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Satyrinae
Heinäperhoset Satyrider, Gräsfjärilar Graylings, Ringlets, Browns, Heaths
Satyrinae, NSG Voucher Specimen [Wahlberg]
Satyrinae, Butterflies and Moths of North America [BAMONA]
Satyridae ;
Agapedidae ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 27 (incl.)
Satyrinae (Nymphalidae) ; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 171; [Wahlberg]; [NL4A, 205]

The higher classification here is not independent work, but is mostly based on Niklas Wahlberg's ongoing work, most recent alignment is based on November 2005 version. See [Wahlberg].

Tribe Haeterini
Haeterinae (Satyridae) ;
Haeterini (Satyrinae) ; [Wahlberg]; [NL4A, 205]
Cithaerias Hübner, [1819] — 5 spp.
Dulcedo d'Almeida, 1951 — 1 sp.
Haetera Fabricius, 1807 — 2 spp.
Pierella Herrich-Schäffer, 1865 — 11 spp.
Pseudohaetera Brown, 1943 — 2 spp.
Tribe Melanitini
Melanitini (Biinae) ; (Satyridae)
Melanitini (Satyrinae) ; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 171; [Wahlberg]; [AfrL]
Cyllogenes Butler, 1868 Evening Browns — 4 spp.
Melanitis Fabricius, 1807 Evening Browns — 12 spp.
Parantirrhoea Wood-Mason, 1881 — 1 sp.
Gnophodes Doubleday, [1849] Evening Browns — 3 spp.
Manataria Kirby, [1904] — 1 sp.
Aeropetes Billberg, 1820 — 1 sp.
Paralethe van Son, 1955 — 1 sp.
Tribe Elymniini
Papurikot Browns, ...
Elymniinae (Satyridae) ;
Pararginae (Satyridae) ;
Elymniini (Satyrinae) ; [Wahlberg]; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 173; [AfrL]; [NL4A, 206]
Subtribe Lethina
Lethini (Pararginae) ;
Lethina (Elymniini) ; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 176; [Wahlberg]
Parargina (Elymniini) ; [NL4A, 206 (incl.)]
Satyrites Scudder, 1872
Satyrites Scudder, 1872; (Fossil)
Satyrites (Lethina) ; [Wahlberg]
Pseudoneorina Nel & Descimon, 1986
Pseudoneorina Nel & Descimon, 1986; (Fossil)
Pseudoneorina (Lethina) ; [Wahlberg]
Neorinopsis Butler, 1873
Neorinopsis Butler, 1873; (Fossil)
Neorinopsis (Lethina) ; [Wahlberg]
Aphysoneura Karsch, 1894 Painted Ringlets — 2 spp.
Lethe Hübner, [1819] Treebrowns — 114 spp.
Enodia Hübner, [1819] — 3 spp.
Satyrodes Scudder, 1875 — 2 spp.
Neope Moore, [1866] — 20 spp.
Ninguta Moore, [1892] — 1 sp.
Ptychandra C. & R. Felder, 1861 — 9 spp.
Mandarinia Leech, [1892] — 1 sp.
Chonala Moore, 1893 — 3 spp.
Kirinia Moore, 1893 — 5 spp.
fi Lasiommata Westwood, 1841 — 16 spp.
fi Lopinga Moore, 1893 — 5 spp.
Orinoma Gray, 1846 — 2 spp.
fi Pararge Hübner, [1819] — 3 spp.
Rhaphicera Butler, 1867 — 3 spp.
Tatinga Moore, 1893 — 1 sp.
Nosea Koiwaya, 1993 — 1 sp.
Subtribe Zetherina
Zetherini (Pararginae) ;
Zetherina (Elymniini) ; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 175; [Wahlberg]
Callarge Leech, [1892] — 2 spp.
Penthema Doubleday, [1848] — 5 spp.
Zethera Felder, 1861 — 6 spp.
Ethope Moore, [1866] — 4 spp.
Neorina Westwood, [1850] Owls — 5 spp.
Subtribe Elymniina
Elymniini (Pararginae) ;
Elymniina (Elymniini) ; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 174; [Wahlberg]; [NL4A, 206]
Elymnias Hübner, 1818 Palmflies — 45 spp.
Subtribe Mycalesina
Mycalesini (Pararginae) ;
Mycalesina (Elymniini) ; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 176; [Wahlberg]
Bicyclus Kirby, 1871 Bush Browns — 84 spp.
Bletogona C. & R. Felder, [1867] — 2 spp.
Mycalesis Hübner, 1818 Bushbrowns — 97 spp.
Lohora Moore, 1880 — 17 spp.
Nirvanopsis Vane-Wright in Vane-Wright & de Jong, 2003 — 1 sp.
Hallelesis Condamin, 1961 — 2 spp.
Heteropsis Westwood, 1850 — 60 spp.
Orsotriaena Wallengren, 1858 — 2 spp.
Tribe Eritini
Eritinae (Satyridae) ;
Eritini (Satyrinae) ; [Wahlberg]
Coelites Westwood, [1850] — 3 spp.
Erites Westwood, [1851] — 5 spp.
Tribe Ragadiini
Ragadiinae (Satyridae) ;
Ragadiini (Satyrinae) ; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 181; [Wahlberg]
Acrophtalmia C. & R. Felder, 1861 pale ringlets — 3 spp.
Acropolis Hemming, 1934 — 1 sp.
Ragadia Westwood, [1851] — 9 spp.
Tribe Satyrini
Heinäperhoset, Nokiperhoset, Kylmäperhoset Graylings, Ringlets
Satyrinae (Satyridae) ;
Satyrini (Satyrinae) ; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 182; [NL4A, 206]
Subtribe Coenonymphina
Niittyperhoset Heaths
Coenonymphinae (Satyridae) ;
Coenonymphini ; [MRS, 403]
Coenonymphina (Satyrini) ; [Wahlberg]; [NL4A, 223]
fi Coenonympha Hübner, [1819] — 39 spp.
Lyela Swinhoe, 1908 — 3 spp.
Triphysa Zeller, 1850 — 2 spp.
Sinonympha Lee, 1974 — 1 sp.
Subtribe Dirina
Dirini (Satyrinae) ;
Dirina (Satyrini) ; [Wahlberg]; [AfrL]
Tarsocera Butler, [1899] — 7 spp.
Dingana van Son, 1955 — 10 spp.
Dira Hübner, [1819] — 4 spp.
Torynesis Butler, [1899] — 5 spp.
Subtribe Erebiina
Erebini (Satyrinae) ;
Erebiini ; [MRS, 406]
Erebiina (Satyrini) ; [Wahlberg]; [NL4A, 215]
fi Erebia Dalman, 1816 Nokiperhoset Arguses — 98 spp.
Diaphanos Adams & Bernard, 1981 — 3 spp.
Ianussiusa Pyrcz & Viloria 2004 — 1 sp.
Idioneurula Strand, 1932 — 1 sp.
Manerebia Staudinger, 1897 — 37 spp.
Neomaniola Hayward, 1949 — 1 sp.
Stuardosatyrus Herrera & Etcheverry, 1965 — 1 sp.
Sabatoga Staudinger, 1897 — 1 sp.
Tamania Pyrcz, 1995 — 1 sp.
Subtribe Euptychiina
Euptychini (Satyrinae) ;
Euptychiina (Satyrini) ; [EcuL]; [Wahlberg]; [NL4A, 217]
Oressinoma Doubleday, [1849] — 2 spp.
Palaeonympha Butler, 1871 — 1 sp.
Archeuptychia Forster, 1964 — 1 sp.
Caenoptychia Le Cerf, 1919 — 1 sp.
Capronnieria Forster, 1964 — 1 sp.
Cercyeuptychia Miller & Emmel, 1971 — 1 sp.
Chloreuptychia Forster, 1964 — 10 spp.
Cissia Doubleday, 1848 — 15 spp.
Coeruleotaygetis Forster, 1964 — 1 sp.
Cyllopsis R. Felder, 1869 — 29 spp.
Erichthodes Forster, 1964 — 5 spp.
Euptychia Hübner, 1818 — 15 spp.
Cepheuptychia Forster, 1964 — 4 spp.
Pharneuptychia Forster, 1964 — 6 spp.
Caeruleuptychia Forster, 1964 — 19 spp.
Euptychoides Forster, 1964 — 10 spp.
Forsterinaria Gray, 1973 — 23 spp.
Godartiana Forster, 1964 — 2 spp.
Harjesia Forster, 1964 — 5 spp.
Hermeuptychia Forster, 1964 — 8 spp.
Magneuptychia Forster, 1964 — 29 spp.
Megeuptychia Forster, 1964 — 2 spp.
Megisto Hübner, [1819] — 2 spp.
Moneuptychia Forster, 1964 — 5 spp.
Neonympha Hübner, 1818 — 5 spp.
Paramacera Butler, 1868 — 4 spp.
Paryphthimoides Forster, 1964 — 13 spp.
Pindis R. Felder, 1869 — 2 spp.
Praefaunula Forster, 1964 — 3 spp.
Pseudeuptychia Forster, 1964 — 2 spp.
Rareuptychia Forster, 1964 — 1 sp.
Splendeuptychia Forster, 1964 — 23 spp.
Taydebis Freitas, 2003 — 1 sp.
Taygetis Hübner, [1819] — 27 spp.
Satyrotaygetis Forster, 1964 — 1 sp.
Pseudodebis Forster, 1964 — 5 spp.
Taygetina Forster, 1964 — 1 sp.
Parataygetis Forster, 1964 — 2 spp.
Pareuptychia Forster, 1964 — 7 spp.
Posttaygetis Forster, 1964 — 1 sp.
Taygetomorpha Miller, 2004 — 2 spp.
Yphthimoides Forster, 1964 — 23 spp.
Zischkaia Forster, 1964 — 4 spp.
Subtribe ?
Setodocis Billberg, 1820 — 1 sp.
Subtribe Hypocystina
Hypocystini (Satyrinae) ;
Hypocystina (Satyrini) ; [Wahlberg]; [NL4A, 216]
Argynnina Butler, 1867 — 2 spp.
Argyronympha Mathew, 1886 — 5 spp.
Argyrophenga Doubleday, 1845 — 3 spp.
Dodonidia Butler, 1884 — 1 sp.
Erebiola Fereday, 1879 — 1 sp.
Erycinidia Rothschild & Jordan, 1905 — 6 spp.
Geitoneura Butler, 1867 — 3 spp.
Harsiesis Fruhstorfer, 1911 — 3 spp.
Heteronympha Wallengren, 1858 — 7 spp.
Hyalodia Jordan, 1924 — 1 sp.
Hypocysta Westwood, [1851] — 12 spp.
Lamprolenis Godman & Salvin, [1881] — 1 sp.
Nesoxenica Waterhouse & Lyell, 1914 — 1 sp.
Oreixenica Waterhouse & Lyell, 1914 — 6 spp.
Paratisiphone Watkins, 1928 — 1 sp.
Percnodaimon Butler, 1876 — 1 sp.
Platypthima Rothschild & Jordan, 1905 — 8 spp.
Altiapa Parsons, 1986 — 4 spp.
Tisiphone Hübner, [1819] — 2 spp.
Zipaetis Hewitson, 1863 Catseyes — 3 spp.
 
Argyrophorus Blanchard, 1852 — 1 sp.
Auca Hayward, 1953 — 4 spp.
Chillanella Herrera, 1966 — 1 sp.
Cosmosatyrus C. & R. Felder, [1867] — 1 sp.
Elina Blanchard, 1852 — 2 spp.
Etcheverrius Herrera, 1965 — 2 spp.
Faunula C. & R. Felder, [1867] — 2 spp.
Haywardella Herrera, 1966 — 1 sp.
Homoeonympha C. & R. Felder, [1867] — 4 spp.
Nelia Hayward, 1953 — 2 spp.
Neomaenas Wallengren, 1858 — 8 spp.
Neosatyrus Wallengren, 1858 — 1 sp.
Palmaris Herrera, 1965 — 4 spp.
Pampasatyrus Hayward, 1953 — 9 spp.
Pamperis Heimlich, 1959 — 1 sp.
Punargentus Heimlich, 1963 — 2 spp.
Quilaphoetosus Herrera, 1966 — 2 spp.
Spinantenna Hayward, 1953 — 1 sp.
Tetraphlebia C. & R. Felder, [1867] — 1 sp.
Subtribe Maniolina
Häränsilmat, Tesmaperhonen Meadow Browns, Ringlet
Maniolinae (Satyridae) ;
Maniolina (Satyrini) ; [Wahlberg]
fi Aphantopus Wallengren, 1853 — 3 spp.
Cercyonis Scudder, 1875 — 4 spp.
fi Hyponephele Muschamp, 1915 Meadowbrowns — 62 spp.
fi Maniola Schrank, 1801 Meadowbrowns — 7 spp.
Proterebia Roos & Arnscheid, 1980 — 1 sp.
Pyronia Hübner, [1819] — 5 spp.
Subtribe Melanargiina
Ruutuperhoset Marbled Whites
Melanargiinae (Satyridae) ;
Melanargiina (Satyrini) ; [Wahlberg]
fi Melanargia Meigen, 1828 — 24 spp.
Subtribe Pronophilina
Pronophilini (Satyrinae) ;
Pronophilina (Satyrini) ; [Wahlberg]; [NL4A, 206]
Altopedaliodes Forster, 1964 — 9 spp.
Apexacuta Pyrcz, 2004 — 4 spp.
Arhuaco Adams & Bernard, 1977 — 2 spp.
Calisto Hübner, 1823 — 42 spp.
Cheimas Thieme, 1907 — 1 sp.
Corades Hewitson, 1849 — 19 spp.
Daedalma Hewitson, 1858 — 8 spp.
Dangond Adams & Bernard, 1979 — 1 sp.
Drucina Butler, 1872 — 2 spp.
Druphila Pyrcz, 2004 — 1 sp.
Eretris Thieme, 1905 — 17 spp.
Eteona Doubleday, 1848 — 1 sp.
Foetterleia Viloria, 2004 — 1 sp.
Junea Hemming, 1964 — 2 spp.
Lasiophila C. & R. Felder, 1859 — 13 spp.
Lymanopoda Westwood, [1851] — 54 spp.
Mygona Thieme, 1907 — 4 spp.
Neopedaliodes Viloria, Miller & Miller, 2004 — 10 spp.
Oxeoschistus Butler, 1867 — 11 spp.
Panyapedaliodes Forster, 1964 — 14 spp.
Paramo Adams & Bernard, 1977 — 1 sp.
Parapedaliodes Forster, 1964 — 1 sp.
Pedaliodes Butler, 1867 — 158 spp.
Praepedaliodes Forster, 1964 — 4 spp.
Antopedaliodes Forster, 1964 — 1 sp.
Pherepedaliodes Forster, 1964 — 3 spp.
Physcopedaliodes Forster, 1964 — 1 sp.
Corderopedaliodes Forster, 1964 — 3 spp.
Praepronophila Forster, 1964 — 3 spp.
Proboscis Thieme, 1907 — 2 spp.
Pronophila Doubleday, [1849] — 18 spp.
Protopedaliodes Viloria & Pyrcz, 1994 — 3 spp.
Pseudomaniola Röber, [1889] — 10 spp.
Punapedaliodes Forster, 1964 — 2 spp.
Redonda Adams & Bernard, 1981 — 2 spp.
Sierrasteroma Adams & Bernard, 1977 — 1 sp.
Steremnia Thieme, 1905 — 7 spp.
Steroma Westwood, [1850] — 3 spp.
Steromapedaliodes Forster, 1964 — 4 spp.
Thiemeia Weymer, 1912 — 1 sp.
Subtribe Satyrina
Satyrini (Satyrinae) ;
Ypthimina (Satyrini) ; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 182
Satyrina (Satyrini) ; [NL4A, 224]; [Wahlberg]
Arethusana de Lesse, 1951 — 1 sp.
Argestina Riley, 1923 — 7 spp.
Aulocera Butler, 1867 Banded Satyrs — 10 spp.
Berberia de Lesse, 1951 — 2 spp.
Boeberia Prout, 1901 — 1 sp.
Brintesia Fruhstorfer, 1911 — 1 sp.
Callerebia Butler, 1867 — 15 spp.
Cassionympha van Son, 1955 — 3 spp.
Chazara Moore, 1893 — 10 spp.
Coenyra Hewitson, 1865 — 3 spp.
Coenyropsis van Son, 1958 — 3 spp.
Davidina Oberthür, 1879 — 2 spp.
fi Hipparchia Fabricius, 1807 — 33 spp.
Kanetisa Moore, 1893 — 1 sp.
Karanasa Moore, 1893 — 33 spp.
Loxerebia Watkins, 1925 — 19 spp.
Mashuna van Son, 1955 — 2 spp.
Melampias Hübner, [1819] — 1 sp.
Minois Hübner, [1819] — 4 spp.
Neita van Son, 1955 — 6 spp.
Neocoenyra Butler, [1886] — 15 spp.
Neominois Scudder, 1875 — 1 sp.
fi Oeneis Hübner, [1819] — 33 spp.
Paralasa Moore, 1893 — 27 spp.
Paroeneis Moore, 1893 — 6 spp.
Physcaeneura Wallengren, 1857 — 5 spp.
Pseudochazara de Lesse, 1951 — 33 spp.
Pseudonympha Wallengren, 1857 — 17 spp.
Satyrus Latreille, 1810 — 13 spp.
Strabena Mabille, 1877 — 37 spp.
Stygionympha van Son, 1955 — 9 spp.
Ypthima Hübner, 1818 Rings Ringlets — 126 spp.
Austroypthima Holloway, 1974 — 1 sp.
Ypthimomorpha van Son, 1955 — 1 sp.
Subtribe ?
? (Satyrina) ; [NL4A, 224]
Amphidecta Butler, 1867 — 3 spp.
Gyrocheilus Butler, 1867 — 1 sp.
Neorinella Martins Neto & al., 1993 — 1 sp.
Tribe ?
Dyndirus Capronnier, 1874
Dyndirus Capronnier, 1874; Ann. Soc. ent. Belg. 17 : 32 (nom. nud.)
Dyndirus (?) ; [NL4A, 224]; [Wahlberg]


Unknown or unplaced taxa

Euptychia albiplaga J. & W. Zikán, 1968; (nom. nud.), TL: Brazil, Rio de Janeiro; Minas Gerais
albiplaga ; [NL4A, 224]

Euptychia ambarina J. & W. Zikán, 1968; (nom. nud.), TL: Brazil, Rio de Janeiro
ambarina ; [NL4A, 224]

Euptychia apamela Hübner, [1819]; (nom. nud.)
apamela ; [NL4A, 224]

Euptychia chilensis Ménétriés, 1857; (nom. nud.), TL: Chile
chilensis ; [NL4A, 224]

Euptychia elisae J. & W. Zikán, 1968; (nom. nud.), TL: Brazil, Rio de Janeiro; Minas Gerais
elisae ; [NL4A, 224]

Euptychia furcifera J. & W. Zikán, 1968; (nom. nud.), TL: Brazil, Rio de Janeiro
furcifera ; [NL4A, 224]

Euptychia fractifasciata J. & W. Zikán, 1968; (nom. nud.), TL: Brazil, Minas Gerais
fractifasciata ; [NL4A, 224]

Euptychia herluga J. & W. Zikán, 1968; (nom. nud.), TL: Brazil, Minas Gerais
herluga ; [NL4A, 224]

Euptychia itatiayensis J. & W. Zikán, 1968; (nom. nud.), TL: Brazil, Rio de Janeiro
itatiayensis ; [NL4A, 224]

Satyrus janirula Ménétriés, 1857; (nom. nud.), TL: Chile
janirula ; [NL4A, 224]

Euptychia lioba J. & W. Zikán, 1968; (nom. nud.), TL: Brazil, Minas Gerais
lioba ; [NL4A, 224]

Euptychia mantiqueirensis J. & W. Zikán, 1968; (nom. nud.), TL: Brazil, Minas Gerais
mantiqueirensis ; [NL4A, 224]

Euptychia montana J. & W. Zikán, 1968; (nom. nud.), TL: Brazil, Minas Gerais
montana ; [NL4A, 224]

Dyndirus nerces Capronnier, 1874; (nom. nud.), TL: Brazil
nerces ; [NL4A, 224]

Euptychia ochripenne J. & W. Zikán, 1968; (nom. nud.), TL: Brazil, Minas Gerais
ochripenne ; [NL4A, 224]

Euptychia otomara J. & W. Zikán, 1968; (nom. nud.), TL: Brazil, Rio de Janeiro
otomara ; [NL4A, 224]

Euptychia punctatolineata J. & W. Zikán, 1968; (nom. nud.), TL: Brazil, Minas Gerais
punctatolineata ; [NL4A, 224]

Euptychia simillima J. & W. Zikán, 1968; (nom. nud.), TL: Brazil, Minas Gerais
simillima ; [NL4A, 224]

Euptychia stellifera J. & W. Zikán, 1968; (nom. nud.), TL: Brazil, Rio de Janeiro; Minas Gerais
stellifera ; [NL4A, 224]

Neonympha subina Capronnier, 1874; (nom. nud.), TL: Brazil
subina ; [NL4A, 224]

Euptychia virginiae J. & W. Zikán, 1968; (nom. nud.), TL: Brazil, Minas Gerais
virginiae ; [NL4A, 224]

Euptychia wilburga J. & W. Zikán, 1968; (nom. nud.), TL: Brazil, Minas Gerais
wilburga ; [NL4A, 224]

Euptychia wordhilda J. & W. Zikán, 1968; (nom. nud.), TL: Brazil, Rio de Janeiro; Minas Gerais
wordhilda ; [NL4A, 224]


18.9.2013 (2284)

References:
[AfrL]
Mark C. Williams; Checklist of Afrotropical Papilionoidea and Hesperoidea; Compiled by Mark C. Williams, 7th ed. (2008) (April 2007); Afrototropical Butterflies; e-mail: Mark C. Williams
[BAMONA]
Opler, Pavulaan, Stanford & Pogue; Butterflies and Moths of North America; Mountain Prairie Information Node
[EcuL]
Tommaso & Racheli, 2001; Fragmenta entomologica 33 (2): 213-380; An annotated list of Ecuadorian Butterflies (Lepidoptera: Papilionidae, Pieridae, Nymphalidae)
[MRS]
Chou Io (Ed.); Monographia Rhopalocerum Sinensium, 1-2
[NL4A]
Gerardo Lamas, 2004; Atlas of Neotropical Lepidoptera; Checklist: Part 4A; Hesperioidea - Papilionoidea
[Wahlberg]
Niklas Wahlberg; The NSG's voucher specimen database; Nymphalidae Systematics Group


Some related literature:
Capronnier, 1874
Notice sur les époques d'apparition des lépidoptères diurnes du Brésil recueillis par M. C. van Volxem, dans son voyage en 1872 Ann. Soc. ent. Belg. 17 : 5-39, pl. 1
Dyar, 1903
A List of North American Lepidoptera and Key to the Literature of this Order of Insects Bull. U.S. nat. Mus. 52 : 1-723
Vane-Wright & de Jong, 2003
The butterflies of Sulawesi: annotated checklist for a critical island faunda Zool. Verh. Leiden 343: 3-268, pl. 1-16


If you have corrections, comments or information to add into these pages, just send mail to Markku Savela
Keep in mind that the taxonomic information is copied from various sources, and may include many inaccuracies. Expert help is welcome.