl. 11, f. 6
Y. s. tonkiniana Fruhstorfer, 1908
N.Vietnam Ypthima ? tonkiniana Fruhstorfer, 1908; Ent. Zs. 22 (31) : 127, TL: Tonkin, Chiem-Hoa
Ypthima savara tonkiniana Fruhstorfer, 1911; in Seitz, Gross-Schmett. Erde 9 : 292, pl. 99, f. 2, TL: Chiem Hoa, N.Vietnam
Ypthima savara tonkiniana, Butterflies in Indo-China [Yutaka Inayoshi]
Ypthima savara tonkiniana ; [BMP: 136, pl. 17, f. 20-21]

Ypthima sakra Moore, 1857
Murree - Assam, Karen Hills. See [About maps] Himalayan Five-Ring
Yphthima[sic] sakra Moore, 1857; in Horsfield & Moore, Cat. lep. Ins. Mus. East India Coy 1 : 236
Yphthima[sic] sakra ; Hewitson, 1865, Trans. ent. Soc. Lond. (3) 2 (4) : 290, pl. 18, f. 18
Ypthima sakra ; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1) : 40; [BOW: pl. 173, f. 9]; [BOR, 472]; [MRS, 392]
Ypthima (Thymipa) sakra ; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80
[sakra-1u.jpg] 512x341 (~37Kb) upperside India: Guwahati, 17.11.2005, Photo © Kedar Tokekar

Larva on Gramineae [MRS]


Sikkim, Bhutan Ypthima sakra sakra ; [BOR, 472]
Y. s. austeni (Moore, 1893)
Assam, N.Burma, SE.Tibet (Yigong), Gooligonshan Mts. Thymipa austeni Moore, 1893; Lepidoptera Indica 2 (15): 69, pl. 109, f. 3, 3a
Ypthima sakra austeni, Butterflies in Indo-China [Yutaka Inayoshi]
Ypthima austeni ; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1) : 51
Ypthima sakra austeni ; [BOR, 472]; Huang, 2001; Huang, 2003, Neue Ent. Nachr. 55: 98 (note)
Y. s. nujiangensis Huang, 2001
SE.Tibet Ypthima sakra nujiangensis Huang, 2001; Neue Entomologische Nachrichten, 51: 94, pl. 7, f. 51, TL: Longpo to Nidadan
Ypthima sakra nujiangensis ; Huang, 2003, Neue Ent. Nachr. 55: 98 (note)
Y. s. leechi Forster, [1948]
Sichuan Ypthima sakra leechi Forster, [1948]; Mitt. Münch. ent. Ges. 34 (2) : 483, pl. 30-31, f. 16
Ypthima sakra leechi, Butterflies in Indo-China [Yutaka Inayoshi]
Ypthima sakra leechi ; Huang, 2001, Neue Entomologische Nachrichten, 51: 94 (note)
Y. s. matinia Fruhstorfer, 1911²
NW.India sakra matinia Fruhstorfer, 1911²; in Seitz, Gross-Schmett. Erde 9 : 292
Ypthima sakra matinia ; [BOR, 472]

Ypthima pseudosavara Uémura & Monastyrskii, 2000
Vietnam. See [About maps] Ypthima pseudosavara Uémura & Monastyrskii, 2000; Trans. Lepid. Soc. Japan 51 (2) : 153, f. 1d-e, 3, 10-13, TL: Vietnam, Thanh Hoa Prov., Nhu Xuan, Ben En Nat. Park (105°21'-105°35'E, 19°30'-19°40'N)
Ypthima pseudosavara, Butterflies in Indo-China [Yutaka Inayoshi]

Ypthima nikaea Moore, [1875]
W.Himalayas, Nepal. See [About maps] Yphthima[sic] nikaea Moore, [1875]; Proc. zool. Soc. Lond. 1874 (4) : 567, TL: NW.Himalayas
= Ypthima sakra ; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1) : 40
= ?; Huang, 2001, Neue Entomologische Nachrichten, 51: 94
Ypthima nikaea ; [BOR, 471]; [MRS, 392]
Ypthima (Thymipa) nikaea ; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80

Ypthima hannyngtoni Eliot, 1967
NW.India. See [About maps] Ypthima hannyngtoni Eliot, 1967; Entomologist 100 (1246) : 54, TL: Masuri
Ypthima (Thymipa) hannyngtoni ; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80

Ypthima methorina Oberthür, 1891
China (Kweichow, Ta-tsien-lo). See [About maps] Ypthima methorina Oberthür, 1891; Étud. d'Ent. 15 : 15, pl. 2, f. 15
Ypthima methorina completa Oberthür, 1911; Étud. Lépid. Comp. 5 (1) : 317, pl. 60, f. 551, TL: Thibet
= ; [NHM]
Ypthima methorina ; Forster, [1948], Mitt. Münch. ent. Ges. 34 (2) : 484; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1) : 39, pl. 2, f. 33; [BOW: pl. 173, f. 9 (text only)]
Ypthima (Thymipa) methorina ; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80

Ypthima medusa Leech, [1892]
Szechwan. See [About maps] Ypthima medusa Leech, [1892]; Butts China Japan Corea (1): 84, pl. 10, f. 6, TL: Szechwan
Ypthima medusa ; Forster, [1948], Mitt. Münch. ent. Ges. 34 (2) : 485; [MRS, 392]
Ypthima (Thymipa) medusa ; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80

Larva on Gramineae [MRS]


Ypthima davidsoni Eliot, 1967
Assam. See [About maps] Ypthima davidsoni Eliot, 1967; Entomologist 100 (1246) : 54, TL: Assam, Khasias
Ypthima (Thymipa) davidsoni ; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80

Ypthima formosana Fruhstorfer, 1908
Taiwan Ypthima methora formosana Fruhstorfer, 1908; Ent. Zs. 22 (29) : 119
Ypthima conjuncta formosana ; Forster, [1948], Mitt. Münch. ent. Ges. 34 (2) : 481; [BOR, 471]
Ypthima formosana ; [MRS, 391]
Ypthima (Thymipa) formosana ; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80
Ypthima formosana ; Takahashi, 2000, Trans. lepid. Soc. Japan 51 (1) : 2

Larva on Brachiaria mutica, Imperata cylindrica, Cynodon dactylon, Miscanthus sinensis, M. floridulus, Setaria palmifolia, Isachne globosa [MRS]


Ypthima atra Cantlie & Norman, 1959
atra Cantlie & Norman, 1959; J. Bombay nat. Hist. Soc. 56 (1) : 69
Ypthima atra, Butterflies in Indo-China [Yutaka Inayoshi]
Ypthima atra ; [NHM]
Ypthima (Thymipa) atra ; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80

Ypthima persimilis Elwes & Edwards, 1893
Bhutan, Assam, Manipur. See [About maps] Ypthima persimilis Elwes & Edwards, 1893; Trans. Ent. Soc. Lond. 1893 (1) : 39
Ypthima persimilis, Butterflies in Indo-China [Yutaka Inayoshi]
Ypthima persimilis ; [BOR, 470]
Ypthima (Thymipa) persimilis ; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80

Ypthima evansi Eliot, 1967
Thailand, Burma. See [About maps] Ypthima evansi Eliot, 1967; Entomologist 100 (1246) : 56, TL: North Shan States
Ypthima evansi evansi, Butterflies in Indo-China [Yutaka Inayoshi]
Ypthima (Thymipa) evansi ; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80
Y. e. haka Eliot, 1967
Burma Ypthima evansi haka Eliot, 1967; Entomologist 100 (1246) : 58, TL: Burma North Chin Hills

Ypthima dohertyi (Moore, [1893])
Manipur - S.Burma. See [About maps] Great Five-Ring
Thymipa dohertyi Moore, [1893]; Lepidoptera Indica 2 (15): 65, pl. 108, f. 1, 1a, TL: Upper Tenasserim
Ypthima dohertyi dohertyi, Butterflies in Indo-China [Yutaka Inayoshi]
Ypthima dohertyi ; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1) : 38, pl. 2, f. 31; [BOR, 470]
Ypthima (Thymipa) dohertyi ; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80
Burma, Thailan, Laos
Y. d. khasia Eliot, 1967
Assam Ypthima dohertyi khasia Eliot, 1967; Entomologist 100 (1246) : 58, TL: Assam, Khasia
Y. d. mossmani Eliot, 1967
Ypthima dohertyi mossmani Eliot, 1967; Entomologist 100 (1246) : 59, pl. 2
Ypthima dohertyi mossmani, Butterflies in Indo-China [Yutaka Inayoshi]
Ypthima dohertyi mossmani ; [BMP: 136, pl. 17, f. 22]

Ypthima methora Hewitson, 1865
Sikkim - Assam, N.Burma, Bhutan, Nepal. See [About maps] Variegated Five-Ring
Yphthima[sic] methora Hewitson, 1865; Trans. ent. Soc. Lond. (3) 2 (4) : 291, pl. 18, f. 20-21, TL: N.India
gela Fruhstorfer, 1911²; in Seitz, Gross-Schmett. Erde 9 : 292
= ; [NHM]
Ypthima methora ; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1) : 37, pl. 2, f. 29; [BOW: pl. 173, f. 3]; [BOR, 470]; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80
Nepal, ... Ypthima methora methora ; Morishita & Inomata, 1998, Trans. lepid. Soc. Japan 49 (1): 3

Ypthima conjuncta Leech, 1891
W.China, C.China, Taiwan. See [About maps] Ypthima conjuncta Leech, 1891; Entomologist 24 (Suppl.) : 66, TL: Ta-chien-lu, Moupin; Chia-Kow-Ho, Omei-Shan; Wa-shan; huang-mu-chang; Chang-Yang; Ichang
Ypthima conjuncta luxurians Forster, [1948]; Mitt. Münch. ent. Ges. 34 (2) : 482, pl. 32-33, f. 1-2, TL: Fukien, Kuatun, 2300m
= ; [NHM]
Ypthima conjuncta, NSG Voucher Specimen [Wahlberg]
Ypthima conjuncta ; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1) : 39, pl. 2, f. 32; Forster, [1948], Mitt. Münch. ent. Ges. 34 (2) : 481; [BOW: pl. 205, f. 1]; [BOR, 471]; [MRS, 391]
Ypthima (Thymipa) conjuncta ; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80
C.China, W.China Ypthima conjuncta conjuncta ; [BOR, 471]
Y. c. monticola Uemura & Koiwaya, 2000
Yunnan Ypthima conjuncta monticola, Butterflies in Indo-China [Yutaka Inayoshi]
Ypthima conjuncta monticola ; Huang, 2003, Neue Ent. Nachr. 55: 100 (note)

Ypthima microphthalma Forster, [1948]
Ypthima methora microphthalma Forster, [1948]; Mitt. Münch. ent. Ges. 34 (2) : 484, pl. 32-33, f. 3, TL: N.Yunnan, Likiang, Yülingshan, ca. 3000m
Ypthima microphthalma ; Huang & Wu, 2003, Neue Ent. Nachr. 55: 122 (note)
Ypthima (Thymipa) microphthalma ; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80

The tappana species-group
tappana ; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80

Ypthima yayeyamana Nire, 1920
Japan (Iriomotejima, Ishigakijima). See [About maps] Ypthima (Thymipa) yayeyamana ; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80

Ypthima tappana Matsumura, 1909
Taiwan. See [About maps] Ypthima tappana Matsumura, 1909; Ent. Zs. 23 (19) : 92, TL: Formosa, Tappan, 4000ft
Ypthima tappana ; [BOR, 472 (note)]; [MRS, 392]; Uemura, 1984, Tyô to Ga 35 (3): 137
Ypthima (Thymipa) tappana ; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80
Y. t. continentalis Murayama, 1981
Yunnan, Sichuan Ypthima tappana continentalis Murayama, 1981; , TL: Sichuan, Qingchenshan
Ypthima tappana continentalis ; Huang, 2003, Neue Ent. Nachr. 55: 100 (note)

Ypthima cantlei Norman, 1958
Assam (Kangpokpi). See [About maps] cantlei Norman, 1958; J. Bombay nat. Hist. Soc. 55 (1): 180, TL: Assam
Ypthima cantliei[sic] ; [NHM]
Ypthima cantlei ; [BOR, 465]
Ypthima (Thymipa) cantlei ; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80

The philomela species-group
philomela ; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80

Ypthima philomela (Linnaeus, 1763)
India (Nilgiri-Wynaad), N.Burma, ?Peninsular Malaya, Java, Sumatra. See [About maps] Baby Fourring
Papilio philomela Linnaeus, 1763; Amoenitates Acad., 6: 404, TL: Java
= ?Ypthima baldus ; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1) : 14
Ypthima laroides Westwood, [1851]; Gen. diurn. Lep. (2): 395
= ; [NHM]
Ypthima tabella Marshall & de Nicéville, 1883; Butts India Burmah Ceylon 1 (2): 234
= Ypthima baldus ; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1) : 14
= ; [NHM]
peguana Evans, 1924²; J. Bombay nat. Hist. Soc. 29 (3) (3): 786, TL: N.Burma
= ; [NHM]
Ypthima philomela ; [BOW: pl. 173, f. 6]; [BOR, 465]
Ypthima (Thymipa) philomela ; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80

Ypthima baldus (Fabricius, 1775)
Himalayas (Chamba - Assam), Burma, India, Thailand, China, Korea, Japan, Ussuri, Kuriles. See [About maps] Common Five-Ring
Papilio baldus Fabricius, 1775; Syst. Ent.: 829, no. 202-3, TL: "India"
Ypthima argus Butler, 1866; J. Linn. Soc. Zool. Lond. 9 (34) : 56, TL: Japan, Hakodate
Ypthima prattii Elwes & Edwards, 1893; Trans. Ent. Soc. Lond. 1893 (1) : 35, pl. 3, f. 55, TL: Ichang
= Yptihma argus ; [NHM]
Ypthima baldus? ishigakina Fruhstorfer, 1908; Ent. Zs. 22 (31) : 127, TL: Ishigaki
= ; [NHM]
gallienus Fruhstorfer, 1911²; in Seitz, Gross-Schmett. Erde 9 : 288, TL: Hainan
= ; [NHM]
scota Fruhstorfer, 1911²; in Seitz, Gross-Schmett. Erde 9 : 289, TL: Hongkong
= ; [NHM]
morus Fruhstorfer, 1911²; in Seitz, Gross-Schmett. Erde 9 : 289, TL: Sumatra
= ; [NHM]
satpura Evans, 1924²; J. Bombay nat. Hist. Soc. 29 (3) (3): 787
= ; [NHM]
madrasa Evans, 1924²; J. Bombay nat. Hist. Soc. 29 (3) (3): 787, TL: S.India
= ; [NHM]
narensis Sugitani, 1932; Zephyrus 4 (?): 90, TL: Japan
= Yptihma argus ; [NHM]
Ypthima baldus kangeana Kalis, 1933; Tijdschr. Ent. 76 (1-2) : 55, TL: Kangean
= ; [NHM]
Ypthima baldus, NSG Voucher Specimen [Wahlberg]
Ypthima baldus baldus, Butterflies in Indo-China [Yutaka Inayoshi]
Yphthima[sic] baldus ; Hewitson, 1865, Trans. ent. Soc. Lond. (3) 2 (4) : 286
Ypthima argus ; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1) : 35, pl. 2, f. 28; Forster, [1948], Mitt. Münch. ent. Ges. 34 (2) : 476
Ypthima (Thymipa) argus ; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80
Ypthima baldus ; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1) : 14, pl. 1, f. 15-16; [BRU, 192]; [BOW: pl. 172, f. 15]; [BOR, 466]
Ypthima balda ; [MRS, 390]
Ypthima (Thymipa) baldus ; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80
[baldus-1.jpg] 664x564 (~92Kb) Korea, 1989, S.W. Cheong col. & det., Photo © Enrique Garcia-Barros [baldus-2v.jpg] 1000x699 (~120Kb) underside An edge of a shady bamboo forest at Dragonfly Biotope Ponds, Hiraoka, Ikazaki-cho, Ehime-ken, Shikoku-Chiho, Japan. 7th August 2002 Photo © Oleg Kosterin [baldus-3u.jpg] 740x572 (~69Kb) upperside Japan: Tokyo, 1.9.2007, Photo © Peter Payzant
[baldus-3v.jpg] 760x893 (~82Kb) underside Japan: Tokyo, 1.9.2007, Photo © Peter Payzant [baldus-4u.jpg] 512x341 (~33Kb) upperside India: Guwahati, 17.11.2005, Photo © Kedar Tokekar [baldus-4v.jpg] 512x341 (~41Kb) underside India: Guwahati, 17.11.2005, Photo © Kedar Tokekar

Larva on Poa annua, Echinochloa crusgalli, Digitaria, Elytrigia, (in Japan) [BRU, Fukuda et al., 1983] Microstegium ciliatum, Pogonatherum crinitinum [MRS]


India - Indo China, Burma (Tenasserim), Thailand, S.Yunnan Ypthima baldus baldus ; [BOR, 466]
Y. b. newboldi Distant, 1882
Peninsular Malaya, Langkawi Is., Singapore Ypthima newboldi Distant, 1882; Ann. Mag. nat. Hist. (5) 9 (53) : 396, TL: Prov. Wellesley, Perak, Malaysia
= Ypthima marshalli ; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1) : 26
Ypthima baldus newboldi, Butterflies in Indo-China [Yutaka Inayoshi]
Ypthima baldus newboldi ; [BMP: 135, pl. 17, f. 25-27]; [BOR, 466]
Y. b. luoi Huang, 1999
Yunnan Ypthima baldus luoi ; Huang, 2003, Neue Ent. Nachr. 55: 98 (note)
Y. b. moerus Fruhstorfer²
Sumatra Ypthima baldus moerus ; [BOR, 466]
Y. b. selinuntius Fruhstorfer, 1911²
Borneo, Natuna Is. baldus selinuntius Fruhstorfer, 1911²; in Seitz, Gross-Schmett. Erde 9 : 289, pl. 99 d, TL: Natuna Is.
Ypthima baldus selinuntius ; [BOR, 466]
Y. b. pasitelides Fruhstorfer, 1911²
Bawean baldus pasitelides Fruhstorfer, 1911²; in Seitz, Gross-Schmett. Erde 9 : 289, TL: Bawean
Ypthima baldus pasitelides ; [BOR, 466]
Y. b. zodina Fruhstorfer, 1911
Taiwan baldus zodina Fruhstorfer, 1911; in Seitz, Gross-Schmett. Erde 9 : 289
Ypthima baldus zodina ; [BOR, 466]
Ypthima balda zodina ; [MRS, 390]

Larva on Miscanthus sinensis, M. floridulum, Isachne globosa, Brachiaria mutica, Oplismenus undulatifolius, Microstegium ciliatum, Pogonatherum crinitum [MRS]

Y. b. marshalli Butler, 1882
Ypthima marshalli Butler, 1882; Ann. Mag. Nat. Hist. (5) 10 : 373, TL: Thaungyin Valley, Meplay
?Ypthima evanescens Butler, 1881; Ann. Mag. Nat. Hist. (5) 7 (38) : 134
= Ypthima marshalli ; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1) : 26
= Ypthima argus ; [NHM]
Ypthima marshalli ; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1) : 25, pl. 1, f. 1, 1a, 11
Ypthima baldus marshalli ; [MRS, 390]
Y. b. hyampeia Fruhstorfer, 1911²
S.Ussuri, Korea baldus hyampeia Fruhstorfer, 1911²; in Seitz, Gross-Schmett. Erde 9 : 290, pl. 99 e, TL: Amur
= Ypthima argus ; [NHM]
?hampeia Seitz, 1930; Gross-Schmett. Erde (Suppl. 1): 131
= Ypthima argus ; [NHM]
Ypthima argus hyampeia ; Forster, [1948], Mitt. Münch. ent. Ges. 34 (2) : 476, pl. 30-31, f. 7-8
Y. b. jezoensis Matsumura, 1919
Kuriles Ypthima argus var. jezoensis Matsumura, 1919; Thous. Ins. Japan. Addit. 3 : 728, TL: Hokkaido
= Ypthima argus argus ; Uemura, 1984, Tyô to Ga 35 (3): 136
= Ypthima argus ; [NHM]
?Ypthima argus jezoensis ; Matsumura, 1928, Insecta Matsumurana 2 (4) : 191
Y. b. okurai Okano, 1962
Formosa Ypthima argus okurai Okano, 1962; Tyô to Ga 12 (3): 52, TL: Sankakuhò (ca. 2370m), Central Formosa

Ypthima zodia Butler, 1871
Hainan. See [About maps] Yphthima zodia Butler, 1871; Trans. ent. Soc. Lond. 1871 (3) : 402
zodiaca Moore, 1878²; Proc. zool. Soc. Lond. 1878 (3) : 696
= Ypthima zodiaca ; [NHM]
Ypthima albescens Poujade, 1885; Bull. Soc. Ent. Fr. (6) 5 : xli
= ; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1) : 28
Ypthima zodia tapaishani Forster, [1948]; Mitt. Münch. ent. Ges. 34 (2) : 474, pl. 30-31, f. 2-3, TL: Tapaishan
= Ypthima zodiaca ; [NHM]
Ypthima zodia septentrionalis Forster, [1948]; Mitt. Münch. ent. Ges. 34 (2) : 474, pl. 30-31, f. 4, TL: Mienshan
= Ypthima zodiaca ; [NHM]
Ypthima melli Forster, [1948]; Mitt. Münch. ent. Ges. 34 (2) : 475, pl. 30-31, f. 10-11, TL: N.Yunna, Likiang
= Ypthima zodiaca ; [NHM]
pseudodromon Forster, [1948]; Mitt. Münch. ent. Ges. 34 (2) : 488
= Ypthima zodiaca ; [NHM]
Ypthima zodia albescens ; Forster, [1948], Mitt. Münch. ent. Ges. 34 (2) : 473
?Ypthima (Thymipa) melli ; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80
Ypthima zodia ; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1) : 27, pl. 1, f. 12-13; Forster, [1948], Mitt. Münch. ent. Ges. 34 (2) : 473, pl. 30, 31, f. 1; [MRS, 391]
Ypthima (Thymipa) zodia ; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80

Ypthima horsfieldii Moore, 1884
-. See [About maps] Ypthima horsfieldii Moore, 1884; J. asiat. Soc. Bengal 53 Pt.II (1) : 18, TL: Java
Ypthima horsfieldii ; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1) : 28
Ypthima (Thymipa) horsfieldi ; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80
Java ?Ypthima baldus horsfieldi ; [BOR, 466]
Y. h. humei Elwes & Edwards, 1893
Ypthima humei Elwes & Edwards, 1893; Trans. Ent. Soc. Lond. 1893 (1) : 32, pl. 3, f. 46
Ypthima horsfieldii humei ; [BMP: 136, pl. 17, f. 28-29]

Ypthima cerealis Watson, 1897
-. See [About maps] Ypthima cerealis Watson, 1897; J. Bombay nat. Hist. Soc. 10 (4) : 646, pl. A, f. 3-4, TL: Myingyan, Upper Burma
Ypthima (Thymipa) cerealis ; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80
Burma, Thailand, Laos
Y. c. singorensis Aoki & Uémura, 1984
Thailand Ypthima cerealis singorensis Aoki & Uémura, 1984; Tokurana 6/7: 79, f. 2j, 6, 8B, 26, pl. 5, f. 3, 7, TL: Thailand

Ypthima nebulosa Aoki & Uémura, 1982
Sumatra, Thailand, Laos. See [About maps] Ypthima nebulosa Aoki & Uémura, 1982; Memoirs of the Tsukada Coll. No. 4: 13, pl. 5, f. 22-23, pl. 6, f. 22-23, TL: Sumatra
Ypthima nebulosa, Butterflies in Indo-China [Yutaka Inayoshi]
Ypthima (Thymipa) nebulosa ; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80

Ypthima yunosukei Aoki & Uémura, 1984
Thailand. See [About maps] Ypthima yunosukei Aoki & Uémura, 1984; Tokurana (6/7): 81-82, pl. 5, f. 1-2, 5-6, TL: Phukadung, Thailand
Ypthima yunosukei, Butterflies in Indo-China [Yutaka Inayoshi]
Ypthima (Thymipa) yunosukei ; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80

Ypthima fasciata Hewitson, 1865
Malaysia, Sumatra. See [About maps] Yphthima[sic] fasciata Hewitson, 1865; Trans. ent. Soc. Lond. (3) 2 (4) : 287, TL: Sarawak, Sumatra
Ypthima fasciata, NSG Voucher Specimen [Wahlberg]
Ypthima fasciata ; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1) : 44; [BOW: pl. 173, f. 1]; [BOR, 468]
Ypthima (Thymipa) fasciata ; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80
Borneo, Natuna Is. Ypthima fasciata fasciata ; [BOR, 468]
Y. f. torone Fruhstorfer, 1911
Peninsular Malaya, Sumatra Ypthima fasciata torone Fruhstorfer, 1911; in Seitz, Gross-Schmett. Erde 9 : 288, (75) , TL: Padang, Sumatra
Ypthima fasciata torone, Butterflies in Indo-China [Yutaka Inayoshi]
Ypthima fasciata torone ; [BOR, 468]; [BMP: 136, pl. 17, f. 23-24]

Ypthima indecora Moore, 1882
Kashmir - Kumaon. See [About maps] Western Five-Ring
Ypthima indecora Moore, 1882; Proc. zool. Soc. Lond. 1882 (1) : 238, pl. 12, f. 7
Ypthima indecora ; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1) : 13, pl. 1, f. 10, 14; [BOR, 465]
Ypthima (Thymipa) indecora ; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80

Ypthima nynias Fruhstorfer, 1911²
Sulawesi. See [About maps] nigricans nynias Fruhstorfer, 1911²; in Seitz, Gross-Schmett. Erde 9 : 290
Ypthima baldus nynias ; [BOR, 466]
Ypthima (Thymipa) nynias ; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 183
Sulawesi, Muna, Buton Ypthima (Thymipa) nynias nynias ; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 183
Y. n. aretas Fruhstorfer, 1911²
Salayar nynias aretas Fruhstorfer, 1911²; in Seitz, Gross-Schmett. Erde 9 : 290, TL: Saleyer
Ypthima baldus aretas ; [BOR, 466]
Ypthima (Thymipa) nynias aretas ; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 183
Y. n. gadames Fruhstorfer, 1911²
Banggai nynias gadames Fruhstorfer, 1911²; in Seitz, Gross-Schmett. Erde 9 : 290, TL: Bangkai
Ypthima gadames ; [BOR, 472 (note)]
Ypthima (Thymipa) nynias gadames ; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 183

Ypthima stellera (Eschscholtz, 1821)
Philippines. See [About maps] Hipparchia stellera Eschscholtz, 1821; in Kotzebue, Entdeck. Reise Süd-See 3 (App. 5): 216, pl. 10, f. 24
galeria Fruhstorfer, 1911²; in Seitz, Gross-Schmett. Erde 9 : 290, TL: Palawan
= ; [NHM]
Yphthima[sic] stellera ; Hewitson, 1865, Trans. ent. Soc. Lond. (3) 2 (4) : 285
Ypthima stellera ; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1) : 26, pl. 3, f. 45; [BOR, 472 (note)]
?Ypthima (Thymipa) sterella[sic] ; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80
Y. s. sepyra Hewitson, 1865
Bachan, Ternate, Halmahera Yphthima[sic] sepyra Hewitson, 1865; Trans. ent. Soc. Lond. (3) 2 (4) : 292, pl. 17, f. 10, TL: Gilolo, Batchian
= Ypthima sepyra ; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1) : 27
Ypthima stellera sepyra ; [BAUR, 265 (text)]

Ypthima aphnius (Godart, [1824])
Celebes, Timor. See [About maps] Satyrus aphnius Godart, [1824]; Encyclopédie Méthodique. 9 (2): 551, TL: Timor
Yphthima leuce Doherty, 1891; J. Asiat. Soc. Bengal 60 Pt.II (2) : 169, TL: Sumba; Sumbawa
= ; [NHM]
caratonus Fruhstorfer, 1911²; in Seitz, Gross-Schmett. Erde 9 : 289, TL: Bali
= ; [NHM]
saravus Fruhstorfer, 1911²; in Seitz, Gross-Schmett. Erde 9 : 290, TL: Lombok
= ; [NHM]
glabrius Fruhstorfer, 1911²; in Seitz, Gross-Schmett. Erde 9 : 290, TL: Flores
= ; [NHM]
budinus Fruhstorfer, 1911²; in Seitz, Gross-Schmett. Erde 9 : 290, TL: Alor
= ; [NHM]
Yphthima[sic] aphnius ; Hewitson, 1865, Trans. ent. Soc. Lond. (3) 2 (4) : 292, pl. 17, f. 8-9
Ypthima leuce ; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1) : 28, pl. 1, f. 21-22
Ypthima aphnius ; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1) : 15; [BOW: pl. 172, f. 13]
Ypthima (Thymipa) aphnius ; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80
Timor, Wetar Ypthima aphnius aphnius ; [BAUR, 265 (text)]

Ypthima lisandra (Cramer, [1780])
Hong Kong, Tonking, Peninsular Malaya - Indo China, S.China, Taiwan. See [About maps] Papilio lisandra Cramer, [1780]; Uitl. Kapellen 4 (25-26a): 11, pl. 293, f. G,H, TL: S.China
Yphthima[sic] micrommatus Holland, 1887; Trans. Amer. Ent. Soc. 14 : 115, pl. 2, f. 3
= ; [NHM]
Ypthima lisandra, NSG Voucher Specimen [Wahlberg]
Ypthima lisandra lisandra, Butterflies in Indo-China [Yutaka Inayoshi]
Ypthima micrommatus ; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1) : 49
Ypthima lisandra ; [BOW: pl. 205, f. 5]; [BOR, 472 (note)]; [MRS, 392]
Ypthima (Thymipa) lisandra ; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80
N.Vietnam, S.China
Y. l. bara Evans, 1923
Sikkim - Burma, Thailand, Laos Ypthima avanta bara Evans, 1923; J. Bombay nat. Hist. Soc. 29 (3) (3): 786, 797, No. D. 14/12b
Ypthima lisandra bara, Butterflies in Indo-China [Yutaka Inayoshi]

Ypthima singala Felder, 1868
Ceylon. See [About maps] Ypthima singala Felder, 1868; Verh. zool.-bot. Ges. Wien 18 (1-2) : 283, TL: Bholahat, Ceylon, Kumaon
Ypthima singala ; Moore, [1880], Lepid. Ceylon 1 (1): 24, pl. 12, f. 3, 3a; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1) : 31, pl. 1, f. 26, 26a; [BOR, 466]
Ypthima (Thymipa) singala ; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80

Ypthima avanta Moore, [1875]
Ceylon, Peninsular India, Himalayas - Kashmir, Assam. See [About maps] Jewel Fourring
Yphthima[sic] avanta Moore, [1875]; Proc. zool. Soc. Lond. 1874 (4) : 567, TL: Bagh Valley, Cashmere
Ypthima thora Moore, [1880]; Lepid. Ceylon 1 (1): 24, pl. 12, f. 4, 4a, TL: Ceylon
= ?Ypthima singala ; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1) : 32
= ; [NHM]
Ypthima ordinata Butler, 1880; Proc. zool. Soc. Lond. 1880 : 148, pl. 15, f. 3
= ; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1) : 33
Ypthima avanta ; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1) : 33, pl. 1, f. 27; Forster, [1948], Mitt. Münch. ent. Ges. 34 (2) : 472; [BOW: pl. 172, f. 10]; [BOR, 466]
Ypthima (Thymipa) avanta ; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80

Subgenus Ypthima Hübner, 1818
Ypthima Hübner, 1818; Zuträge Samml. exot. Schmett. 1 : 17 [missid. TS Papilio philomela Linnaeus], TS: Ypthima huebneri Kirby
Ypthima, NSG Voucher Specimen [Wahlberg]
Ypthima ; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80

The arctous species-group
arctous ; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80
arctoa ; Shirôzu & Shima, 1979, (incl.)
serasa ; Shirôzu & Shima, 1979, (incl.)

Ypthima arctous (Fabricius, 1775)
Australia. See [About maps] Dingy Ring
Papilio arctous Fabricius, 1775; Syst. Ent.: 489
Yphthima[sic] arctoides Hewitson, 1865; Trans. ent. Soc. Lond. (3) 2 (4) : 284
Ypthima arctoa, NSG Voucher Specimen [Wahlberg]
Ypthima arctous, Lepidoptera Larvae of Australia [Don Herbison-Evans]
Yphthima[sic] arctous ; Hewitson, 1865, Trans. ent. Soc. Lond. (3) 2 (4) : 283
Ypthima arctoa ;
Ypthima arctous ; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1) : 14, pl. 3, f. 49
Ypthima (Ypthima) arctous ; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80
Misidentified/misapplied:
Papilio acrctous; Donovan, 1805, Epitome nat. Hist. Ins. New Holland: pl. 24

Larva on Digitaria didactyla, Imperata cylindrica, I. sp., Themeda triandra, (Unidentified grasses) K. L. & L. E. Dunn, 1991, Review Austr. Butts. (1-4): (120-140)

Northern Territory, Torres Straits Is., Cape York - S.New South Wales Xois arctous arctous
Xois arctous arctous ; [BAUR, 265]
Y. a. subarctua Tryon, 1890
Papua, New Guinea papuana Fruhstorfer, 1911; in Seitz, Gross-Schmett. Erde 9 : 286
Xois arctous papuana ; [BAUR, 265]

Ypthima sesara (Hewitson, 1865)
Fiji Xois sesara Hewitson, 1865; Trans. ent. Soc. Lond. (3) 2 (4) : 282, pl. 17, f. 3-4, TL: Fiji
Acrophthalmia? diophthalma Prittwitz, 1867; Stettin ent. Ztg 28 (7-9): 274
= ; [NHM]
Xois sesara ; [BOW: pl. 172, f. 4]; [BAUR, 265]
Ypthima (Ypthima) sesara ; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80

Ypthima fulvida (Butler, 1883)
Fiji Xois fulvida Butler, 1883; Ann. Mag. nat. Hist. (5) 11 (66) : 411, TL: Viti-Levu
vitiensis Fruhstorfer, 1911²; in Seitz, Gross-Schmett. Erde 9 : 294
= ; [NHM]
Xois fulvida ; [BAUR, 265]
Ypthima (Ypthima) fulvida ; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80

The nareda species-group
nareda ; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80

Ypthima nareda (Kollar, [1844])
Pakistan, NW.India, Nepal, Bhutan, Sikkim. See [About maps] Large Three-Ring
Satyrus nareda Kollar, [1844]; in Hügel, Kaschmir und das Reich der Siek, 4: 451
Ypthima nareda, NSG Voucher Specimen [Wahlberg]
Ypthima nareda ; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1) : 20, pl. 1, f. 2, pl. 2, f. 40; [BOW: pl. 173, f. 4]; [BOR, 470]; [MRS, 394]
Ypthima (Ypthima) nareda ; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80
Ypthima nareda ; Huang & Wu, 2003, Neue Ent. Nachr. 55: 122 (note)
Misidentified/misapplied:
Ypthima newara; Lee, 1982, in Huang, Insects of Xizang, 2: 135, No. 23 (65) part (missid.)

Larva on Gramineae [MRS]

W.Himalayas Ypthima nareda nareda ; [BOR, 470]
Y. n. sarcaposa Fruhstorfer, 1911²
Burma, Thailand, Indo China nareda sarcaposa Fruhstorfer, 1911²; in Seitz, Gross-Schmett. Erde 9 : 291, TL: Burma
Ypthima newara sarcaposa ; Forster, [1948], Mitt. Münch. ent. Ges. 34 (2) : 477, pl. 32-33, f. 4
Ypthima nareda sarcaposa ; [BOR, 470]; [MRS, 394]
Y. n. fusca Elwes & Edwards²
Assam

= Ypthima fusca Elwes & Edwards, 1893?


The asterope species-group
asterope ; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80

Ypthima asterope (Klug, 1832)
Lebanon, Syria, Himalayas, Kashmir, Baluchistan, India, Tropical Africa. See [About maps] Common Three-Ring
Hipparchia asterope Klug, 1832; in Ehrenberg, Symbolae phys., Ins. 3: pl. 29, f. 11-14
Ypthima alemola Swinhoe, 1885; Proc. Zool. Soc. Lond. 1885 : 127
= ; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1) : 12
Ypthima complexiva Swinhoe, 1886; Proc. zool. Soc. Lond. 1886 (4) : 423, pl. 40, f. 2
= ; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1) : 12
hereroica Grünberg, 1910²; Denkschr. med.-naturw. Ges. Jena 16 (4): 101
= ; [NHM]
merlenii Kattulas & Koutsaftikis, 1976; Biologia gallo-hellen 7 (1-2): 156, TL: Greece
= ; [NHM]
Ypthima asterope, Moths and Butterflies of Europe [MBE]
Yphthima[sic] asterope ; Hewitson, 1865, Trans. ent. Soc. Lond. (3) 2 (4) : 283
Ypthima asterope ; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1) : 11, pl. 1, f. 4, pl. 3, f. 44b; Kielland, 1982, Tijdschr. Ent. 125 (5) : 104; [BOW: pl. 204, f. 18]; [BOR, 464]; [AfrL]; [Otakar Kudrna]
Ypthima (Ypthima) asterope ; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80
[asterope-1av.jpg] 902x657 (~199Kb) underside Ethiopia, Oromia, Debre Libanos env., at German Ethio Lodge, the Gur River right bank. 30th July 2012, Photo © Oleg Kosterin [asterope-1bv.jpg] 880x615 (~228Kb) underside Ethiopia, Oromia, Debre Libanos env., at German Ethio Lodge, the Gur River right bank. 30th July 2012, Photo © Oleg Kosterin [asterope-1xu.jpg] 1100x1100 (~375Kb) upperside Ethiopia, Oromia, Debre Libanos env., at German Ethio Lodge, the Gur River right bank. 30th July 2012, Photo © Oleg Kosterin

Larva on Cynodon [BK]


India, Arabia, N.Tropical Africa ?Ypthima asterope ab. binucholata Strand, 1909; Ent. Zs. Stutt. 24: 112, TL: Sadani, Tanzania
= ?Ypthima granulosa ; Kielland, 1982, Tijdschr. Ent. 125 (5) : 104
?Ypthima asterope ab. biocelligera Strand, 1909; Ent. Zs. Stutt. 24: 112, TL: Sadani, Tanzania
= ?Ypthima granulosa ; Kielland, 1982, Tijdschr. Ent. 125 (5) : 104
Ypthima asterope asterope ; Kielland, 1982, Tijdschr. Ent. 125 (5) : 104, pl. 1, f. 1-8; [BK: 274, pl. 30, f. 428]; [BAFR, 192]; [AfrL]
Y. a. mahratta Moore, 1884
NW.India - Assam Ypthima mahratta Moore, 1884; J. asiat. Soc. Bengal 53 Pt.II (1) : 16, TL: Deccan
= Ypthima asterope ; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1) : 12
Ypthima asterope mahratta ; [BOR, 464]
Y. a. hereroica van Son, 1955
SW.Africa, Nambibia Ypthima asterope hereroica van Son, 1955; Transv. Mus. Mem. 8: 156, f. 165
Yphtima asterope ab. hereroica Grünberg, 1910; Denkschr. med.-naturw. Ges. Jena 16 (4): 101, TL: Damaraland
Ypthima asterope hereroica ; Kielland, 1982, Tijdschr. Ent. 125 (5) : 107, pl. 2, f. 1-2; [BAFR, 192 (text)]; [AfrL]

Ypthima norma (Westwood, 1851)
-. See [About maps] Iphthima norma Westwood, 1851; Gen. diurn. Lep. (2): pl. 67, f. 1
Ypthima norma ; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1) : 51
Ypthima (Ypthima) norma ; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 183

Larva on Bambusa, Cynodon, Digitaria, Ehrharta, Imperata, Microstegium, Oplismenus, Paspalum, Pogonatherum Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 183

Y. n. burmana Evans, 1923
Burma, Thailand, Laos Ypthima asterope burmana Evans, 1923; J. Bombay nat. Hist. Soc. 29 (3) (3): 785, TL: N.Burma
Ypthima norma burmana, Butterflies in Indo-China [Yutaka Inayoshi]
Ypthima asterope burmana ; [BOR, 464]
Y. n. annamitica Fruhstorfer, 1911
S.Vietnam Ypthima asterope annamitica Fruhstorfer, 1911; in Seitz, Gross-Schmett. Erde 9 : 286, TL: Annam
Ypthima norma annamitica, Butterflies in Indo-China [Yutaka Inayoshi]
Ypthima asterope annamitica ; [BOR, 464]
Y. n. pusilla Fruhstorfer, 1911²
Sulawesi, Buton, Buru, Ambon norma pusilla Fruhstorfer, 1911²; in Seitz, Gross-Schmett. Erde 9 : 287
Ypthima pusilla ; [BAUR, 265 (text)]
Ypthima (Ypthima) norma pusilla ; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 184

Ypthima cataractae van Son, 1955
Victoria Falls, Zambesi Ypthima cataractae van Son, 1955; Transv. Mus. Mem. 8: 154 [wet-season]
Ypthima cataractae f. irvingi van Son, 1955; Transv. Mus. Mem. 8: 154 [dry season]

Ypthima yatta Kielland, 1982
Ethiopia, Kenya. See [About maps] Yatta Three-Ring
Ypthima yatta Kielland, 1982; Tijdschr. Ent. 125 (5) : 108, pl. 2, f. 3-6, TL: Kenya, Thika, Yatta
Ypthima yatta ; [BK: 275]; [AfrL]

Ypthima congoana Overlaet, 1955
Zaire (Katanga). See [About maps] Ypthima asterope congoana Overlaet, 1955; Miss. de Witte 27: 85, f. 12a-b, TL: Upemba, Katanga
Ypthima congoana ; Kielland, 1982, Tijdschr. Ent. 125 (5) : 109, pl. 8, f. 1-2, 4-5; [AfrL]

Ypthima rhodesiana Carcasson, 1961
Zambia, Zimbabwe, S.Zaire, Tanzania, SE.Kenya. See [About maps] Ypthima rhodesiana Carcasson, 1961; Occ. Pap. Coryndon meml Mus. (7): 14, f. 11, TL: Choma, Zambia
Ypthima rhodesiana ; Kielland, 1982, Tijdschr. Ent. 125 (5) : 111, pl. 4, f. 1-6; [BK: 275, pl. 30, f. 430]; [AfrL]

Ypthima simplicia Butler, 1876
Ethiopia, S.Sudan, Kenya, N.Tanzania. See [About maps] Ypthima simplicia Butler, 1876; Ann. Mag. nat. Hist. (4) 18 (108) : 481, TL: Ethiopia
Ypthima simplicia ; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1) : 13, pl. 3, f. 54; Kielland, 1982, Tijdschr. Ent. 125 (5) : 113, pl. 2, f. 7-8, pl. 3, f. 1-6; [BK: 275]; [AfrL]

Ypthima condamini Kielland, 1982
-. See [About maps] Ypthima condamini Kielland, 1982; Tijdschr. Ent. 125 (5) : 115, TL: Tanzania, MPanda, Sibweza
Ypthima condamini condamini ; [BK: 276]
Ypthima condamini ; [AfrL]
Ethiopia, S.Sudan, E.Africa, Zambia, Mozambique, Transvaal, W.Zaire, N.Angola Ypthima condamini condamini ; Kielland, 1982, Tijdschr. Ent. 125 (5) : 115, pl. 4, f. 7-14; [AfrL]
Y. c. nigeriae Kielland, 1982
Nigeria - Ghana, Upper Volta, Senegal Ypthima condamini nigeriae Kielland, 1982; Tijdschr. Ent. 125 (5) : 116, pl. 4, f. 15-18, TL: Nigeria, Farniso near Kano
Ypthima condamini nigeriae ; [AfrL]

Ypthima antennata van Son, 1955
E.Africa, S.Africa, W.Kenya, S.Sudan, Nigeria, Ghana. See [About maps] Ypthima antennata van Son, 1955; Transv. Mus. Mem. 8: 156, pl. 5-6, f. 86-87, TL: Munnik, N. Transvaal
Ypthima antennata ; Kielland, 1982, Tijdschr. Ent. 125 (5) : 118; [AfrL]
E.Africa - S.Africa, Zambia, Uganda, S.Sudan Ypthima antennata antennata ; Kielland, 1982, Tijdschr. Ent. 125 (5) : 118, pl. 5, f. 1-4; [BK: 276, pl. 30, f. 433]; [AfrL]
Y. a. cornesi Kielland, 1982
Nigeria, Ghana Ypthima antennata cornesi Kielland, 1982; Tijdschr. Ent. 125 (5) : 120, TL: N. Nigeria, Benue Plateau, Jos
Ypthima antennata cornesi ; [AfrL]

Ypthima jacksoni Kielland, 1982
Kenya (coast) - Ethiopia. See [About maps] Ypthima jacksoni Kielland, 1982; Tijdschr. Ent. 125 (5) : 120, pl. 3, f. 5-8, TL: Kenya, Mombasa, Mtwapa Creek
Ypthima jacksoni ; [BK: 276]; [AfrL]

Ypthima vuattouxi Kielland, 1982
Ivory Coast, Ghana, Nigeria, Cameroun. See [About maps] Ypthima vuattouxi Kielland, 1982; Tijdschr. Ent. 125 (5) : 121, pl. 5, f. 5-8, TL: Ivory Coast, Lamto (Toumodi)
Ypthima vuattouxi ; [AfrL]

Ypthima lamto Kielland, 1982
Ivory Coast, Cameroun. See [About maps] Ypthima lamto Kielland, 1982; Tijdschr. Ent. 125 (5) : 123, pl. 5, f. 9-14, TL: Ivory Coast, Lamto (Toumodi)
Ypthima lamto ; [AfrL]

Ypthima recta Overlaet, 1955
W.Africa - W.Kenya, Zaire, Angola. See [About maps] Ypthima recta Overlaet, 1955; Miss. de Witte 27: 87, f. 25a-b, TL: Upemba, Zaire
Ypthima recta ; Kielland, 1982, Tijdschr. Ent. 125 (5) : 124, pl. 8, f. 3, 6; [BK: 276]; [AfrL]

Ypthima granulosa Butler, 1883
Moçambique, Malawi, Zambia, Rhodesia, N.Botswana, E.Africa. See [About maps] Ypthima granulosa Butler, 1883; Ann. Mag. nat. Hist. (5) 12 (68) : 101, TL: Victoria Nyanza
Ypthima cataracta van Son, 1955; Transv. Mus. Mem. 8: 154, pl. 5-6, f. 81
= ; Kielland, 1982, Tijdschr. Ent. 125 (5) : 126
Ypthima asterope ab. inocellata Strand, 1909; Ent. Zs. Stutt. 24: 112, TL: Uluna-Langenburg
= ; Kielland, 1982, Tijdschr. Ent. 125 (5) : 126
Ypthima asterope ab. triocellata Strand, 1909; Ent. Zs. Stutt. 24: 112, TL: Langenburg, N.Nyass)
= ; Kielland, 1982, Tijdschr. Ent. 125 (5) : 126
Ypthima asterope ab. biocellata Strand, 1909; Ent. Zs. Stutt. 24: 112, TL: Dar es Salaam, Tanzania
= ; Kielland, 1982, Tijdschr. Ent. 125 (5) : 126
Ypthima asterope ab. interrupta Strand, 1909; Ent. Zs. Stutt. 24: 113, TL: Dar es Salaam
= ; Kielland, 1982, Tijdschr. Ent. 125 (5) : 126
Ypthima asterope ab. uniocellata Strand, 1909; Ent. Zs. Stutt. 24: 113, TL: Dar es Salaam
= ; Kielland, 1982, Tijdschr. Ent. 125 (5) : 126
Ypthima granulosa ; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1) : 12, pl. 3, f. 44, 44a; Kielland, 1982, Tijdschr. Ent. 125 (5) : 126, pl. 6, f. 1-8; [BAFR, 192]; [BK: 277, pl. 30, f. 336]; [AfrL]

Larva on Poaceaea [BK]


Ypthima pupillaris Butler, 1888
-. See [About maps] Ypthima pupillaris Butler, 1888; Proc. Zool. Soc. Lond. 1888 : 59, TL: River Dangu, Ganyese Tambu, Eq. Afr.
Ypthima pupillaris ; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1) : 20; Kielland, 1982, Tijdschr. Ent. 125 (5) : 133; [AfrL]
Ivory Coast - Nigeria, W.Zaire, Rhodesia Ypthima gazana van Son, 1955; Transv. Mus. Mem. 8: 151, pl. 5, f. 79-80
= ; Kielland, 1982, Tijdschr. Ent. 125 (5) : 133
Ypthima pupillaris ab. depupillata Strand, 1909; Ent. Zs. Stutt. 24: 113, TL: Loma
= ; Kielland, 1982, Tijdschr. Ent. 125 (5) : 133
Ypthima pupillaris ab. macrocellata Strand, 1913; Archiv Naturg. 79 A (7) : 150, TL: Cameroun
= ; Kielland, 1982, Tijdschr. Ent. 125 (5) : 133
Ypthima pupillaris pupillaris ; Kielland, 1982, Tijdschr. Ent. 125 (5) : 133, pl. 6, f. 9-12; [AfrL]
Y. p. obscurata Kielland, 1982
E.Africa Ypthima pupillaris obscurata Kielland, 1982; Tijdschr. Ent. 125 (5) : 134, pl. 6, f. 13-18, TL: Tanzania, MPanda, Ngondo, 1500m
Ypthima pupillaris obscurata ; [BK: 277]; [AfrL]

Ypthima impura Elwes & Edwards, 1893
-. See [About maps] Impure Ringlet
Ypthima impura Elwes & Edwards, 1893; Trans. Ent. Soc. Lond. 1893 (1) : 23, pl. 3, f. 48, TL: Angola
Ypthima impura ; [AfrL]
Ivory Coast - Nigeria, Cameroun, Zaire, Angola Ypthima impura impura ; Kielland, 1982, Tijdschr. Ent. 125 (5) : 135, pl. 7, f. 1-2; [BAFR, 192 (text)]; [AfrL]
Y. i. paupera Ungemach, 1932
Ethiopia, E.Africa - Transvaal, Natal Ypthima pupillaris paupera Ungemach, 1932; Mém. Soc. Sci. nat. phys. Maroc 32 : 51, TL: Ethiopia
Ypthima pupillaris paupera f. hiemis Ungemach, 1932; Mém. Soc. Sci. nat. phys. Maroc 32 : 51, TL: Ethiopia
= ; Kielland, 1982, Tijdschr. Ent. 125 (5) : 137
Ypthima impura f. badhami van Son, 1955; Transv. Mus. Mem. 8: 150, pl. 56, f. 77, 79, TL: Rhodesia
= ; Kielland, 1982, Tijdschr. Ent. 125 (5) : 137
Ypthima impura paupera ; Kielland, 1982, Tijdschr. Ent. 125 (5) : 137, pl. 7, f. 3-8; [BAFR, 192 (text)]; [BK: 277]; [AfrL]

Ypthima pulchra Overlaet, 1954
S.Zaire (Shaba), Tanzania - Uganda. See [About maps] Ypthima pulchra Overlaet, 1954; Lambillionea 54 : 41, f. 1a-b, TL: Katanga, Zaire
Ypthima pulchra ; Kielland, 1982, Tijdschr. Ent. 125 (5) : 138, pl. 8, f. 7-8, 10-11; [BAFR, 192 (text)]; [AfrL]

Ypthima praestans Overlaet, 1954
S.Zaire (Shaba), Angola, W.Zambia. See [About maps] Ypthima praestans Overlaet, 1954; Lambillionea 54 : 43, f. 2, TL: Sankuru
Ypthima praestans ; Kielland, 1982, Tijdschr. Ent. 125 (5) : 139, pl. 7, f. 9-12, pl. 8, f. 12; [BAFR, 192]; [AfrL]

Ypthima doleta Kirby, 1880
Sierra Leone - Ivory Coast, Ghana, Togo, Nigeria - Zaire, Angola, Uganda, NW.Tanzania. See [About maps] Ypthima doleta Kirby, 1880; Proc. R. Dublin Soc. (2) 2: 335, TL: Sierra Leone
Ypthima doleta ab. tripunctata Strand, 1909; Ent. Zs. Stutt. 24: 114, TL: Cameroun
= ; Kielland, 1982, Tijdschr. Ent. 125 (5) : 141
Ypthima doleta ab. quadripunctata Strand, 1909; Ent. Zs. Stutt. 24: 114, TL: Cameroun
= ; Kielland, 1982, Tijdschr. Ent. 125 (5) : 141
Ypthima doleta ab. quinquepunctata Strand, 1909; Ent. Zs. Stutt. 24: 114, TL: Cameroun
= ; Kielland, 1982, Tijdschr. Ent. 125 (5) : 141
Ypthima doleta ab. semptemocellata Strand, 1909; Ent. Zs. Stutt. 24: 114, TL: Cameroun
= ; Kielland, 1982, Tijdschr. Ent. 125 (5) : 141
Ypthima doleta ab. strigata Strand, 1913; Archiv Naturg. 79 A (7) : 150
= ; Kielland, 1982, Tijdschr. Ent. 125 (5) : 141
Ypthima doleta ab. defecta Strand, 1913; Archiv Naturg. 79 A (7) : 150, TL: Uelleburg
= ; Kielland, 1982, Tijdschr. Ent. 125 (5) : 141
?Ypthima doleta ab. asteropina Strand, 1913; Archiv Naturg. 79 A (7) : 150, TL: Alen
= ?; Kielland, 1982, Tijdschr. Ent. 125 (5) : 141
Ypthima doleta ab. dschangensis Strand, 1914; Arch. Naturgesch. 80 A (2) : 153, TL: Cameroun
= ; Kielland, 1982, Tijdschr. Ent. 125 (5) : 141
Ypthima doleta, NSG Voucher Specimen [Wahlberg]
Ypthima doleta ; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1) : 24, pl. 3, f. 51; Kielland, 1982, Tijdschr. Ent. 125 (5) : 141, pl. 7, f. 13-16; [BAFR, 192]; [BOW: pl. 118, f. 23]; [AfrL]

Ypthima diplommata Overlaet, 1954
Katanga, W.Zambia, Angola. See [About maps] Ypthima diplommata Overlaet, 1954; Lambillionea 54 : 44, f. 3, TL: Kafakumba, Katanga
Ypthima diplommata ; Kielland, 1982, Tijdschr. Ent. 125 (5) : 142, pl. 7, f. 17-18, pl. 8, f. 13-14; [AfrL]

Ypthima albida Butler, 1888
Nigeria, Cameroons. See [About maps] Silver Ringlet
Ypthima albida Butler, 1888; Proc. Zool. Soc. Lond. 1888 : 59, TL: Fóda, Equatorial Africa
Ypthima albida ; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1) : 48; Kielland, 1982, Tijdschr. Ent. 125 (5) : 143; [BOW: pl. 118, f. 6]; [BK: 277, pl. 30, f. 439]; [AfrL]
W.Kenya, Uganda, NW.Tanzania, E.Zaire, Rwanda, Burundi Ypthima albida argentata Bartel, 1905; Novit. Zool. 12 : 134, TL: Niarugungu, Ruanda
= ; Kielland, 1982, Tijdschr. Ent. 125 (5) : 143
Ypthima albida ab. conradsi Strand, 1909; Ent. Zs. Stutt. 24: 113, TL: Ukerewe I., L. Victoria
= ; Kielland, 1982, Tijdschr. Ent. 125 (5) : 143
Ypthima albida argentata ; [BAFR, 193 (text)]
Ypthima albida albida ; Kielland, 1982, Tijdschr. Ent. 125 (5) : 143; [BAFR, 193]; [AfrL]
Y. a. uniformis Bartel, 1905
E.Zaire (Kivu) Ypthima albida uniformis Bartel, 1905; Novit. Zool. 12 : 134, TL: Central Africa
Ypthima albida uniformis ab. pseudalbida Dufrane, 1945; 94, TL: Kamitunga
= ; Kielland, 1982, Tijdschr. Ent. 125 (5) : 144
Ypthima albida uniformis ab. impunctata Dufrane, 1945; 94, TL: Kamitunga
= ; Kielland, 1982, Tijdschr. Ent. 125 (5) : 144
Ypthima albida uniformis ; Kielland, 1982, Tijdschr. Ent. 125 (5) : 144; [BAFR, 193]; [AfrL]
Y. a. occidentalis Bartel, 1905
Camoroun - N.Nigeria Ypthima albida occidentalis Bartel, 1905; Novit. Zool. 12 : 134, TL: N. Cameroun
Ypthima albida ab. argentoides Strand, 1914; Arch. Naturgesch. 80 A (2) : 153, TL: Dschang, Cameroun
= ; Kielland, 1982, Tijdschr. Ent. 125 (5) : 143
Ypthima albida occidentalis ; Kielland, 1982, Tijdschr. Ent. 125 (5) : 143; [BAFR, 193 (text)]; [AfrL]

The chenu species-group
chenu ; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80
praenubila ; Shirôzu & Shima

Ypthima chenu (Guérin-Méneville, 1843)
Coorg, S.India. See [About maps] Nilgiri Fourring
Satyrus chenu Guérin-Méneville, 1843; in Delessert, Souvenirs Voy. Inde 2: 77, pl. 21, f. 2
Yphthima[sic] chenu ; Hewitson, 1865, Trans. ent. Soc. Lond. (3) 2 (4) : 285
Ypthima chenui[sic, recte chenu] ; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1) : 43; [BIR]; [BOW: pl. 172, f. 14]
Ypthima chenu ; [BOR, 466]
Ypthima (Ypthima) chenu ; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80

Ypthima ypthimoides (Moore, 1881)
S.India - S.Nilgiris. See [About maps] Palni Fourring
Callerebia ypthimoides Moore, 1881; Trans. ent. Soc. Lond. 1881 (3) : 307
Ypthima robinsoni Distant, 1882; Ann. Mag. nat. Hist. (5) 10 (59) : 406
= ; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1) : 43
Ypthima ypthimoides ; [BOR, 468]
Ypthima (Ypthima) ypthimoides ; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80

Ypthima praenubila Leech, 1891
W.China, C.China. See [About maps] Ypthima praenubila Leech, 1891; Entomologist 24 (Suppl.) : 66, TL: ;Ta-chien-lu, Chia-Kow-Ho, Moupin; Omei-Shan; Wa-Shan; Kiukiang
Ypthima praenubila praenubila, Butterflies in Indo-China [Yutaka Inayoshi]
Ypthima praenubila ; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1) : 18, pl. 3, f. 47; Forster, [1948], Mitt. Münch. ent. Ges. 34 (2) : 481; [BOW: pl. 205, f. 2]; [BOR, 468]; [MRS, 393]
Ypthima (Ypthima) praenubila ; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80

Larva on Pogonatherum crinitinum [MRS]

Y. p. kanonis Matsumura, 1929
Taiwan Ypthima praenubila kanonis Matsumura, 1929; Insecta Matsumurana 3: 141, TL: Formosa
Ypthima praenubila kanonis ; Uemura, 1984, Tyô to Ga 35 (3): 136; [MRS, 393]

The huebneri species-group
huebneri ; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80

Ypthima huebneri Kirby, 1871
NW.India - Peninsular Malaya, Langkawi Is., Singapore, Thailand, S.Yunnan. See [About maps] Common Fourring
Ypthima hübneri Kirby, 1871; Syn. Cat. diurn. Lep. : 95 (repl. Ypthima philomela Hübner, [1813])
Ypthima philomela Hübner, [1813]; Zuträge Samml. exot. Schmett. 1 : pl. [15], f. 83-84
Ypthima howra Moore, 1884; J. asiat. Soc. Bengal 53 Pt.II (1) : 17, TL: Calcutta
= ; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1) : 9
Ypthima apicalis Moore, 1884; J. asiat. Soc. Bengal 53 Pt.II (1) : 17, TL: "Deyra Dhoon"
= ; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1) : 9
Ypthima kasmira Moore, 1884; J. asiat. Soc. Bengal 53 Pt.II (1) : 17, TL: Cashmere
Ypthima catharina Butler, 1886; Ann. Mag. nat. Hist. (5) 18 (105) : 183, TL: Upper Burma
= ; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1) : 9
Ypthima jocularia Swinhoe, 1889; Proc. zool. Soc. Lond. 1889 (4) : 396, TL: Mahabalesvar
honora Fruhstorfer, 1911²; in Seitz, Gross-Schmett. Erde 9 : 287
= ; [NHM]
Ypthima huebneri, Butterflies in Indo-China [Yutaka Inayoshi]

Yphthima[sic] philomela ; Hewitson, 1865, Trans. ent. Soc. Lond. (3) 2 (4) : 284
Ypthima hübneri ; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1) : 9, pl. 1, f. 5
Ypthima kasmira ; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1) : 10, pl. 1, f. 6
Ypthima (Ypthima) kasmira ; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80
Ypthima jocularia ; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1) : 50
Ypthima ceylonica huebneri ; [BOR, 464]
Ypthima huebneri ; [EBW]; [BOW: pl. 173, f. 2]; [BMP: 135, pl. 17, f. 30-31]
Ypthima (Ypthima) huebneri ; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80
[huebneri-1v.jpg] 769x513 (~84Kb) underside Thailand, Krabi 24.12.2003-7.1.2004, Photo © Olli Vesikko


Ypthima ceylonica Hewitson, 1865
Ceylon. See [About maps] White Fourring
Yphthima[sic] ceylonica Hewitson, 1865; Trans. ent. Soc. Lond. (3) 2 (4) : 288, pl. 18, f. 14-15, TL: Ceylon
gellia Fruhstorfer, 1911²; in Seitz, Gross-Schmett. Erde 9 : 287, pl. 99 e
= ; [NHM]
Ypthima ceylonica ; Moore, [1880], Lepid. Ceylon 1 (1): 25, pl. 12, f. 5, 5a; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1) : 11, pl. 1, f. 8; [BOW: pl. 172, f. 12]
Ypthima ceylonica ceylonica ; [BOR, 464]
Ypthima (Ypthima) ceylonica ; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80

The megalomma species-group
megalomma ; Shirozu & Shima, 1979; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80; Huang & Wu, 2003, Neue Ent. Nachr. 55: 121
akbar ; Aoki & Uémura, 1984

Ypthima chinensis Leech, 1892
C.China. See [About maps] Ypthima newara var. chinensis Leech, [1892]; Butts China Japan Corea (1): 89, pl. 10, f. 5
Ypthima (Ypthima) chinensis ; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80
Ypthima chinensis ; Forster, [1948], Mitt. Münch. ent. Ges. 34 (2) : 477; [MRS, 394]; Huang & Wu, 2003, Neue Ent. Nachr. 55: 120 (note)

Ypthima iris Leech, 1891
-. See [About maps] Ypthima iris Leech, 1891; Entomologist 24 (Suppl.) : 57, TL: Wau-ssu-kou; Chow-pin-sa
Ypthima dromonides Oberthür, 1891; Étud. d'Ent. 15 : 15, pl. 2, f. 14
= ; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1) : 42
hygrophilus Mell, 1942; Arch. Naturgesch. (N.F.) 11 : 261
= ; [NHM]
Ypthima iris hygrophilus ; Forster, [1948], Mitt. Münch. ent. Ges. 34 (2) : 485
Ypthima iris ; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1) : 42; Forster, [1948], Mitt. Münch. ent. Ges. 34 (2) : 485; [BOW: pl. 205, f. 3]
Ytphima iris ; [MRS, 395]
Ypthima (Ypthima) iris ; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80
Sichuan Ypthima iris iris ; Huang, 2003, Neue Ent. Nachr. 55: 53 (note)
Y. i. microiris Uémura & Koiwaya, 2000
NE.Tibet Ypthima iris microiris Uémura & Koiwaya, 2000; , TL: Qamdo, NE.Tibet
Y. i. paradromon Uémura & Koiwaya, 2000
N.Yunnan Ypthima iris paradromon Uémura & Koiwaya, 2000; , TL: N.Yunnan
Y. i. naqialoa Huang, 2003
Yunnan Ypthima iris naqialoa Huang, 2003; Neue Ent. Nachr. 55: 53, pl. 7, f. 4, TL: on path between Naqialo and Longpo, Nujiang valley, NW.Yunnan

Ypthima dromon Oberthür, 1891
Yunnan. See [About maps] Ypthima dromon Oberthür, 1891; Étud. d'Ent. 15 : 15, pl. 2, f. 12, TL: Yunnan
Ypthima dromon ; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1) : 51; Forster, [1948], Mitt. Münch. ent. Ges. 34 (2) : 488, pl. 32-33, f. 5-6; [MRS, 395]
Ypthima (Ypthima) dromon ; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80

Ypthima megalomma Butler, 1874
-. See [About maps] Ypthima megalomma Butler, 1874; Cistula ent. 1 (9) : 236
Ypthima megalomma ; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1) : 44; Forster, [1948], Mitt. Münch. ent. Ges. 34 (2) : 491; [BOW: pl. 204, f. 4 (text only)]; [MRS, 395]
Ypthima (Ypthima) megalomma ; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80
Y. m. megalia de Nicéville, 1897
N.Shan States Ypthima megalia de Nicéville, 1897; J. Asiat. Soc. Bengal 66 Pt.II (3) : 546, pl. 1, f. 5
Ypthima megalia ; [BOR, 472 (note)]

Ypthima beautei Oberthür, 1884
-. See [About maps] Ypthima beautei Oberthür, 1884; Étud. d'Ent. 9 : 18, pl. 2, f. 1, TL: Ta-tsien-lo, West China
Ypthima (Ypthima) beautei ; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80
Ypthima beautei ; [BOW: pl. 205, f. 4 (text only)]; Huang & Wu, 2003, Neue Ent. Nachr. 55: 115 (note)
W.China Ypthima beautei beautei ; Huang & Wu, 2003, Neue Ent. Nachr. 55: 118 (note), f. 8, 9, 11, pl. 10, f. 1
Y. b. qinghaiensis Huang & Wu, 2003
Qinghai Ypthima beautei qinghaiensis Huang & Wu, 2003; Neue Ent. Nachr. 55: 118, f. 5-7, 12, pl. 10, f. 2, TL: Make valley, E.Qinghai, China

Ypthima pseudodromon Forster, [1948]
NW.Yunnan. See [About maps] Ypthima pseudodromon Forster, [1948]; Mitt. Münch. ent. Ges. 34 (2) : 488, pl. 32-33, f. 7-8, TL: Likiang, N.Yunnan
Ypthima (Ypthima) pseudodromon ; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80
Ypthima pseudodromon ; Huang & Wu, 2003, Neue Ent. Nachr. 55: 116 (ntoe)

Ypthima akbar Talbot, 1947
Burma, Thailand, Laos, S.Yunnan. See [About maps] Ypthima akbar Talbot, 1947; Fauna Br. Ind., Butt. (2nd ed.) 2: 324, TL: Karen Hills
Ypthima akbar, Butterflies in Indo-China [Yutaka Inayoshi]
Ypthima (Ypthima) akbar ; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80

Ypthima insolita Leech, 1891
W.China. See [About maps] Ypthima insolita Leech, 1891; Entomologist 24 (Suppl.) : 66, TL: Wa-ssu-Kou
Ypthima insolita ; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1) : 45; [BOW: pl. 205, f. 4]
Ypthima (Ypthima) insolita ; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80

Ypthima putamdui South, 1913
Sichuan. See [About maps] Ypthima putamdui South, 1913; J. Bomb. nat. Hist. Soc. 22 (2): 348, TL: Nya Chua, Sichuan
Ypthima (Ypthima) putamdui ; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80
Ypthima putamdui ; Huang & Wu, 2003, Neue Ent. Nachr. 55: 116 (note)

Ypthima baileyi South, 1913
Sichuan. See [About maps] Ypthima baileyi South, 1913; J. Bomb. nat. Hist. Soc. 22 (2): 348, TL: Batang, W.Sichuan
Ypthima baileyi ; Huang & Wu, 2003, Neue Ent. Nachr. 55: 118, f. 7, 14, 18-20, pl. 10, f. 4-5

Ypthima frontierii Uémura & Monastyrskii, 2000
N.Vietnam, Guanxi. See [About maps] Ypthima frontierii Uémura & Monastyrskii, 2000; Trans. Lepid. Soc. Japan 51 (2) : 150, f. 1a-c, 2, 4-9, TL: N.Vietnam, Lao Cai Prov., Sa Pa, ~2000m (103°46'E, 22°15'N)
Ypthima frontierii, Butterflies in Indo-China [Yutaka Inayoshi]
Ypthima frontierii ; Huang & Wu, 2003, Neue Ent. Nachr. 55: 120, f. 21, pl. 10, f. 7

Ypthima yoshinobui Huang & Wu, 2003
Qinghai Ypthima yoshinobui Huang & Wu, 2003; Neue Ent. Nachr. 55: 115, f. 3, 4, 7, 10, pl. 10, f. 3, TL: Make valley, E.Qinghai, China

Ypthima lihongxingi Huang & Wu, 2003
Hubei. See [About maps] Ypthima lihongxingi Huang & Wu, 2003; Neue Ent. Nachr. 55: 120, f. 13, 15-17, pl. 10, f. 6, TL: Xing-dou-shan, Lichuan, Hubei

The newara species-group
newara ; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80; Huang, 2001, Neue Entomologische Nachrichten, 51: 92

Ypthima newara Moore, [1875]
Nepal. See [About maps] Yphthima[sic] newara Moore, [1875]; Proc. zool. Soc. Lond. 1874 (4) : 567, TL: Nepal
Ypthima nareda newara ; [BOR, 470]
Ypthima (Ypthima) newara ; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80
Ypthima newara ; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1) : 21, pl. 2, f. 42; Huang, 2001, Neue Entomologische Nachrichten, 51: 92 (note), f. 81, pl. 9, f. 72-right
Misidentified/misapplied:
Yphthima nareda; Hewitson, 1865, Trans. ent. Soc. Lond. (3) 2 (4) : pl. 17, f. 7 (nec. f. 6)
Y. n. yaluzangbui Huang, 1999
SE.Tibet Ypthima newara yaluzangbui Huang, 1999; , TL: Metok
Ypthima newara yaluzangbui ; Huang, 2001, Neue Entomologische Nachrichten, 51: 92 (note), f. 80 (gen); Huang, 2003, Neue Ent. Nachr. 55: 52

Ypthima dengae Huang, 2001
SE.Tibet. See [About maps] Ypthima dengae Huang, 2001; Neue Entomologische Nachrichten, 51: 92, f. 79, 115, 117, pl. 7. f. 49, 53, TL: Ba-an-tang, Chayu area Tibet
Ypthima dengae ; Huang, 2003, Neue Ent. Nachr. 55: 52

Ypthima confusa Shirôzu & Shima, 1977
Nepal, N.Thailand, Laos, Yunnan. See [About maps] Ypthima confusa Shirôzu & Shima, 1977; Kontyu 45 (4): 504, f. 1c-d, 1g, TL: Kalapani, Nepal
Ypthima confusa, NSG Voucher Specimen [Wahlberg]
Ypthima confusa, Butterflies in Indo-China [Yutaka Inayoshi]
Ypthima (Ypthima) confusa ; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80
Ypthima confusa ; Huang, 2001, Neue Entomologische Nachrichten, 51: 92 (note); Huang, 2003, Neue Ent. Nachr. 55: 49, f. 71, pl. 6, f. 8-10
Ypthima confusa confusa ; Huang, 2001, Neue Entomologische Nachrichten, 51: f. 119 (gen.), pl. 9, f. 72-left

Ypthima muotuoensis Huang, 2000
-. See [About maps] Ypthima confusa muotuoensis Huang, 2000; , TL: Metok
Ypthima confusa muotuoensis ; Huang, 2001, Neue Entomologische Nachrichten, 51: f. 84 (gen.)
Ypthima muotuoensis ; Huang, 2003, Neue Ent. Nachr. 55: 48 (note)
SE.Tibet Ypthima muotuoensis muotuoensis ; Huang, 2003, Neue Ent. Nachr. 55: 49, f. 70, pl. 6, f. 4-7
Y. m. dulongae Huang, 2003
Yunnan Ypthima muotuoensis dulongae Huang, 2003; Neue Ent. Nachr. 55: 48, 50, f. 67, 69, pl. 6, f. 1-3, TL: Xiandjiudang to Longyuan, Dulong valley, NW.Yunnan

Ypthima pemakoi Huang, 1998
SE.Tibet. See [About maps] Ypthima pemakoi Huang, 1998; , TL: Metok
Ypthima pemakoi ; Huang, 2001, Neue Entomologische Nachrichten, 51: 92 (note), f. 85, 116; Huang, 2003, Neue Ent. Nachr. 55: 49, f. 74, pl. 7, f. 1-2

Ypthima sinica Uémura & Koiwaya, 2000
Sichuan, E.China. See [About maps] Ypthima sinica Uémura & Koiwaya, 2000; , TL: Qingchengshan, Sichuan
Ypthima sinica ; Huang, 2001, Neue Entomologische Nachrichten, 51: 92 (note); Huang, 2003, Neue Ent. Nachr. 55: 52, f. 75

Ypthima tiani Huang & Liu, 2000
-. See [About maps] Ypthima tiani ; Huang, 2001, Neue Entomologische Nachrichten, 51: 92 (note)
Sichuan
Y. t. nuae Huang, 2001
Yunnan Ypthima tiani nuae Huang, 2001; Neue Entomologische Nachrichten, 51: 93, f. 82, 114, 120, pl. 7, f. 50, 54, TL: Longpo to Ninadan, Nujiang Valley, SE.Tibet
Ypthima tiani nuae ; Huang, 2003, Neue Ent. Nachr. 55: 51, f. 68, 72-73, pl. 6, f. 11-18

The pandocus species-group
pandocus ; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80

Ypthima pandocus Moore, [1858]
-. See [About maps] Ypthima pandocus Moore, [1858]; in Horsfield & Moore, Cat. lep. Ins. Mus. East India Coy 1 : 235, TL: Java
Ypthima pandocus var. mentawica Hagen, 1902; Abh. Senckenb. Nat. Ges. 20 : 330, TL: Mentawej
= ; [NHM]
naerius Fruhstorfer, 1911²; in Seitz, Gross-Schmett. Erde 9 : 293, TL: Nias
= ; [NHM]
nacrianus Fruhstorfer, 1911²; in Seitz, Gross-Schmett. Erde 9 : 293, TL: Minahassa
= ; [NHM]
dyna Fruhstorfer, 1911²; in Seitz, Gross-Schmett. Erde 9 : 293
= ; [NHM]
andokides Fruhstorfer, 1911²; in Seitz, Gross-Schmett. Erde 9 : 293, TL: Java
= ; [NHM]
Ypthima pandocus barissanus Martin, 1929; Mitt. Munchen ent. Ges. 19 (5-9) : 125, TL: Battak
= ; [NHM]
Ypthima pandocus minimus Martin, 1929; Mitt. Munchen ent. Ges. 19 (5-9) : 126, TL: SE.Borneo
= ; [NHM]
Ypthima pandocus balianus Martin, 1929; Mitt. Munchen ent. Ges. 19 (5-9) : 127, TL: Bali I.
= ; [NHM]
kosong Cowan, 1956; Bull. Raffles Mus. 27: 39, TL: Malaya
= ; [NHM]
microocellata Miller & Miller, 1978; Bull. Allyn Mus. 49: 10, TL: Batjan
= ; [NHM]
Yphthima[sic] pandocus ; Hewitson, 1865, Trans. ent. Soc. Lond. (3) 2 (4) : 290, pl. 18, f. 12; van Eecke, 1913, Notes Leyden Mus. 35 : 243
Ypthima pandocus ; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1) : 22, pl. 2, f. 38; [BOR, 468]
Ypthima (Ypthima) pandocus ; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80
Java Ypthima pandocus pandocus ; [BOR, 468]
Y. p. sertorius Fruhstorfer, 1911²
Borneo pandocus sertorius Fruhstorfer, 1911²; in Seitz, Gross-Schmett. Erde 9 : 293, pl. 99 e
Ypthima pandocus sertorius ; [BOR, 468]
Y. p. corticaria Butler, 1879
Sumatra, Peninsular Malaya Common Three-ring
Ypthima corticaria Butler, 1879; Trans. linn. Soc. Lond. (2) 1 (8) : 537, TL: Malacca
= Ypthima pandocus ; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1) : 22
Ypthima pandocus emporialis Martin, 1929; Mitt. Munchen ent. Ges. 19 (5-9) : 126, TL: "Bukit Fima", Singapore
Ypthima pandocus f. alada Fruhstorfer, 1911; in Seitz, Gross-Schmett. Erde 9 : 293, TL: Battaker Plateau, NE.Sumatra.
Ypthima pandocus corticaria, Butterflies in Indo-China [Yutaka Inayoshi]
Ypthima pandocus corticaria ; [BMP: 135, pl. 17, f. 16-17]; [BOR, 468]
Y. p. tahanensis Pendlebury, 1933
Peninsular Malaya (Gunong Tahan) Ypthima tahanensis Pendlebury, 1933; J. Fed. Malay St. Mus. 17 (2): 384
Ypthima pandocus tahanensis ; Eliot, 1959, Bull. Br. Mus. nat. Hist. (Ent.) 7 (8) : 372; [BMP: 135, pl. 17, f. 18-19]; [BOR, 468]
Y. p. moenus Fruhstorfer, 1911²
Natuna Is. pandocus moenus Fruhstorfer, 1911²; in Seitz, Gross-Schmett. Erde 9 : 293, TL: Natuna Is.
Ypthima pandocus moenus ; [BOR, 468]
Y. p. aquillius Fruhstorfer, 1911²
Palawan pandocus aquillius Fruhstorfer, 1911²; in Seitz, Gross-Schmett. Erde 9 : 293, TL: Palawan
Ypthima pandocus aquillius ; [BOR, 468]
Y. p. calamus Fruhstorfer, 1911²
?Sulu Is. pandocus calamus Fruhstorfer, 1911²; in Seitz, Gross-Schmett. Erde 9 : 293
Ypthima pandocus calanus[sic?] ; [BOR, 468]
Y. p. jamaeus Fruhstorfer, 1911²
Banggai pandocus jamaeus Fruhstorfer, 1911²; in Seitz, Gross-Schmett. Erde 9 : 293
Ypthima pandocus jamaeus ; [BOR, 468]
Y. p. chaboras Fruhstorfer, 1911²
Philippines (Basilan, Mindanao) pandocus chaboras Fruhstorfer, 1911²; in Seitz, Gross-Schmett. Erde 9 : 293, TL: Bazilan
Ypthima pandocus chaboras ; [BOR, 468]
Y. p. sempera C. & R. Felder, 1863
Philippines (Luzon, ...) Ypthima sempera C. & R. Felder, 1863; Wien. ent. Monats. 7 (4) : 125
= Ypthima pandocus ; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1) : 22
Ypthima (Ypthima) sempera ; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80
Ypthima pandocus sempera ; [BOR, 468]

Ypthima hanburyi Holloway, 1984
Sarawak (Gunong Mulu) Ypthima hanburyi Holloway, 1984; Sarawak Mus. J. (Suppl. II): -
Ypthima hanburyi ; [BOR, 471]
Ypthima (Ypthima) hanburyi ; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80

Ypthima nigricans Snellen, 1892
Java. See [About maps] Ypthima nigricans Snellen, 1892; Tijdschr. Ent. 35 (3/4) : 135, TL: W.Java
phasis Fruhstorfer, 1911²; in Seitz, Gross-Schmett. Erde 9 : 294
= ; [NHM]
Ypthima nigricans ; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1) : 52; [BOR, 468]
Ypthima (Ypthima) nigricans ; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80

Ypthima kalelonda Westwood, 1888
-. See [About maps] Ypthima kalelonda Westwood, 1888; Trans. Ent. Soc. Lond. 1888 (3) : 475, pl. 12, f. 4, TL: Kalelonda
= Ypthima pandocus ; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1) : 22
Ypthima kaledonda[sic] ; [NHM]
Ypthima kalelonda ; [BOR, 468]
Ypthima (Ypthima) kalelonda ; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 184
N.Sulawesi, Talisei Ypthima kalelonda kalelonda ; [BOR, 468]
Ypthima (Ypthima) kalelonda kalelonda ; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 184
Y. k. celebensis Rothschild, 1892
Sulawesi Ypthima celebensis Rothschild, 1892; Dt. ent. Z. Iris 5 (2) : 433, pl. 5, f. 3-4, TL: S.Celebes
= Ypthima loryma ; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1) : 23
Ypthima (Ypthima) kalelonda celebensis ; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 184
Y. k. mangolina Uemura, 1982
Sula Ypthima (Ypthima) kalelonda mangolina ; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 184

Ypthima risompae Uémura, 1982
Sulawesi. See [About maps] Ypthima (Ypthima) risompae ; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 184

Larva on Miscanthus, (in captivity) Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 184


Ypthima ancus Fruhstorfer, 1911²
S.Sulawesi (Bonthain Peak). See [About maps] ancus Fruhstorfer, 1911²; in Seitz, Gross-Schmett. Erde 9 : 293, pl. 99 g, TL: Bonthein
Ypthima ancus ; [BOR, 472 (note)]
Ypthima (Ypthima) ancus ; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 184

Ypthima gavalisi Martin, 1913
Sulawesi. See [About maps] Ypthima gavalisi Martin, 1913; Dt. ent. Z. Iris 27 (2): 109, (3): 121, TL: Celebes
?pandocoides Miller & Miller, 1978; Bull. Allyn Mus. 49: 11, TL: Celebes
Ypthima gavalisi ; [BOR, 466]
?Ypthima pandocoides ; [NHM]
Ypthima gavalisi f. pandocoides ; [BOR, 467, f.]
Ypthima gawalisi ; (missp.)
Ypthima (Ypthima) gavalisi ; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 184

Ypthima loryma Hewitson, 1865
Sulawesi. See [About maps] Yphthima[sic] loryma Hewitson, 1865; Trans. ent. Soc. Lond. (3) 2 (4) : 289, pl. 18, f. 16-17, TL: Macassar, Celebes
= Ypthima pandocus ; [NHM]
Ypthima loryma, NSG Voucher Specimen [Wahlberg]
Ythima loryma ; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1) : 23, pl. 3, f. 50
Ypthima loryma ; [BOR, 466]; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80
Ypthima (Ypthima) loryma ; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 184

Ypthima sensilis Kashiwai, 1982
Philippines (Mindanao). See [About maps] Ypthima sensilis Kashiwai, 1982; Tyô to Ga 33 (1,2): 44, TL: Penangudloton, Upper region of Upian River, Davao City Prov., Mindanao
Ypthima sensilis ; [BOR, 468]
Ypthima (Ypthima) sensilis ; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80

The motschulskyi species-group
motschulskyi ; Shirôzu & Shima; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80; Huang, 2001, Neue Entomologische Nachrichten, 51: 91
sordida ; Shirôzu & Shima
obscura ; Shirôzu & Shima

Ypthima motschulskyi (Bremer & Grey, 1853)
E.China, Hong Kong, Amur, Korea, Japan. See [About maps] Satyrus motschulskyi Bremer & Grey, 1853; Schmett. N. China: 8, pl. 2, f. 2
Ypthima akragas var. takamukuana Matsumura, 1919; Thous. Ins. Japan. Addit. 3 : 526, pl. 37, f. 4, 4a, TL: Formosa
= ; [NHM]
= Ypthima perfecta akragas ; Uemura, 1984, Tyô to Ga 35 (3): 136
formosicola Matsumura, 1929; Ill. comm. Ins. Japan 1: 10
= ; [NHM]
Ypthima motschulskyi ab. yoshisakai Murayama, 1961; Tyô to Ga 11 (4): 57, f. 5, TL: Mikage-cho, Kobe
Yphthima motschulskiji[sic] ; Hewitson, 1865, Trans. ent. Soc. Lond. (3) 2 (4) : 290
Ypthima motschulskyi ; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1) : 16, pl. 2, f. 34; Forster, [1948], Mitt. Münch. ent. Ges. 34 (2) : 479; [BRU, 192]; [BOW: pl. 205, f. 6]; [MRS, 394]
Ypthima (Ypthima) motschulskyi ; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80
[motschulskyi-1.jpg] 668x560 (~77Kb) Korea, 1989, S.W. Cheong col. & det., Photo © Enrique Garcia-Barros

Larva on Carex [BRU, Fukuda et al., 1983]


Ypthima obscura Elwes & Edwards, 1893; Trans. Ent. Soc. Lond. 1893 (1) : 17, pl. 2, f. 35, TL: Gensan, Korea
= ; Dubatolov & Lvovsky, 1997, Trans. lepid. Soc. Japan 48 (4): 194
Ypthima elongatum Matsumura, 1929; Insecta Matsumurana 3: 142, TL: Korea
= Ypthima amphithea obscura ; Uemura, 1984, Tyô to Ga 35 (3): 135
= ; Dubatolov & Lvovsky, 1997, Trans. lepid. Soc. Japan 48 (4): 194
Ypthima obscura ; Forster, [1948], Mitt. Münch. ent. Ges. 34 (2) : 480, pl. 32-33, f. 11-12
Ypthima motschulskyi motschulskyi ; Dubatolov & Lvovsky, 1997, Trans. lepid. Soc. Japan 48 (4): 194
Y. m. amphithea Ménétriés, 1859
Ypthima amphithea Ménétriés, 1859; in Schrenck, Reise Forschungen Amur-Lande 2 (1) : 41, pl. 3, f. 10, TL: Amoor-land, Sula
= Ypthima motschulskyi ; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1) : 16
Yphthima[sic] amphithea ; Hewitson, 1865, Trans. ent. Soc. Lond. (3) 2 (4) : 289
Ypthima (Ypthima) amphithea ; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80
Ypthima motschulskyi amphithea ; Dubatolov & Lvovsky, 1997, Trans. lepid. Soc. Japan 48 (4): 194; Huang, 2001, Neue Entomologische Nachrichten, 51: 90 (note)

Ypthima yangjiahei Huang, 2001
Yunnan. See [About maps] Ypthima yangjiahei Huang, 2001; Neue Entomologische Nachrichten, 51: 91, f. 78, 113, pl. 6, f. 45, TL: Langpo to Ninadan, Nujiang Valley, SE.Tibet
Ypthima yangjiahei ; Huang, 2003, Neue Ent. Nachr. 55: 100 (note), f. 161, pl. 7, f. 5-6

Ypthima multistriata Butler, 1883
Taiwan. See [About maps] Ypthima multistriata Butler, 1883; Ann. Mag. nat. Hist. (5) 12 (67) : 50, TL: Ichang
Ypthima arcuata Matsumura, 1919; Thous. Ins. Japan. Addit. 3 : 699, pl. 53, f. 17, TL: Formosa, Horisha
= ; Uemura, 1984, Tyô to Ga 35 (3): 135; Dubatolov & Lvovsky, 1997, Trans. lepid. Soc. Japan 48 (4): 194
Ypthima multistriata ; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1) : 18, pl. 2, f. 36; Forster, [1948], Mitt. Münch. ent. Ges. 34 (2) : 480; [BOR, 466]; [MRS, 395]; Dubatolov & Lvovsky, 1997, Trans. lepid. Soc. Japan 48 (4): 195; Huang, 2001, Neue Entomologische Nachrichten, 51: 91 (note)

Larva on Miscanthus sinensis, Setaria palmifolia, Isachne globosa, Paspalum conjugatum [MRS]

Y. m. ganus Fruhstorfer, 1911
N.China, C.China multistriata ganus Fruhstorfer, 1911; in Seitz, Gross-Schmett. Erde 9 : 291, TL: Qingdao
Ypthima multistriata koreana Dubatolov & Lvovsky, 1997; Trans. lepid. Soc. Japan 48 (4): 195, TL: Korea
= ; Huang, 2001, Neue Entomologische Nachrichten, 51: 91
Ypthima multistriata koreana, Lepidoptera collection [SZMN]
Ypthima motschulskyi ganus ; Forster, [1948], Mitt. Münch. ent. Ges. 34 (2) : 479
Ypthima multistriata ganus ; Huang, 2001, Neue Entomologische Nachrichten, 51: 91 (note)
Y. m. niphonica Murayama, 1969
Japan (Honshu) Ypthima niphonica Murayama, 1969; Tohoku Konchu Kenkyu 4 (2): 18
Ypthima motschulskyi niphonica ;
Ypthima multistriata niphonica ; Dubatolov & Lvovsky, 1997, Trans. lepid. Soc. Japan 48 (4): 196
Y. m. tsushimana Murayama, 1969
Japan Ypthima niphonica tsushimana Murayama, 1969; Tohoku Konchu Kenkyu 4 (2): 20
Ypthima multistriata tsushimana ; Dubatolov & Lvovsky, 1997, Trans. lepid. Soc. Japan 48 (4): 196

Ypthima perfecta Leech, 1892
-. See [About maps] Ypthima motschulskyi var. perfecta Leech, [1892]; Butts China Japan Corea (1): 88, pl. 10, f. 7
Ypthima (Ypthima) perfecta ; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80
Ypthima perfecta ; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1) : 19, pl. 2, f. 37; Forster, [1948], Mitt. Münch. ent. Ges. 34 (2) : 477; [MRS, 393]; Huang, 2001, Neue Entomologische Nachrichten, 51: 91 (note)
W.China, C.China
Y. p. akragas Fruhstorfer, 1911
Taiwan perfecta akragas Fruhstorfer, 1911; in Seitz, Gross-Schmett. Erde 9 : 291
Ypthima perfecta akragas ; [BOR, 472 (note)]; [MRS, 393]

Larva on Miscanthus sinensis, M. transmorrisonensis, Oplismenus undulatifolius [MRS]


Ypthima esakii Shirôzu, 1960
Taiwan Ypthima esakii Shirôzu, 1960; Butterflies of Formosa in Colour: 128
Ypthima esakii ; [BOR, 472 (note)]; [MRS, 394]
Ypthima (Ypthima) esakii ; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80

Larva on Miscanthus sinensis, Setaria palmifolia, Isachne globosa, Oplismenus undulatifolius [MRS]


Ypthima sordida Elwes & Edwards, 1893
China (N.Yunnan, Shensi, ...). See [About maps] Ypthima sordida Elwes & Edwards, 1893; Trans. Ent. Soc. Lond. 1893 (1) : 19, pl. 3, f. 52, TL: Kiukiang
Ypthima sordida tsinlingi Forster, [1948]; Mitt. Münch. ent. Ges. 34 (2) : 478, pl. 30-31, f. 14-15, TL: Tapaishan, Tsinlingshan, S.Shensi
= ; [NHM]
Ypthima (Ypthima) sordida ; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80
Ypthima sordida ; Forster, [1948], Mitt. Münch. ent. Ges. 34 (2) : 478, pl. 30-31, f. 12-13; Huang, 2001, Neue Entomologische Nachrichten, 51: 91 (note)

Ypthima lycus de Nicéville, 1889
Assam, Bhutan, Sikkim. See [About maps] Plain Three-Ring
Ypthima lycus de Nicéville, 1889; J. Bombay nat. Hist. Soc. 4 (3) : 165, pl. A, f. 2
?Ypthima lycoides Watson, 1897; J. Bombay nat. Hist. Soc. 10 (4) : 648, TL: North Chin Hills, Burma
Ypthima (Ypthima) lycus ; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80
Ypthima lycus ; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1) : 19, pl. 2, f. 39; Forster, [1948], Mitt. Münch. ent. Ges. 34 (2) : 479, pl. 32-33, f. 9-10; [BOR, 470]; Huang, 2001, Neue Entomologische Nachrichten, 51: 91 (note)

Ypthima riukiuana Matsumura, 1906
Japan. See [About maps] Ypthima riukiuana Matsumura, 1906; Annot. Zool. Jap. 6 (1) : 11, pl. 1, f. 2, TL: Okinawa, The Loochoos
Ypthima riukiuana ; Uemura, 1984, Tyô to Ga 35 (3): 136
Ypthima (Ypthima) riukiuana ; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80
Ypthima riukiuana ; Huang, 2001, Neue Entomologische Nachrichten, 51: 91 (note)

Ypthima imitans Elwes & Edwards, 1893
Hainan. See [About maps] Ypthima imitans Elwes & Edwards, 1893; Trans. Ent. Soc. Lond. 1893 (1) : 17, pl. 3, f. 53, TL: Hainan
Ypthima imitans, Butterflies in Indo-China [Yutaka Inayoshi]
Ypthima (Ypthima) imitans ; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80
Ypthima imitans ; [MRS, 394]; Huang, 2001, Neue Entomologische Nachrichten, 51: 91 (note)

Ypthima phania (Oberthür, 1891)
Yunnan. See [About maps] phania (Oberthür, 1891) (Epinephele); Étud. d'Ent. 15 : 17, pl. 2, f. 17, TL: Yunnan
Ypthima phania ; Forster, [1948], Mitt. Münch. ent. Ges. 34 (2) : 492, pl. 32-33, f. 13-14; Huang, 2001, Neue Entomologische Nachrichten, 51: 91 (note)

Ypthima watsoni (Moore, [1893])
Assam, Manipur - S.Burma. See [About maps] Looped Three-Ring
Pandima watsoni Moore, [1893]; Lepidoptera Indica 2 (16): 89, (15): pl. 113, f. 4, 4a-b, TL: Burma
Ypthima watsoni watsoni, Butterflies in Indo-China [Yutaka Inayoshi]
Ypthima watsoni ; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1) : 47, pl. 1, f. 25; [BOR, 470]; Huang, 2001, Neue Entomologische Nachrichten, 51: 91 (note)
Assam - Burma, Thailand Ypthima watsoni var. howarthi Cantlie & Norman, 1959; J. Bombay nat. Hist. Soc. 56 (1) : 66, TL: Sebong, Manipur
Y. w. peninsulae Aoki & Uémura, 1984
Peninsular Thailand Ypthima watsoni peninsulae Aoki & Uémura, 1984; Tokurana 6/7: 87, f. 16i-j, 17, 43, pl. 5, f. 4, 8, TL: Thailand
Ypthima watsoni peninsulae, Butterflies in Indo-China [Yutaka Inayoshi]
Y. w. inouei Shirôzu & Shima, 1977
S.Vietnam Ypthima inouei Shirôzu & Shima, 1977; Kontyu 45 (4): 507, f. 1e-f, TL: Trang Bom, S.Vietnam
Ypthima watsoni inouei, Butterflies in Indo-China [Yutaka Inayoshi]
Ypthima (Ypthima) inouei ; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80
?Ypthima watsoni r. inouei ; Huang, 2001, Neue Entomologische Nachrichten, 51: 91

Ypthima fusca Elwes & Edwards, 1893
Ypthima fusca Elwes & Edwards, 1893; Trans. Ent. Soc. Lond. 1893 (1) : 21
Ypthima (Ypthima) fusca ; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80
Ypthima fusca ; Huang, 2001, Neue Entomologische Nachrichten, 51: 91 (note)

Ypthima masakii Ito, 1947
Japan. See [About maps] Ypthima masakii Ito, 1947; Zephyrus 9: 272, TL: Loochoo I.
Ypthima (Ypthima) masakii ; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80
Ypthima masakii ; Huang, 2001, Neue Entomologische Nachrichten, 51: 91 (note)

Ypthima ciris Leech, 1891
W.China. See [About maps] Ypthima ciris Leech, 1891; Entomologist 24 (Suppl.) : 4, TL: Wa-Shan; Chia-Ting-Fu; Ta-Tsien-Lu; Huang-Mu-Chang
Ypthima clinia Oberthür, 1891; Étud. d'Ent. 15 : 16, pl. 2, f. 13
= ; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1) : 41
Ypthima civis[sic] ; [NHM]
Ypthima (Ypthima) ciris ; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80
Ypthima ciris ; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1) : 41; Forster, [1948], Mitt. Münch. ent. Ges. 34 (2) : 486; [BOW: pl. 205, f. 3 (text only)]; [MRS, 393]; Huang, 2001, Neue Entomologische Nachrichten, 51: 91 (note)
Y. c. clinioides Oberthür, 1891
Yunnan, ... Ypthima clinioides Oberthür, 1891; Étud. d'Ent. 15 : 16
Ypthima clinioides ; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1) : 51
Ypthima ciris clinioides ; Huang, 2003, Neue Ent. Nachr. 55: 100 (note)

The ? species-group
? ; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80

Ypthima bolanica Marshall, 1882
Baluchistan. See [About maps] Desert Fourring
Ypthima bolanica Marshall, 1882; Proc. zool. Soc. Lond. 1882 (4) : 759, TL: Bolan Pass
Ypthima bolanica ; [AfrL]

Ypthima inica Hewitson, 1865
Punjab - C.India, Bengal, Assam. See [About maps] Lesser Three-Ring
Yphthima[sic] inica Hewitson, 1865; Trans. ent. Soc. Lond. (3) 2 (4) : 284, pl. 17, f. 5, TL: N.India [?Bengal]
Yphthima[sic] ariaspa Moore, [1875]; Proc. zool. Soc. Lond. 1874 (4) : 568, TL: Punjab
Ypthima rara Butler, 1883; Proc. zool. Soc. Lond. 1883 (2) : pl. 24, f. 1
= Ypthima ariaspa ; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1) : 11
Ypthima alkibie Swinhoe, 1886; Proc. zool. Soc. Lond. 1886 (4) : 422
= ; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1) : 13
Ypthima daedalea Swinhoe, 1886; Proc. zool. Soc. Lond. 1886 (4) : 423
= Ypthima ariaspa ; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1) : 11
Ypthima ariaspa ; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1) : 11
Ypthima inica ; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1) : 13, pl. 1, f. 9; [BOR, 464]

Ypthima abnormis Shelford, 1904²
Borneo (Sarawak). See [About maps] abnormis Shelford, 1904²; Straits asiat. Soc. 41: 98, TL: Sarawak
Ypthima abnormis ; [BOR, 472]

Ypthima junkoae Uémura, 1999
C.Sulawesi. See [About maps] Ypthima (Ypthima) junkoae ; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 184

Subgenus ?
? ; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80

Ypthima argillosa Snellen, 1892
Java. See [About maps] Ypthima argillosa Snellen, 1892; Tijdschr. Ent. 35 (3/4) : 133, TL: Java
Ypthima argillosa ; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1) : 52; [BOR, 472 (note)]

Ypthima florensis Snellen, 1891
Flores. See [About maps] Yphthima[sic] florensis Snellen, 1891; Tijdschr. Ent. 34 : 235, pl. 14, f. 3, TL: Flores
Ypthima florensis ; [BOR, 472 (note)]

Ypthima angustipennis Takahashi, 2000
Taiwan Ypthima angustipennis Takahashi, 2000; Trans. lepid. Soc. Japan 51 (1) : 4, TL: Taiwan, Kaohsiung, Tengzhi

Ypthima hyagriva Moore, 1857
N.India, Kashmir - Kumaon, Sikkim. See [About maps] Brown Argus
Yphthima[sic] hyagriva Moore, 1857; in Horsfield & Moore, Cat. lep. Ins. Mus. East India Coy 1 : 236, TL: Darjeeling
Yphthima[sic] hyagriva ; Hewitson, 1865, Trans. ent. Soc. Lond. (3) 2 (4) : pl. 18, f. 11
Dallachia hyagriva ;
?Erebia hyagriva ; [BOW: pl. 200, f. 13]

Ypthima albipuncta Lee, 1985
Yunnan. See [About maps] Ypthima albipuncta Lee, 1985; , TL: Yi-liang, Yunnan, 1800m
Ypthima albipuncta ; Huang & Wu, 2003, Neue Ent. Nachr. 55: 122, f. 1-2, pl. 10, f. 8-9

Ypthima posticalis Matsumura, 1909
Taiwan. See [About maps] Ypthima posticalis Matsumura, 1909; Ent. Zs. 23 (19) : 92, TL: Formosa, Horisha
Ypthima minuta Matsumura, 1909; Ent. Zs. 23 (19) : 92, TL: Formosa, Hokulo
= ; Uemura, 1984, Tyô to Ga 35 (3): 136
= Ypthima lisandra ; [NHM]
Ypthima norma posticalis ; Uemura, 1984, Tyô to Ga 35 (3): 136
Ypthima posticalis ; [BOR, 472 (note)]
Y. p. aei Shirôzu & Shima, 1977
Philippines (Luzon) Ypthima posticalis aei Shirôzu & Shima, 1977; Kontyu 45 (4): 501, TL: Baguio City, Luzon Is, Philippines
Ypthima posticalis aeigma[sic] ; [NHM]

Ypthima parasakra Eliot, 1987
-. See [About maps] Ypthima parasakra Eliot, 1987; Tyô to Ga 38 (1): 21, f. 1a-d, 2-5, TL: Nepal, Mustang District (28°40'N, 83°35'E), Larjung
Nepal, Bhutan Ypthima parasakra parasakra ; Huang, 2003, Neue Ent. Nachr. 55: 52 (note); Huang & Wu, 2003, Neue Ent. Nachr. 55: 124
Misidentified/misapplied:
Ypthima methorina; Lee, 1982, in Huang, Insects of Xizang, 2: 135, No. 23 (66) (missid.)
Y. p. menpae Huang, 1999
SE.Tibet Ypthima parasakra menpae ; Huang, 2003, Neue Ent. Nachr. 55: 52 (note)
Y. p. mabiloa Huang, 2003
Yunnan Ypthima parasakra mabiloa Huang, 2003; Neue Ent. Nachr. 55: 52, pl. 7, f. 3, TL: Xiongdang, Dulong valley, NW.Yunnan

Ypthima striata Hampson, 1888
S.India (Nilgiri Hills). See [About maps] Ypthima striata Hampson, 1888; J. asiat. Soc. Bengal 57 (4) : 349, TL: Nilgiri Hills
Ypthima striata ; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1) : 34, pl. 1, f. 23-24; [BOR, 466]

Ypthima yamanakai Sonan, 1938
Taiwan Ypthima yamanakai Sonan, 1938; Trans. nat. Hist. Soc. Formosa 28: 250, pl. 20, f. 8, TL: Formosa
Ypthima yamanakai ; [BOR, 472 (note)]; [MRS, 391]; Takahashi, 2000, Trans. lepid. Soc. Japan 51 (1) : 9

Ypthima zyzzomacula Chou & Li, 1994
Yunnan. See [About maps] Ypthima zyzzomacula Chou & Li, 1994; in Chou, Monographia Rhopalocerum Sinensium 1-2: 759, 395, f. 30-31, TL: Gengma, Yunnan

Ypthima wenlungi Takahashi, 2007
S.Taiwan Ypthima wenlungi Takahashi, 2007; Trans. lepid. Soc. Japan 58 (3) : 277, f. 1-4, TL: Feng Gang Shan (1400-1600m), Kaohsiung, Taiwan


Unknown or unplaced taxa

selinuntioides (Mell, 1942) (Ypthima); Arch. Naturgesch. (N.F.) 11 : 261
Ypthima selinuntioides ; [NHM]

Papilo lysandra Cramer, [1780]; Uitl. Kapellen 4 (25-26a): pl. 293
Yphthima[sic] lysandra ; Hewitson, 1865, Trans. ent. Soc. Lond. (3) 2 (4) : 287

praenobilia
neobilia (Murayama, 1980) (Ypthima); New Entomologist 29 (2): 51
Ypthima praenobilia ; [NHM]

Satyrus corynetes Boisduval, 1847; in Delegorgue, Voyage Afr. australe, 2: 294 (nom. nud)

anana Fruhstorfer, 1911²; in Seitz, Gross-Schmett. Erde 9 : 293, TL: E.Celebes
Ypthima anana ; [NHM]


7.5.2013 (126)

References:
[About maps]
Warning! The maps are automatically generated from the textual information, and the process does not always produce acceptable result; See about maps for more info.
[AfrL]
Mark C. Williams; Checklist of Afrotropical Papilionoidea and Hesperoidea; Compiled by Mark C. Williams, 7th ed. (2008) (April 2007); Afrototropical Butterflies; e-mail: Mark C. Williams
[BAFR]
D'Abrera, 1980; Butterflies of the Afrotropical Region
[BAUR]
D'Abrera, 1977; Butterflies of the Australian Region (2nd edn)
[BHL]
The; Biodiversity Heritage Library; and on; Internet Archive; Note: The linking to the content of Internet Archive is somewhat delicate -- notify me when links break!
[BIR]
Wynter-Blyth, 1957; Butterflies of the Indian Region; (1982 Reprint)
[BK]
Larsen, 1996; The Butterflies of Kenya and their natural history
[BMP]
Eliot (Ed.), Corbet & Pendlebury, 1992; The Butterflies of The Malay Peninsula; 4th edition; ISBN 983-9681-05-2
[BOR]
D'Abrera, 1982, 1985, 1986; Butterflies of the Oriental Region, Part I-III
[BOW]
Lewis, 1974; Butterflies of the World; ISBN 0 245 52097 X
[BRU]
Tuzov, Bogdanov, Devyatkin, Kaabak, Korolev, Murzin, Samodurov, Tarasov, 1997; Guide to the Butterflies of Russia and adjacent territories; Hesperiidae, Papilionidae, Pieridae, Satyridae; Volume 1
[CiNii]
http://ci.nii.ac.jp/
[Don Herbison-Evans]
and Stella A. Crossley; Lepidoptera Larvae of Australia; e-mail: Don Herbison-Evans; e-mail: Don Herbison-Evans
[EBW]
Smart, Paul, 1976; The Illustrated Encyclopedia of the Butterfly World; ISBN 0 600 31381 6
[Enrique Garcia-Barros]
Enrique García-Barros; Dept. Biol. (Zool.); Univ. Autonoma de Madrid
[HBN]
Francis Hemming, 1937; HÜBNER, A bibliographical and systematic account of the entomological works of Jacob Hübner and of the supplements thereto by Carl Geyer, Gottfried Franz von Frölich and Gottlieb August Wilhelm Herrich-Schäffer, Vol 1 & 2
[Kedar Tokekar]
India
[LAMAS-2005]
Gerardo Lamas; Entomofauna 26 (6): 57-100; A Bibliography of the Zoological Publications of Hans Fruhstorfer (1886* - 1922+)
[MBE]
Moths and Butteflies of Europe; Paolo Mazzei, Diego Reggianti and Ilaria Pimpinelli; http://www.leps.it/
[MRS]
Chou Io (Ed.); Monographia Rhopalocerum Sinensium, 1-2
[NHM]
Natural History Museum; The Global Lepidoptera Names Index
[Oleg Kosterin]
Institue of Cytology & Genetics; Siberian Division of Russian Academy of Sciences; Novosibirsk; e-mail: Oleg Kosterin; http://pisum.bionet.nsc.ru/kosterin/
[Olli Vesikko]
http://koti.welho.com/ovesikko/
[Peter Payzant]
Canada; http://www.pbase.com/ppayzant/profile
[SZMN]
Lepidoptera; Siberian Zoological Museum
[Wahlberg]
Niklas Wahlberg; The NSG's voucher specimen database; Nymphalidae Systematics Group
[Yutaka Inayoshi]
A Check List of Butterflies in Indo-China; Chiefly from THAILAND, LAOS & VIETNAM
[²]
This may require parentheses or not. I don't have the necessary information for this taxon.


Some related literature:
Aoki & Uémura, 1982
Description of new species and new subspecies of South-East Asian diurnal Lepidoptera belonging to the family Satyridae, with some taxonomic notes Memoirs of the Tsukada Coll. No. 4: 66pp., 8pls
Aoki & Uémura, 1984
Studies on the genus Ypthima in Thailand Tokurana 6/7: 71-103
Bartel, 1905
Neue Aethiopische Rhopalocera des Kgl. Museums für Naturkunde in Berlin Novit. Zool. 12 : 129-152
Boisduval, 1847
Catalogue des Lépidoptères recuellis par M. Delegorgue, 1833-1844 in Delegorgue, Voyage Afr. australe, 2: 585-614
Bremer & Grey, 1853
Beiträge zur Schmetterlings-Fauna des nördlichen China Schmett. N. China: 23pp, [2 pls]
Butler, 1866
A list of the diurnal Lepidoptera recently collected by Mr. Whitely in Hakodadi (North Japan) J. Linn. Soc. Zool. Lond. 9 (34) : 50-58
Butler, 1871
Descriptions of five new species, and a new genus , of diurnal Lepidoptera, from Shanghai Trans. ent. Soc. Lond. 1871 (3) : 401-403
Butler, 1874
Descriptions of four new Asiatic Butterflies Cistula ent. 1 (9) : 235-236
Butler, 1876
On a Collection of Lepidoptera recently received from Abyssinia Ann. Mag. nat. Hist. (4) 18 (108) : 480-490
Butler, 1879
The butterflies of Malacca Trans. linn. Soc. Lond. (2) 1 (8) : 533-568, pl. 68-69
Butler, 1880
On new and little-known Butterflies from India Proc. zool. Soc. Lond. 1880 : 147-152, pl. 15
Butler, 1881
On a Collection of Butterflies from Nikko, Central Japan Ann. Mag. Nat. Hist. (5) 7 (38) : 132-140
Butler, 1882
Descriptions of new Species of Lepidoptera from Tenasserim Ann. Mag. Nat. Hist. (5) 10 : 372-376
Butler, 1883
The Lepidoptera collected during the recent expedition of H.M.S. "Challenger." Ann. Mag. nat. Hist. (5) 11 (66) : 402-428
Butler, 1883
On a Third Collection of Lepidoptera made by Mr. H. E. Hobson in Formosa Ann. Mag. nat. Hist. (5) 12 (67) : 50-52
Butler, 1883
On some Lepidoptera from the Victoria Nyanza Ann. Mag. nat. Hist. (5) 12 (68) : 101-107
Butler, 1883
On a collection of Indian Lepidoptera received from Lieut.-Colonel Charles Swinhoe; with numerous notes by the collector Proc. zool. Soc. Lond. 1883 (2) : 144-175, pl. 24
Butler, 1886
On a Collection of Lepidoptera made by Commander Alfred Carpenter, R. N., in Upper Burma, in the Winter of 1885-86 Ann. Mag. nat. Hist. (5) 18 (105) : 182-191
Butler, 1888
On the Lepidoptera received from Dr. Emin Pascha [from Equatorial Africa] Proc. Zool. Soc. Lond. 1888 : 56-85
Cantlie & Norman, 1959
Notes on the butterfly genus Ypthima J. Bombay nat. Hist. Soc. 56 (1) : 66-71
Carcasson, 1961
New and Little-known African Lepidoptera Occ. Pap. Coryndon meml Mus. (7): 3-23, 5pls
Cramer, [1780]
Uitlandsche Kapellen (Papillons exotiques) Uitl. Kapellen 4 (25-26a): 1-28, pl. 289-304 (1780)
de Nicéville, 1889
On new and little-known Butterflies from the Indian Region, with Revision of the Genus Plesioneura of Felder and of Authors J. Bombay nat. Hist. Soc. 4 (3) : 163-194, pl. A-B
de Nicéville, 1895
On new and little-known Butterflies from the Indo-Malayan Region J. Bombay nat. Hist. Soc. 10 (1): 13-40, (2): 169-194
de Nicéville, 1897
On new or little-known Butterflies from the Indo- and Austro-Malayan Regions J. Asiat. Soc. Bengal 66 Pt.II (3) : 543-577, pl. 1-4
Distant, 1882
Undescribed Rhopalocera from the Malay Peninsula Ann. Mag. nat. Hist. (5) 9 (53) : 396-397
Distant, 1882
On some apparently undescribed Rhopalocera Ann. Mag. nat. Hist. (5) 10 (59) : 405-407
Doherty, 1891
The Butterflies of Sumba and Sumbawa, with some Account of the Island of Sumba J. Asiat. Soc. Bengal 60 Pt.II (2) : 141-197, pl. 2
Donovan, 1805
An Epitome of the Natural History of the Insects of New Holland, New Zealand, New Guinea, Otaheite and other Islands in the Indian, Sothern and Pacific Oceans ... Epitome nat. Hist. Ins. New Holland: 41pls
Westwood, 1851
The genera of diurnal Lepidoptera, comprising their generic characters, a notice of their habitats and transformations, and a catalogue of the species of each genus; illustrated with 86 plates by W. C. Hewitson Gen. diurn. Lep. (1): 1-18, pl. A, 1-4 (1-2) (1846), (1): 19-132, pl. 4*, 5-25, 28 (3-14) (1847), (1): 133-200, pl. 1*, 26, 27, 29, 31-44 (15-23) (1848), (1): 201-242, pl. 30, 45-52, 56-58, 60-62, 64 (24-31) (1849), (1): 243-250 [?], (2): 243-326, pl. 53-55, 63, 65, 66 (32-38) (1850), (2): 327-466, pl. 59, 67-77 (39-50) (1851), (2): 467-534, 143, 144, pl. 78-80 (Index) (51-54) (1852)
Dubatolov & Lvovsky, 1997
What is true Ypthima motschulskyi (Lepidoptera, Satyridae)? Trans. lepid. Soc. Japan 48 (4): 191-198
K. L. & L. E. Dunn, 1991
Review of Australian Butterflies: distribution, life history and taxonomy. Pushlished by authors, Melbourne Review Austr. Butts. (1-4):
Eliot, 1959
New or little known Butterflies from Malaya Bull. Br. Mus. nat. Hist. (Ent.) 7 (8) : 371-391, 7fgs., pl. 10
Eliot, 1967
The Sakra Moore, 1857 Section of the Satyrid Genus Ypthima Hübner Entomologist 100 (1246) : 49-61, 1+22figs, pl. 2
Eliot, 1987
On some butteflies from Nepal, Bhutan and Thailand Tyô to Ga 38 (1): 21-29
van Eecke, 1913
Fauna Simalurensis. Lepidoptera Rhopalocera, fam. Satyridae, Morphidae and Nymphalidae Notes Leyden Mus. 35 : 243-247
Elwes & Edwards, 1893
A revision of the genus Ypthima, with especial reference to the characters afforded by the male genitalia Trans. Ent. Soc. Lond. 1893 (1) : 1-54, pl. 1-3
Eschscholtz, 1821
Beschreibung neuer ausländischer Schmetterling in Kotzebue, Entdeck. Reise Süd-See 3 (App. 5): 201-219, 11pls
Evans, 1912
A list of Indian butterflies J. Bombay nat. Hist. Soc. 21 (2): 553-584, (3): 969-1008
Evans, 1923; Evans, 1924
The Identification of Indian Butterflies (3-4) J. Bombay nat. Hist. Soc. 29 (3) (3): 780-797, pl. 13-16, : 890-907
Fabricius, 1775
Systema Entomologiae, sistens Insectorum Classes, Ordines, Genera, Species, Adiectis Synonymis, Locis, Descriptionibus, Observationibus. Syst. Ent.: 832pp
Felder, 1868
Diagnose neuer von E. Baron v. Ransonnet in Vorder-Indien gesammelter Lepidopteren Verh. zool.-bot. Ges. Wien 18 (1-2) : 281-286
C. & R. Felder, 1863
Lepidoptera nova a Dr. Carolo Semper in Insulis Philippinis collecta diagnosibus. (3) Wien. ent. Monats. 7 (4) : 105-127
Forster, [1948]
Beitrage zur Kenntnis der ostasiatischen Ypthima-Arten Mitt. Münch. ent. Ges. 34 (2) : 472-492, pl. 30-33
Fruhstorfer, 1908
Lepidopterologisches Pêle-Mêle. VI. Neue Rhopaloceren von Formosa Ent. Zs. 22 (29) : 118-119 (17 October)
Fruhstorfer, 1908
Neue ostasiatische Satyriden Ent. Zs. 22 (31) : 127 (31 October)
Fruhstorfer, 1911
4. Familie: Satyridae in Seitz, Gross-Schmett. Erde 9 : 285-288 (14 February, 1911), : 289-296 (15 February, 1911), : 297-304 (22 February, 1911), : 305-312 (23 February, 1911), : 313-328 (before 25 April, 1911), : 329-336 (before 2 May, 1911), : 337-344 (before 9 May, 1911), : 345-352 (before 23 May, 1911), : 353-360 (before 30 May, 1911), : 361-368 (before 7 June, 1911), : 369-376 (before 13 June, 1911), : 377-384 (before 20 June, 1911), : 385-392 (before 27 June, 1911), : 393-400 (24 July), : 401 (15 August)
Godart, [1824]
in Latreille & Godart, Encyclopédie Méthodique. Histoire naturelle Entomologie, ou histoire naturelle des crustacés, des arachnides et des insectes Encyclopédie Méthodique. 9 (1): 3-328 (1819), (2): 329-828 ([1824])
Grose-Smith, 1887
Descriptions of eight new Species of Asiatic Butterflies Ann. Mag. nat. Hist. (5) 20 : 265-268
Grünberg, 1910
in Schultze (Ed), Zoologische und anthropologische Ergebnisse einer Forschungsreise in Westlichen und Zentralen Süd-Afrika Denkschr. med.-naturw. Ges. Jena 16 (4): 91-146
Hagen, 1902
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Keep in mind that the taxonomic information is copied from various sources, and may include many inaccuracies. Expert help is welcome.