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Vanessa Fabricius, 1807
Vanessa Fabricius, 1807; Magazin f. Insektenk. (Illiger) 6: 281, TS: Papilio atalanta Linnaeus
Nymphalis Latreille, 1804; Nouv. Dict. Hist. nat. 24 (6): 184, 199 (preocc. Nymphalis Kluk, 1780), TS: Papilio atalanta Linnaeus
= ; [NL4A, 250]
Cynthia Fabricius, 1807; Magazin f. Insektenk. (Illiger) 6: 281, TS: Papilio cardui Linnaeus
= ; [NL4A, 250]
Pyrameis Hübner, [1819]; Verz. bek. Schmett. (3): 33, TS: Papilio atalanta Linnaeus
= ; [NL4A, 250]
Bassaris Hübner, [1821]; Samml. exot. Schmett. 2 : pl. [24], TS: Papilio itea Fabricius
= ; [NL4A, 250]
Ammiralis Rennie, 1832; Conspectus Butts. Moths: 10, TS: Papilio atalanta Linnaeus
= ; [NL4A, 250]
Phanessa Sodoffsky, 1837; Bull. Soc. imp. Nat. Moscou 1837 (6) : 80 (unj. emend. of Vanessa Fabricius, 1807)
= ; [NL4A, 250]
Neopyrameis Scudder, 1889; Butts eastern U.S. Canada 1: 434, TS: Papilio cardui Linnaeus
= ; [NL4A, 250]
Fieldia Niculescu, 1979; Revue Verv. Hist. nat. 36 (1-3): 3, TS: Vanessa carye Hübner
= ; [NL4A, 250]
Vanessa, NSG Voucher Specimen [Wahlberg]

Pyrameides ; Hübner, [1826], Verz. bek. Schmett. (Anz.) (1-9): 7 (missp.)
Vanessa (Nymphalinae) ; Moore, [1881], Lepid. Ceylon 1 (2): 48
Pyrameis (Nymphalinae) ; Moore, [1881], Lepid. Ceylon 1 (2): 49
Cynthia (Nymphalinae) ; Moore, [1881], Lepid. Ceylon 1 (2): 53
Vanessa ; Godman & Salvin, [1882], Biol. centr.-amer., Lep. Rhop. 1: 214
Pyrameis ; Godman & Salvin, [1882], Biol. centr.-amer., Lep. Rhop. 1: 217
Cynthia (Nymphalini) ; [MadL]
Vanessa (Nymphalidae) ; [BRU2, 25]
Vanessa (Nymphalini) ; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 207; [Wahlberg]; [AfrL]; [NL4A, 250]

fiVanessa atalanta (Linnaeus, 1758)
Azores, Canary Islands, NAF, EU, AM, Iran, NA, Guatemala, Haiti, New Zealand, Baluchistan, Hawaii. See [About maps] Amiraali Red Admiral Amiralfjäril Amiral Admiral Le Vulcain Numerada Rusałka admirał
Papilio atalanta Linnaeus, 1758; Syst. Nat. (Edn 10) 1 : 478, TL: Sweden
admiralis (Retzius, 1783)
Pyrameis ammiralis Godart, 1821
Pyrameis atalanta italica Stichel, 1900
Pyrameis atalanta ab. umbrosa Fischer, 1908; Soc. Ent. 23 (17) : 130
Pyrameis atalanta minutior Verity & Querci, 1924
?Pyrameis atalanta ab. rubra Fritsch, 1913; Ent. Rundschau 30 : 25
Vanessa atalanta, E.H. Strickland Ent. Mus, U of [ALBERTA]
Vanessa atalanta, NSG Voucher Specimen [Wahlberg]
Vanessa atalanta, Butterflies and Moths of North America [BAMONA]
Vanessa atalanta, Larvae of North-European Lepidoptera [Kimmo Silvonen]
Vanessa atalanta, Svenska fjärilar [SVF]
Vanessa atalanta, Norges Sommerfugler [NORLEP]
V. atalanta, Georgia Lepidoptera (USA) [GALEPS]
Vanessa atalanta, Moths and Butterflies of Europe [MBE]
Vanessa atalanta, Europaische Schmetterlinge [Mario Maier]
Vanessa atalanta, Captain's European Butterfly Guide [Simon Coombes]
Pyrameis atalanta ; Godman & Salvin, [1882], Biol. centr.-amer., Lep. Rhop. 1: 219; Godman & Salvin, [1901], Biol. centr.-amer., Lep. Rhop. 2: 684
Vanessa atalanta ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 23; [BOW: pl. 3, f. 18]; [NACL, #4437]; [BMAT: 47, pl. 14, f. 7-10]; [BRU2: 25, pl. 14, f. 1-3]; [Opler]; [NL4A, #2039]; [AfrL]; [Otakar Kudrna]
[atalanta-1u.jpg] 500x390 (~46Kb) upperside Hoe lake, Devon, UK 15/08/96, Photo © Simon Coombes [atalanta-1v.jpg] 450x356 (~46Kb) underside Hoe lake, Devon, UK 15/08/96, Photo © Simon Coombes [atalanta-2v.jpg] 500x512 (~50Kb) underside Finland, Kangasala, Savo, 6814:338, 25.8.1996, Photo © Tero Piirainen
[atalanta-3l.jpg] 409x615 (~60Kb) larva Germany, Baden-Württemberg, Maulbronn (8°47'E, 48°59'N, 275m), 02.09.1999 (on Urtica dioica), Photo © Karl Hofsäß [atalanta-3p.jpg] 409x615 (~61Kb) pupa Germany, Baden-Württemberg, Schönenberg (8°49'E, 48°57'N, 225m), 01.09.1999, Photo © Karl Hofsäß [atalanta-4.jpg] 1000x741 (~77Kb) FINLAND, Ab: Suomusjärvi, 669:31, 6.9.1998, Photo © Markku Savela
[atalanta-4u.jpg] 620x834 (~67Kb) upperside FINLAND, Ab: Suomusjärvi, 669:31, 13.9.1998, Photo © Markku Savela [atalanta-4v.jpg] 900x739 (~98Kb) underside FINLAND, Ab: Suomusjärvi, 669:31, 13.9.1998, Photo © Markku Savela [atalanta-5a.jpg] 886x805 (~131Kb) Finland, Espoo, Otaniemi (CSC:n piha), 16.8.2000, Photo © Pekka Järveläinen
[atalanta-5b.jpg] 665x995 (~136Kb) Finland, Espoo, Otaniemi (CSC:n piha), 16.8.2000, Photo © Pekka Järveläinen [atalanta-5l.jpg] 596x400 (~83Kb) larva Central Europe 1990-2000, Photo © Michael Borsch [atalanta-5p.jpg] 400x600 (~48Kb) pupa Central Europe 1990-2000, Photo © Michael Borsch
[atalanta-5u.jpg] 596x400 (~41Kb) upperside Central Europe 1990-2000, Photo © Michael Borsch [atalanta-6v.jpg] 600x800 (~65Kb) underside Sweden, Ekebo, Julita, 7.9.2003, Photo © Leif Wahlberg [atalanta-7u.jpg] 594x840 (~99Kb) upperside Sweden, Ekebo, Julita, 7.9.2003, Photo © Leif Wahlberg
[atalanta-8u.jpg] 820x551 (~142Kb) upperside Russia: Moscow area (36°25'E, 56°00'N), 02.09.2007, Photo © D. Smirnov [atalanta-8v.jpg] 800x698 (~142Kb) underside Russia, Moscow area, August 2004, Photo © D. Smirnov [atalanta-9v.jpg] 476x592 (~46Kb) underside Czech Republic, sw Bohemia, Sumava Mtns., Bucina, 9.IX.2005, Photo © L.Dvorák
[atalanta-a1v.jpg] 1200x1600 (~284Kb) underside USA: Alabama, 1.6.2003, Photo © Vitaly Charny

Larva on Urtica, U. urens, U. dioica, Humulus lupulus [SPRK], Urtica membranacea, U. pilulifera, Parietaria officinalis [BMAT], (Europe and elsewhere) Parietaria diffusa, P. alsinifolia, P. debilis, Salix sp. [BMAT]


Vanessa (Vanessa) atalanta atalanta ; Vane-Wright & Hughes, 2007, J. Lep. Soc. 61 (4): 212
V. a. rubria (Fruhstorfer, 1909)
Mexico, California atalanta rubria (Fruhstorfer, 1909) (Pyrameis); Int. ent. Zs. 3 (17): 94, TL: Mexico
Vanessa atalanta var. edwardsi Grinnell, 1918; , TL: California
= ; [NL4A, #2039a]
Vanessa atalanta rubria, Listing of Mexican Butterflies [ILMB]
Vanessa atalanta rubria, Butterflies of America [BOA]
Vanessa atalanta rubria, Butterflies of Southeastern Arizon [Bruce Walsh]
Vanessa atalanta rubria ; [NL4A, #2039a]; [NACL, #4437a]
Vanessa (Vanessa) atalanta rubria ; Vane-Wright & Hughes, 2007, J. Lep. Soc. 61 (4): 212

Vanessa tameamea (Eschscholtz, 1821)
Hawaii. See [About maps] Kamehameha
Pyrameis[?] tameamea Eschscholtz, 1821; in Kotzebue, Entdeck. Reise Süd-See 3 (App. 5): 207, pl. 5, f. 8a-b
Pyrameis cordelia Doubleday, 1847; Gen. diurn. Lep. (1): pl. 25, f. 3
Pyrameis tammeamea[sic, recte tameamea] ; Butler, 1874, Proc. Zool. Soc. Lond. 1874 : 284; [BOW: pl. 15, f. 3]
Vanessa (Vanessa) tameamea ; Vane-Wright & Hughes, 2007, J. Lep. Soc. 61 (4): 212

Vanessa indica (Herbst, 1794)
Ceylon, S.India (hills), Coorg, Himalayas - Kashmir, NE.India (hills), N.Burma. See [About maps] Indian Red Admiral Asian Admiral
Papilio indica Herbst, 1794; in Jablonsky, Naturs. Schmett. 7: pl.180. f. 1,2, TL: India
?Hamadryas calliroe Hübner, 1808
Pyrameis[?] callirhoe Millière, 1868; Ann. Soc. Linn. Lyon 16: 26, pl. 88, f. 1-2
Pyrameis indica v. asakurae Matsumura, 1908; Ent. Zs. 22 (39) : 158, TL: Formosa
horishanus (Nire, 1917) (Pyrameis); Dobuts. Zasahi, Tokyo 29: 145-148
Vanessa indica, NSG Voucher Specimen [Wahlberg]
Vanessa indica indica, Butterflies in Indo-China [Yutaka Inayoshi]
Pyrameis calliroe ; Hübner, [1819], Verz. bek. Schmett. (3): 33
Vanessa indica ab. horishana ; Murayama & Shimonoya, 1963, Tyô to Ga 13 (3): 53, f. 31-34
Pyrameis indica ; Moore, [1881], Lepid. Ceylon 1 (2): 50, pl. 27, f. 2; [BOW: pl. 151, f. 7]
Vanessa indica ; [MRS, 569]; [BRU2: 25, pl. 14, f. 4-6]
Vanessa (Vanessa) indica ; Vane-Wright & Hughes, 2007, J. Lep. Soc. 61 (4): 212
[indica-1v.jpg] 951x624 (~91Kb) underside Sukayu, Aomori, Japan, 8-90, Photo © S. Shuichi Haupt [indica-2mu.jpg] 400x300 (~41Kb) upperside male The hill Kuruntei 15 km upstream of the village Nizhnii Tsasuchei, Onon district, Chita Province, S Transbaikalia (Dahuria), Siberia, Russia. 6th July 1996, Photo © Oleg Kosterin [indica-2mv.jpg] 600x800 (~114Kb) underside male The hill Kuruntei 15 km upstream of the village Nizhnii Tsasuchei, Onon district, Chita Province, S Transbaikalia (Dahuria), Siberia, Russia. 6th July 1996, Photo © Oleg Kosterin
[indica-3.jpg] 425x347 (~38Kb) Jamori [West Arunachal Pradesh], 28 October 2003, Photo © Kedar Tokekar [indica-3v.jpg] 325x327 (~29Kb) underside Jamori [West Arunachal Pradesh], 28 October 2003, Photo © Kedar Tokekar

Larva on Urtica [BIR], Girardinia heterophylla, Boehmeria densiflora, B. nivea, Urtica thunbergiana [MRS]


Sikkim - C.India, China, Taiwan Vanessa indica indica ; [BOR, 276]
Vanessa (Vanessa) indica indica ; Vane-Wright & Hughes, 2007, J. Lep. Soc. 61 (4): 212
V. i. nubicola (Fruhstorfer, 1898)
Ceylon ?nubicola (Fruhstorfer, 1898) (Pyrameis); Ent. Nachr. 24 (4) : 61 [ ?? ]
Vanessa indica nubicola ; [BOR, 276]
Vanessa (Vanessa) indica nubicola ; Vane-Wright & Hughes, 2007, J. Lep. Soc. 61 (4): 212
V. i. pholoe (Fruhstorfer, 1912)
S.India (Nilgiris) pholoe (Fruhstorfer, 1912) (Pyrameis[?]); in Seitz, Gross-Schmett. Erde 9 : 525
Vanessa indica pholoe ; [BOR, 276]
Vanessa (Vanessa) indica pholoe ; Vane-Wright & Hughes, 2007, J. Lep. Soc. 61 (4): 212

Vanessa vulcania (Godart, 1819)
Canary Islands, Madeira Canary Red Admiral
Pyrameis[?] vulcania Godart, 1819; Encyclopédie Méthodique. 9 (1): 320, TL: Tenerife, Canary Islands
Pyrameis callirhoe occidentalis C. Felder, 1862; Verh. zool.-bot. Ges. Wien 12 (1/2) : 473
Vanessa vulcania, NSG Voucher Specimen [Wahlberg]
Vanessa vulcania, Moths and Butterflies of Europe [MBE]
Vanessa indica vulcania, Europaische Schmetterlinge [Mario Maier]
Vanessa vulcania, Captain's European Butterfly Guide [Simon Coombes]
Vanessa indica vulcania ;
Vanessa vulcania ; [Otakar Kudrna]
Vanessa (Vanessa) vulcania ; Vane-Wright & Hughes, 2007, J. Lep. Soc. 61 (4): 212
[vulcania-1.jpg] 416x363 (~39Kb) Tenerife, Puerto de la Cruz, 2/4/97, Photo © Simon Coombes [vulcania-1u.jpg] 398x312 (~35Kb) upperside Tenerife, Puerto de la Cruz, 2/4/97, Photo © Simon Coombes


Vanessa abyssinica (C. & R. Felder, [1867])
Kenya, Abyssinia. See [About maps] Abyssinian Admiral
Pyrameis abyssinica C. & R. Felder, [1867]; Reise Fregatte Novara, Bd 2 (Abth. 2) (3): 397, TL: Abyssinia
Vanessa abyssinica, NSG Voucher Specimen [Wahlberg]
Antanartia abyssinica ; [EBW]; [BOW: pl. 92, f. 1]
Vanessa abyssinica ; Wahlberg, Brower & Nylin, 2005, Biol. J. Linn. Soc. 86: 236; [Joseph Belicek, 2006]
Vanessa (Vanessa) abyssinica ; Vane-Wright & Hughes, 2007, J. Lep. Soc. 61 (4): 212
Vanessa abyssinica ; [AfrL]

Larva on Australina, Obetia, Pouzolzia, Urtica [BK]

Ethiopia Antanartia abyssinica abyssinica
Antanartia abyssinica abyssinica ; [BAFR, 239]
Vanessa (Vanessa) abyssinica abyssinica ; Vane-Wright & Hughes, 2007, J. Lep. Soc. 61 (4): 212
Vanessa abyssinica abyssinica ; [AfrL]
V. a. jacksoni (Howarth, 1966)
highlands (Kenya, N.Tanzania) Antanartia abyssinica jacksoni Howarth, 1966; Bull. Br. Mus. nat. Hist. (Ent.) 18 (2) : 31
Antanartia abyssinica jacksoni ; [BAFR, 239 (text)]; [BK, 354]
Vanessa (Vanessa) abyssinica jacksoni ; Vane-Wright & Hughes, 2007, J. Lep. Soc. 61 (4): 212
Vanessa abyssinica jacksoni ; [AfrL]
V. a. vansomereni (Howarth, 1966)
W.Uganda, Rwanda, Zaire (Kivu) Antanartia abyssinica vansomereni Howarth, 1966; Bull. Br. Mus. nat. Hist. (Ent.) 18 (2) : 31
Antanartia abyssinica vansomereni ; [BAFR, 239 (text)]
Vanessa (Vanessa) abyssinica vansomereni ; Vane-Wright & Hughes, 2007, J. Lep. Soc. 61 (4): 212
Vanessa abyssinica vansomereni ; [AfrL]

Vanessa dejeanii Godart, [1824]
Java. See [About maps] Vanessa dejeanii Godart, [1824]; Encyclopédie Méthodique. 9 (2): 821
Vanessa dejeanii, NSG Voucher Specimen [Wahlberg]
Vanessa dejeani ; [EBW]; [BOR, 276]
Pyrameis dejeani ; [BOW: pl. 151, f. 3]
Vanessa (Vanessa) dejeanii ; Vane-Wright & Hughes, 2007, J. Lep. Soc. 61 (4): 212
Java, ?Bali, Lombok Pyrameis dejeani sambaluna Fruhstorfer, 1898; Dt. ent. Z. Iris 11 (1): 150
Vanessa dejeanii dejeanii ; [BOR, 276]
Vanessa dejeani dejeani ; [BOR, 276]
Vanessa dejeani sambulana[sic?] ; [BOR, 276]
Vanessa (Vanessa) dejeanii dejeanii ; Vane-Wright & Hughes, 2007, J. Lep. Soc. 61 (4): 212
V. d. mounseyi (Talbot, 1936)
Philippines (Mindanao, Samar) dejeanii mounseyi (Talbot, 1936); Entomologist 69: 112
Vanessa (Vanessa) dejeanii mounseyi ; Vane-Wright & Hughes, 2007, J. Lep. Soc. 61 (4): 212

Vanessa samani (Hagen, 1895)
Sumatra (Mt. Korintji, Karo Mts.). See [About maps] Pyrameis samani Hagen, 1895; Dt. ent. Z. Iris 7 (2) : 359, TL: Sumatra
Vanessa samani, NSG Voucher Specimen [Wahlberg]
Pyrameis samani ; [BOW: pl. 151, f. 3 (text only)]
Vanessa samani ; [BOR, 276]
Vanessa (Vanessa) samani ; Vane-Wright & Hughes, 2007, J. Lep. Soc. 61 (4): 212

Vanessa buana (Fruhstorfer, 1898)
S.Sulawesi. See [About maps] Lompobatang Lady
Pyrameis indica buana Fruhstorfer, 1898; Ent. Nachr. 24 (4) : 60
Vanessa buana, NSG Voucher Specimen [Wahlberg]
Vanessa indica buana ; [BOR, 276]
Vanessa (Vanessa) buana ; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 207; Vane-Wright & Hughes, 2007, J. Lep. Soc. 61 (4): 212

Vanessa dilecta Hanafusa, 1992
W.Timor. See [About maps] Vanessa dilecta Hanafusa, 1992; Futao (10): ?, TL: W.Timor
Vanessa (Vanessa) dilecta ; Vane-Wright & Hughes, 2007, J. Lep. Soc. 61 (4): 212

Vanessa itea (Fabricius, 1775)
Australia, New Zealand. See [About maps] Papilio itea Fabricius, 1775; Syst. Ent.: 498
Vanessa itea, NSG Voucher Specimen [Wahlberg]
Vanessa itea, Lepidoptera Larvae of Australia [Don Herbison-Evans]
Vanessa itea ; [EBW]
Pyrameis itea ; [BOW: pl. 151, f. 5]
Vanessa itea ; [BAUR, 212]
Bassaris itea ;
[itea-1u.jpg] 478x324 (~30Kb) upperside New Zealand: Dunedin, March 2007, Photo © Christopher A. Rickards [itea-1v.jpg] 324x487 (~29Kb) underside New Zealand: Dunedin, March 2007, Photo © Christopher A. Rickards

Larva on Urtica [BAUR], Parietaria debilis, Soleirolia soleirolii, Urtica incisa, U. urens K. L. & L. E. Dunn, 1991, Review Austr. Butts. (1-4): (120-140)



Vanessa gonerilla (Fabricius, 1775)
New Zealand Papilio gonerilla Fabricius, 1775; Syst. Ent.: 498
Vanessa gonerilla, NSG Voucher Specimen [Wahlberg]
Pyrameis gonarilla ; [BOW: pl. 151, f. 6 (missp.)]
Vanessa gonerilla ; [EBW]
Bassaris gonerilla ;
New Zealand Vanessa gonerilla gonerilla ; [BAUR, 212]
Bassaris gonerilla gonerilla ;
[gonerilla_gonerilla-1u.jpg] 487x324 (~26Kb) upperside New Zealand: Dunedin, December 2005, Photo © Christopher A. Rickards [gonerilla_gonerilla-1v.jpg] 476x324 (~31Kb) underside New Zealand: Dunedin, March 2007, Photo © Christopher A. Rickards

V. g. ida (Alfken, 1899)
Chathan Is. Pyrameis ida Alfken, 1899; Zool. Anz.: 5
Pyrameis ida var. argentata Alfken, 1899; Zool. Anz.: 5 [?]
Vanessa gonerilla ida ; [BAUR, 212 (text)]
Bassaris gonerilla ida ;
[ida-1u.jpg] 324x487 (~22Kb) upperside New Zealand: Clatham Islands, December 2007, Photo © Christopher A. Rickards [ida-1va.jpg] 487x324 (~23Kb) underside New Zealand: Clatham Islands, December 2007, Photo © Christopher A. Rickards [ida-1vb.jpg] 486x324 (~40Kb) underside New Zealand: Clatham Islands, December 2007, Photo © Christopher A. Rickards


fiVanessa cardui (Linnaeus, 1758)
EU, Africa, Asia, AU, NA, Hawaii. See [About maps] Ohdakeperhonen Painted Lady Tistelfjäril Tistelfuks Distelfalter La Belle Dame Bella Dama o Cardero Rusałka osetnik
Papilio cardui Linnaeus, 1758; Syst. Nat. (Edn 10) 1 : 475, TL: Sweden
Papilio belladonna Linnaeus, 1758; (unavail.)
= ; [NL4A, #2041]
?Papilio carduelis Seba, 1765
Papilio carduelis Cramer, 1775; , TL: "Cap de Bonne Espérance"
= ; [NL4A, #2041]
Vanessa elymi Rambur, 1829; , TL: France
= ; [NL4A, #2041]
var. leachiana (Doubleday, 1849) (Pyrameis); (nom. nud.)
= ; [NL4A, #2041]
ab. ate (Strecker, 1878) (Pyrameis); , TL: Ohio
= ; [NL4A, #2041]
ab. pallens Noel, 1881; , TL: France
= ; [NL4A, #2041]
var. pallida Schøyen, 1881; , TL: Norway
= ; [NL4A, #2041]
ab. inornata Bramson, 1886; , TL: Rusia
= ; [NL4A, #2041]
ab. minor Failla-Tedaldi, 1887; , TL: Italy
= ; [NL4A, #2041]
cardui semisuffusa Cockerell, 1889; , TL: ?
= ; [NL4A, #2041]
f. minor (Cockerell, 1890) (Pyrameis); , TL: Colorado
= ; [NL4A, #2041]
ab. wiskotti Standfuss, 1895; , TL: Switzerland
= ; [NL4A, #2041]
ab. minor (Cannviello, 1900) (Pyrameis); , TL: Italy
= ; [NL4A, #2041]
var. huntera (Lowe, 1902) (Pyrameis); (preocc. Fabricius, 1775), TL: Italy
= ; [NL4A, #2041]
ab. carduelis (Schultz, 1905) (Pyrameis); , TL: ?
= ; [NL4A, #2041]
ab. priameis (Schultz, 1905) (Pyrameis); , TL: ?
= ; [NL4A, #2041]
ab. carduelina (Alphéraky, 1908) (Pyrameis); , TL: Russia
= ; [NL4A, #2041]
Pyrameis cardui japonica Stichel, 1908; , TL: Japan,China
= ; [NL4A, #2041]
ab. ocellata (Rebel, 1910) (Pyrameis); , TL: ?
= ; [NL4A, #2041]
ab. carnea (Fritsch, 1912) (Pyrameis); , TL: Germany
= ; [NL4A, #2041]
ab. melanosa (Cabeau, 1913) (Pyrameis); , TL: France
= ; [NL4A, #2041]
f. brunnea-albimaculata (Reuss, 1916) (Pyrameis); , TL: Germany
= ; [NL4A, #2041]
f. rosacea (Reuss, 1916) (Pyrameis); , TL: Germany
= ; [NL4A, #2041]
f. ochracea (Reuss, 1918) (Pyrameis); , TL: Germany
= ; [NL4A, #2041]
Pyrameis cardui universa Verity, 1919; , TL: Italy
= ; [NL4A, #2041]
f. conjuncta (Verity, 1919) (Pyrameis); , TL: "Europe"
= ; [NL4A, #2041]
f. emiolymi (Verity, 1919) (Pyrameis); , TL: Italy
= ; [NL4A, #2041]
f. infrabrunnea (Verity, 1919) (Pyrameis); (nom. nud.)
= ; [NL4A, #2041]
f. infraflava (Verity, 1919) (Pyrameis); (nom. nud.)
= ; [NL4A, #2041]
f. infragrisea (Verity, 1919) (Pyrameis); (nom. nud.)
= ; [NL4A, #2041]
f. infranigrans (Verity, 1919) (Pyrameis); (nom. nud.)
= ; [NL4A, #2041]
f. infaochracea (Verity, 1919) (Pyrameis); (nom. nud.)
= ; [NL4A, #2041]
f. inops (Verity, 1919) (Pyrameis); , TL: [Italy]
= ; [NL4A, #2041]
f. sexispupillata (Verity, 1919) (Pyrameis); , TL: [Italy]
= ; [NL4A, #2041]
f. septiespupillata (Verity, 1919) (Pyrameis); , TL: [Italy]
= ; [NL4A, #2041]
?Pyrameis cardui inops Verity & Querci, 1924
ab. maria (Stephan, 1924) (Pyrameis); , TL: Germany
= ; [NL4A, #2041]
ab. martha-maria (Stephan, 1924) (Pyrameis); , TL: Germany
= ; [NL4A, #2041]
ab. schoenfellneri (Hoffmann, 1925) (Pyrameis); , TL: Austria
= ; [NL4A, #2041]
ab. subfracta (Stach, 1925) (Pyrameis); , TL: Poland
= ; [NL4A, #2041]
ab. takesakiana (Kato, 1925) (Pyrameis); , TL: Japan
= ; [NL4A, #2041]
cardui littoralis (Souza, 1926) (Pyrameis); , TL: Portugal
= ; [NL4A, #2041]
ab. rosea (Pionneau, 1926) (Pyrameis); , TL: France
= ; [NL4A, #2041]
Cynhia cardui jacksoni Clark, 1927; , TL: Massachusetts/California),Mongolia,France
= ; [NL4A, #2041]
ab. browni (Meilhan, 1928) (Pyrameis); , TL: [France]
= ; [NL4A, #2041]
ab. rogeri (Meilhan, 1928) (Pyrameis); , TL: France
= ; [NL4A, #2041]
ab. varini (Meilhan, 1928) (Pyrameis); , TL: [France]
= ; [NL4A, #2041]
ab. flava (Bandermann, 1928) (Pyrameis); , TL: Germany
= ; [NL4A, #2041]
ab. johni (Fischer, 1932) (Pyrameis); , TL: Switzerland
= ; [NL4A, #2041]
ab. pallida Frohawk, 1938; , TL: ?
= ; [NL4A, #2041]
Vanessa cardui ushuwaia Bryk, 1944; , TL: "Argentina"
= ; [NL4A, #2041]
Vanessa cardui japonica f. koreana Bryk, 1946; , TL: Korea
= ; [NL4A, #2041]
ab. albicans Verity, 1950; , TL: ?
= ; [NL4A, #2041]
ab. infralutea Verity, 1950; , TL: ?
= ; [NL4A, #2041]
f. albipuncta Lempke, 1956; , TL: Netherlands
= ; [NL4A, #2041]
f. nigripuncta Lempke, 1956; , TL: Netherlands
= ; [NL4A, #2041]
minor (Cockerell, 1890)
Pyrameis cardui litoralis de Souza, 1926
Pyrameis cardui rogeri Gaede, 1931
Vanessa cardui, E.H. Strickland Ent. Mus, U of [ALBERTA]
Vanessa cardui, Listing of Mexican Butterflies [ILMB]
Vanessa cardui, Butterflies of America [BOA]
Vanessa cardui, NSG Voucher Specimen [Wahlberg]
Vanessa cardui, Butterflies and Moths of North America [BAMONA]
Vanessa cardui cardui, Butterflies in Indo-China [Yutaka Inayoshi]
Vanessa cardui, Larvae of North-European Lepidoptera [Kimmo Silvonen]
Vanessa cardui, Svenska fjärilar [SVF]
Vanessa cardui, Norges Sommerfugler [NORLEP]
Vanessa cardui, Lepidoptera Larvae of Australia [Don Herbison-Evans]
V. cardui, Georgia Lepidoptera (USA) [GALEPS]
Vanessa cardui, Moths and Butterflies of Europe [MBE]
Vanessa cardui, Butterflies of Southeastern Arizon [Bruce Walsh]
Vanessa cardui, Lepidoptera of French Antilles [Pierre Zagatti]
Vanessa cardui, Europaische Schmetterlinge [Mario Maier]
Vanessa cardui, Captain's European Butterfly Guide [Simon Coombes]
Vanessa kardui ab. takesakiana ; Murayama & Shimonoya, 1966, Tyô to Ga 15 (3/4) : 61, f. 24, 27
Pyrameis hemielymi ; Gaede, 1930, (missp.)
Pyrameis cardui ; Moore, [1881], Lepid. Ceylon 1 (2): 50, pl. 27, f. 1, 1a; Godman & Salvin, [1882], Biol. centr.-amer., Lep. Rhop. 1: 217; Godman & Salvin, [1901], Biol. centr.-amer., Lep. Rhop. 2: 683
Vanessa cardui ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 24; [BOW: pl. 3, f. 19]; [EBW]; [NACL, #4435]; [BCR: 180, pl. 28]; [BOR, 276]; [MRS, 570]; [BMAT: 48, pl. 14, f. 16-17]; [BAFR, 238]; [BRU2: 25, pl. 14, f. 7-9]; [Opler]; [NL4A, #2041]; [MadL]; [AfrL]; [Otakar Kudrna]
Vanessa (Cynthia) cardui ; [BMP: 161, pl. 23, f. 1]
Vanessa cardui cardui ; [BK, 353, pl. 50]
[cardui-1u.jpg] 500x385 (~65Kb) upperside Turunc, SW Turkey 14/04/95, Photo © Simon Coombes [cardui-1v.jpg] 500x408 (~54Kb) underside Turunc, SW Turkey 14/04/95, Photo © Simon Coombes [cardui-2u.jpg] 500x416 (~35Kb) upperside Finland, A: Lemland, Herrö, 6/1995, Photo © Tero Piirainen
[cardui-2v.jpg] 500x424 (~31Kb) underside Sweden, Gotland, Roma, 24.7.1996, Photo © Tero Piirainen [cardui-3u.jpg] 500x424 (~32Kb) upperside Finland, Kangasala, Saarenmaa, 681:33, 23.6.1996, Photo © Tero Piirainen [cardui-4l.jpg] 597x400 (~65Kb) larva Central Europe 1990-2000, Photo © Michael Borsch
[cardui-4u.jpg] 597x400 (~50Kb) upperside Central Europe 1990-2000, Photo © Michael Borsch [cardui-5.jpg] 600x435 (~34Kb) Morocco, Aït-Baha, Anti-Atlas, Photo © Jean Delacre [cardui-6u.jpg] 1600x1200 (~182Kb) upperside Japan: Lake Kawaguchiko, Yamanashi Prefecture, 23.8.2007, Photo © Peter Payzant
[cardui-7u.jpg] 2048x1536 (~255Kb) upperside CANADA: July 2005 Edmonton, Alberta, Photo © Lois Maksymuik [cardui-8u.jpg] 727x624 (~120Kb) upperside Russia: Moscow area (36°25'E, 56°00'N), 29.08.2007, Photo © D. Smirnov [cardui-8v.jpg] 536x486 (~84Kb) underside Russia: Moscow area (36°25'E, 56°00'N), 29.08.2007, Photo © D. Smirnov
[cardui-9u.jpg] 1600x1200 (~213Kb) upperside USA: Alabama, 9.8.2003, Photo © Vitaly Charny [cardui-9v.jpg] 1600x1200 (~222Kb) underside USA: Alabama, 9.8.2003, Photo © Vitaly Charny [cardui-u.jpg] 900x595 (~106Kb) upperside FINLAND: Ka: Virolahti, Virojoki 671:53 (Lavakallio), 13.6.1998, Photo © Markku Savela
[cardui-v.jpg] 700x955 (~111Kb) underside USA: End of FR 6520 /Johnson Ridge, Mt. Baker Snoqualmie Nat. Forest (47°47'48N 121°15'45W), Snohomish Co., WA, 7.8.2001, Photo © Markku Savela

Larva on Urtica dioeca, Arctium minus, A. tomentosum, Carduus crispus, Cirsium heterophyllum, C. vulgare, C. arvense, Achillea millefolium, Echium vulgare [SPRK]

Larva on Carduus, Artotis grandis, Gnaphalium purpureum, Stobaea discolor, Gazania spp., Boehmeria nivea, Phaseolus vulgaris, (and many others) [PBSA] Urtica spp., Debregeasia bicolor, Carduus spp., Artemisia spp., Anchusa spp., Blumea spp., Zornia diphylla [MRS], Citrullus colocynthis, Vitis vinifera, Medicago sativa, Phaseolus vulgaris, Echium plantagineum, Micropus bombicinus, M. erectus, (in Egypt) Malva parviflora [BMAT]


'japonica'
[japonica-1u.jpg] 1600x1321 (~222Kb) upperside Japan: Aiuchi, Ichiuramua, Aomori P., 8.8.2004, Photo © Norbert Kondla [japonica-1v.jpg] 1600x1307 (~236Kb) underside Japan: Aiuchi, Ichiuramua, Aomori P., 8.8.2004, Photo © Norbert Kondla



Vanessa kershawi (McCoy, 1868)
Australia. See [About maps] Australian Painted Lady
Cynthia kershawi McCoy, 1868; Ann. Mag. nat. Hist. 4 (1) : 76
Pyrameis suffusa Olliff, 1888
Pyrameis lucasii Miskin, 1889
Cynthia austral Meredith, 1891
Vanessa kershawi, NSG Voucher Specimen [Wahlberg]
Vanessa kershawi, Lepidoptera Larvae of Australia [Don Herbison-Evans]
kershawii ; Kirby, 1871, (missp.)
lucasi ; Fruhstorfer, 1912, (missp.)
Vanessa cardui kershawi ; [BAUR, 212]
Misidentified/misapplied:
?Pyrameis kershawi; Godman & Salvin, [1882], Biol. centr.-amer., Lep. Rhop. 1: 218
[kershawi-1u.jpg] 488x324 (~42Kb) upperside New Zealand: Nelson, February 2006, Photo © Christopher A. Rickards [kershawi-1v.jpg] 487x324 (~35Kb) underside New Zealand: Nelson, February 2006, Photo © Christopher A. Rickards

Larva on Helichrysum, Cryptostemma, Artemisia [BAUR], Chrysocephalum semipapposum Dunn, 1997, Victorian Ent. 27 (6): (114-118), Lavandula officinalis, Ammobium alatum, Arctotheca calendula, Artemisia sp., Carduus spp., Gnaphalium spicatum, G. sp., Helichrysum apiculatum, H. bracteatum, H. spp., Helipterum roseum, Onopordum acanthium, O. illyricum K. L. & L. E. Dunn, 1991, Review Austr. Butts. (1-4): (120-140)



Vanessa virginiensis (Drury, [1773])
Canary Islands, Madeira, Temperate NA, CA, Guatemala, Cuba, Hawaii, Galapagos. See [About maps] American Painted Lady
Papilio virginiensis Drury, [1773]; Illust. Nat. Hist. Exot. Insects 1 : 10, TL: New York; Maryland; Virginia
Papilio huntera Fabricius, 1775; Syst. Ent.: 499, TL: America
= ; [NL4A, #2045]
Papilio iole Cramer, 1775; (preocc. Denis & Schiffermüller, 1775), TL: Jamaica
= ; [NL4A, #2045]
Pyrameis huntera ab. fulvia Dodge, 1900; Can. Ent. 32 (3) : 92, TL: [Missouri; Louisiana]
= ; [NL4A, #2045]
Vanessa virginiensis var. ahwashtee Fox, 1921; , TL: California
= ; [NL4A, #2045]
Vanessa virginiensis ab. ♂ massachusettensis Gunder, 1925; Ent. News 36 (7) : 198, pl. 5, f. 10, TL: Massachusetts
= ; [NL4A, #2045]
Cynthia virginiensis tr.f. simmsi Gunder, 1927; , TL: Canada
= ; [NL4A, #2045]
Cynthia gnaphalii Fabricius, 1938; (repl. name), TL: America
= ; [NL4A, #2045]
Papilio jole Cramer, 1776; (missp.?)
Vanessa virginiensis, E.H. Strickland Ent. Mus, U of [ALBERTA]
Vanessa virginiensis, Listing of Mexican Butterflies [ILMB]
Vanessa virginiensis, Butterflies of America [BOA]
Vanessa virginiensis, NSG Voucher Specimen [Wahlberg]
Vanessa virginiensis, Butterflies and Moths of North America [BAMONA]
Vanessa virginiensis, Larvae of North-European Lepidoptera [Kimmo Silvonen]
Vanessa virginiensis, Georgia Lepidoptera (USA) [GALEPS]
Vanessa virginiensis, Moths and Butterflies of Europe [MBE]
hunteri ; Hübner, 1819, (missp.?)
Pyrameis huntera ; Godman & Salvin, [1882], Biol. centr.-amer., Lep. Rhop. 1: 218; Godman & Salvin, [1901], Biol. centr.-amer., Lep. Rhop. 2: 683; Williams, 1911, Proc. Calif. Acad. Sci. 1: 299
Vanessa huntera ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 23
Vanessa huntera fulvia ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 23
Vanessa virginiensis ; Linsley & Usinger, 1966, Proc. Cal. Acad. Sci. (4) 33: 156; [NACL, #4434]; [BOW: pl. 16, f. 16]; [BCR: 179, pl. 28]; [Opler]; Roque-Albelo, 2004, J. Lep. Soc. 58 (1): 41, f. 13-14; [NL4A, #2045]; [Otakar Kudrna]
[virginiensis-1.jpg] 900x616 (~108Kb) USA: Horsefall Beach, Oregon Dunes, (43°27'09N 124°16'41W ±1km), Douglas Co. OR, 1.8.2001, Photo © Markku Savela [virginiensis-1u.jpg] 900x645 (~76Kb) upperside USA: Horsefall Beach, Oregon Dunes, (43°27'09N 124°16'41W ±1km), Douglas Co. OR, 1.8.2001, Photo © Markku Savela [virginiensis-1v.jpg] 500x630 (~38Kb) underside USA: Horsefall Beach, Oregon Dunes, (43°27'09N 124°16'41W ±1km), Douglas Co. OR, 1.8.2001, Photo © Markku Savela
[virginiensis-2u.jpg] 1200x779 (~184Kb) upperside USA: Alabama, 19.6.1999, Photo © Vitaly Charny [virginiensis-3v.jpg] 1600x1200 (~232Kb) underside USA: Alabama, 1.6.2003, Photo © Vitaly Charny [virginiensis-4.jpg] 1600x1200 (~567Kb) USA: Alabama, 6.6.2004, Photo © Vitaly Charny
[virginiensis-5.jpg] 1320x1711 (~204Kb) USA: Alabama, 24.7.2005, Photo © Vitaly Charny [virginiensis-6v.jpg] 1483x1224 (~191Kb) underside USA: Alabama, 30.9.2006, Photo © Vitaly Charny

Larva on Compositae, Gnaphalium [H&R]

Larva on Gnaphalium, Antennaria, Anaphalis [NAB], Senecio, Antirrhinum, Malva [BCR]



Vanessa altissima (Rosenberg & Talbot, 1914)
Ecuador, Peru. See [About maps] Pyrameis huntera altissima Rosenberg & Talbot, 1914; Trans. ent. Soc. Lond. 1913 (4) : 675, TL: Agualani, SE Peru & Baños, Rio Pastaza
Vanessa altissima, NSG Voucher Specimen [Wahlberg]
Vanessa altissima ; [EcuL]; [NL4A, #2037]

Vanessa braziliensis (Moore, 1883)
Brazil(São Paulo), Bolivia, Argentina. See [About maps] Pyrameis braziliensis Moore, 1883; , TL: [Brazil, São Paulo]
Pyrameis virginiensis var. rubia Staudinger, 1894; Dt. Ent. Z. Iris 7 (1) : 70, TL: Bolivia
= ; [NL4A, #2040]
Pyrameis huntera ab. dallasi Köhler, 1945; , TL: Argentina
= ; [NL4A, #2040]
Vanessa braziliensis, NSG Voucher Specimen [Wahlberg]
Vanessa virginiensis brasiliensis[sic] ; Brown & Mielke, 1967, J. Lep. Soc. 21 (2): 98
Vanessa braziliensis ; [EcuL]; [NL4A, #2040]

Vanessa terpsichore Philippi, 1859
Chile. See [About maps] Vanesja[sic] terpsichore Philippi, 1859; , TL: Chile
Vanessa terpsichore, NSG Voucher Specimen [Wahlberg]
Vanessa terpsichore ; [NL4A, #2044]

Vanessa myrinna (Doubleday, 1849)
Brazil(Minas Gerais, Rio de Janeiro), S.Ecuador, Peru, Colombia. See [About maps] Pyrameis myrinna Doubleday, 1849; , TL: Brazil
Vanessa syngenesiae Capronnier, 1874; (unavail.)
= ; [NL4A, #2043]
Pyrameis myrinna ab. incarnata Seitz, 1914; , TL: Brazil, Minas Gerais
= ; [NL4A, #2043]
Pyrameis myrinna ab. eunice Hall, 1917; , TL: Colombia
= ; [NL4A, #2043]
Pyrameis myrinna ab. merlinde Zikán, 1937; , TL: Brazil, Rio de Janeiro
= ; [NL4A, #2043]
Pyrameis myrinna; [BOW: pl. 42, f. 3], TL: 3
Vanessa myrinna, NSG Voucher Specimen [Wahlberg]
Vanessa myrinna ; Brown & Mielke, 1967, J. Lep. Soc. 21 (2): 98; [EcuL]; [NL4A, #2043]
[myrinna-1.jpg] 500x385 (~64Kb) Venezuela, Parai Tepui near Mount Roraima in the Gran Sabana, July 2002, Photo © Simon Coombes


Vanessa annabella × atalanta
? ; Dimock, 1973, J. Lep. Soc. 27 (4): (274-278)
[x_annabella_atalanta-1m.jpg] 900x721 (~128Kb) male California, Ventura Co., ex. l. 1972, Photo © Thomas Dimock


Vanessa annabella (Field, 1971)
California, .... See [About maps] West Coast Lady
Cynthia annabella Field, 1971
Cynthia annabella Field, 1971; , TL: California
Pyrameis carye var. muelleri Letcher, 1898; , TL: California
= ; [NL4A, #2038]
Pyrameis caryae[sic] var. intermedia Grinnel, 1918; , TL: California
= ; [NL4A, #2038]
Pyrameis caryae[sic] var. letcheri Grinnel, 1918; California
= ; [NL4A, #2038]
Cynthia carye tr.f. nivosa Gunder, 1927; , TL: California
= ; [NL4A, #2038]
Cynthia carye tr.f. schraderi Gunder, 1929; , TL: California
= ; [NL4A, #2038]
Vanessa annabella, E.H. Strickland Ent. Mus, U of [ALBERTA]
Vanessa annabella, Listing of Mexican Butterflies [ILMB]
Vanessa annabella, Butterflies of America [BOA]
Vanessa annabella, NSG Voucher Specimen [Wahlberg]
Vanessa annabella, Butterflies and Moths of North America [BAMONA]
Vanessa annabella, Butterflies of Southeastern Arizon [Bruce Walsh]
Vanessa annabella ; [NACL, #4436]; [Opler]
Vanessa anabella ; [EBW, (text only, missp.)]
Vanessa annabella ; [NL4A, #2038]
Misidentified/misapplied:
Vanessa carye; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 24

Larva on Malvaceae, Malva parviflora, Sidalcea, Althaea, Sphaeralcea, less Urtica [NAB]


Vanessa carye (Hübner, [1812])
Colombia, Ecuador, Peru, Bolivia, Paraguay, Uruguay, Argentina, Patagonia, Galapagos, Easter Island, Juan Fernandez, Tuoamotu Archipelago. See [About maps] Hamadryas carye Hübner, [1812]; Samml. exot. Schmett. 1: pl. 45. f. 1-2
Pyrameis caryoides Giacomelli, 1922; , TL: Argentina
= ; [NL4A, #2042]
Pyrameis carye f. minuscula Hayward, 1931; , TL: Argentina
= ; [NL4A, #2042]
Pyrameis caryae[sic] ab. bruchi Köhler, 1945; , TL: Argentina
= ; [NL4A, #2042]
Vanessa carye, NSG Voucher Specimen [Wahlberg]
Pyrameis carye ; Godman & Salvin, [1882], Biol. centr.-amer., Lep. Rhop. 1: 219; Godman & Salvin, [1901], Biol. centr.-amer., Lep. Rhop. 2: 684
Pyrameis caryae[sic, recte carye] ; Williams, 1911, Proc. Calif. Acad. Sci. 1: 300
Vanessa caryae[sic, recte carye] ; Linsley & Usinger, 1966, Proc. Cal. Acad. Sci. (4) 33: 156
Vanessa carye ; [BOW: pl. 16, pl. 17]; [EcuL]; Roque-Albelo, 2004, J. Lep. Soc. 58 (1): 41, f. 15-16; [NL4A, #2042]


Unknown or unplaced taxa

Vanessa aequatorialis Wagner, 1870; SB. bayer Akad. 1870 (2): 171 (nom. nud.), TL: Ecuador
aequatorialis ; [NL4A, 251]

Pyrameis chilena Herrich-Schäffer, 1865; (nom. nud.)
chilena ; [NL4A, 251]

Pyrameis venezuelae Herrich-Schäffer, 1865; (nom. nud.)
venezuelae ; [NL4A, 251]

Pyrameis vulpecula Herrich-Schäffer, 1865; (nom. nud.)
vulpecula ; [NL4A, 251]


20.2.2013 (20)

References:
[ALBERTA]
University of Alberta, E.H. Strickland Entomological Museum; Browse Entomology Collection
[About maps]
Warning! The maps are automatically generated from the textual information, and the process does not always produce acceptable result; See about maps for more info.
[AfrL]
Mark C. Williams; Checklist of Afrotropical Papilionoidea and Hesperoidea; Compiled by Mark C. Williams, 7th ed. (2008) (April 2007); Afrototropical Butterflies; e-mail: Mark C. Williams
[BAFR]
D'Abrera, 1980; Butterflies of the Afrotropical Region
[BAMONA]
Opler, Pavulaan, Stanford & Pogue; Butterflies and Moths of North America; Mountain Prairie Information Node
[BAUR]
D'Abrera, 1977; Butterflies of the Australian Region (2nd edn)
[BCA]
electronic; BIOLOGIA CENTRALI-AMERICANA
[BCR]
DeVries; The Butterflies of Costa Rica 1; Papilionidae, Pieridae, Nymphalidae
[BHL]
The; Biodiversity Heritage Library; and on; Internet Archive; Note: The linking to the content of Internet Archive is somewhat delicate -- notify me when links break!
[BIR]
Wynter-Blyth, 1957; Butterflies of the Indian Region; (1982 Reprint)
[BK]
Larsen, 1996; The Butterflies of Kenya and their natural history
[BMAT]
Tennent, 1996; The butterflies of Morocco, Algeria and Tunisia; ISBN 0 906802 05 9
[BMP]
Eliot (Ed.), Corbet & Pendlebury, 1992; The Butterflies of The Malay Peninsula; 4th edition; ISBN 983-9681-05-2
[BOA]
Interactive Listing of American Butterflies; Listado Interactivo de la Mariposas Americanas; Butterflies of America
[BOR]
D'Abrera, 1982, 1985, 1986; Butterflies of the Oriental Region, Part I-III
[BOW]
Lewis, 1974; Butterflies of the World; ISBN 0 245 52097 X
[BRU2]
Tuzov (ed.), Bogdanov, Churkin, Devyatkin, Dantchenko, Murzin, Samodurov, Zhdanko, 2000; Guide to the Butterflies of Russia and adjacent territories; Libytheidae, Danaidae, Nymphalidae, Riodinidae, Lycaenidae; Volume 2
[Bruce Walsh]
Bruce Walsh; e-mail: Bruce Walsh; The Moths of Southeastern Arizona; Butterflies of Southeastern Arizona
[Christopher A. Rickards]
Ann Arbor, Mi 48105, USA.; e-mail: Christopher A. Rickards
[CiNii]
http://ci.nii.ac.jp/
[D. Smirnov]
Russia, Moscow; e-mail: D. Smirnov
[Don Herbison-Evans]
and Stella A. Crossley; Lepidoptera Larvae of Australia; e-mail: Don Herbison-Evans; e-mail: Don Herbison-Evans
[EBW]
Smart, Paul, 1976; The Illustrated Encyclopedia of the Butterfly World; ISBN 0 600 31381 6
[EcuL]
Tommaso & Racheli, 2001; Fragmenta entomologica 33 (2): 213-380; An annotated list of Ecuadorian Butterflies (Lepidoptera: Papilionidae, Pieridae, Nymphalidae)
[GALEPS]
Moths and Butterflies of Georgia and the Southeastern United States; James K. Adams; Dept. of Natural Science, Dalton State College, Dalton, GA ; http://www.daltonstate.edu/galeps/index.htm
[H&R]
Higgins, L.G, Riley, N.D, 1970; A Field Guide to the Butterflies of Britain and Europe
[HBN]
Francis Hemming, 1937; HÜBNER, A bibliographical and systematic account of the entomological works of Jacob Hübner and of the supplements thereto by Carl Geyer, Gottfried Franz von Frölich and Gottlieb August Wilhelm Herrich-Schäffer, Vol 1 & 2
[ILMB]
Interactive Listing of Mexican Butterflies
[JLS]
Journal of the Lepidopterists' Society; the original J. Lep. Soc. articles
[Jean Delacre]
http://homepage.mac.com/jdelacre
[Joseph Belicek]
e-mail: Joseph Belicek
[Karl Hofsäß]
Karl Hofsaess; e-mail: Karl Hofsäß
[Kedar Tokekar]
India
[Kimmo Silvonen]
Espoo, Finland; Kimmo's Lep Site
[LAMAS-2005]
Gerardo Lamas; Entomofauna 26 (6): 57-100; A Bibliography of the Zoological Publications of Hans Fruhstorfer (1886* - 1922+)
[LAMAS-NEO]
Gerardo Lamas; Bibliography of butterflies; An Annocated Bibliography of the Neotropical Butterflies and Skipper (Lepididoptera: Papilionoidea and Hesperioidea); Revised Electronic Edition (2007)
[Leif Wahlberg]
Julita, Sweden; e-mail: Leif Wahlberg
[MBE]
Moths and Butteflies of Europe; Paolo Mazzei, Diego Reggianti and Ilaria Pimpinelli; http://www.leps.it/
[MRS]
Chou Io (Ed.); Monographia Rhopalocerum Sinensium, 1-2
[MadL]
Lees, D.C., Kremen, C. and Raharitsimba, H. 2003; in S.M. Goodman & J. P. Benstead (eds.), The Natural History of Madagascar; Classification, Diversity and Endemism of the Butterflies (Papilionoidea and Hesperioidea): A Revised Species Checklist: : pp. 762-793; University of Chicago Press. 1709 pp.
[Mario Maier]
Europäische Schmetterlinge; e-mail: Mario Maier
[Michael Borsch]
Geesthacht, Germany; contact through; e-mail: Francisco Welter-Schultes
[NAB]
Pyle, 1981; Field Guide to North American Butterflies, National Audubon Society; ISBN 0-394-51914-0
[NACL]
Hodges, 1983; Check List of the Lepidoptera of America North of Mexico
[NL4A]
Gerardo Lamas, 2004; Atlas of Neotropical Lepidoptera; Checklist: Part 4A; Hesperioidea - Papilionoidea
[NORLEP]
Norges Sommerfugler
[Norbert Kondla]
Canada, BC; e-mail: Norbert Kondla
[Oleg Kosterin]
Institue of Cytology & Genetics; Siberian Division of Russian Academy of Sciences; Novosibirsk; e-mail: Oleg Kosterin; http://pisum.bionet.nsc.ru/kosterin/
[Opler]
Opler & Warren, 2003 (April); Butterflies of North America. 2. Scientific Names List for Butterfly Species of North America, north of Mexico.
[PBSA]
Dickson, C.G.C (ed.), Kroon, D.M.; Pennington's Butterflies of Southern Africa; AD. DONKER 1978
[Peter Payzant]
Canada; http://www.pbase.com/ppayzant/profile
[Pierre Zagatti]
WWW pages; Catalogue of the Lepidoptera of the French Antilles; e-mail: Pierre Zagatti
[S. Shuichi Haupt]
e-mail: S. Shuichi Haupt; http://www.tu-berlin.de/fb7/ioeb/biologie/stefan.htm
[SPRK]
Seppänen, E. J, 1970; Suomen suurperhostoukkien ravintokasvit, Animalia Fennica 14
[SVF]
Svenska fjärilar; Naturhistoriska riksmuseet; (Sweden)
[Simon Coombes]
1 Park Street, Plymouth, PL3 4BL, UK; 01752 607854; e-mail: Simon Coombes; http://www.butterfly-guide.co.uk
[Tero Piirainen]
Tampere, Finland; e-mail: Tero Piirainen
[Thomas Dimock]
California, USA; e-mail: Thomas Dimock
[Vitaly Charny]
Birmingham, Alabama, USA
[Wahlberg]
Niklas Wahlberg; The NSG's voucher specimen database; Nymphalidae Systematics Group
[Yutaka Inayoshi]
A Check List of Butterflies in Indo-China; Chiefly from THAILAND, LAOS & VIETNAM


Some related literature:
Brown & Mielke, 1967
Lepidoptera of the Central Brazil Plateau. I. Preliminary list of Rhopalocera J. Lep. Soc. 21 (2): 77-106
Butler, 1874
List of Diurnal Lepidoptera of the South-Sea Islands Proc. Zool. Soc. Lond. 1874 : 274-291, pl. 44
Dimock, 1973
Three natural hybrids of Vanessa atalanta rubria x Cynthia annabella (Nymphalidae) J. Lep. Soc. 27 (4): 274-278
Dodge, 1900
Pyrameis huntera, n. var. fulvia Can. Ent. 32 (3) : 92
Doubleday, 1847
The genera of diurnal Lepidoptera, comprising their generic characters, a notice of their habitats and transformations, and a catalogue of the species of each genus; illustrated with 86 plates by W. C. Hewitson Gen. diurn. Lep. (1): 1-18, pl. A, 1-4 (1-2) (1846), (1): 19-132, pl. 4*, 5-25, 28 (3-14) (1847), (1): 133-200, pl. 1*, 26, 27, 29, 31-44 (15-23) (1848), (1): 201-242, pl. 30, 45-52, 56-58, 60-62, 64 (24-31) (1849), (1): 243-250 [?], (2): 243-326, pl. 53-55, 63, 65, 66 (32-38) (1850), (2): 327-466, pl. 59, 67-77 (39-50) (1851), (2): 467-534, 143, 144, pl. 78-80 (Index) (51-54) (1852)
Drury, [1773]
Illustrations of natural history; wherein are exhibited ... Illust. Nat. Hist. Exot. Insects 1 : 1-130, pl. 1-50 (1770), (1773)
Dunn, 1997
Biological notes on some eastern Australian butterflies. Part II Victorian Ent. 27 (6): 114-118
K. L. & L. E. Dunn, 1991
Review of Australian Butterflies: distribution, life history and taxonomy. Pushlished by authors, Melbourne Review Austr. Butts. (1-4):
Dyar, 1903
A List of North American Lepidoptera and Key to the Literature of this Order of Insects Bull. U.S. nat. Mus. 52 : 1-723
Eschscholtz, 1821
Beschreibung neuer ausländischer Schmetterling in Kotzebue, Entdeck. Reise Süd-See 3 (App. 5): 201-219, 11pls
Fabricius, 1775
Systema Entomologiae, sistens Insectorum Classes, Ordines, Genera, Species, Adiectis Synonymis, Locis, Descriptionibus, Observationibus. Syst. Ent.: 832pp
Fabricius, 1807
Systema Glossatorum. Die neueste Gattungs-Eintheilung der Schmetterlinge aus den Linnéischen Gattungen Papilio und Sphinx Magazin f. Insektenk. (Illiger) 6: 277-295
C. Felder, 1862
Verzeichniss der von den Naturforschern der k. k. Fregatte Novara gesammelten Macropepidoteren Verh. zool.-bot. Ges. Wien 12 (1/2) : 473-496
C. & R. Felder, [1867]
Reise der österreichischen Fregatte Novara um die Erde in den Jahren 1857, 1858, 1859 unter den Behilfen des Commodore B. von Wüllerstorf-Urbair. Zoologischer Theil. Band 2. Abtheilung 2. Lepidoptera. Rhopalocera Reise Fregatte Novara, Bd 2 (Abth. 2) (1): 1-136, pl. 1-21 (Rhop.) (C. & R. Felder, [1865]), (2): 137-378, pl. 22-47 (Rhop.) (C. & R. Felder, 1865), (3): 379-536, pl. 48-74 (Rhop.) (C. & R. Felder, 1867), (4): 537-548, pl. 75-120, (Inhalts-Verz.) 1-9 (pl. 1-74), (Felder & Rogenhofer, 1874) , (5): pl. 121-140, (erklärung) 1-10 (pl. 75-107), (Inhalts-Verz.) 1-20 (pl. 108-140), (Felder & Rogenhofer, 1875)
Fischer, 1908
Neue Tagfalterformen meiner Sammlung Soc. Ent. 23 (17) : 129-130
Fritsch, 1913
Natürliche Farbenspiele von Pyrameis Atalanta Ent. Rundschau 30 : 25-26
Fruhstorfer, 1898
Neue Rhopaloceren aus dem Malayischen Archipel Ent. Nachr. 24 (4) : 60-62 (15 February)
Fruhstorfer, 1898
Neue Rhopaloceren aus dem malayischen Archipel Dt. ent. Z. Iris 11 (1): 149-152 (10 August)
Fruhstorfer, 1909
Neue Vanessa- und Pyrameis-Rassen Int. ent. Zs. 3 (17): 94 (24 July)
Fruhstorfer, 1912
6. Familie: Nymphalidae in Seitz, Gross-Schmett. Erde 9 : 453-456 (31 January, 1912), : 457-488 (18 March, 1912), : 489-496 (25 May, 1912), : 489-496 (10 May, 1912), : 497-504 (15 April, 1912), : 505-512 (10 May, 1912), : 513-528 (11 April, 1912), : 529-552 (20 April, 1912), : 553-592 (15 May, 1912), : 593-616 (25 September, 1912), : 617-632 (15 October, 1912), : 633-640 (25 October, 1912), : 641-656 (10 January, 1913), : 657-664 (12 January, 1913), : 665-680 (13 January, 1913), : 681-688 (25 January, 1913), : 689-696 (16 June, 1913), : 697-704 (5 August, 1913), : 705-712 (20 January, 1914), : 713-720 (20 November, 1913), : 721-744 (9 March, 1914), : 745-751 (30 July, 1915), : 753-760 (28 March, 1914), : 761-766 (14 April, 1914)
Godart, 1819; Godart, [1824]
in Latreille & Godart, Encyclopédie Méthodique. Histoire naturelle Entomologie, ou histoire naturelle des crustacés, des arachnides et des insectes Encyclopédie Méthodique. 9 (1): 3-328 (1819), (2): 329-828 ([1824])
Godman & Salvin, [1882]
Biologia Centrali-Americana. Rhopalocera. Vol. 1. (1879-1886) Biol. centr.-amer., Lep. Rhop. 1: 1-487, 3: pl. 1-24,24a,25-112
Godman & Salvin, [1901]
Biologia Centrali-Americana. Rhopalocera. (1887-1901) Biol. centr.-amer., Lep. Rhop. 2: 1-782, 3: pl. 1-112
Gunder, 1925
Several New Aberrant Lepidoptera (Rhopalocera) Ent. News 36 (7) : 193-200, pl. 5
Hagen, 1895
Pyrameis Samani n. sp. Dt. ent. Z. Iris 7 (2) : 359-360
Hanafusa, 1992
Three new nymphalid butteflies from Indonesia and Philippines (Lep: Nymphalidae) Futao (10): 1-2, 7-8, 15-16
Howarth, 1966
Revisional notes on the genus Antanartia (Lepidoptera; Nymphalidae) Bull. Br. Mus. nat. Hist. (Ent.) 18 (2) : 21-43, pl. 1-5
Hübner, [1812]
Sammlung exotischer Schmetterlinge, Vol. 1 (1806 - [1819]) Samml. exot. Schmett. 1 : [1-4] (1807), : [5-12] (1807-1809), : [13-36] (1823-1824), : pl. [1-2], [5-8], [12], [15], [29], [47] (1806), : pl. [9], [11], [17], [21], [25-27], [32], [35], [43-44], [56], [60], [79], [102], [104], [116], [141], [148], [150-151], [155], [159], [200] (1807), : pl. [19], [39], [57], [73], [82], [114], [157], [167], [170], [173], [187] (1807), : pl. [3], [31], [37], [46], [54], [58], [62], [76], [84], [92], [95], [105], [112-113], [117], [137-138], [146], [154], [158], [180], [209] (1808), : pl. [16], [28], [65], [89], [142], [160], [162], [166], [182], [188], [193] (1809), : pl. [4], [22], [42], [53], [61], [88], [90], [97], [100], [107], [110], [118], [120], [140], [143], [149], [161], [171], [178], [184], [196], [202] (1809), : pl. [10], [18], [30], [41], [51], [80], [144], [172], [189] (1810), : pl. [36], [70], [87], [91], [101], [124], [129], [177], [183], [207], [210] (1811), : pl. [20], [38], [45], [55], [85], [98], [103], [108], [136], [139], [152-153], [156], [163], [179], [186], [190], [205-206] (1812), : pl. [13-14], [40], [48], [50], [67], [74], [86], [119], [121], [130], [135], [175-176], [197-199], [204], [212] (1813), : pl. [72], [75], [77-78], [83], [93], [109], [111], [115], [128], [131], [165], [174], [194] (1813), : pl. [24], [49], [52], [63-64], [66], [68-69], [71], [81], [123], [145], [181], [192] (1814), : pl. [126], [132-134], [168] (1815), : pl. [23], [96], [122], [125], [169], [195] (1816), : pl. [33], [99], [127], [187], [185], [213] (1817), : pl. [59], [106], [147], [164], [208], [211] (1818), : pl. [34], [94], [191], [201] (1819), : (Index Systematicus) [1-4] (1819 - April 1823)
Hübner, [1821]
Sammlung exotischer Schmetterlinge, Vol. 2 ([1819] - [1827]) Samml. exot. Schmett. 2 : pl. [90], [180], [223] (1819), : pl. [2], [12], [16], [78], [141], [182] (1820), : pl. [11], [13], [24], [77], [83], [97], [102-103], [108-109], [116-117], [122], [147], [150], [164], [183], [195], [197], [212-214] (1821), : pl. [14], [18-19], [26], [38-39], [43], [68], [70-71], [74], [84], [86], [92], [94-95], [106], [114], [121], [161], [169], [174], [177-178], [184], [194], [200], [204-205], [207-208], [210] (1821), : pl. [33], [72], [76], [115], [131], [138-139], [153], [198], [211], [218], [220] (1822), : pl. [32], [75], [81], [93], [105], [129] (1822), : pl. [1], [6], [22-23], [25], [27], [29-31], [34-37], [40-42], [48], [50-53], [56], [58-59], [64-66], [69], [73], [87], [89], [91], [96], [98-101], [110], [124], [128], [130], [132-137], [142], [155-158], [160], [162-163], [165-167], [170-173], [175-176], [187-188], [196], [199], [201-202], [206], [215-216], [219], [222], [224] (1823), : pl. [54], [118], [143], [149], [159] [179], [181], [185], [203], [217] (1824), : pl. [3-5], [8-10], [15], [20], [45], [49], [55], [60], [62-63], [67], [79-80], [88], [104], [111-113], [120], [125], [127], [140], [144-146], [148], [151-152], [154], [168], [189-193], [221], [225] (1825), : pl. [7], [17], [21], [28], [44], [46-47], [57], [61], [82], [123], [126] (1826), : pl. [85], [119], [186], [209] (1827), : (Index Systematicus) [1-4] (1827-1832)
Hübner, [1819]
Verzeichniss bekannter Schmettlinge, 1816-[1826] Verz. bek. Schmett. (1): [1-3], 4-16 (1816), (2): 17-32 (1819), (3): 33-48 (1819), (4): 49-64 (1819), (5): 65-80 (1819), (6): 81-96 (1819), (7): 97-112 (1819), (8): 113-128 (1819), (9): 129-144 (1819), (10): 145-160 (1819), (11): 161-176 (1819), (12): 177-192 (1820), (13): 193-208 (1820), (14): 209-224 (1821), (15): 225-240 (1821), (16): 241-256 (1821), (17): 257-272 (1823), (18): 273-288 (1823), (19): 289-304 (1823), (20): 305-320 (1825), (21): 321-336 (1825), (22): 337-352 (1825), (23-27): 353-431 ([1825])
Hübner, [1826]
Anzeiger der im Verzeichniss bekannter Schmettlinge angenommenen Benennungen ihrer Horden, Rotten, Stämme, Familien, Vereine und Gattungen Verz. bek. Schmett. (Anz.) (1-9): 1-72
Linnaeus, 1758
Systema Naturae per Regna Tria Naturae, Secundum Clases, Ordines, Genera, Species, cum Characteribus, Differentiis, Symonymis, Locis. Tomis I. 10th Edition Syst. Nat. (Edn 10) 1 : 1-338, : 339-824
Linsley & Usinger, 1966
Insects of Galápagos islands Proc. Cal. Acad. Sci. (4) 33: 113-196
Matsumura, 1908
Die Nymphaliden Japans Ent. Zs. 22 (39) : 157-158
McCoy, 1868
The Australian Representative of Cynthia cardui Ann. Mag. nat. Hist. 4 (1) : 76
Millière, 1868
Iconographie et description de chenilles et Lépidopteres inédits Ann. Soc. Linn. Lyon 16: 1-82, pl. 85-92
Moore, [1881]
The Lepidoptera of Ceylon Lepid. Ceylon 1 (1): 1-40, pl. 1-18 (Dec 1880), (2): 41-80, pl. 19-36 (Jan 1881), (3): 81-136, pl. 37-54 (Jun 1881), (4): 137-190, pl. 55-71 (Dec 1881)
Murayama & Shimonoya, 1963
On some interesting butterflies from Formosa, with descriptions of 2 new species, 2 new races, and 7 new aberrant forms [in Japanese] Tyô to Ga 13 (3): 51-59
Murayama & Shimonoya, 1966
Some Formosan butterflies of new species, new race, new aberrant forms or little known species and forms. [in Japanese] Tyô to Ga 15 (3/4) : 58-67
Roque-Albelo, 2004
The butterflies (Papilionoidea, Hesperoidea) of the Galápagos Islands, Ecuador: distribution, hostplants and biology J. Lep. Soc. 58 (1): 33-43
Rosenberg & Talbot, 1914
New South American butterflies Trans. ent. Soc. Lond. 1913 (4) : 671-682
Scudder, 1889
The Butterflies of the Eastern United States and Canada, with special reference to New England Butts eastern U.S. Canada 1: 1-766, 2: 767-1774, 3: 1775-1958, pl. 1-89
Sodoffsky, 1837
Entomologische untersuchungen ueber die Gattungsnamen der Schmetterlinge Bull. Soc. imp. Nat. Moscou 1837 (6) : 76-97
Staudinger, 1894
Hochlandine Lepidoptera Dt. Ent. Z. Iris 7 (1) : 43-100, pl. 1-2
Talbot, 1936
A new species of Danaidae (Lep. Rhop.) and other new butterflies Entomologist 69: 112-114
Vane-Wright & Hughes, 2007
Did a member of the Vanessa indica complex (Nymphalidae) formerly occur in North America J. Lep. Soc. 61 (4): 199-212
Vane-Wright & de Jong, 2003
The butterflies of Sulawesi: annotated checklist for a critical island faunda Zool. Verh. Leiden 343: 3-268, pl. 1-16
Wahlberg, Brower & Nylin, 2005
Phylogenetic relationships and historical biogeography of tribes and genera in the subfamily Nymphalinae (Lepidoptera: Nymphalidae) Biol. J. Linn. Soc. 86: 227-251
Williams, 1911
Expedition of the California Academy of Sciences to the Galápagos Islands, 1905-1906. II. The butterflies and hawk-moths of the Galápagos Islands Proc. Calif. Acad. Sci. 1: 289-322


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