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Pierella Herrich-Schäffer, 1865
Pierella Herrich-Schäffer, 1865; CorrespBl. zool.-min. Ver. Regensburg 19 (5): 65, TS: Papilio nereis Drury
Pieris Hübner, [1819]; (preocc. Schrank, 1801)
= ; [NL4A, 206]
Pierella Westwood, [1851]; Gen. diurn. Lep. (2): 365 (invalid)
Antirrhaea Westwood, 1851; (unavail.)
= ; [NL4A, 206]
Pierella, NSG Voucher Specimen [Wahlberg]
Pierella ; Godman & Salvin, [1880], Biol. centr.-amer., Lep. Rhop. 1: 66
Pierella (Haeterini) ; [Wahlberg]; [NL4A, 206]

Pierella amalia Weymer, 1885
Peru. See [About maps] Pierella amalia Weymer, 1885; , TL: Peru, Pebas
Pierella lena ab. leucospila Staudinger, 1887; , TL: "Upper Amazon"
= ; [NL4A, #517]
Pierella amalia ; [EcuL, (note)]; [NL4A, #517]

Pierella astyoche (Erichson, [1849])
Guianas. See [About maps] Hetaera[sic] astyoche Erichson, [1849]; , TL: Guyana
Haetera astyoche var. ; Hewitson, 1860, Ill. exot. Butts [4] (Haetera): [40], pl. [21], f. 5
Pierella astyoche ; [BOW: pl. 63, f. 12]; [NL4A, #518]
Guyana, Brazil(Pará) Haetera larymna Doubleday, [1849]; Gen. diurn. Lep. (1): pl. 62, TL: Brazil, Pará
= ; [NL4A, #518a]
Pierella astyoche astyoche ; [NL4A, #518a]
P. a. stollei Ribeiro, 1931
Brazil(Rondônia) astyoche stollei Ribeiro, 1931; , TL: Brazil, Rondônia
Pierella stollei ;
Pierella astyoche stollei ; [NL4A, #518c]
Brazil(Amazonas) astyoche bernhardina Bryk, 1953; , TL: Brazil, Amazonas
Pierella astyoche f. obscura Zikán & Wygodzinsky, 1948; (nom. nud.), TL: Brazil, Amazonas
= ; [NL4A, #518b]
Pierella astyoche bernhardina ; [NL4A, #518b]

Pierella helvina (Hewitson, 1860)
-. See [About maps] Haetera helvina Hewitson, 1860; Ill. exot. Butts [4] (Haetera): [40], pl. [21], f. 4, TL: Colombia
Pierella helvetia ; D'Abrera, 1988, (missp.)
Pierella helvina ; [BOW: pl. 63, f. 5]; [NL4A, #519]
Colombia Pierella helvina helvina ; [NL4A, #519a]
P. h. ocreata Godman & Salvin, 1868
Panama, Ecuador? Pierella ocreata Godman & Salvin, 1868; Ann. Mag. nat. Hist. (4) 2 (8) : 143, TL: Panama, Calobre
Pierella ocreata Butler, 1868; (nom. nud.), TL: Panama
= ; [NL4A, #519d]
Pierella ocreata ; Godman & Salvin, [1880], Biol. centr.-amer., Lep. Rhop. 1: 68, 3: pl. 6, f. 7-9
Pierella helvina ocreata ; [EcuL]; [NL4A, #519d]
P. h. incanescens Godman & Salvin, 1877
Nicaragua, Costa Rica, Panama Pierella incanescens Godman & Salvin, 1877; Proc. zool. Soc. Lond. 1877 (1) : 61, TL: Nicaragua,Costa Rica,Panama
Pierella incanescens costarica Niepelt, 1927; , TL: Costa Rica
= ; [NL4A, #519c]
Pierella incanescens ; Godman & Salvin, [1880], Biol. centr.-amer., Lep. Rhop. 1: 68, 3: pl. 6, f. 5-6; Godman & Salvin, [1901], Biol. centr.-amer., Lep. Rhop. 2: 651
Pierella helvetia incanescens ; [BCR: 262, pl. 47]
Pierella helvina incanescens ; [NL4A, #519c]

Larva on Heliconia, Calathea [BCR]

P. h. hymettia Staudinger, [1886]
Colombia helvina hymettia Staudinger, [1886]; , TL: Rio San Juan, W.Colombia
Pierella incanescens werneri Hering & Hopp, 1925; , TL: [W. Colombia], Rio Micay
= Pierella helvina ocreata ; [EcuL]
= ; [NL4A, #519b]
Pierella helvina hymettia ; [NL4A, #519b]
P. h. pacifica Niepelt, 1924
Colombia helvina pacifica Niepelt, 1924; , TL: Colombia, "Pacifico, Süd Columbia"
= Pierella helvina ocreata ; [EcuL]
Pierello[sic] ocreata johnsoni Talbot, 1932; , TL: Colombia
= ; [NL4A, #519e]
Pierella pacifica ; [BOW: pl. 63, f. 5 (text only)]
Pierella helvina pacifica ; [NL4A, #519e]

Pierella hortona (Hewitson, 1854)
Ecuador, .... See [About maps] Haetera hortona Hewitson, 1854; , TL: Brazil, Ega
Pierella hortona, NSG Voucher Specimen [Wahlberg]
Pierella hortona ; [EBW]; [BOW: pl. 63, f. 8]; [EcuL]; [NL4A, #520]
Brazil(Amazonas), Peru Haetera hortensia C. & R. Felder, 1862; Wien. ent. Monats. 6 (4): 124, TL: "Rio Negro" [= Peru, San Martin]
= ; [NL4A, #520a]
Pierella hortona f. albopunctata Brown, 1948; , TL: Peru
= ; [NL4A, #520a]
Pierella hortona f. ocellata Smart, 1975; , TL: Peru
= ; [NL4A, #520a]
Pierella hortona hortona ; [NL4A, #520a]
P. h. albofasciata Rosenberg & Talbot, 1914
Peru, Bolivia Pierella albofasciata Rosenberg & Talbot, 1914; Trans. ent. Soc. Lond. 1913 (4) : 678, TL: Peru
Pierella albofaciata[sic] decepta Brown, 1948; , TL: Bolivia
= ; [NL4A, #520b]
Pierella hortona albofasciata ; [NL4A, #520b]
Peru Pierella hortona ssp. ; Lamas, MS, [NL4A, #520c]

Pierella hyalinus (Gmelin, [1790])
Surinam, Trinidad, Amazon. See [About maps] Papilio hyalinus Gmelin, [1790]; , TL: Surinam
Pierella hyalina ; [EcuL, (note)]
Pierella hyalinus ; [NL4A, #521]
Surinam, Trinidad Pieris dracontis Hübner, [1819]; , TL: Surinam
= ; [NL4A, #521a]
Pierella hyalinus fusimaculata Brown, 1948; , TL: Trinidad
= ; [NL4A, #521a]
Pierella dracontis ; [EBW]
Pierella hyalinus hyalinus ; [NL4A, #521a]
P. h. extincta Weymer, 1910
Brazil(Amazonas) hyalinus extincta Weymer, 1910; , TL: Brazil, Amazonas
Pierella hyalinus extincta ; [NL4A, #521b]
P. h. schmidti Constantino, 1995
Colombia hyalinus schmidti Constantino, 1995; , TL: Colombia
Pierella hyalinus schmidti ; [NL4A, #521c]
P. h. velezi Constantino, 1995
Colombia hyalinus velezi Constantino, 1995; , TL: Colombia
Pierella hyalinus velezi ; [NL4A, #521d]

Pierella hyceta (Hewitson, 1860)
Amazon, Bolivia. See [About maps] Haetera hyceta Hewitson, 1860; Ill. exot. Butts [4] (Haetera): [39], pl. [21], f. 1, TL: [Peru] Nauta, Amazons
Pierella hyceta, NSG Voucher Specimen [Wahlberg]
Pierella hyceta ; [EBW]; [BOW: pl. 63, f. 9]; [NL4A, #522]
Peru Pierella hyceta hyceta ; [NL4A, #522a]
P. h. ceryce (Hewitson, 1874)
Bolivia Pierella ceryce Hewitson, 1874; , TL: Bolivia
Pierella cercye[sic] ; [BOW: pl. 63, f. 9 (text only)]
Pierella ceryce ; [EcuL, (note)]
Pierella hyceta ceryce ; [NL4A, #522b]
P. h. latona (C. & R. Felder, 1867)
Colombia, Bolivia? hyceta latona (C. & R. Felder, 1867) (Haetera); , TL: Colombia
Pierella latona ; [BOW: pl. 63, f. 9 (text only)]
Pierella hyceta latona ; [EcuL]; [NL4A, #522c]
Peru Pierella hyceta ssp. ; Lamas, MS, [NL4A, #522d]
Peru Pierella hyceta ssp. ; Lamas, MS, [NL4A, #522e]

Pierella lamia (Sulzer, 1776)
Surinam, Colombia, Cayenne, Amazon. See [About maps] Papilio lamia Sulzer, 1776; , TL: America
Pierella lamia, NSG Voucher Specimen [Wahlberg]
Pierella lamia ; [NL4A, #523]
Surinam, French Guiana Papilio rhea Fabricius, 1775; (preocc. Papilio rhea Poda, 1761), TL: "Indiis et Caienne"
= ; [NL4A, #523a]
Papilio dyndimene Cramer, 1779; , TL: Surinam
= ; [NL4A, #523a]
Pierella luna ab. albina Oberthür, 1896; , TL: French Guiana
= ; [NL4A, #523a]
?Papilio fabricius Ebert, 1965; (unnec. repl. Papilio rhea Fabricius, 1775), TL: Cayenne
f. fabriciana Ebert, 1965; (repl. nema), TL: "Indiis"
= ; [NL4A, #523a]
Pierella rhea ; [BOW: pl. 64, f. 1]
Pierella lamia lamia ; [NL4A, #523a]
P. l. chalybaea Godman, 1905
Brazil(Mato Grosso), Colombia lamia chalybaea Godman, 1905; , TL: Brazil, Chapada [Mato Grosso]
Haetera lamia f. columbina Krüger, 1925; , TL: Colombia
= ; [NL4A, #523c]
Pierella lamia chalybaea ; [EcuL]; [NL4A, #523c]
P. l. boliviana Brown, 1948
Bolivia lamia boliviana Brown, 1948; , TL: Bolivia
Pierella lamia boliviana ; [NL4A, #523b]

Pierella lena (Linnaeus, 1767)
Surinam, Guianas, .... See [About maps] Papilio lena Linnaeus, 1767; , TL: "America meridionali"
Pierella lena, NSG Voucher Specimen [Wahlberg]
Papilio lena ; Cramer, [1779], Uitl. Kapellen 3 (17-21): pl. 198
Haetera lena ; Hewitson, 1860, Ill. exot. Butts [4] (Haetera): [39], pl. [21], f. 2
Pierella lena ; [NL4A, #524]; [BOW: pl. 63, f. 10]; [EBW]
[lena-1.jpg] 500x397 (~70Kb) Venezuela, Parai Tepui near Mount Roraima in the Gran Sabana, July 2002, Photo © Simon Coombes

Papilio sectator Meerburgh, 1775
= ; [NL4A, #524a]
Pierella lena lena ; [NL4A, #524a]
P. l. brasiliensis C. & R. Felder, 1862
Peru, Bolivia, Brazil(Amazonas) Haetera lena var. brasiliensis C. & R. Felder, 1862; Wien. ent. Monats. 6 (4): 125, TL: "Rio Negro" [= Peru, San Martin]
Pierella lena glaucolena Weymer, 1910; , TL: Bolivia
= ; [NL4A, #524b]
Pierella lena f. melanosa Zikán & Wygodzinsky, 1948; (nom. nud.), TL: Brazil, Amazonas
= ; [NL4A, #524b]
f. obsoleta Brown, 1948; , TL: Brazil, Amazonas
= ; [NL4A, #524b]
Pierella lena browni Forster, 1964; , TL: Brazil, Amazonas
= ; [NL4A, #524b]
Pierella lena brasiliensis ; [EcuL]; [NL4A, #524b]

Pierella lucia Weymer, 1885
Peru. See [About maps] Pierella lucia Weymer, 1885; , TL: Peru, Pebas
Pierella astyoche var. albomaculata Staudinger, 1887; , TL: "Upper Amazon"
= ; [NL4A, #525]
Pierella lucia, NSG Voucher Specimen [Wahlberg]
Pierella astyoche lucia ;
Pierella lucia ; [EcuL]; [NL4A, #525]

Pierella luna (Fabricius, 1793)
-. See [About maps] Papilio luna Fabricius, 1793; Ent. Syst. 3 (1): 109, TL: "Surinam"
Haetera luna ; Hewitson, 1860, Ill. exot. Butts [4] (Haetera): [39], pl. [21], f. 3
Pierella luna ; Godman & Salvin, [1880], Biol. centr.-amer., Lep. Rhop. 1: 66; Godman & Salvin, [1901], Biol. centr.-amer., Lep. Rhop. 2: 650; [NL4A, #526]
[luna-1m.jpg] 539x547 (~44Kb) male Belize, Central America, Photo © John Shuey

Nicaragua - Colombia f. rubra Brown, 1948; , TL: Colombia
= ; [NL4A, #526a]
Pierella luna luna ; [BCR: 262, pl. 47]; [NL4A, #526a]

Larva on Heliconia [BCR]

P. l. pallida (Salvin & Godman, 1868)
Nicaragua Hetaera[sic] pallida Salvin & Godman, 1868; Ann. Mag. nat. Hist. (4) 2 (8) : 142, TL: Nicaragua
Pierella luna pallida ; [NL4A, #526c]
P. l. rubecula Salvin & Godman, 1868
Mexico, Guatemala, Honduras Pierella rubecula Salvin & Godman, 1868; Ann. Mag. nat. Hist. (4) 2 (8) : 142, TL: Guatemala
Haetera heracles Boisduval, 1870; , TL: Honduras
= ; [NL4A, #526d]
Pierella luna rubecula, Listing of Mexican Butterflies [ILMB]
Pierella luna rubecula, Butterflies of America [BOA]
Pierella luna heracles ;
Pierella rubecula ; Godman & Salvin, [1880], Biol. centr.-amer., Lep. Rhop. 1: 67, 3: pl. 6, f. 10-11; Godman & Salvin, [1901], Biol. centr.-amer., Lep. Rhop. 2: 651
Pierella luna rubecula ; [NL4A, #526d]
P. l. lesbia Staudinger, 1887
Colombia Pierella lesbia Staudinger, 1887; , TL: Rio San Juan, W. Colombia
Pierella luna lesbia ; [EcuL]; [NL4A, #526b]

Pierella nereis (Drury, [1782])
S.Brazil(Rio de Janeiro). See [About maps] Papilio nereis Drury, [1782]; Illust. Nat. Hist. Exot. Insects 3: index, : 48, pl. 35, f. 2-3, TL: Brazil, Rio de Janeiro
Pierella nereis, NSG Voucher Specimen [Wahlberg]
Pierella nereis ; [EBW]; [BOW: pl. 63, f. 11]; [NL4A, #527]


15.2.2013 (11)

References:
[About maps]
Warning! The maps are automatically generated from the textual information, and the process does not always produce acceptable result; See about maps for more info.
[BCA]
electronic; BIOLOGIA CENTRALI-AMERICANA
[BCR]
DeVries; The Butterflies of Costa Rica 1; Papilionidae, Pieridae, Nymphalidae
[BHL]
The; Biodiversity Heritage Library; and on; Internet Archive; Note: The linking to the content of Internet Archive is somewhat delicate -- notify me when links break!
[BOA]
Interactive Listing of American Butterflies; Listado Interactivo de la Mariposas Americanas; Butterflies of America
[BOW]
Lewis, 1974; Butterflies of the World; ISBN 0 245 52097 X
[EBW]
Smart, Paul, 1976; The Illustrated Encyclopedia of the Butterfly World; ISBN 0 600 31381 6
[EcuL]
Tommaso & Racheli, 2001; Fragmenta entomologica 33 (2): 213-380; An annotated list of Ecuadorian Butterflies (Lepidoptera: Papilionidae, Pieridae, Nymphalidae)
[ILMB]
Interactive Listing of Mexican Butterflies
[John Shuey]
e-mail: jshuey at tnc.org; Director of Conservation Science; Indiana Office of The Nature Conservancy; http://www.nature.org
[LAMAS-NEO]
Gerardo Lamas; Bibliography of butterflies; An Annocated Bibliography of the Neotropical Butterflies and Skipper (Lepididoptera: Papilionoidea and Hesperioidea); Revised Electronic Edition (2007)
[NL4A]
Gerardo Lamas, 2004; Atlas of Neotropical Lepidoptera; Checklist: Part 4A; Hesperioidea - Papilionoidea
[Simon Coombes]
1 Park Street, Plymouth, PL3 4BL, UK; 01752 607854; e-mail: Simon Coombes; http://www.butterfly-guide.co.uk
[Wahlberg]
Niklas Wahlberg; The NSG's voucher specimen database; Nymphalidae Systematics Group


Some related literature:
Cramer, [1779]
Uitlandsche Kapellen (Papillons exotiques) Uitl. Kapellen 3 (17-21): 1-104, pl. 193-252 (1779), (22): 105-128, pl. 253-264 ([1780]), (23-24): 129-176, pl. 265-288 (1780)
Doubleday, [1849]; Westwood, [1851]
The genera of diurnal Lepidoptera, comprising their generic characters, a notice of their habitats and transformations, and a catalogue of the species of each genus; illustrated with 86 plates by W. C. Hewitson Gen. diurn. Lep. (1): 1-18, pl. A, 1-4 (1-2) (1846), (1): 19-132, pl. 4*, 5-25, 28 (3-14) (1847), (1): 133-200, pl. 1*, 26, 27, 29, 31-44 (15-23) (1848), (1): 201-242, pl. 30, 45-52, 56-58, 60-62, 64 (24-31) (1849), (1): 243-250 [?], (2): 243-326, pl. 53-55, 63, 65, 66 (32-38) (1850), (2): 327-466, pl. 59, 67-77 (39-50) (1851), (2): 467-534, 143, 144, pl. 78-80 (Index) (51-54) (1852)
Drury, [1782]
Illustrations of natural history; wherein are exhibited ... Illust. Nat. Hist. Exot. Insects 3 : 1-76, pl. 1-50
C. & R. Felder, 1862
Specimen faunae lepidopterologicae riparum fluminis Negro superioris in Brasilia septentrionali Wien. ent. Monats. 6 (3): 65-80, (4): 109-126, (6): 175-192, (7): 229-235
Godman & Salvin, 1868; Salvin & Godman, 1868
On some new species of diurnal Lepidoptera from South America Ann. Mag. nat. Hist. (4) 2 (8) : 141-152
Godman & Salvin, 1877
Description of twelwe new species and a new genus of Rhopalocera from Central America Proc. zool. Soc. Lond. 1877 (1) : 60-64
Godman & Salvin, [1880]
Biologia Centrali-Americana. Rhopalocera. Vol. 1. (1879-1886) Biol. centr.-amer., Lep. Rhop. 1: 1-487, 3: pl. 1-24,24a,25-112
Godman & Salvin, [1901]
Biologia Centrali-Americana. Rhopalocera. (1887-1901) Biol. centr.-amer., Lep. Rhop. 2: 1-782, 3: pl. 1-112
Herrich-Schäffer, 1865
Lepidopterorum index systematicus CorrespBl. zool.-min. Ver. Regensburg 18 (11/12): 175-181 (1865), 19 (5): 63-76, 19 (6): 84-92, 19 (7): 100-108
Hewitson, 1860
Illustrations of new species of exotic Butterflies selected chiefly from the collections of W. Wilson Saunders and William C. Hewitson Ill. exot. Butts [4] : (Systematic Index) IV, (Clerome): [1-2], pl. [1] (1863), (Drusilla & Hyantis): [3-4], pl. [2] (1862), (Dasyophthalma & Thaumantis): [5-6], pl. [3] (1862), (Morpho I): [7-8], pl. [4] (1856), (Pavonia I-II): [9-10], pl. [5-6] (1877), (Opsiphanes): [11-12], pl. [7] (1873), (Corades): [13-14], pl. [8] (1863), (Pronophila): [15-16], pl. [9] (1857), (Pronophila II): [17-18], pl. [10] (1860), (Pronophila III-IV): [19-22], pl. [11-12] (1868), (Pronophila V): [23-24], pl. [13] (1871), (Pronophila VI-VII): [25-28], pl. [14-15] (1872), (Pronophila VIII-IX): [29-32], pl. [16-17] (1874), (Debis I): [33-34], pl. [18] (1862), (Debis II-III): [35-38], pl. [19-20] (1863), (Haetera): [39-40], pl. [21] (1860), (Daedalma): [41-42], pl. [22] (1858), (Euptychia & Ragadia): [43-44], pl. [23] (1862), (Hypocista): [45-46], pl. [24] (1863), (Mycalesis I-IV): [47-54], pl. [25-28] (1862), (Mycalesis V-VI): [55-58], pl. [29-30] (1864), (Mycalesis VII-VIII): [59-62], pl. [31-32] (1866), (Mycalesis IX): [63-64], pl. [33] (1873), (Mycalesis & Idiomorphus): [65-66], pl. [34] (1877), (Idiomorphus): [67-68], pl. [35] (1865), (Melanitis): [69-70], pl. [36] (1863), (Taxila): [71-72], pl. [37] (1861), (Taxila II): [73-74], pl. [38] (1862), (Sospita): [75-76], pl. [39] (1861), (Sospita II): [77-78], pl. [40] (1862), (Dodona Sospita): [79-80], pl. [41] (1866), (Mesosemia I): [81-82], pl. [42] (1857), (Mesosemia II-III): [83-86], pl. [43-44] (1858), (Mesosemia IV-VI): [87-92], pl. [45-47] (1859), (Mesosemia VII-VIII): [93-96], pl. [48-49] (1860), (Mesosemia IX-X): [97-100], pl. [50-51] (1870), (Mesosemia XI): [101-102], pl. [52] (1871), (Mesosemia XII): [103-104], pl. [53] (1873), (Eurygona I-II): [105-108], pl. [54-55] (1853), (Eurygona III-IV): [109-112], pl. [56-57] (1855), (Eurygona V-VII): [113-118], pl. [58-60] (1856), (Eurygona VIII): [119-120], pl. [61] (1860), (Eurygona IX): [121-122], pl. [62] (1870), (Eurygona X): [123-124], pl. [63] (1872)
Rosenberg & Talbot, 1914
New South American butterflies Trans. ent. Soc. Lond. 1913 (4) : 671-682


If you have corrections, comments or information to add into these pages, just send mail to Markku Savela
Keep in mind that the taxonomic information is copied from various sources, and may include many inaccuracies. Expert help is welcome.