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Photo      Graylings, Ringlets, Browns, Heaths
Satyrinae
Heinäperhoset Satyrider, Gräsfjärilar Graylings, Ringlets, Browns, Heaths
Satyrinae, NSG Voucher Specimen [Wahlberg]
Satyrinae, Butterflies and Moths of North America [BAMONA]
 Satyridae ;
Agapedidae ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 27 (incl.)
Satyrinae (Nymphalidae) ; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 171; [Wahlberg]; [NL4A, 205]

The higher classification here is not independent work, but is mostly based on Niklas Wahlberg's ongoing work, most recent alignment is based on November 2005 version. See [Wahlberg].

Tribe Haeterini
Haeterinae (Satyridae) ;
Haeterini (Satyrinae) ; [Wahlberg]; [NL4A, 205]
Cithaerias Hübner, [1819]
Dulcedo d'Almeida, 1951
Haetera Fabricius, 1807
Pierella Herrich-Schäffer, 1865
Pseudohaetera Brown, 1943
Tribe Melanitini
Melanitini (Biinae) ; (Satyridae)
Melanitini (Satyrinae) ; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 171; [Wahlberg]; [AfrL]
Cyllogenes Butler, 1868 Evening Browns
Melanitis Fabricius, 1807 Evening Browns
Parantirrhoea Wood-Mason, 1881
Gnophodes Doubleday, [1849] Evening Browns
Manataria Kirby, [1904]
Aeropetes Billberg, 1820
Paralethe van Son, 1955
Tribe Elymniini
Papurikot Browns, ...
Elymniinae (Satyridae) ;
Pararginae (Satyridae) ;
Elymniini (Satyrinae) ; [Wahlberg]; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 173; [AfrL]; [NL4A, 206]
Subtribe Lethina
Lethini (Pararginae) ;
Lethina (Elymniini) ; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 176; [Wahlberg]
Parargina (Elymniini) ; [NL4A, 206 (incl.)]
Satyrites Scudder, 1872
Satyrites Scudder, 1872; (Fossil)
Satyrites (Lethina) ; [Wahlberg]
Pseudoneorina Nel & Descimon, 1986
Pseudoneorina Nel & Descimon, 1986; (Fossil)
Pseudoneorina (Lethina) ; [Wahlberg]
Neorinopsis Butler, 1873
Neorinopsis Butler, 1873; (Fossil)
Neorinopsis (Lethina) ; [Wahlberg]
Aphysoneura Karsch, 1894 Painted Ringlets
Lethe Hübner, [1819] Treebrowns
Enodia Hübner, [1819]
Satyrodes Scudder, 1875
Neope Moore, [1866]
Ninguta Moore, [1892]
Ptychandra C. & R. Felder, 1861
Mandarinia Leech, [1892]
Chonala Moore, 1893
Kirinia Moore, 1893
fi Lasiommata Westwood, 1841
fi Lopinga Moore, 1893
Orinoma Gray, 1846
fi Pararge Hübner, [1819]
Rhaphicera Butler, 1867
Tatinga Moore, 1893
Nosea Koiwaya, 1993
Subtribe Zetherina
Zetherini (Pararginae) ;
Zetherina (Elymniini) ; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 175; [Wahlberg]
Callarge Leech, [1892]
Penthema Doubleday, [1848]
Zethera Felder, 1861
Ethope Moore, [1866]
Neorina Westwood, [1850] Owls
Subtribe Elymniina
Elymniini (Pararginae) ;
Elymniina (Elymniini) ; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 174; [Wahlberg]; [NL4A, 206]
Elymnias Hübner, 1818 Palmflies
Subtribe Mycalesina
Mycalesini (Pararginae) ;
Mycalesina (Elymniini) ; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 176; [Wahlberg]
Bicyclus Kirby, 1871 Bush Browns
Bletogona C. & R. Felder, [1867]
Mycalesis Hübner, 1818 Bushbrowns
Lohora Moore, 1880
Nirvanopsis Vane-Wright in Vane-Wright & de Jong, 2003
Hallelesis Condamin, 1961
Heteropsis Westwood, 1850
Orsotriaena Wallengren, 1858
Tribe Eritini
Eritinae (Satyridae) ;
Eritini (Satyrinae) ; [Wahlberg]
Coelites Westwood, [1850]
Erites Westwood, [1851]
Tribe Ragadiini
Ragadiinae (Satyridae) ;
Ragadiini (Satyrinae) ; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 181; [Wahlberg]
Acrophtalmia C. & R. Felder, 1861 pale ringlets
Acropolis Hemming, 1934
Ragadia Westwood, [1851]
Tribe Satyrini
Heinäperhoset, Nokiperhoset, Kylmäperhoset Graylings, Ringlets
Satyrinae (Satyridae) ;
Satyrini (Satyrinae) ; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 182; [NL4A, 206]
Subtribe Coenonymphina
Niittyperhoset Heaths
Coenonymphinae (Satyridae) ;
Coenonymphini ; [MRS, 403]
Coenonymphina (Satyrini) ; [Wahlberg]; [NL4A, 223]
fi Coenonympha Hübner, [1819]
Lyela Swinhoe, 1908
Triphysa Zeller, 1850
Sinonympha Lee, 1974
Subtribe Dirina
Dirini (Satyrinae) ;
Dirina (Satyrini) ; [Wahlberg]; [AfrL]
Tarsocera Butler, [1899]
Dingana van Son, 1955
Dira Hübner, [1819]
Torynesis Butler, [1899]
Subtribe Erebiina
Erebini (Satyrinae) ;
Erebiini ; [MRS, 406]
Erebiina (Satyrini) ; [Wahlberg]; [NL4A, 215]
fi Erebia Dalman, 1816 Nokiperhoset Arguses
Diaphanos Adams & Bernard, 1981
Ianussiusa Pyrcz & Viloria 2004
Idioneurula Strand, 1932
Manerebia Staudinger, 1897
Neomaniola Hayward, 1949
Stuardosatyrus Herrera & Etcheverry, 1965
Sabatoga Staudinger, 1897
Tamania Pyrcz, 1995
Subtribe Euptychiina
Euptychini (Satyrinae) ;
Euptychiina (Satyrini) ; [EcuL]; [Wahlberg]; [NL4A, 217]
Oressinoma Doubleday, [1849]
Palaeonympha Butler, 1871
Archeuptychia Forster, 1964
Caenoptychia Le Cerf, 1919
Capronnieria Forster, 1964
Cercyeuptychia Miller & Emmel, 1971
Chloreuptychia Forster, 1964
Cissia Doubleday, 1848
Coeruleotaygetis Forster, 1964
Cyllopsis R. Felder, 1869
Erichthodes Forster, 1964
Euptychia Hübner, 1818
Cepheuptychia Forster, 1964
Pharneuptychia Forster, 1964
Caeruleuptychia Forster, 1964
Euptychoides Forster, 1964
Forsterinaria Gray, 1973
Godartiana Forster, 1964
Harjesia Forster, 1964
Hermeuptychia Forster, 1964
Magneuptychia Forster, 1964
Megeuptychia Forster, 1964
Megisto Hübner, [1819]
Moneuptychia Forster, 1964
Neonympha Hübner, 1818
Paramacera Butler, 1868
Paryphthimoides Forster, 1964
Pindis R. Felder, 1869
Praefaunula Forster, 1964
Pseudeuptychia Forster, 1964
Rareuptychia Forster, 1964
Splendeuptychia Forster, 1964
Taydebis Freitas, 2003
Taygetis Hübner, [1819]
Satyrotaygetis Forster, 1964
Pseudodebis Forster, 1964
Taygetina Forster, 1964
Parataygetis Forster, 1964
Pareuptychia Forster, 1964
Posttaygetis Forster, 1964
Taygetomorpha Miller, 2004
Yphthimoides Forster, 1964
Zischkaia Forster, 1964
Subtribe ?
Setodocis Billberg, 1820
Subtribe Hypocystina
Hypocystini (Satyrinae) ;
Hypocystina (Satyrini) ; [Wahlberg]; [NL4A, 216]
Argynnina Butler, 1867
Argyronympha Mathew, 1886
Argyrophenga Doubleday, 1845
Dodonidia Butler, 1884
Erebiola Fereday, 1879
Erycinidia Rothschild & Jordan, 1905
Geitoneura Butler, 1867
Harsiesis Fruhstorfer, 1911
Heteronympha Wallengren, 1858
Hyalodia Jordan, 1924
Hypocysta Westwood, [1851]
Lamprolenis Godman & Salvin, [1881]
Nesoxenica Waterhouse & Lyell, 1914
Oreixenica Waterhouse & Lyell, 1914
Paratisiphone Watkins, 1928
Percnodaimon Butler, 1876
Platypthima Rothschild & Jordan, 1905
Altiapa Parsons, 1986
Tisiphone Hübner, [1819]
Zipaetis Hewitson, 1863 Catseyes
 
Argyrophorus Blanchard, 1852
Auca Hayward, 1953
Chillanella Herrera, 1966
Cosmosatyrus C. & R. Felder, [1867]
Elina Blanchard, 1852
Etcheverrius Herrera, 1965
Faunula C. & R. Felder, [1867]
Haywardella Herrera, 1966
Homoeonympha C. & R. Felder, [1867]
Nelia Hayward, 1953
Neomaenas Wallengren, 1858
Neosatyrus Wallengren, 1858
Palmaris Herrera, 1965
Pampasatyrus Hayward, 1953
Pamperis Heimlich, 1959
Punargentus Heimlich, 1963
Quilaphoetosus Herrera, 1966
Spinantenna Hayward, 1953
Tetraphlebia C. & R. Felder, [1867]
Subtribe Maniolina
Häränsilmat, Tesmaperhonen Meadow Browns, Ringlet
Maniolinae (Satyridae) ;
Maniolina (Satyrini) ; [Wahlberg]
fi Aphantopus Wallengren, 1853
Cercyonis Scudder, 1875
fi Hyponephele Muschamp, 1915 Meadowbrowns
fi Maniola Schrank, 1801 Meadowbrowns
Proterebia Roos & Arnscheid, 1980
Pyronia Hübner, [1819]
Subtribe Melanargiina
Ruutuperhoset Marbled Whites
Melanargiinae (Satyridae) ;
Melanargiina (Satyrini) ; [Wahlberg]
fi Melanargia Meigen, 1828
Subtribe Pronophilina
Pronophilini (Satyrinae) ;
Pronophilina (Satyrini) ; [Wahlberg]; [NL4A, 206]
Altopedaliodes Forster, 1964
Apexacuta Pyrcz, 2004
Arhuaco Adams & Bernard, 1977
Calisto Hübner, 1823
Cheimas Thieme, 1907
Corades Hewitson, 1849
Daedalma Hewitson, 1858
Dangond Adams & Bernard, 1979
Drucina Butler, 1872
Druphila Pyrcz, 2004
Eretris Thieme, 1905
Eteona Doubleday, 1848
Foetterleia Viloria, 2004
Junea Hemming, 1964
Lasiophila C. & R. Felder, 1859
Lymanopoda Westwood, [1851]
Mygona Thieme, 1907
Neopedaliodes Viloria, Miller & Miller, 2004
Oxeoschistus Butler, 1867
Panyapedaliodes Forster, 1964
Paramo Adams & Bernard, 1977
Parapedaliodes Forster, 1964
Pedaliodes Butler, 1867
Praepedaliodes Forster, 1964
Antopedaliodes Forster, 1964
Pherepedaliodes Forster, 1964
Physcopedaliodes Forster, 1964
Corderopedaliodes Forster, 1964
Praepronophila Forster, 1964
Proboscis Thieme, 1907
Pronophila Doubleday, [1849]
Protopedaliodes Viloria & Pyrcz, 1994
Pseudomaniola Röber, [1889]
Punapedaliodes Forster, 1964
Redonda Adams & Bernard, 1981
Sierrasteroma Adams & Bernard, 1977
Steremnia Thieme, 1905
Steroma Westwood, [1850]
Steromapedaliodes Forster, 1964
Thiemeia Weymer, 1912
Subtribe Satyrina
Satyrini (Satyrinae) ;
Ypthimina (Satyrini) ; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 182
Satyrina (Satyrini) ; [NL4A, 224]; [Wahlberg]
Arethusana de Lesse, 1951
Argestina Riley, 1923
Aulocera Butler, 1867 Banded Satyrs
Berberia de Lesse, 1951
Boeberia Prout, 1901
Brintesia Fruhstorfer, 1911
Callerebia Butler, 1867
Cassionympha van Son, 1955
Chazara Moore, 1893
Coenyra Hewitson, 1865
Coenyropsis van Son, 1958
Davidina Oberthür, 1879
fi Hipparchia Fabricius, 1807
Kanetisa Moore, 1893
Karanasa Moore, 1893
Loxerebia Watkins, 1925
Mashuna van Son, 1955
Melampias Hübner, [1819]
Minois Hübner, [1819]
Neita van Son, 1955
Neocoenyra Butler, [1886]
Neominois Scudder, 1875
fi Oeneis Hübner, [1819]
Paralasa Moore, 1893
Paroeneis Moore, 1893
Physcaeneura Wallengren, 1857
Pseudochazara de Lesse, 1951
Pseudonympha Wallengren, 1857
Satyrus Latreille, 1810
Strabena Mabille, 1877
Stygionympha van Son, 1955
Ypthima Hübner, 1818 Rings Ringlets
Austroypthima Holloway, 1974
Ypthimomorpha van Son, 1955
Subtribe ?
? (Satyrina) ; [NL4A, 224]
Amphidecta Butler, 1867
Gyrocheilus Butler, 1867
Neorinella Martins Neto & al., 1993
Tribe ?
Dyndirus Capronnier, 1874
Dyndirus Capronnier, 1874; Ann. Soc. ent. Belg. 17 : 32 (nom. nud.)
Dyndirus (?) ; [NL4A, 224]; [Wahlberg]

Unknown or unplaced taxa
Euptychia albiplaga J. & W. Zikán, 1968; (nom. nud.), TL: Brazil, Rio de Janeiro; Minas Gerais
albiplaga ; [NL4A, 224]
Euptychia ambarina J. & W. Zikán, 1968; (nom. nud.), TL: Brazil, Rio de Janeiro
ambarina ; [NL4A, 224]
Euptychia apamela Hübner, [1819]; (nom. nud.)
apamela ; [NL4A, 224]
Euptychia chilensis Ménétriés, 1857; (nom. nud.), TL: Chile
chilensis ; [NL4A, 224]
Euptychia elisae J. & W. Zikán, 1968; (nom. nud.), TL: Brazil, Rio de Janeiro; Minas Gerais
elisae ; [NL4A, 224]
Euptychia furcifera J. & W. Zikán, 1968; (nom. nud.), TL: Brazil, Rio de Janeiro
furcifera ; [NL4A, 224]
Euptychia fractifasciata J. & W. Zikán, 1968; (nom. nud.), TL: Brazil, Minas Gerais
fractifasciata ; [NL4A, 224]
Euptychia herluga J. & W. Zikán, 1968; (nom. nud.), TL: Brazil, Minas Gerais
herluga ; [NL4A, 224]
Euptychia itatiayensis J. & W. Zikán, 1968; (nom. nud.), TL: Brazil, Rio de Janeiro
itatiayensis ; [NL4A, 224]
Satyrus janirula Ménétriés, 1857; (nom. nud.), TL: Chile
janirula ; [NL4A, 224]
Euptychia lioba J. & W. Zikán, 1968; (nom. nud.), TL: Brazil, Minas Gerais
lioba ; [NL4A, 224]
Euptychia mantiqueirensis J. & W. Zikán, 1968; (nom. nud.), TL: Brazil, Minas Gerais
mantiqueirensis ; [NL4A, 224]
Euptychia montana J. & W. Zikán, 1968; (nom. nud.), TL: Brazil, Minas Gerais
montana ; [NL4A, 224]
Dyndirus nerces Capronnier, 1874; (nom. nud.), TL: Brazil
nerces ; [NL4A, 224]
Euptychia ochripenne J. & W. Zikán, 1968; (nom. nud.), TL: Brazil, Minas Gerais
ochripenne ; [NL4A, 224]
Euptychia otomara J. & W. Zikán, 1968; (nom. nud.), TL: Brazil, Rio de Janeiro
otomara ; [NL4A, 224]
Euptychia punctatolineata J. & W. Zikán, 1968; (nom. nud.), TL: Brazil, Minas Gerais
punctatolineata ; [NL4A, 224]
Euptychia simillima J. & W. Zikán, 1968; (nom. nud.), TL: Brazil, Minas Gerais
simillima ; [NL4A, 224]
Euptychia stellifera J. & W. Zikán, 1968; (nom. nud.), TL: Brazil, Rio de Janeiro; Minas Gerais
stellifera ; [NL4A, 224]
Neonympha subina Capronnier, 1874; (nom. nud.), TL: Brazil
subina ; [NL4A, 224]
Euptychia virginiae J. & W. Zikán, 1968; (nom. nud.), TL: Brazil, Minas Gerais
virginiae ; [NL4A, 224]
Euptychia wilburga J. & W. Zikán, 1968; (nom. nud.), TL: Brazil, Minas Gerais
wilburga ; [NL4A, 224]
Euptychia wordhilda J. & W. Zikán, 1968; (nom. nud.), TL: Brazil, Rio de Janeiro; Minas Gerais
wordhilda ; [NL4A, 224]

15.4.2013 (2283)

References:
[AfrL]
Mark C. Williams; Checklist of Afrotropical Papilionoidea and Hesperoidea; Compiled by Mark C. Williams, 7th ed. (2008) (April 2007); Afrototropical Butterflies; e-mail: Mark C. Williams
[BAMONA]
Opler, Pavulaan, Stanford & Pogue; Butterflies and Moths of North America; Mountain Prairie Information Node
[EcuL]
Tommaso & Racheli, 2001; Fragmenta entomologica 33 (2): 213-380; An annotated list of Ecuadorian Butterflies (Lepidoptera: Papilionidae, Pieridae, Nymphalidae)
[MRS]
Chou Io (Ed.); Monographia Rhopalocerum Sinensium, 1-2
[NL4A]
Gerardo Lamas, 2004; Atlas of Neotropical Lepidoptera; Checklist: Part 4A; Hesperioidea - Papilionoidea
[Wahlberg]
Niklas Wahlberg; The NSG's voucher specimen database; Nymphalidae Systematics Group

If you have corrections, comments or information to add into these pages, just send mail to Markku Savela
Keep in mind that the taxonomic information is copied from various sources, and may include many inaccuracies. Expert help is welcome.