[HOME] [INDEXES] < Life < Insecta < Lepidoptera < Ditrysia < Papilionoidea < Nymphalidae < Satyrinae
Coenonympha Hübner, [1819]
Coenonympha Hübner, [1819]; Verz. bek. Schmett. (5): 65, TS: Papilio geticus Esper
Chortobius [Dunning & Pickard], 1858; Accentuated List Brit. Lep.: 5, TS: Papilio pamphilus Linnaeus
= ; [NL4A, 223]
Chortobius Doubleday, 1859; Zool. Syn. List Br. Butts Moths (edn 2) : 2 (preocc. [Dunning & Pickard], [1859])
= ; [NL4A, 223]
Sicca Verity, 1953; Le Farfalle diurn. d'Italia 5: 83, TS: Papilio dorus Esper
= ; [NL4A, 223]
Coenonympha, NSG Voucher Specimen [Wahlberg]
Coenonympha (Coenonymphina) ; [Wahlberg]; [NL4A, 223]

fiCoenonympha pamphilus (Linnaeus, 1758)
Palearctic. See [About maps] Keltaniittyperhonen Small Heath Kamgräsfjäril Staggängefjäril Wiesenvögelchen Kleiner Heufalter Kälberauge Le Procris Nispola Strzępotek ruczajnik
Papilio pamphilus Linnaeus, 1758; Syst. Nat. (Edn 10) 1 : 472, TL: Sweden
?Coenonympha pamphiloides Reakirt, 1866; Proc. Ent. Soc. Philad. 6 (1) : 146
= ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 30
?nephela Hufnagel
= ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 30
?menalcas Poda, 1761; Ins. Mus. Graecensis: 78
= ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 30
?Coenonympha pamphilus ab. semilyllus Krulikowsky, 1908; Soc. Ent. 23 (1): 3, TL: Wiatka; Kasan
Coenonympha pamphilus, NSG Voucher Specimen [Wahlberg]
Coenonympha pamphilus, Larvae of North-European Lepidoptera [Kimmo Silvonen]
Coenonympha pamphilus, Svenska fjärilar [SVF]
Coenonympha pamphilus, Norges Sommerfugler [NORLEP]
Coenonympha pamphilus, Moths and Butterflies of Europe [MBE]
Coenonympha pamphilus, Europaische Schmetterlinge [Mario Maier]
Coenonympha pamphilus, Captain's European Butterfly Guide [Simon Coombes]
Coenonympha pamphilius[sic] ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 30
Coenonympha pamphilus ; [BRU, 192]; [EBW]; [BOW: pl. 6, f. 5]; [MRS, 404]; [Otakar Kudrna]
[pamphilus-1v.jpg] 500x376 (~39Kb) underside Hoe lake, Devon, UK 24/06/95, Photo © Simon Coombes [pamphilus-2v.jpg] 1000x1076 (~169Kb) underside Russia, Tatarstan, Varkljed-Bodja (56°13'N 52°49'E), about 20km S of Agriz, 8.8.2004, Photo © Markku Savela [pamphilus-3v.jpg] 700x572 (~77Kb) underside France, Sophia Antipolis, 5.10.2000, Photo © Markku Savela
[pamphilus-4v.jpg] 700x603 (~74Kb) underside France, Sophia Antipolis, 5.10.2000, Photo © Markku Savela [pamphilus-5v.jpg] 470x384 (~99Kb) underside Czech Republic, Blansko, 15.6.2004, Photo © Michal Koupý [pamphilus-6v.jpg] 1200x900 (~187Kb) underside Italy, Ravenna, August 2002, Photo © Maurizio Elia
[pamphilus-f.jpg] 293x244 (~10Kb) female [pamphilus-m.jpg] 282x259 (~10Kb) male [pamphilus-v.jpg] 900x723 (~54Kb) underside FINLAND, Ab: Suomusjärvi, 669:31, 28.6.1998 Photo © Markku Savela

Larva on Poaceae, Poa, Nardus, Stipa, Cynosurus, Dactilis, Festuca, Anthoxanthum, Brachypodium, (etc.) [BRU] Anthoxanthum odoratum, Deschampsia caespitosa, D. flexuosa [SPRK]

Various forms described, real status uncertain [BRU]


C. p. lyllus (Esper, 1806)
Papilio lyllus Esper, 1805; Die Schmett., Th I, Suppl. Th. 2 (11): 23, pl. 122, f. 1-2, TL: Portugal
Coenonympha pamphilus latevittata Verity, 1916; Ent. Rec. J. Var. 28 (7-8): 172, TL: Africa [Algeria]
Coenonympha pamphilus arenosa Verity, 1926; Z. Wiss. InsektBiol. Berlin 21: 198, TL: Fez (Morocco)
Coenonympha pamphilus atlantea Verity, 1926; Z. Wiss. InsektBiol. Berlin 21: 199, TL: Atlas Mts. (Morocco)
Coenonympha pamphilus lyllus ; [BMAT: 84, pl. 28, f. 28-44]
[lyllus-1v.jpg] 600x435 (~30Kb) underside Morocco, Pays Beni-Routen, Rif, Photo © Michel Tarrier

C. p. marginata Heyne, [1894]
pamphilus marginata Heyne, [1894]; in Rühl, Die palaearktischen Grossschmetterlinge und ihre Naturgeschichte 1 (Tagfalter): 619
C. p. fulvolactea Verity, 1926
pamphilus fulvolactea Verity, 1926; Z. Wiss. InsektBiol. Berlin 21: 205
C. p. centralasiae Verity, 1926
pamphilus centralasiae Verity, 1926; Z. Wiss. InsektBiol. Berlin 21: 205
C. p. infrarasa Verity, 1926
pamphilus infrarasa Verity, 1926; Z. Wiss. InsektBiol. Berlin 21: 206
C. p. juldusica Verity, 1926
pamphilus juldusica Verity, 1926; Z. Wiss. InsektBiol. Berlin 21: 206
C. p. ferghana Stauder, 1924
pamphilus ferghana Stauder, 1924; Int. ent. Z. 17: 152
C. p. nitidissima Verity, 1926
pamphilus nitidissima Verity, 1926; Z. Wiss. InsektBiol. Berlin 21: 199
C. p. asiaemontium Verity, 1926
pamphilus asiaemontium Verity, 1926; Z. Wiss. InsektBiol. Berlin 21: 206

Coenonympha thyrsis (Freyer, 1845)
Crete thyrsis (Freyer, 1845); Neuere Beitr. Schmett. 5: pl. 475, f. 1
Coenonympha thyrsis, NSG Voucher Specimen [Wahlberg]
Coenonympha thyrsis ; [Otakar Kudrna]

Coenonympha saadi Kollar, [1849]
Iraq, Iran, Kurdistan. See [About maps] Coenonympha saadi Kollar, [1849]; Denkschr. Akad. Wiss. Wien. 1 : 52, TL: Shiraz, S.Iran
Coenonympha saadi, NSG Voucher Specimen [Wahlberg]
Coenonympha saadi ; [BOW: pl. 200, f. 2]

Larva on Poa, (and other grasses) [BRU, Hesselbarth, 1983]

C. s. iphias Eversmann, 1851
Transcaucasia saadi iphias Eversmann, 1851; Bull. Soc. imp. Nat. Moscou 24 (2) : 618
mesopotamica Heyne, [1894]; in Rühl, Die palaearktischen Grossschmetterlinge und ihre Naturgeschichte 1 (Tagfalter): 617

Coenonympha mangeri O. Bang-Haas, 1927
Afghanistan. See [About maps] mangeri O. Bang-Haas, 1927; Horae Macrolep. Palaearct. 1 : 50

Coenonympha haydenii (Edwards, 1872)
Montana, Colorado. See [About maps] Hayden's Ringlet Wyoming Ringlet Yellowstone Ringlet
haydenii (Edwards, 1872); in Hayden, Geol. Surv. Montana: 467
Coenonympha haydenii, Butterflies and Moths of North America [BAMONA]
Coenonympha haydenii ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 30; [NACL, #4581]
Coenonympha haydeni ; [BOW: pl. 18, f. 14]; [Opler]

Larva on Poaceae?, Cyperaceae? [NAB]


The "tullia" species-group

All of these may just be single C. tullia.


fiCoenonympha tullia (Müller, 1764)
NWEU, Temperate Asia, NA. See [About maps] Saraikkoniittyperhonen Large Heath Common Ringlet Starrgräsfjäril Starrängefjäril Le Daphnis Grosser Heufalter Strzępotek soplaczek
Papilio tullia Müller, 1764; Fauna Ins. Fridr.: 36, TL: Zealand Is., Denmark
Papilio davus Fabricius, 1777; Genera Insectorum : 259
isis Thunberg, 1791; Diss. Ent. sistens Insecta Suecica (2): 31
Coenonympha tullia, E.H. Strickland Ent. Mus, U of [ALBERTA]
Coenonympha tullia, NSG Voucher Specimen [Wahlberg]
Coenonympha tullia, Butterflies and Moths of North America [BAMONA]
Coenonympha tullia, Larvae of North-European Lepidoptera [Kimmo Silvonen]
Coenonympha tullia, Svenska fjärilar [SVF]
Coenonympha tullia, Norges Sommerfugler [NORLEP]
Coenonympha tullia, Moths and Butterflies of Europe [MBE]
Coenonympha tullia, Europaische Schmetterlinge [Mario Maier]
Coenonympha tullia ; [BRU, 193]; [EBW]; [BOW: pl. 6, f. 6]; [Opler]; [Otakar Kudrna]
[tullia-1l.jpg] 599x400 (~31Kb) larva Central Europe 1990-2000, Photo © Michael Borsch [tullia-1p.jpg] 400x599 (~27Kb) pupa Central Europe 1990-2000, Photo © Michael Borsch [tullia-1v.jpg] 400x588 (~38Kb) underside Central Europe 1990-2000, Photo © Michael Borsch
[tullia-2v.jpg] 444x358 (~40Kb) underside USA: Alabama, 2.8.1999, Photo © Vitaly Charny [tullia-isis.jpg] 351x621 (~35Kb) [tullia-muonioensis-f.jpg] 360x285 (~19Kb) female
[tullia-muonioensis-m.jpg] 370x282 (~21Kb) male

Larva on Eriophorum, Carex, Ryhchospora, Festuca, Poa, Stipa [BRU]


C. t. tullia (Müller, 1764)
EU, W.Siberia Coenonympha typhon Rottemburg, 1775; Der Naturforscher 6 : 15
?laidon (Borkhausen, 1788); Naturges. Eur. Schmett., 1: 91
timanensis Sedykh, 1977
?Coenonympha typhon ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 30
?Coenonympha typhon laidon ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 30
C. t. suecica Hemming, 1936
NEU, N.Siberia (Yamal Peninsula) fridolini Kusnezov, 1941; in Davenport
minimus Sedykh, 1977
C. t. bosniae Davenport, 1941
Coenonympha tullia bosniae Davenport, 1941; Bull. Mus. comp. Zool. Harv. 87 (4): 244-245, TL: Lake Jesero, Bosnia
bosniae ; Abadjiev, 2002, Neue Ent. Nachr. 23: 27
C. t. chatiparae Sheljuzhko, 1937
Caucasus Minor
C. t. elwesi Davenport, 1941
Altai Mts.
[elwesi-1mv.jpg] 413x400 (~45Kb) underside male An alpine meadow, at 2200 m above sea level, on the eastern spurs of the Katunskiy Mt. Range, the valley of the Argem (Direntai) stream, Central Altai Mts., West Siberia, Russia. 13th July, 1988, Photo © Oleg Kosterin

C. t. sibirica Davenport, 1941
Amur Region
C. t. viluiensis Ménétriés, 1859
Chukot Peninsula witimensis Davenport, 1941
C. t. mixturata Alpheraky, 1897
Kamchatka Coenonympha kodiak mixturata ; [NACL, #4582a]
C. t. yontocket Porter & Mattoon, 1989
N.California (Del Norte Co.) Coenonympha tullia yontocket Porter & Mattoon, 1989; J. Lep. Soc. 43 (3): 231, f. 1-3, TL: California, Del Norte Co. (dunes N of Crescent City, between Lake Earl and Smith River)

Coenonympha kodiak Edwards, 1869
Alaska, Yukon, NW.Territories, N.Siberia. See [About maps] Kodiak Ringlet
Coenonympha kodiak Edwards, 1869; Trans. amer. ent. Soc. 2 : 375
Coenonympha kodiak ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 30; [NACL, #4582]

Larva on Poaceae?

C. k. yukonensis Holland, 1900
Coenonympha kodiak var. yukonensis Holland, 1900; Ent. News 11 (3): 386
Coenonympha kodiak yukonensis ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 30; [NACL, #4582b]

Coenonympha inornata Edwards, 1861
Newfoundland, Canada, Montana. See [About maps] Prairie Ringlet
quebecensis Barnes & Benjamin, 1926
Coenonympha tullia inornata ; [USGS]
Coenonympha inornata ; [NACL, #4583]; [BOW: pl. 18, f. 7]

Larva on Poaceae [NAB]

C. i. mcisaaci dos Passos, 1935
Coenonympha inornata mcisaaci dos Passos, 1935; Can. Ent. 67: (82-88), TL: Doyles Station, Newfoundland
maciasaaci dos Passos, 1964; (emend.)
Coenonympha inornata mcisaaci ; [NACL, #4583a]
C. i. benjamini McDunnough, 1928
Coenonympha inornata benjamini ; [NACL, #4583b]
C. i. nipisquit McDunnough, 1939
New Brunswick, Quebec Nipisquit Ringlet
Coenonympha inornata[?] nipisquit McDunnough, 1939; Can. ent. 71: 266, TL: near Bathurst, New Brunswick
Coenonympha inornata nipisquit ; [NACL, #4583c]
Coenonympha tullia nipisquit ; Webster, 1998, J. Lep. Soc. 52 (4): 345-355
Coenonympha nipisquit ;
?nipisiquit ; (missp.?)

Larva on Spartina patens, less Festuca rubra Webster, 1998, J. Lep. Soc. 52 (4): 350

C. i. heinemanni F.M. Brown, 1959
Coenonympha inornata heinemanni ; [NACL, #4583d]

Coenonympha ochracea Edwards, 1861
Colorado. See [About maps] Ocher Ringlet
Coenonympha ochracea Edwards, 1861; Proc. Acad. nat. Sci. Philad. 13: 161, TL: Colorado
Coenonympha albescens Field, 1936
Coenonympha phantasma Burdick, 1956
Coenonympha ochracea ; [NL4A, #1602]
ochracia F. & R. Chermock, 1938; (missp.)
Coenonympha ochracea ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 30; [NACL, #4584]
Coenonympha tullia ochracea ; [USGS]

Larva on Poaceae? [NAB]

C. o. brenda Edwards, 1869
California Caenonympha[sic] ochracea brenda Edwards, 1869; , TL: California
Coenonympha ochracea brenda ; [NACL, #4584a]; [NL4A, #1602a]
C. o. mackenziei Davenport, 1936
Coenonympha ochracea mackenziei ; [NACL, #4584b]
C. o. mono Burdick, 1942
Coenonympha ochracea mono ; [NACL, #4584c]
C. o. furcae Barnes & Benjamin, 1926
Coenonympha ochracea furcae ; [NACL, #4584d]
C. o. subfusca Barnes & Benjamin, 1926
Coenonympha ochracea subfusca ; [NACL, #4584e]

Coenonympha ampelos Edwards, 1871
-. See [About maps] Northwest Ringlet
Coenonympha tullia ampelos ; [USGS]
Coenonympha ampelos ; [NACL, #4585]
[ampelos-1vU.jpg] identification uncertain 700x510 (~33Kb) underside USA: Lake Sammamish Park, Issaquah (47°33'36N 122°03'25W), King Co., WA, 11.7.1999, Photo © Markku Savela [ampelos-2vU.jpg] identification uncertain 800x739 (~47Kb) underside USA: Issaquah (47°32'35N 122°04'00W), King Co., WA, 7.7.1999, Photo © Markku Savela [ampelos-3vU.jpg] identification uncertain 500x439 (~34Kb) underside USA: FR9708, Table Mtn, Kittitas Co., WA, 19.7.1999, Photo © Markku Savela
[ampelos-4vU.jpg] identification uncertain 900x1002 (~120Kb) underside USA: near Tyler (47°27'09"N 117°48'41"W), Spokane Co., Washington, 18.8.2005, Photo © Markku Savela

Larva on Poaceae [NAB]


C. a. eunomia Dornfield, 1967
eunomia Field, 1937; (spring form.)
Coenonympha ampelos eunomia ; [NACL, #4585a]
C. a. elko Edwards, 1881
Nevada - Vancouver Coenonympha elko Edwards, 1881; Can. Ent. 13 (3) : 57
Coenonympa sweadneri F. & R. Chermock, 1941
Coenonympha elko ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 30
Coenonympha ampelos elko ; [NACL, #4585b]
Coenonympha elko ; [BOW: pl. 18, f. 6]
C. a. columbiana McDunnough, 1928
Coenonympha ampelos columbiana ; [NACL, #4585c]
C. a. insulana McDunnough, 1928
Coenonympha ampelos insulana ; [NACL, #4585d]

Coenonympha california Westwood, [1851]
WNA. See [About maps] California Ringlet
Coenonympha california Westwood, [1851]; Gen. diurn. Lep. (2): 398, TL: California
californica H. Edwards, 1881; (missp.)
Coenonympha californica ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 29
Coenonympha california ; [NACL, #4586]; [EBW]; [BOW: pl. 18, f. 5]
Coenonympha tullia californica ; [USGS]
Coenonympha california ; [NL4A, #1601]

Larva on Poaceae [NAB]

California Satyrus galactinus Boisduval, 1852; , TL: California
= ; [NL4A, #1601a]
Coenonympha ceres Butler, 1866; , TL: California
= ; [NL4A, #1601a]
Coenonympha californica[sic] var. pulla H. Edwards, 1881; Papilio 1 (4) : 51, TL: San Mateo Co., California
= ; [NL4A, #1601a]
Coenonympha californica pulla ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 29
Coenonympha california california ; [NL4A, #1601a]
C. c. eryngii H. Edwards, 1877
siskiyouensis Comstock, 1925
Coenonympha california eryngii ; [NACL, #4586a]

 

Coenonympha caeca Staudinger, 1886
Pamirs-Alai, Ghissar, Tian-Shan - Altai. See [About maps] Coenonympha caeca Staudinger, 1886; Stett. Ent. Ztg. 47 (7-9): 251, TL: Chatkalsky Mts, Uzbekistan
Coenonympha caeca ; [BRU, 193]

Larva on Carex spp. [BRU]

?C. c. eupompus (Stauder, 1924)
Dzhungarsky Alatau caeca eupompus (Stauder, 1924); , TL: Ili
pamphilus eupompus Stauder, 1924
Coenonympha caeca heptopotamica Sheljuzhko, 1929; Mitt. Münch. Ent. Ges. 19 : 352, TL: Aulie-ata
C. c. subcaeca Heyne, [1894]
Sailyughem Mts

Coenonympha symphita Lederer, 1870
NE.Turkey, SW.Georgia. See [About maps] Coenonympha symphita Lederer, 1870; Ann. Soc. ent. Belg. 13 : 44, TL: Between Akhaltsikhe and the Turkish border, Georgia
Coenonympha symphita karsiana Sheljuzhko, 1929; Mitt. Münch. Ent. Ges. 19 : 351, TL: Transcaucasia
ab. inocellata Sheljuzhko, 1929; Mitt. Münch. Ent. Ges. 19 : 352
Coenonympha symphita ; [BRU, 194]

Larva on Poa annua [BRU, Hesselbarth et al, 1995]


Coenonympha rhodopensis Elwes, 1900
Italy, Romania, Bulgaria, Albania, N.Greece, Yugoslavia. See [About maps] Eastern Large Heath
Coenonympha tiphon var. rhodopensis Elwes, 1900; Trans. ent. Soc. Lond. 1900 (2) : 205, TL: Bulgaria, Rila Mts., Levi Iskar, 5000ft
Coenonympha rhodopensis, NSG Voucher Specimen [Wahlberg]
Coenonympha rhodopensis, Moths and Butterflies of Europe [MBE]
Coenonympha rhodopensis, Captain's European Butterfly Guide [Simon Coombes]
rhodopensis ; Abadjiev, 2002, Neue Ent. Nachr. 23: 27
Coenonympha rhodopensis ; Abadjiev, 2000, Atalanta 31 (3/4): 461-466; [Otakar Kudrna]

Coenonympha corinna (Hübner, 1804)
Corsica, Sardinia, Elba. See [About maps] Corsican Heath
corinna (Hübner, 1804); Samml. eur. Schmett. [1] : f. 536, 537
Coenonympha corinna, NSG Voucher Specimen [Wahlberg]
Coenonympha corinna, Moths and Butterflies of Europe [MBE]
Coenonympha corinna ; [BOW: pl. 5, f. 16]; [Otakar Kudrna]

Coenonympha elbana Staudiger, 1901
Elba, Giglio, Monte Argentario Coenonympha elbana Staudiger, 1901; Cat. Lep. pal. Fauna: 66
Coenonympha elbana, NSG Voucher Specimen [Wahlberg]

Coenonympha fettigii Oberthür, 1874
Morocco, Algeria, Tunisia. See [About maps] Coenonympha fettigii Oberthür, 1874; Petites Nouv. Ent. 1: 412, TL: Province of Oran [Telaghre] (Algeria)
Coenonympha fettigii, Moths and Butterflies of Europe [MBE]
Coenonympha dorus fettigii ;
Coenonympha fettigii fettigii ab. infra-simplex Rothschild, 1917; Novit. Zool. 24 (1): 118, TL: Les Pins (Algeria)
Coenonympha fettigii fettigii ; [BMAT: 84, pl. 29, f. 9-11]
C. f. nicholasi Rothschild, 1925
Algeria (Blida, Chrea, etc.), Tunisia (Aïn Draham, Tunis) Coenonympha dorus nicholasi Rothschild, 1925; Novit. Zool. 32 : 208, TL: Blida-les-Glaciéres (Algeria)
Coenonympha fettigii holli Oberthür, 1910; Ètud. Lépid. Comp. 4 : 42, pl. 48, f. 396-397, TL: Blida-les-Glaciéres (Algeria)
Coenonympha fettigii nicholasi ; [BMAT: 85, pl. 29, f. 12-17]
C. f. inframaculata Oberthür, 1922
W.Morocco (Rif, Midle Atlas, High Atlas) Coenonympha fettigi var. inframaculata Oberthür, 1922; Étud. Lépid. Comp. 19 (1): 87, TL: Fort Toumliline (Morocco)
Coenonympha fettigii inframaculata ; [BMAT: 86, pl. 29, f. 18-31]
[inframaculata-1v.jpg] 600x435 (~33Kb) underside Morocco, Djebel Tisiréne, Rif, Photo © Michel Tarrier


Coenonympha dorus (Esper, 1782)
NAF, SWEU, it. See [About maps] Dusky Heath Velago de negro
Papilio dorus Esper, 1782; Die Schmett. Th. I, Bd. 2 (5): 130, pl. 78 f. 1
Coenonympha mathewi Tutt, 1904; Ent. Rec. J. Var. 16 : 308, TL: Vigo
Coenonympha dorus, NSG Voucher Specimen [Wahlberg]
Coenonympha dorus, Moths and Butterflies of Europe [MBE]
Coenonympha dorus, Europaische Schmetterlinge [Mario Maier]
Coenonympha dorus, Captain's European Butterfly Guide [Simon Coombes]
Coenonympha dorus ; [EBW]; [BOW: pl. 5, f. 17]; [Otakar Kudrna]
[dorus-1.jpg] 568x740 (~90Kb) Albarracin, prov. Teruel, , Spain, August-1988, Photo © Enrique Garcia-Barros

Larva on Agrostis spp., Festuca ovina [BMAT]


C. d. austauti Oberthür, 1881
W.Algeria Austaut's Algerian Heath
Coenonympha dorus var. austauti Oberthür, 1881; Étud. d'Ent. 6: 59, TL: Nemours [Ghazaouet] (Algeria)
Coenonympha dorus austauti ; [BMAT: 86, pl. 29, f. 1-8]

Coenonympha vaucheri Blachier, 1905
Morocco. See [About maps] Vaucher's Heath
Coenonympha vaucheri Blachier, 1905; Bull. Soc. ent. Fr. 1905 : 213, TL: High Atlas Mts. (Morocco)
Coenonympha vaucheri, NSG Voucher Specimen [Wahlberg]
Coenonympha vaucheri, Moths and Butterflies of Europe [MBE]
Coenonympha vaucheri, Captain's European Butterfly Guide [Simon Coombes]
Coenonympha vaucheri ; [BOW: pl. 115, f. 6]
Morocco (High Atlas) Coenonympha vaucheri ab. geminipuncta Blachier, 1905; Bull. Soc. ent. Fr. 1905 : 213, TL: High Atlas Mts. (Morocco)
Coenonympha meade-waldoi Elwes, 1905; Proc. zool. Soc. Lond. 1905 (4): 377
Coenonympha vaucheri f. mediocellis Le Cerf, 1923; Bull. Soc. ent. Fr. 1923 (17) : 224, TL: Grand Atlas, Djebel Tachdirt, 3100-3250m
Coenonympha vaucheri vaucheri ; [BMAT: 86, pl. 29, f. 32-40]
[vaucheri_vaucheri-1v.jpg] 599x434 (~51Kb) underside Morocco, Djebel Azourki, Haut Atlas, Photo © Jean Delacre

C. v. annoceuri Wyatt, 1952
Morocco (Middle Atlas) Coenonympha vaucheri annoceuri Wyatt, 1952; Z. wien. ent. Ges. 37: 175, pl. 21, f. B1-B4, TL: Annoceur (Morocco)
Coenonympha vaucheri annoceuri ; [BMAT: 87, pl. 29, f. 41-48]
C. v. rifensis Weiss, 1979
Morocco (W.Rif Mts.) Coenonympha vaucheri rifensis Weiss, 1979; Entomops 48: 271, f. 1, TL: Dj. Lakraa (Morocco)
Coenonympha vaucheri rifensis ; [BMAT: 87, pl. 29, f. 49-57]
C. v. beraberensis Lay & Rose, 1979
Morocco (Tizi-n'Ouguerd-Zegzaoune, Dj.Aourach, Imlil, Lake Tislit, etc.) Coenonympha vaucheri beraberensis Lay & Rose, 1979; Ent. Z. Frankf. a. M. 89 (13): 143, f. 2, TL: [Tizi-n'Ouguerd-Zegzaoune] (Morocco)
Coenonympha vaucheri beraberensis ; [BMAT: 87, pl. 29, f. 58-66]
[beraberensis-1v.jpg] 600x435 (~34Kb) underside Morocco, Tizi-n-Inezgane (Imilchil), Haut Atlas, Photo © Michel Tarrier


Coenonympha arcania (Linnaeus, 1761)
EU, Asia Minor, S.Russia, S.Urals. See [About maps] Pearly Heath Pärlgräsfjäril Perlgrasfalter Mancha Leonada Strzępotek perełkowiec
Papilio arcania Linnaeus, 1761; Fauna Suecica (Edn 2): 273, TL: Sweden
Papilio amyntas Poda, 1761; Ins. Mus. Graecensis: 79
caucasica Jachontov, 1914²
Coenonympha arcania, NSG Voucher Specimen [Wahlberg]
Coenonympha arcania, Larvae of North-European Lepidoptera [Kimmo Silvonen]
Coenonympha arcania, Svenska fjärilar [SVF]
Coenonympha arcania, Norges Sommerfugler [NORLEP]
Coenonympha arcania, Moths and Butterflies of Europe [MBE]
Coenonympha arcania, Europaische Schmetterlinge [Mario Maier]
Coenonympha arcania, Captain's European Butterfly Guide [Simon Coombes]
Coenonympha arcania ; [BRU, 195]; [EBW]; [BOW: pl. 5, f. 15]; [MRS, 405]; [Otakar Kudrna]
[arcania-1v.jpg] 500x471 (~45Kb) underside Axat, Pyrenees orientales, France 24/07/96, Photo © Simon Coombes [arcania-2mv.jpg] 652x452 (~27Kb) underside male Poland, Tresta near Tomaszow Mazowiecki, 07.1997, Photo © Marek Michalski leg. [arcania-3v.jpg] 309x300 (~24Kb) underside An opening in an oak forest, the town Krymsk environs, Krymsk District, Krasnodarskii Krai Province, S Russia. 26th May 1990, Photo © Oleg Kosterin
[arcania-4v.jpg] 517x689 (~57Kb) underside Sweden, Ekebo, Julita, 5.7.2003, Photo © Leif Wahlberg [arcania-5v.jpg] 479x639 (~66Kb) underside Sweden, Ekebo, Julita, 20.6.2004, Photo © Leif Wahlberg [arcania-6v.jpg] 470x441 (~93Kb) underside Czech Republic, Blansko, 15.6.2004, Photo © Michal Koupý

Larva on Gramineae, Melica [H&R]



Coenonympha gardetta (de Prunner, 1798)
Alps, N.Balkans. See [About maps] Alpine Heath
gardetta (de Prunner, 1798); Lepid. Pedemont.: 73
= Coenonympha pamphilus ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 30
Papilio satyrion Esper, 1805; Die Schmett., Th I, Suppl. Th. 2 (11): 24, pl. 122, f. 2
philea (Hübner, 1800)
neoclides (Hübner, 1805)
Coenonympha gardetta, NSG Voucher Specimen [Wahlberg]
Coenonympha gardetta, Moths and Butterflies of Europe [MBE]
Coenonympha gardetta, Europaische Schmetterlinge [Mario Maier]
Coenonympha gardetta, Captain's European Butterfly Guide [Simon Coombes]
Coenonympha gardetta ; [Otakar Kudrna]

Coenonympha darwiniana Staudinger, 1871
SEU. See [About maps] Coenonympha darwiniana, NSG Voucher Specimen [Wahlberg]
Coenonympha darwiniana, Captain's European Butterfly Guide [Simon Coombes]

Coenonympha arcanioides (Pierret, 1837)
Morocco, Algeria, Tunisia. See [About maps] Moroccan Pearly Heath
Satyrus arcanioides Pierret, 1837; Ann. Soc. ent. Fr. 6 : 306, TL: Province of Oran (Algeria)
?Satyrus marginalis Butler, 1868; Cat. diurn. Lep. Satyridae Brit. Mus. : 42
= ?Coenonympha arcania ;
Coenonympha arcanioides f. major Seitz, 1908; Gross-Schmett. Erde 1 : 143, pl. 48[e], TL: Algeria, Morocco
Coenonympha arcanioides gen. aest. holli Oberthür, 1910; Ètud. Lépid. Comp. 4 : 20, TL: Algeria
Coenonympha arcanioides ab. biocellata Oberthür, 1910; Ètud. Lépid. Comp. 4 : 21, TL: [Algeria]
Coenonympha arcanioides f. pluriocellata Stetter-Stättermayer, 1933; Int. ent. Z. 27: 340, TL: Bône [Annaba] (Algeria)
Coenonympha arcanioides f. inocellata Stetter-Stättermayer, 1933; Int. ent. Z. 27: 340, TL: Bône [Annaba] (Algeria)
Coenonympha arcanioides f. obliterata Stetter-Stättermayer, 1933; Int. ent. Z. 27: 340, TL: Bône [Annaba] (Algeria)
Coenonympha arcanioides f. nigro-ocellata Stetter-Stättermayer, 1933; Int. ent. Z. 27: 340, TL: Bône [Annaba] (Algeria)
Coenonympha arcanioides, NSG Voucher Specimen [Wahlberg]
Coenonympha arcanioides, Moths and Butterflies of Europe [MBE]
Coenonympha arcanioides ; [BOW: pl. 5, f. 12]; [BMAT: 88, pl. 28, f. 45-49]
[arcanioides-1v.jpg] 600x435 (~30Kb) underside Morocco, Chefchaouen, Rif, Photo © Michel Tarrier [arcanioides-2v.jpg] 599x434 (~19Kb) underside Morocco, Chefchaouen, Rif, Photo © Jean Delacre


Coenonympha leander (Esper, 1784)
Hungary, Bulgaria, S.Russia, Asia Minor, Armenia, Iran. See [About maps] Russian Heath
Papilio leander Esper, 1784; Die Schmett. Th. I, Bd. 2 (9): 176, pl. 89, f. 5, TL: Volga region, Russia
Coenonympha leander, NSG Voucher Specimen [Wahlberg]
Coenonympha leander, Captain's European Butterfly Guide [Simon Coombes]
Coenonympha leander ; [BRU, 195]; [BOW: pl. 6, f. 3]; [Otakar Kudrna]

Larva on Poa, Melica, Anthoxanthum, Lolium [BRU, Koenig, 1959, Hesselbarth et al, 1995, etc.]

C. l. obscura Heyne, 1894
Caucasus Minor, Armenian Highland, Talysh

fiCoenonympha hero (Linnaeus, 1761)
CEU, NEU, Temperate Asia, Amur, Korea, Japan. See [About maps] Ruskoniittyperhonen Scarce Heath Wald-Wiesenvögelchen Brun gräsfjäril Madängefjäril Strzępotek hero
Papilio hero Linnaeus, 1761; Fauna Suecica (Edn 2): 274, TL: Sweden
Coenonympha hero, NSG Voucher Specimen [Wahlberg]
Coenonympha hero, Larvae of North-European Lepidoptera [Kimmo Silvonen]
Coenonympha hero, Svenska fjärilar [SVF]
Coenonympha hero, Norges Sommerfugler [NORLEP]
Coenonympha hero, Moths and Butterflies of Europe [MBE]
Coenonympha hero, Europaische Schmetterlinge [Mario Maier]
Coenonympha hero ; [BRU, 196]; [EBW]; [BOW: pl. 6, f. 1]; [MRS, 404]; [Otakar Kudrna]

Larva on Gramineae, Elymus arenarius [H&R], Elymus, Poa, Hordeum, Calamagrostis [BRU, Lang, 1884, Henriksen, Kreutzer, 1982]

C. h. sabaeus Fabricius, 1775
E.Europe, Urals, W.Siberia
C. h. perseis Lederer, 1853
up to Pacific sibirica Fuchs, 1899
Coenonympha hero perseis ; [MRS, 404]
[perseis-1muv.jpg] 478x640 (~92Kb) upperside underside male Russia, N Tuva, Piykhemskiy distr., h=800., 06.1997 Vashchenko S.N. leg, Photo © D. Smirnov

Larva on Cyperaceae [MRS]


C. h. pilwonis Matsumura, 1925
Sakhalin
C. h. latefasciata Matsumura, 1925
Kunashir
C. h. coreana Matsumura, 1927
Korea Coenonympha hero coreana Matsumura, 1927; Insecta Matsumurana 1 (4) : 164, TL: Corea

fiCoenonympha glycerion (Borkhausen, 1788)
EU, S.Siberia, Japan. See [About maps] Idänniittyperhonen Chestnut Heath Rostbraunes Wiesvögelchen Darrgräsfjäril Darrängefjäril Strzępotek glicerion
Papilio glycerion Borkhausen, 1788; Naturges. Eur. Schmett., 1: 90
Papilio iphis Denis & Shiffermüller, 1775; (preocc.)
Coenonympha iphis var. anaxagoras Assmus, 1857; Breslau Ent. Z. 10 : 9
Papilio tiphon Esper, 1777; Die Schmett. Th. I, Bd. 1 (9): 341, (6): pl. 35, f. 3-4 (preocc.)
Coenonympha glycerion glycerion, Lepidoptera collection [SZMN]
Coenonympha glycerion, NSG Voucher Specimen [Wahlberg]
Coenonympha glycerion, Larvae of North-European Lepidoptera [Kimmo Silvonen]
Coenonympha glycerion, Moths and Butterflies of Europe [MBE]
Coenonympha glycerion, Europaische Schmetterlinge [Mario Maier]
Coenonympha glycerion, Captain's European Butterfly Guide [Simon Coombes]
Coenonympha glycerion ; [BOW: pl. 5, f. 18]; [MRS, 405]; [Otakar Kudrna]
[glycerion-1v.jpg] 500x430 (~28Kb) underside Finland, Ylöjärvi, Teivaala, 6828:320, 16.7.1996, Photo © Tero Piirainen [glycerion-2v.jpg] 800x540 (~51Kb) underside Finland, Vihti, Nummela, ?July 2002, Photo 10072002_24 © Timo Viitanen [glycerion-3av.jpg] 470x443 (~80Kb) underside Czech Republic, Blansko, 15.6.2004, Photo © Michal Koupý
[glycerion-3bv.jpg] 470x379 (~76Kb) underside Czech Republic, Blansko, 22.7.2004, Photo © Michal Koupý [glycerion-4v.jpg] 600x400 (~60Kb) underside Estonia, Hiiumaa, Kootsaare nina. 24.07.2004, Photo © Hannu Tanner [glycerion-5v.jpg] 724x724 (~116Kb) underside Russia, Moscow area, 10.06.2007, Photo © D. Smirnov
[glycerion-5vm.jpg] 710x522 (~96Kb) underside male Russia: Moscow area (36°25'E, 56°00'N), 14.7.2008, Photo © D. Smirnov [glycerion-6fv.jpg] 811x660 (~113Kb) underside female Russia, Moscow area, 18.8.2010 (36°25'E, 56°00'N), Photo © D. Smirnov [glycerion-f.jpg] 333x278 (~15Kb) female
[glycerion-m.jpg] 320x275 (~23Kb) male

Larva on Gramineae, Deschampsia caespitosa [SPRK]

Larva on Briza, Poa, Brachypodium, Melica, Cynosurus, (and other grasses) [BRU, Eckstein, 1913, Nekrutenko, 1990, Gorbuno, Korshunov, 1995]


C. g. alta Sheljuzhko, 1937
Caucasus
C. g. korshunovi Nekrutenko, 1978
Crimea
C. g. iphicles Staudinger, 1892
N.Alai, Sayan, Transbaikalia, Amur, Ussuri heroides Christoph, 1893
Coenonympha glycerion iphicles, Lepidoptera collection [SZMN]
C. g. iphina Staudinger, 1892
Saur, Tarbagatai
C. g. wutaica Murayama
Coenonympha glycerion wutaica ; [MRS, 405]

Coenonympha mongolica Alphéraky, 1881
Ili River Basin Coenonympha mongolica Alphéraky, 1881; Horae Soc. ent. Ross. 16 (3-4): 426, 17 (1-2): pl. 1, f. 26, TL: Near Kuldzha, China
Coenonympha mongolica, Lepidoptera collection [SZMN]
Coenonympha mongolica ; [BRU, 197]; [BOW: pl. 200, f. 10]

Larva on Achnaterum [BRU]


Coenonympha mahometana Alphéraky, 1881
Tian-Shan. See [About maps] Coenonympha mahometana Alphéraky, 1881; Horae Soc. ent. Ross. 16 (3-4): 428, TL: Kuldzha, Kunges Valley, W. China
Coenonympha mahometana, NSG Voucher Specimen [Wahlberg]
Coenonympha mahometana ; [BRU, 197]

Coenonympha sunbecca Eversmann, 1843
Ghissar - Tarbagatai. See [About maps] Coenonympha sunbecca Eversmann, 1843; Bull. Soc. Imp. Nat. Moscou 16 (3) : 538, pl. 7, f. 4a-b, TL: Zaisan, SE. Kazakhstan
?sunbecca var. alexandra Heyne, [1894]; , TL: Kirghizsky Mts.
Coenonympha sunbecca, NSG Voucher Specimen [Wahlberg]
Coenonympha sunbecca alexandra ;
Coenonympha sunbecca ; [BRU, 197]; [EBW]; [BOW: pl. 200, f. 7]; [MRS, 404]
[sunbecca-1v.jpg] 300x400 (~28Kb) underside A steppefied meadow in the middle flow of the Kora River, 30 km NE of the Tekeli City (1900 m above sea level), the North Dzhungarskiy Alatau mountain chain (NE part of Tien Shan), Taldy-Kurgan Province, SE Kazakhstan. 15th June, 1993 (on Artemisia santolinifolia Turcz. ex Besser), Photo © Oleg Kosterin


Coenonympha mahometana × sunbecca
?decolorata Wagner, 1913
mahometana × sunbecca ; [BRU, 197]

Coenonympha iphioides Staudinger, 1870
Spain. See [About maps] Spanish Heath Castaneo morena
Coenonympha iphioides Staudinger, 1870; Berl. Ent. Z. 14 : 101
Coenonympha iphioides, NSG Voucher Specimen [Wahlberg]
Coenonympha iphioides, Moths and Butterflies of Europe [MBE]
Coenonympha iphioides ; [BOW: pl. 6, f. 2]; [Otakar Kudrna]
[iphioides-1v.jpg] 612x520 (~45Kb) underside Barbatona (prov. Guadalajara), Spain 1-august-1989, Photo © Enrique Garcia-Barros


Coenonympha oedippus (Fabricius, 1787)
CEU, S.Siberia - Ussuri, China, Korea, Japan. See [About maps] False Ringlet Moor-Wiesenvögelchen Lindos ojos Strzępotek edypus
Papilio oedippus Fabricius, 1787; Mantissa Insectorum 2: 31, TL: S.Russia
Papilio geticus Esper, 1794; Die Schmett., Suppl. Th 1 (3-4): 51, (5-6): pl. 102, f. 2
Coenonympha annulifer Butler, 1877; Ann. Mag. nat. Hist. (4) 19 (109) : 91, TL: 370 miles from Tokei (Yedo)
= ; [NHM]
Coenonympha oedippus; [BRU, 196]
Coenonympha oedippus, NSG Voucher Specimen [Wahlberg]
Coenonympha oedippus, Moths and Butterflies of Europe [MBE]
Coenonympha oedippus, Europaische Schmetterlinge [Mario Maier]
Coenonympha oedippus ; [BOW: pl. 6, f. 4]; [MRS, 403]; [Otakar Kudrna]
[oedippus-1v.jpg] 1200x1565 (~150Kb) underside Hungary: Ocsa, June 2005, Photo © Szabolcs Safian

Larva on Lolium, Carex, Iris pseudacorus [H&R], Poa [BRU], Phragmites communis [MRS]


C. o. magna Heyne, 1895
Amur, Ussuri oedippus amurensis Heyne, 1895; in Ruhl
Coenonympha oedippus amurensis Rühl; [MRS, 403]
[magna-1mfv.jpg] 600x800 (~97Kb) underside male female A meadow steppe on the hill Malyi Batur on the left bank of the Onon River 7 km upstream of the village Nizhnii Tsasuchei, Onon District, Chita Province, SE Transbaikalia (Dahuria), Siberia, Russia. 11th July 1996, Photo © Oleg Kosterin

C. o. taibaica Maurayama
Coenonympha oedippus taibaica ; [MRS, 403]

Coenonympha nolckeni Erschoff, 1874
Ghissar, Alai, Tian-Shan, N.Pamirs, W.Pamirs. See [About maps] Coenonympha nolckeni Erschoff, 1874; in Fedschenko, Travels in Turkestan. 2 (5) : 23, pl. 2, f. 17, TL: Mt. Naubid, Zeravshan Valley, Uzbekistan
Coenonympha nolckeni, NSG Voucher Specimen [Wahlberg]
Coenonympha nolckeni ; [BRU, 196]; [BOW: pl. 200, f. 9]

Coenonympha amaryllis (Stoll, [1782])
S.Urals, Altai Mts, W.Siberia, E.Siberia, Transbaikalia, Amur, Ussuri, China, Mongolia, Korea. See [About maps] Papilio amaryllis Stoll, [1782]; in Cramer, Uitl. Kapellen 4 (32-32): 210, pl. 391, f. A, B, TL: "...de la Sibérie"
rinda Ménétriés, 1859; (infrasubsp.) - Amur
accrescens Staudinger, 1901; (infrasubsp.) - Ussuri
borisovi Korshunov & Ivonin, 1996; (infrasubsp.) - Baikal region
Coenonympha amaryllis, Lepidoptera collection [SZMN]
Coenonympha amaryllis, NSG Voucher Specimen [Wahlberg]
Coenonympha amaryllis ; [BRU, 194]; [BOW: pl. 200, f. 5]; [MRS, 403]; [Otakar Kudrna]
[amaryllis-1v.jpg] 400x398 (~41Kb) underside The valley of the Shivilig-Khem River just upstream its leaving the the southern foot of the East Tannu-Ola mounatin range for the Ubsu-Nur Hollow, Tes-Khem District, Tuva Republic, Siberia, Russia. 11th July 1990, Photo © Oleg Kosterin [amaryllis-2u.jpg] 600x800 (~94Kb) upperside A steppefied meadow on the Onon River right bank terrace at a pine forest edge, 5 km upstream of the village Nizhnii Tsasuchei, Onon District, Chita Province, Transbaikalia, Siberia, Russia, 30th June 1996 (Two individuals catched by a Thomisidae spider), Photo © Oleg Kosterin

Larva on Poa [BRU]


Coenonympha amaryllis simingica ; [MRS, 403]

Coenonympha xinjiangensis Chou & Huang, 1994
Xinjiang. See [About maps] Coenonympha xinjiangensis Chou & Huang, 1994; in Chou, Monographia Rhopalocerum Sinensium 1-2: 759, 403, f. 32-33, TL: Xinjiang, Jiandengyu

Coenonympha tydeus Leech, [1892]
Tibet. See [About maps] Coenonympha tydeus Leech, [1892]; Butts China Japan Corea (1): 96
Coenonympha tydeus ; [MRS, 405]

Coenonympha sinica Alphéraky, 1888
Kashmir, Tibet. See [About maps] Coenonympha sinica Alphéraky, 1888; Stettin Ent. Ztg 49 : 66, TL: Nian-Chian-Kette, Djin-ta-sy
Coenonympha sinica ; [MRS, 405]

Coenonympha pavonina Alphéraky, 1888
China. See [About maps] Coenonympha pavonina Alphéraky, 1888; Stettin Ent. Ztg 49 : 66, TL: R. Hei-ho

Coenonympha semenovi Alphéraky, 1887
Tibet, W.China. See [About maps] Coenonympha semenovi Alphéraky, 1887; in Romanoff, Mem. lép. 3 : 405
Coenonympha semenovi, NSG Voucher Specimen [Wahlberg]
?Coenonympha semenori[sic?] ; [BOW: pl. 200, f. 6]
Coenonympha semenovi ; [MRS, 404]
[semenovi-1fv.jpg] 1000x1058 (~190Kb) underside female China, Qinghai prov, 20 km S Tianjun (3900m), Photo © A. Timchenko leg. [semenovi-1mu.jpg] 1000x915 (~154Kb) upperside male China, Qinghai prov, 20 km S Tianjun (3900m), Photo © A. Timchenko leg. [semenovi-1mv.jpg] 1000x1062 (~176Kb) underside male China, Qinghai prov, 20 km S Tianjun (3900m), Photo © A. Timchenko leg.

C. s. jiadengyuica Huang & Murayama, 1992
Xianjing Coenonympha semenovi jiadengyuica Huang & Murayama, 1992; Tyô to Ga 43 (1) : 5, f. 15, TL: Jiadengyu, Altai, 1380m
Coenonympha semenovi jiadengyuica ; [MRS, 404]

Coenonympha tyderes Leech²
Tibet. See [About maps] Coenonympha tyderes ; [BOW: pl. 200, f. 5 (text only)]


22.9.2013 (39)

References:
[A. Timchenko]
Andrew Timchenko; Beijing, China (ex-Moscow, Russia)
[ALBERTA]
University of Alberta, E.H. Strickland Entomological Museum; Browse Entomology Collection
[About maps]
Warning! The maps are automatically generated from the textual information, and the process does not always produce acceptable result; See about maps for more info.
[BAMONA]
Opler, Pavulaan, Stanford & Pogue; Butterflies and Moths of North America; Mountain Prairie Information Node
[BHL]
The; Biodiversity Heritage Library; and on; Internet Archive; Note: The linking to the content of Internet Archive is somewhat delicate -- notify me when links break!
[BMAT]
Tennent, 1996; The butterflies of Morocco, Algeria and Tunisia; ISBN 0 906802 05 9
[BOW]
Lewis, 1974; Butterflies of the World; ISBN 0 245 52097 X
[BRU]
Tuzov, Bogdanov, Devyatkin, Kaabak, Korolev, Murzin, Samodurov, Tarasov, 1997; Guide to the Butterflies of Russia and adjacent territories; Hesperiidae, Papilionidae, Pieridae, Satyridae; Volume 1
[BnF]
Gallica / Bibliothèque nationale de France; Annales de la Société entomologique de France; Bulletin de la Sociéte entomologique de France
[CiNii]
http://ci.nii.ac.jp/
[D. Smirnov]
Russia, Moscow; e-mail: D. Smirnov
[EBW]
Smart, Paul, 1976; The Illustrated Encyclopedia of the Butterfly World; ISBN 0 600 31381 6
[ESP]
Heppner, 1981; The dates of E. J. C. Esper's Die Schmetterlinge in Abblidungen... 1776-[1830]; Archives of Natural History (1981) 10 (2): 251-254
[Enrique Garcia-Barros]
Enrique García-Barros; Dept. Biol. (Zool.); Univ. Autonoma de Madrid
[H&R]
Higgins, L.G, Riley, N.D, 1970; A Field Guide to the Butterflies of Britain and Europe
[HBN]
Francis Hemming, 1937; HÜBNER, A bibliographical and systematic account of the entomological works of Jacob Hübner and of the supplements thereto by Carl Geyer, Gottfried Franz von Frölich and Gottlieb August Wilhelm Herrich-Schäffer, Vol 1 & 2
[Hannu Tanner]
Hannu Tanner, Siuntio, Finland; e-mail: Hannu Tanner; http://www.kolumbus.fi/hannu.j.tanner/coleoptera/
[Jean Delacre]
http://homepage.mac.com/jdelacre
[Kimmo Silvonen]
Espoo, Finland; Kimmo's Lep Site
[Leif Wahlberg]
Julita, Sweden; e-mail: Leif Wahlberg
[MBE]
Moths and Butteflies of Europe; Paolo Mazzei, Diego Reggianti and Ilaria Pimpinelli; http://www.leps.it/
[MRS]
Chou Io (Ed.); Monographia Rhopalocerum Sinensium, 1-2
[Marek Michalski]
Lodz, Poland; e-mail: Marek Michalski
[Mario Maier]
Europäische Schmetterlinge; e-mail: Mario Maier
[Maurizio Elia]
Ravenna, Italy; e-mail: Maurizio Elia
[Michael Borsch]
Geesthacht, Germany; contact through; e-mail: Francisco Welter-Schultes
[Michal Koupý]
Czech Republic
[Michel Tarrier]
Michel R. Tarrier; Malaga, Spain; http://www.tarrier.org
[NAB]
Pyle, 1981; Field Guide to North American Butterflies, National Audubon Society; ISBN 0-394-51914-0
[NACL]
Hodges, 1983; Check List of the Lepidoptera of America North of Mexico
[NHM]
Natural History Museum; The Global Lepidoptera Names Index
[NL4A]
Gerardo Lamas, 2004; Atlas of Neotropical Lepidoptera; Checklist: Part 4A; Hesperioidea - Papilionoidea
[NORLEP]
Norges Sommerfugler
[Oleg Kosterin]
Institue of Cytology & Genetics; Siberian Division of Russian Academy of Sciences; Novosibirsk; e-mail: Oleg Kosterin; http://pisum.bionet.nsc.ru/kosterin/
[Opler]
Opler & Warren, 2003 (April); Butterflies of North America. 2. Scientific Names List for Butterfly Species of North America, north of Mexico.
[SPRK]
Seppänen, E. J, 1970; Suomen suurperhostoukkien ravintokasvit, Animalia Fennica 14
[SVF]
Svenska fjärilar; Naturhistoriska riksmuseet; (Sweden)
[SZMN]
Lepidoptera; Siberian Zoological Museum
[Simon Coombes]
1 Park Street, Plymouth, PL3 4BL, UK; 01752 607854; e-mail: Simon Coombes; http://www.butterfly-guide.co.uk
[Szabolcs Safian]
Hungary; Hungarian Lepidopterologial Society; Butterflies of Ghana
[Tero Piirainen]
Tampere, Finland; e-mail: Tero Piirainen
[Timo Viitanen]
http://www.kuvarkisto.com/
[USGS]
Northern Prairie Wildlife Research Center; http://www.npwrc.usgs.gov/index.htm
[Vitaly Charny]
Birmingham, Alabama, USA
[Wahlberg]
Niklas Wahlberg; The NSG's voucher specimen database; Nymphalidae Systematics Group
[²]
This may require parentheses or not. I don't have the necessary information for this taxon.


Some related literature:
Abadjiev, 2000
Type of Balkan butterflies. II. Eastern Large Heath, Coenonympha rhodopensis Elwes, 1900: type locality and lectotype designation Atalanta 31 (3/4): 461-466
Abadjiev, 2002
Types of Balkan butterflies in the collection of The Natural History Museum, London Neue Ent. Nachr. 23: 3-53
Alphéraky, 1881
Lépidoptères du district de Kouldjà et des montagnes environnantes. (1-3) Horae Soc. ent. Ross. 16 (3-4): 334-435, pl. 14-15 (1881), 17 (1-2): 15-103, pl. 1-3 (1882), 17 (3-4): 156-227, pl. 8-9 (1883)
Alphéraky, 1887
Diagnoses de quelques lépidoptères inédits du Thibet in Romanoff, Mem. lép. 3 : 403-406
Alphéraky, 1888
Neue Lepidopteren Stettin Ent. Ztg 49 : 66-69
Assmus, 1857
Coenonympha anaxagoras eine neue Varietät der Coenonympha iphis und ein Beitrag zur Naturgeschichte von Larentia Pyropata Hüb. Breslau Ent. Z. 10 : 9-10
O. Bang-Haas, 1927
Horae Macrolepidopterologicae Regionis Palaearcticae Horae Macrolep. Palaearct. 1 : 1-128, pl. 1-11
Blachier, 1905
Descriptions de Lépidoptères nouveaux de Maroc Bull. Soc. ent. Fr. 1905 : 212-215
Borkhausen, 1788
Naturgeschichte der europäischen Schmetterlinge nach Systematischer Ordnung. 1 Naturges. Eur. Schmett., 1: 36+288pp, 1pl
Butler, 1868
A catalogue of diurnal Lepidoptera of the family Satyridae in the collection of the British Museum Cat. diurn. Lep. Satyridae Brit. Mus. : 1-211, pl. 1-5
Butler, 1877
On Rhopalocera from Japan and Shanghai, with Descriptions of new Species Ann. Mag. nat. Hist. (4) 19 (109) : 91-97
Le Cerf, 1923
Lépidoptères nouveaux du Maroc Bull. Soc. ent. Fr. 1923 (17) : 224
Davenport, 1941
The Butterflies of the Satyrid Genus Coenonympha Bull. Mus. comp. Zool. Harv. 87 (4): 215-349, 10pls
dos Passos, 1935
Some butterflies of southern Newfoundland with descriptions of new subspecies Can. Ent. 67: 82-88
Doubleday, 1859
Zoologist Synonyic List of British Butterfles and Moths Zool. Syn. List Br. Butts Moths (edn 2) : 1-40
Westwood, [1851]
The genera of diurnal Lepidoptera, comprising their generic characters, a notice of their habitats and transformations, and a catalogue of the species of each genus; illustrated with 86 plates by W. C. Hewitson Gen. diurn. Lep. (1): 1-18, pl. A, 1-4 (1-2) (1846), (1): 19-132, pl. 4*, 5-25, 28 (3-14) (1847), (1): 133-200, pl. 1*, 26, 27, 29, 31-44 (15-23) (1848), (1): 201-242, pl. 30, 45-52, 56-58, 60-62, 64 (24-31) (1849), (1): 243-250 [?], (2): 243-326, pl. 53-55, 63, 65, 66 (32-38) (1850), (2): 327-466, pl. 59, 67-77 (39-50) (1851), (2): 467-534, 143, 144, pl. 78-80 (Index) (51-54) (1852)
Dyar, 1903
A List of North American Lepidoptera and Key to the Literature of this Order of Insects Bull. U.S. nat. Mus. 52 : 1-723
Edwards, 1861
Descriptions of certain Species of Diurnal Lepidoptera found within the limits of the United States and of British America. - No. [1]-3 Proc. Acad. nat. Sci. Philad. 13: 160-164 (1861), 14: 54-58, 221-226 (1862)
Edwards, 1869
Descriptions of new species of diurnal Lepidoptera found within the United States Trans. amer. ent. Soc. 2 : 369-376
Edwards, 1872
List of species of butterflies collected by Campbell Carrington and Wm. B. Logan, of the expedition [to Montana], in 1871. Preliminary report [5th Ann. rep.] U. S. Geological Survey of Montana, by F. V. Hayden in Hayden, Geol. Surv. Montana: 466-467
Edwards, 1881
Coenonympha elko Can. Ent. 13 (3) : 57-58
H. Edwards, 1881
On some apparently new forms of diurnal lepidoptera Papilio 1 (4) : 50-55
Elwes, 1900
On the Butterflies of Bulgaria Trans. ent. Soc. Lond. 1900 (2) : 183-206, pl. 4
Esper, 1777
Die Schmetterlinge in Abbildungen nach der Natur mit Beschreibungen. Theil I. Die Tagschmetterlinge. Band 1 Die Schmett. Th. I, Bd. 1 (1): 1-40, pl. 1-6 (1776), (2): 41-72, pl. 7-12 (1776), (3): 73-104, pl. 13-18 (1777), (4): 105-120, pl. 19-24 (1777), (5): 121-144, pl. 25-30 (1777), (6): 145-176, pl. 31-36 (1777), (7): 177-200, pl. 37-46 (1778), (8): 201-216, pl. 43-48 (1778), (9): 217-388, pl. 49-50 (1779)
Esper, 1782; Esper, 1784
Die Schmetterlinge in Abbildungen nach der Natur mit Beschreibungen. Theil I. Die Tagschmetterlinge. Fortsetzung. Band 2 Die Schmett. Th. I, Bd. 2 (1): 1-36, pl. 51-56 (1780), (2): 37-68, pl. 57-62 (1781), (3): 69-100, pl. 63-68 (1781), (4): 101-124, pl. 69-74 (1781), (5): 125-140, pl. 75-80 (1782), (7): 141-156, pl. 81-82 (1783), (8): 157-172, pl. 83-88 (1783), (9): 173-184, pl. 89-92 (1784), (28): 185-190, pl. 93 (1786)
Esper, 1794
Die Schmetterlinge in Abbildungen nach der Natur mit Beschreibungen. Theil I. Die Tagschmetterlinge. Supplement Theil 1. Abschnitt 1 Die Schmett., Suppl. Th 1 (1-2): 1-8, pl. 94 (1789), (3-4): 9-60, pl. 95-101 (1789), (5-6): 61-68, pl. 102-106 (1794), ([7]): [69-88], pl. [107-109] (1798), (8-9): 89-104, pl. 110-112 (1800), (10): 105-120, pl. 113-116 ([1803-1804])
Esper, 1805
Die Schmetterlinge in Abbildungen nach der Natur mit Beschreibungen. Theil I. Die Tagschmetterlinge. Supplement Theil 2 Die Schmett., Th I, Suppl. Th. 2 (11): 1-24, pl. 117-122 (1805), (12): 25-48, pl. 123-126 [1830]
Erschoff, 1874
Travels in Turkestan. Volume 2. Zoogeographical Investigations. Lepidoptera [in Russian] in Fedschenko, Travels in Turkestan. 2 (5) : 1-128, pl. 1-6
Eversmann, 1843
Quaedam lepidopterorum species novae in montibus Uralensibus et Altaicus habitantes nunc descriptae et depictae Bull. Soc. Imp. Nat. Moscou 16 (3) : 535-553, 4pls
Eversmann, 1851
Description de quelques nouvelles espéce de Lépidoptéres de la Russie Bull. Soc. imp. Nat. Moscou 24 (2) : 610-644
Fabricius, 1777
Genera Insectorum eorumque characters naturales secundum numerum, figuram, situm et proportionem ... Genera Insectorum : 1-310
Freyer, 1845
Neuere Beiträge zur Schmetterlingskunde mit Abbildungen nach der Natur. (65-80) Neuere Beitr. Schmett. 5 : 1-40 (1842), : 41-92 (1844), : 93-164 (1845)
Heyne, [1894]
Satyridae, Lycaenidae in Rühl, Die palaearktischen Grossschmetterlinge und ihre Naturgeschichte 1 (Tagfalter): 385-857
Holland, 1900
Alaska Insects Ent. News 11 (3): 381-389 (part I), (4): 416-423 (part II)
Huang & Murayama, 1992
Butterflies of Xinjiang province, China Tyô to Ga 43 (1) : 1-22
Hübner, 1804
Sammlung europäischer Schmetterlinge. I. Papiliones - Falter ("Erste Band") Samml. eur. Schmett. [1] : 3-74, : pl. 1-88, f. 1-446 (1799-1800), : pl. 89-96, f. 447-490 (1800-1803), : pl. 97-114, f. 491-588 (1803-1804), : pl. 115-119, f. 589-612 (1805), : pl. 120-124, f. 613-636 (1805-1806), : pl. 125-128, f. 637-652 (1806-1808), : pl. 129-144, f. 653-727 (1808-1813), : pl. 145-147, f. 728-741 (1814-1816), : pl. 148-150, f. 742-756 (1816-1817), : pl. 151, f. 757-762 (1818-1819), : pl. 152-154, f. 757-774 (1819-1822), : pl. 155, f. 775-778 (1822-1823), : pl. 156-161, f. 779-806 (1823), : pl. 162-175, f. 807-874 (1823-1824), : pl. 176, f. 869-874 (1824-1825), : pl. 177-181, f. 875-895 (1825-1826), : pl. 182-187, f. 896-981 (1827-1828), : pl. 188-195, f. 922-968 (1828-1832), : pl. 196, f. 969-974 (1832-1833), : pl. 197-201, f. 975-997 (1834-1836), : pl. 202-203, f. 998-1009 (1836-1838), : pl. 204-207, f. 1010-1029 (1837-1838)
Hübner, [1819]
Verzeichniss bekannter Schmettlinge, 1816-[1826] Verz. bek. Schmett. (1): [1-3], 4-16 (1816), (2): 17-32 (1819), (3): 33-48 (1819), (4): 49-64 (1819), (5): 65-80 (1819), (6): 81-96 (1819), (7): 97-112 (1819), (8): 113-128 (1819), (9): 129-144 (1819), (10): 145-160 (1819), (11): 161-176 (1819), (12): 177-192 (1820), (13): 193-208 (1820), (14): 209-224 (1821), (15): 225-240 (1821), (16): 241-256 (1821), (17): 257-272 (1823), (18): 273-288 (1823), (19): 289-304 (1823), (20): 305-320 (1825), (21): 321-336 (1825), (22): 337-352 (1825), (23-27): 353-431 ([1825])
Krulikowsky, 1908
Einige neue Varietäten und Aberrationen der Lepidopteren des östlichen Russlands Soc. Ent. 23 (1): 2-3, (2): 11-12, (3): 18
Lederer, 1870
Contributions a la faune des Lepidopteres de la Transcaucasie Ann. Soc. ent. Belg. 13 : 17-54, pl. 1-2
Leech, [1892]
Butterflies from China, Japan, and Corea Butts China Japan Corea (1): 1-297, pl. 1-28 (1892-1894), (2): 297-681, pl. 29-43 (1893-1894)
Linnaeus, 1758
Systema Naturae per Regna Tria Naturae, Secundum Clases, Ordines, Genera, Species, cum Characteribus, Differentiis, Symonymis, Locis. Tomis I. 10th Edition Syst. Nat. (Edn 10) 1 : 1-338, : 339-824
Matsumura, 1927
A list of the butterflies of Corea, with description of new species, subspecies and aberrations Insecta Matsumurana 1 (4) : 159-170
McDunnough, 1939
A new Coenonympha race from northeastern New Brunswick Can. ent. 71: 266
Müller, 1764
Fauna Insectorum Fridrichsdalina sive Methodica Descriptio Insectorum Agri Fridrichsdalensis cum Characteribus Genericis et Specificis, Nominibus Trivialibus, Locis Natalibus, Iconibus Allegatis, Novsque Pluribus Speciebus Additis Fauna Ins. Fridr.: 96pp
Oberthür, 1874
Lépidoptères nouveaux d'Algérie. 1. Diurnes Petites Nouv. Ent. 1: 412-413
Oberthür, 1910
Notes pour servir à établir la Faune Française et Algérienne des Lépidoptères Ètud. Lépid. Comp. 4 : 15-691, pl. 35-58
Oberthür, 1922
Les Lépidoptères du Maroc Étud. Lépid. Comp. 19 (1): 13-402, (2): pl. 530-546
Pierret, 1837
Description de deux nouvelles espèces du genre Satyre (Latr.) Ann. Soc. ent. Fr. 6 : 303-307, pl. 12
Poda, 1761
Insecta Musei Graecensis, quae in ordines, genera et species juxta Systema Naturae Caroli Linnaei digessit Ins. Mus. Graecensis: 8+127, 2pls
Porter & Mattoon, 1989
A new subspecies of Coenonympha tullia (Müller) confined to the coastal dunes of northern California J. Lep. Soc. 43 (3): 229-238
de Prunner, 1798
Lepidoptera Pedemontana illustrata. Turin. 1 Lepid. Pedemont.: 52+124
Reakirt, 1866
Coloradian butteflies Proc. Ent. Soc. Philad. 6 (1) : 122-151
Rothschild, 1917
Supplemental notes to Mr. Charles Oberthür's Faune des Lépidoptères de la Barbarie, with lists of the specimens contained in the Tring Museum Novit. Zool. 24 (1): 61-120, : 325-373, : 393-409
Rothschild, 1925
Critical List of the collection of Algerian Lepidoptera of the late Captain N. J. E. Holl. Novit. Zool. 32 : 195-229
Rottemburg, 1775
Unmertungen zu den Hufnagelifchen Tabellen der Schmetterlinge Der Naturforscher 6 : 1-34
Seitz, 1908
Die Großschmetterlinge des Palaearktischen Faunengebietes. 1. Die Palaearktischen Tagfalter Gross-Schmett. Erde 1 : 1-379
Sheljuzhko, 1929
Einige neue palaearktische Lepidopteren-Formen Mitt. Münch. Ent. Ges. 19 : 347-362, pl. 27-28
Staudinger, 1870
Beschreibung neuer Lepidopteren des europäischen faunegebiets Berl. Ent. Z. 14 : 97-132, 193-208, (3/4): 273-330 (1871)
Staudinger, 1886
Centralasiatische Lepidopteren Stett. Ent. Ztg. 47 (4-6): 193-215, (7-9): 225-256
Stoll, [1782]
Uitlandsche Kapellen (Papillons exotiques) in Cramer, Uitl. Kapellen 4 (26b-28): 29-90, pl. 305-336 (1780), (29-31): 91-164, pl. 337-372 (1781), (32-32): 165-252, 1-29, pl. 373-400 (1782)
Thunberg, 1791
Dissertio Entomologica sistens Insecta Suecica, ... Diss. Ent. sistens Insecta Suecica (1): 1-24 (1 plate) (1784), (2): [25-31] ??], 32-46 (9 unnumbered) (1791), (3): [47],48-52 (2 unnumbered) (1792), (4): 53-62 (1792)
Tutt, 1904
Coenonympha mathewi, n.sp. Ent. Rec. J. Var. 16 : 308-309
Verity, 1916
The Brish Races of Butterflies: their relationships and nomeclature Ent. Rec. J. Var. 28 (5): 97-102, (6): 128-133, (7-8): 165-174
Webster, 1998
The life history of the Maritime Ringlet, Coenonympha tullia nipisquit J. Lep. Soc. 52 (4): 345-355


If you have corrections, comments or information to add into these pages, just send mail to Markku Savela
Keep in mind that the taxonomic information is copied from various sources, and may include many inaccuracies. Expert help is welcome.