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Euphydryas Scudder, 1872
Euphydryas Scudder, 1872; 4th Ann. Rep. Peabody Acad. Sci. (1871): 48, TS: Papilio phaeton Drury
Lemonias Hübner, [1806]; Tentamen determinationis digestionis ...: [1] (suppr.), TS: Papilio maturna Linnaeus
= ; [NL4A, 252]
Occidryas Higgins, 1978; , TS: anicia Doubleday
= ; [NL4A, 252]
Hypodryas Higgins, 1978; , TS: Papilio maturna Linnaeus
= ; [NL4A, 252]
Eurodryas Higgins, 1978; , TS: Papilio aurinia Rottenburg
= ; [NL4A, 252]
Euphydryas, NSG Voucher Specimen [Wahlberg]
Euphydryas (Eyphydryini) ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3) : 165
Hypodryas (Euphydryini) ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3) : 165
Occidryas (Euphydryini) ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3) : 165
Eurodryas (Euphydryini) ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3) : 165
Euphydryas (Euphydryina) ; [Wahlberg]; [NL4A, 252]


[sp-1.jpg] 700x579 (~85Kb) USA: Bridge Creek Trailhead, North Cascades Highway, (48°30'30N 120°43'00W), Chelan Co., WA, 28.7.1999, Photo © Markku Savela [sp-2.jpg] 1000x720 (~149Kb) USA: Near Crystal Mtn, (46°57'51N 121°28'55W), Pierce Co., WA, 27.7.1999, Photo © Markku Savela [sp-3x.jpg] 600x639 (~46Kb) USA: Wenatchee National Forest, (47°04'12N 120°56'10W), Kittitas Co., WA, 29.7.1999, Photo © Markku Savela
[sp-3y.jpg] 900x663 (~71Kb) USA: Wenatchee National Forest, (47°04'12N 120°56'10W), Kittitas Co., WA, 29.7.1999, Photo © Markku Savela [sp-3z.jpg] 900x588 (~60Kb) USA: Wenatchee National Forest, (47°04'12N 120°56'10W), Kittitas Co., WA, 29.7.1999, Photo © Markku Savela [sp-4amf.jpg] 700x919 (~101Kb) male female USA: Corral Pass region (47°00'15N 121°29'02W), Pierce Co., WA, 18.7.2000, Photo © Markku Savela
[sp-4bmf.jpg] 660x784 (~82Kb) male female USA: Corral Pass region (47°00'15N 121°29'02W), Pierce Co., WA, 18.7.2000, Photo © Markku Savela

The exact identification of these species is still unknown, but tentatively assumed to belong into this group.


The "Euphydryas" species-group
Euphydrini euphydryas ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3) : 165

Euphydryas phaeton (Drury, [1773])
ENA. See [About maps] Baltimore
Papilio phaeton Drury, [1773]; Illust. Nat. Hist. Exot. Insects 1: index, : 42, pl. 21, f. 3-4, TL: New York
phaedon (Herrich-Schäffer, 1865); (misspel.)
Euphydryas phaeton phaeton, Butterflies of America [BOA]
Euphydryas phaeton, NSG Voucher Specimen [Wahlberg]
Euphydryas phaeton, Butterflies and Moths of North America [BAMONA]
Euphydryas phaeton, Georgia Lepidoptera (USA) [GALEPS]
Euphydryas phaeton ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 17; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3) : 165 (name); [NACL, #4516]; [EBW]; [BOW: pl. 13, f. 28]; [Opler]; [NL4A, #2064]
[phaeton-1.jpg] 432x289 (~30Kb) USA: Ann Arbor, Michigan, May 1999, Photo © Christopher A. Rickards [phaeton-2.jpg] 1036x991 (~183Kb) USA: Alabama, Jeff. Co., 13.5.2000, Photo © Vitaly Charny [phaeton-2l.jpg] 1157x771 (~131Kb) larva USA: Alabama, 22.4.2001, Photo © Vitaly Charny
[phaeton-3la.jpg] 1091x867 (~104Kb) larva USA: Alabama, 5.5.2001, Photo © Vitaly Charny [phaeton-3lb.jpg] 800x1238 (~239Kb) larva USA: Alabama, 5.5.2001, Photo © Vitaly Charny [phaeton-3u.jpg] 1300x927 (~250Kb) upperside USA: Alabama, 5.5.2001, Photo © Vitaly Charny
[phaeton-3v.jpg] 967x1339 (~220Kb) underside USA: Alabama, 5.5.2001, Photo © Vitaly Charny [phaeton-3x.jpg] 828x633 (~95Kb) USA: Alabama, 13.5.2001, Photo © Vitaly Charny [phaeton-4u.jpg] 1049x766 (~209Kb) upperside USA: Alabama, 6.5.2006, Photo © Vitaly Charny

Larva on Chelone glabra, Gerardia grandiflora, G. pedicularia, Plantago lanceolata, Fraxinus americana [NAB]


New York, Maryland, Maine, ... Melitaea phaetaena Hübner, [1819]
Melitaea phaëtaena Hübner, [1819]; (repl. name), TL: New York
= ; [NL4A, #2064a]
Argynnis phaetontea Godart, 1819; (repl. name), TL: New York
= ; [NL4A, #2064a]
ab. superba (Strecker, 1878) (Melitaea); , TL: New York
= ; [NL4A, #2064a]
Melitaea phaethusa Hulst, 1881; , TL: New York
= ; [NL4A, #2064a]
Melitaea streckeri Ellsworth, 1902; Ent. News. 13 : 104, TL: New York
= ; [NL4A, #2064a]
Euphydryas phaëton schausi Clark, 1927; Proc. U.S. Nat. Mus. 71 (2683): (1-22) , TL: Maryland
= ; [NL4A, #2064a]
Euphydryas phaëton schausi var. magnifica Clark, 1927; Proc. U.S. Nat. Mus. 71 (2683): (1-22) , TL: Maryland
= ; [NL4A, #2064a]
Euphydryas phaeton borealis F. & R. Chermock, 1940; , TL: Maine
= ; [NL4A, #2064a]
Euphydryas phaeton phaeton ; [NL4A, #2064a]
E. p. ozarkae Masters, 1968
Euphydryas phaeton ozarkae, Butterflies of America [BOA]
Euphydryas phaeton ozarkae ; [NACL, #4516a]

The "Hypodryas" species-group
Euphydrini hypodryas ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3) : 165

fiEuphydryas maturna (Linnaeus, 1758)
NW.China, Mongolia. See [About maps] Kirjoverkkoperhonen Scarce Fritillary Broknätfjäril Eschenscheckenfalter Kleiner Maivogel Przeplatka maturna
Papilio maturna Linnaeus, 1758; Syst. Nat. (Edn 10) 1 : 480, TL: Sweden
Papilio agrotera Bergtrasser, 1780
Euphydryas maturna, NSG Voucher Specimen [Wahlberg]
Euphydryas maturna, Larvae of North-European Lepidoptera [Kimmo Silvonen]
Euphydryas maturna, Svenska fjärilar [SVF]
Euphydryas maturna, Moths and Butterflies of Europe [MBE]
Euphydryas maturna, Europaische Schmetterlinge [Mario Maier]
Hypodryas maturna, Captain's European Butterfly Guide [Simon Coombes]
Hypodryas maturna ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3) : 165 (name)
Euphydryas maturna ; [BOW: pl. 2, f. 11]; [EBW]; [BARU, #152]; [BRU2: 57, pl. 37, f. 16-18]; [Wahlberg]; [Otakar Kudrna]
[maturna-1mu.jpg] 800x612 (~60Kb) upperside male FINLAND: Kb: Kiihtelysvaara, 692:68, 1.7.1998 Photo © Markku Savela [maturna-2v.jpg] 1000x765 (~67Kb) underside FINLAND: Sa: Ristiina, 6817:528, 2.7.1998, Photo © Markku Savela [maturna-3l.jpg] 594x400 (~49Kb) larva Central Europe 1990-2000, Photo © Michael Borsch
[maturna-3u.jpg] 595x400 (~62Kb) upperside Central Europe 1990-2000, Photo © Michael Borsch

Larva on Betula spp., Salix spp., Fraxinus excelsior, Veronica longifolia, Melampyrum sylvaticum, Viburnum opulus, Lonicera xylosteum, Lonicera coerulea, L. caprifolium [SPRK], Scabiosa, Veronica, Salvia, Plantago, Viola, Digitalis [BRU2, 58]


NEU, CEU Euphydryas maturna maturna ; [BRU2, 57]
E. m. staudingeri (Wnukowsky, 1929)
Altai, Siberia, Transbaikalia, Dzhungarski Alatau, Mongolia?, NW.China? uralensis (Staudinger, 1871) (Melitaea); in Staudinger & Wocke, Cat. Lepid. eur. Faunegeb. (Edn. 2) : 17 (preocc.)
Euphydryas idunoides Fruhstorfer, 1916
Melitaea maturna var. uralensis ; Staudinger, 1881, Stettin ent. Ztg 42 (7-9): 287
Euphydryas maturna staudingeri ; [BARU, (note)]; [BRU2, 57]
[staudingeri-1fv.jpg] 392x393 (~14Kb) underside female A hillock swamped meadow at a riparian spruce wood at the vllage Topuchaya, Shebalino District, Altai Republic, the Altai Mts., S Siberia, Russia. 8th July 1998 (on Pentaphylloides fruticosa), Photo © Oleg Kosterin [staudingeri-1mu.jpg] 480x408 (~41Kb) upperside male A forb meadowy edge of a birch forest, the lower part of the Kora River valley, the Mynchukur mountain range, the northern chain of the Dzhungarian Alatau Mts., Taldy-Kurgan Province, SE Kazakhstan. 13th June 1994, Photo © Oleg Kosterin

E. m. coreanica Murayama, 1955
Korea Euphidryas[sic] maturna coreanica Murayama, 1955; Tyô to Ga 6 (1) : 1, f. 4, TL: Kosho, Heian-kokude, N-Korea

Euphydryas ichnea (Boisduval, [1833])
-. See [About maps] Melitaea ichnea Boisduval, [1833]; Icon. hist. Lépid. Europe 1 (11-12): 112, pl. 23, f. 5-6, TL: "le nord de la Laponiae et de la Sibirie"
Euphydryas ichnea ; [BARU, #153]; [BRU2: 58, pl. 37, f. 19-21]
S.Siberia, Transbaikalia, Far East, Amur, Ussuri Asian Fritillary
Melitaea intermedia Ménétriés, 1859; in Schrenck, Reise Forschungen Amur-Lande 2 (1) : 22, TL: Amurland
Euphydryas intermedia, NSG Voucher Specimen [Wahlberg]
Euphydryas intermedia, Moths and Butterflies of Europe [MBE]
Hypodryas intermedia, Captain's European Butterfly Guide [Simon Coombes]
Euphydryas ichnea ichnea ; [BRU2, 58]
Hypodryas intermedia ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3) : 165 (name)
Euphydryas intermedia ; [BOW: pl. 2, f. 11 (text only)]; [MRS, 585]; [Wahlberg]; [Otakar Kudrna]

Larva on Lonicera maacki [BRU2, 58]


[intermedia-1mu.jpg] 547x400 (~46Kb) upperside male Dwarf pine/dwarf alder forest-tundra at about 1000 m above sea level, the mountains of the Koni Peninsula, Ola District, S Magadan Province, Russia. 23rd July 1989 (on Ledum palustre), Photo © Oleg Kosterin

E. i. mongolica (Staudinger, 1892)
Sayan mongolica (Staudinger, 1892); , TL: ["Kentei-gebirge"]
Euphydryas intermedia mongolica ; [MRS, 585]
Euphydryas ichnea mongolica ; [BRU2, 58]
E. i. altaiana (Wnukowsky, 1929)
Altai altaica (Seitz, [1909]); (preocc.)
Euphydryas intermedia altaica ;
Euphydryas ichnea altaiana ; [BRU2, 58]
Euphydrays ichnea altaica ; [BARU, (note)]

[altaica-1fu.jpg] 906x600 (~84Kb) upperside female Dwarf birch (Betula rotundifolia) thickets on the left bank of the Dzhazator River in its upper flow, between the falls of its right tributaries Chikty and Akbul, 2000 m above sea level, Kosh-Agach District, Altai Republic, West Siberia, Russia. 19th July 1998, Photo © Oleg Kosterin

E. i. konumensis (Matsumura, 1927)
Sakhalin Melitaea maturna konumensis Matsumura, 1927; Ins. Matsumurana 2 (2) : 117, pl. 3, f. 5, TL: Saghalien
Euphydryas ichnea konumensis ; [BRU2, 58]
Alps (Maritime Alps) Euphydryas intermedia wolfensbergeri ;
Euphydryas ichnea wolfensbergeri ; [BRU2, 58]

Euphydryas cynthia ([Schiffermüller], 1775)
Alps, Bulgaria. See [About maps] Cynthia's Fritillary Veilchen-Scheckenfalter
Papilio cynthia [Schiffermüller], 1775; Ankündung eines systematischen Werkes von den Schmetterlingen der Wienergegend: 179
Papilio mysia Hübner, 1800
Euphydryas cynthia, NSG Voucher Specimen [Wahlberg]
Euphydryas cynthia, Moths and Butterflies of Europe [MBE]
Hypodryas cynthia, Captain's European Butterfly Guide [Simon Coombes]
Hypodryas cynthia ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3) : 165 (name)
Euphydryas cynthia ; [BOW: pl. 2, f. 8]; [EBW]; [Wahlberg]; [Otakar Kudrna]

Larva on Plantago, Plantago alpina, Alchemilla [H&R]


fiEuphydryas iduna (Dalman, 1816)
. See [About maps] Lapinverkkoperhonen Lapland Fritillary Lappnätfjäril
Melitaea iduna Dalman, 1816; K. VetenskAcad. Handl. 1816 (1): 75, TL: Sweden
Euphydryas iduna, Lepidoptera collection [SZMN]
Euphydryas iduna, NSG Voucher Specimen [Wahlberg]
Euphydryas iduna, Larvae of North-European Lepidoptera [Kimmo Silvonen]
Euphydryas iduna, Svenska fjärilar [SVF]
Euphydryas iduna, Norges Sommerfugler [NORLEP]
Euphydryas iduna, Moths and Butterflies of Europe [MBE]
Euphydryas iduna, Europaische Schmetterlinge [Mario Maier]
Hypodryas iduna ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3) : 165 (name)
Euphydryas iduna ; [BOW: pl. 2, f. 10]; [EBW]; [BARU, #154]; [MRS, 585]; [BRU2: 58, pl. 37, f. 22-30]; [Wahlberg]; [Otakar Kudrna]
[iduna-1p.jpg] 400x520 (~50Kb) pupa Finland, Kilpisjärvi 767:25, 06.07.1982, Photo © Pertti Pakkanen [iduna-1u.jpg] 860x520 (~80Kb) upperside (info missing), Photo © Oleg Kosterin

Larva on Bartsia alpina?, Vaccinium uliginosum?, V. myrtillus? [SPRK], Plantago, Veronica, Vaccinium [BRU2, 59]


Arctic Europe, Arctic Asia Euphydryas iduna iduna ; [BRU2, 58]
E. i. inexpectata (Sheljuzhko, 1934)
Caucasus Major Euphydryas iduna inexpectata ; [BRU2, 58]
E. i. alferakyi Korshunov, 1996
Far East Euphydryas iduna alferakyi ; [Oleg Kosterin]; [BRU2, 58]
[alferakyi-1v.jpg] 533x400 (~58Kb) underside A subalpine meadow at the dwarf pine (Pinus pumila (Pall.) Regel) thickets on the southen slope of a pass between Khindzha and Burgauli valleys, in a valley of a left tributary of the latter, about 500 m above sea level, the mountains of the Koni Peninsula, Ola District, S Magadan Province, Russia. 12th July, 1989, Photo © Oleg Kosterin

E. i. semenovi Korshunov & Ivonin, 1996
Kuznetsky Alatau Mts. Euphydryas iduna semenovi, Lepidoptera collection [SZMN]
Euphydryas iduna semenovi ; [BRU2, 58]
E. i. sajana Higgins, 1950
Altai, Sayan sajana Gaede, 1932; (nom. nud.)
Euphydryas iduna sajana ; [BRU2, 58]
[sajana-1v.jpg] 457x500 (~42Kb) underside Highland dwarf birch (Betula rotundifolia) tundras on a ledge of the southerm principle slope of the Yuzhno-Chuiskii mountain range between the Chikty and Akbul Rivulets, 2300 m above sea level, the upper Dzhazator River basin, Kosh-Agach District, Altai Republic, West Siberia, Russia. 15th July 1998, Photo © Oleg Kosterin


Euphydryas gillettii (Barnes, 1897)
Wyoming. See [About maps] Gillette's Checkerspot
Melitaea gillettii Barnes, 1897; Can. Ent. 29 (2) : 40, TL: Yellowstone Park, Wyoming
Melitaea glacialis Skinner, 1921; Ent. News 32 (3) : 89, TL: Montana
herri Gunder, 1929; Pan-Pacific Entomol. 6: 8
Euphydryas gillettii, E.H. Strickland Ent. Mus, U of [ALBERTA]
Euphydryas gillettii, Butterflies of America [BOA]
Euphydryas gillettii, NSG Voucher Specimen [Wahlberg]
Euphydryas gillettii, Butterflies and Moths of North America [BAMONA]
gracialis[sic, recte glacialis] ; McDunnough, 1938
Lemonias gillettei[sic, recte gillettii] ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 17
Hypodryas gillettii ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3) : 165 (name); [NACL, #4521]
Euphydryas gillettii ; [Wahlberg]; [Opler]

Larva on Lonicera involucrata, Symphoricarpos albus [NAB]


The "Occidryas" species-group
Euphydrini occidryas ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3) : 165

Euphydryas anicia (Doubleday, [1847])
British Columbia. See [About maps] Anicia Checkerspot
Melitaea anicia Doubleday, [1847]; Gen. diurn. Lep. (1): 179
mayi Gunder, 1932²
Euphydryas anicia, E.H. Strickland Ent. Mus, U of [ALBERTA]
Euphydryas anicia, NSG Voucher Specimen [Wahlberg]
Lemonias anicia ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 17
Occidryas anicia ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3) : 165 (name); [NACL, #4519]
Euphydryas anicia ; [BOW: pl. 13, f. 23]; [Opler]; [NL4A, #2061]
Euphydryas chalcedona anicia ;
[anicia-1U.jpg] identification uncertain 1300x1013 (~240Kb) USA: Cabin Mountain, (1408m), (47°14'15N 121°16'27W), Kittitas Co., WA, 29.7.2006, Photo © Markku Savela [anicia-2U.jpg] identification uncertain 1200x1685 (~234Kb) USA: Cabin Mountain, (1408m), (47°14'15N 121°16'27W), Kittitas Co., WA, 29.7.2006, Photo © Markku Savela

Larva on Castilleja, Penstemon, Scrophulariaceae, Plantaginaceae, Boraginaceae [NAB]


E. a. alena Barnes & Benjamin, 1926
Euphydryas anicia alena, Butterflies of America [BOA]
Occidryas anicia alena ; [NACL, #4519f]
E. a. bakeri Stallings & Turner, 1945
Occidryas anicia bakeri ; [NACL, #4519o]
E. a. bernadetta Leussler, 1920
leussleri Gunder, 1927²
belli Gunder, 1929²
Occidryas anicia bernadetta ; [NACL, #4519m]
E. a. capella Barnes, 1897
rubrolimbata Comstock, 1918²
Euphydryas anicia var. capella ab. ♂ oslari Gunder, 1925; Ent. News 36 (7) : 196, pl. 5, f. 7, TL: Platte Canyon, Colorado
Occidryas anicia capella ; [NACL, #4519c]
E. a. carmentis Barnes & Benjamin, 1926
charlottae Gunder, 1928²
Occidryas anicia carmentis ; [NACL, #4519g]
E. a. chuskae Ferris & R. Holland, 1980
E. a. cloudcrofti Ferris & R. Holland, 1980
E. a. effi Stallings & Turner, 1945
Occidryas anicia effi ; [NACL, #4519k]
E. a. eurytion Mead, 1875
Melitaea brucei Edwards, 1888; Can. Ent. 20 (5) : 81
melanodisca Comstock, 1918²
Lemonias anicia brucei ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 17
Occidryas anicia eurytion ; [NACL, #4519i]
E. a. helvia Scudder, 1869
Alaska anicia helvia Scudder, 1869; Proc. Boston Soc. nat. Hist. 12 : 405
Lemonias helvia ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 18
Occidryas anicia helvia ; [NACL, #4519a]
E. a. hermosa (Wright, 1905)
Arizona anicia hermosa (Wright, 1905) (Melitaea); , TL: Arizona
venusta Gunder, 1932²
Euphydryas anicia hermosa, Listing of Mexican Butterflies [ILMB]
Euphydryas anicia hermosa, Butterflies of America [BOA]
Euphydryas anicia hermosa, Butterflies of Southeastern Arizon [Bruce Walsh]
Occidryas anicia hermosa ; [NACL, #4519d]
Euphydryas anicia hermosa ; [NL4A, #2061a]
E. a. hopfingeri Gunder, 1934
andersoni Gunder, 1934²
Occidryas anicia hopfingeri ; [NACL, #4519k]
E. a. howlandi Stallings & Turner, 1947
Occidryas anicia howlandi ; [NACL, #4519b]
E. a. irelandi Gunder, 1929
E. a. macyi Fender & Jewett, 1953
Occidryas anicia macyi ; [NACL, #4519p]
E. a. magdalena Barnes & McDunnough, 1918
Euphydryas anicia magdalena, Butterflies of America [BOA]
Occidryas anicia magdalena ; [NACL, #4519e]
E. a. maria (Skinner, 1899)
Utah Melitaea maria Skinner, 1899; Can. Ent. 31 (1) : 12
skinneri Gunder, 1928²
spaldingi Gunder, 1928²
Lemonias maria ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 17
Occidryas anicia maria ; [NACL, #4519j]
E. a. morandi Gunder, 1928
Occidryas anicia morandi ; [NACL, #4519r]
E. a. veazieae Fender & Jewett, 1953²
anicia veazieae Fender & Jewett, 1953²
Occidryas anicia veazieae ; [NACL, #4519n]
E. a. wheeleri (H. Edwards, 1881)
Melitaea anicia var. wheeleri H. Edwards, 1881; Papilio 1 (4) : 52
duncani Gunder, 1934²
Lemonias nubigena wheeleri ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 17
Occidryas anicia wheeleri ; [NACL, #4519p]
E. a. windi Gunder, 1932
Occidryas anicia windi ; [NACL, #4519h]

Euphydryas chalcedona (Doubleday, [1847])
W.USA. See [About maps] Chalcedon Checkerspot
Melitaea chalcedona Doubleday, [1847]; Gen. diurn. Lep. (1): 180, TL: "Haiti"
Euphydryas chalcedona chalcedona, Listing of Mexican Butterflies [ILMB]
Euphydryas chalcedona chalcedona, Butterflies of America [BOA]
Euphydryas chalcedona, NSG Voucher Specimen [Wahlberg]
Euphydryas chalcedona, Butterflies and Moths of North America [BAMONA]
Lemonias chalcedon[sic] ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 17
Occidryas chalcedona ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3) : 165 (name); [NACL, #4517]
Euphydryas chalcedon[sic] ; [BOW: pl. 13, f. 25 (missp.)]
Euphydryas chalcedona ; [EBW]; [Wahlberg]; [Opler]; [NL4A, #2062]

Larva on Plantago, Lonicera, Symphoricarpos ablus [NAB]

California Melitaea cooperi Behr, 1863; Proc. Calif. Acad. Sci. 3 (1) : 90, TL: California
= ; [NL4A, #2062a]
Melitaea chalcedon[sic] ab. fusimacula Barnes, 1900; Can. Ent. 32 (2) : 42, TL: California
= ; [NL4A, #2062a]
Melitaea chalcedon[sic] ab. mariana Barnes, 1900; Can. Ent. 32 (2) : 42, TL: California
= ; [NL4A, #2062a]
Lemonias chalcedon[sic] ab. grundeli Coolidge, 1908; , TL: California
= ; [NL4A, #2062a]
Melitaea chalcedon[sic] ab. lorquini Oberthür, 1914; , TL: California
= ; [NL4A, #2062a]
Euphydryas chalcedona ab. omniluteofuscus Gunder, 1925; Ent. News 36 (7) : 195, pl. 5, f. 4, TL: California
= ; [NL4A, #2062a]
Euphydryas chalcedona ab. ♂ hemiluteofuscus Gunder, 1925; Ent. News 36 (7) : 196, pl. 5, f. 5, TL: California
= ; [NL4A, #2062a]
ab. supranigrella Comstock, 1926; , TL: California
= ; [NL4A, #2062a]
ab. suprafusa Comstock, 1926; , TL: California
= ; [NL4A, #2062a]
ab. hemimelanica Comstock, 1926; , TL: California
= ; [NL4A, #2062a]
ab. fusisecunda Comstock, 1926; , TL: California
= ; [NL4A, #2062a]
Lemonias cooperi ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 17
Lemonias chalcedon[sic] fusimacula ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 17
Lemonias chalcedon[sic] mariana ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 17
Euphydryas chalcedona chalcedona ; [NL4A, #2062a]
E. c. corralensis Emmel & Emmel, 1973
Occidryas chalcedona corralensis ; [NACL, #4517d]
E. c. dwinnellei H. Edwards, 1881
Melitaea chalcedon[sic] dwinnellei H. Edwards, 1881; Papilio 1 (4) : 51
sperryi F. & R. Chermock, 1945²
Lemonias chalcedon[sic] dwinellei[sic] ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 17
Occidryas chalcedona dwinnellei ; [NACL, #4517a]
E. c. hennei Scott, 1981
California Euphydryas chalcedona hennei Scott, 1981; , TL: California
Euphydryas chalcedona quino tr.f. hennei Gunder, 1932; , TL: California
= ; [NL4A, #2062b]
Euphydryas chalcedona hennei, Listing of Mexican Butterflies [ILMB]
Euphydryas chalcedona hennei, Butterflies of America [BOA]
Euphydryas chalcedona hennei ; [NL4A, #2062b]
E. c. huellemanni Comstock, 1926
E. c. kingstonensis Emmel & Emmel, 1973
Euphydryas chalcedona kingstonensis, Butterflies of America [BOA]
Occidryas chalcedona kingstonensis ; [NACL, #4517e]
E. c. klotsi dos Passos, 1938
Arizona Euphydryas chalcedona klotsi dos Passos, 1938; Can. Ent. 70: (199-200), TL: Roosevelt Lake, Arizona
Euphydryas chalcedona klotsi, Butterflies of Southeastern Arizon [Bruce Walsh]
Occidryas chalcedona klotsi ; [NACL, #4517c]
E. c. macglashanii (Rivers, 1888)
California Melitaea macglashanii Rivers, 1888; Proc. Cal. Acad. Sci. (2) 1 : 103
truckeensis Gunder, 1928²
georgei Gunder, 1928²
hilli Gunder, 1928²
Occidryas mcglashanii ; McDunnough, 1938, (missp.)
Limonias chalcedon macglashani ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 17
Occidryas chalcedona macglashanii ; [NACL, #4517f]
Euphydryas macglashanii ; [BOW: pl. 13, f. 26]
E. c. olancha Wright, 1905
malcolmi Gunder, 1927²
Occidryas chalcedona olancha ; [NACL, #4517g]
E. c. paradoxa McDunnough, 1927
Occidryas colon paradoxa; [NACL, #4518b]
E. c. perdiccas Edwards, 1880
nigrisupernipennis Gunder, 1926²
svihlae Gunder, 1932²
perdiceas ; Edwards, 1880, (missp.)
Occidryas colon perdiccas ; [NACL, #4518a]
E. c. sierra Wright, 1905
Euphydryas sierra ab. ♂ magdelenae Gunder, 1925; Ent. News 36 (7) : 197, pl. 5, f. 8, TL: Glen Alpine Creek, El Dorao Co., California
umbrobasana Comstock, 1925²
irelandi Gunder, 1929²
Occidryas chalcedona sierra ; [NACL, #4517h]
E. c. sperryi Chermock & Chermock, 1945
E. c. colon (Edwards, 1881)
Washington, Oregon Colon Checkerspot
Melitaea colon Edwards, 1881; Papilio 1 (4) : 45
mcdunnoughi Gunder, 1928²
fenderi Gunder, 1932²
bakeri Fender, 1945²; (preocc Stallings & Turner, 1945)
macdunnoughi dos Passos, 1964²; (emend.)
Lemonias colon ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 17
Occidryas colon ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3) : 165 (name); [NACL, #4518]
?Euphydryas chalcedona colon ; [Opler]

Larva on Symphoricarpos albus, Symphoricarpos vaccinoides, Penstemon antirrhinoides, P. subserratus, Plantago major [NAB]

E. c. wallacensis Gunder, 1928
Euphydryas colon wallacensis Gunder, 1928
huellemanni Comstock, 1926²; (ab.)
idahoensis Gunder, 1929²
Occidryas colon wallacensis ; [NACL, #4518c]
Euphydryas chalcedona wallacensis ; [Wahlberg]
E. c. nevadensis Bauer, 1975
Euphydryas colon nevadensis Bauer, 1975
Occidryas colon nevadensis ; [NACL, #4518d]
Euphydryas chalcedona nevadensis ; [Wahlberg]

Euphydryas editha (Boisduval, 1852)
-. See [About maps] Edith's Checkerspot
Melitaea editha Boisduval, 1852; Ann. Soc. ent. Fr. (2) 10 : 304
cottlei Gunder, 1928²
Euphydryas editha, E.H. Strickland Ent. Mus, U of [ALBERTA]
Euphydryas editha, NSG Voucher Specimen [Wahlberg]
Euphydryas editha, Butterflies and Moths of North America [BAMONA]
Lemonias editha ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 18
Occidryas editha ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3) : 165 (name); [NACL, #4520]
Euphydryas editha ; [Opler]; [NL4A, #2063]
[editha-1aU.jpg] identification uncertain 1000x1290 (~179Kb) USA: Cabin Mountain, (1408m), (47°14'15N 121°16'27W), Kittitas Co., WA, 29.7.2006, Photo © Markku Savela [editha-1bU.jpg] identification uncertain 1000x1214 (~157Kb) USA: Cabin Mountain, (1408m), (47°14'15N 121°16'27W), Kittitas Co., WA, 29.7.2006, Photo © Markku Savela

Larva on Schrophulariaceae, Castilleja, Orthocarpus, Pedicularis, Plantago [NAB]


E. e. quino (Behr, 1863)
California editha quino (Behr, 1863) (Melitaea); Proc. Calif. Acad. Sci. 3 (1) : 90, TL: California
Melitaea augusta Edwards, 1890; Can. Ent. 22 (2) : 21, TL: California
= ; [NL4A, #2063a]
Euphydryas editha ab. fieldi Gunder, 1924; , TL: California
= ; [NL4A, #2063a]
Euphydryas editha wrighti Gunder, 1929; , TL: California
= ; [NL4A, #2063a]
Euphydryas editha wrighti tr.f. thornei Gunder, 1934; , TL: California
= ; [NL4A, #2063a]
Euphydryas quino tr.f. rubyae Hower, 1935; , TL: California
= ; [NL4A, #2063a]
?kennei Gunder, 1932²
?augustina Wright, 1905²
Euphydryas editha quino, Listing of Mexican Butterflies [ILMB]
Euphydryas editha quino, Butterflies of America [BOA]

Lemonias augusta ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 18
Occidryas editha augusta ; [NACL, #4520n]
Euphydryas editha augusta ; [BOW: pl. 13, f. 24]
Lemonias quino ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 18
Occidryas chalcedona quino ; [NACL, #4517b]
Euphydryas chalcedona quino ;
Euphydryas editha quino ; [Wahlberg]; [NL4A, #2063a]
E. e. alebarki Ferris, 1971
Occidryas editha alebarki ; [NACL, #4520g]
E. e. aurilacus Gunder, 1928
ab. foxi Gunder, 1924²
ab. albiradiata Gunder, 1926²
Occidryas editha aurilacus ; [NACL, #4520p]
E. e. baroni Edwards, 1879
editha baroni Edwards, 1879; , TL: 'Mendocino'
Melitaea baroni H. Edwards, 1881; Papilio 1 (4) : 52
mirabilis Wright, 1905²
dunni Gunder, 1929²
sternizkyi Gunder, 1929²
Lemonias baroni ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 18
Occidryas editha baroni ; [NACL, #4520j]
E. e. bayensis Sternitzky, 1937
Occidryas editha bayensis ; [NACL, #4520c]
E. e. beani Skinner, 1897
editha beani Skinner, 1897; Can. Ent. 29 (6) : 155
blackmorei Gunder, 1926²
Lemonias anicia beani ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 17
Occidryas editha beani ; [NACL, #4520c]
E. e. colonia (Wright, 1905)
Occidryas editha colonia ; [NACL, #4520o]
Euphydryas colonia ; [BOW: pl. 2, f. 11 (text only)]
E. e. edithana Strand, 1914
Occidryas editha edithana ; [NACL, #4520i]
E. e. fridayi Gunder, 1931
Occidryas editha gunnisonensis ; [NACL, #4520h]
E. e. hutchinsi McDunnough, 1928
montanus McDunnough, 1928²
Occidryas editha hutchinsi ; [NACL, #4520f]
E. e. insularis Emmel & Emmel, 1974
Occidryas editha insularis ; [NACL, #4520b]
E. e. lawrencei Gunder, 1931
diamondensis Gunder, 1931²
Euphydryas editha lawrencei, Butterflies of America [BOA]
Occidryas editha lawrencei ; [NACL, #4520r]
E. e. lehmani Gunder, 1929
Occidryas editha lehmani ; [NACL, #4520m]
E. e. luestherae Murphy & Ehrlich, 1981
editha luestherae Murphy & Ehrlich, 1981; , TL: Stanislaus Co., Del Puerto Canyon
E. e. monoensis Gunder, 1928
fridayi Gunder, 1931²
Occidryas editha monoensis ; [NACL, #4520l]
E. e. nubigena Behr, 1863
California editha nubigena Behr, 1863; Proc. Calif. Acad. Sci. 3 (1) : 91
rubrosuffusa Comstock, 1926²
tiogaensis Gunder, 1929²
boharti Gunder, 1929²
nubigeria[sic, recte nubigena] ; Edwards, 1873
nubigena ; Mead, 1875
Occidryas tiogoensis[sic, recte tiogaensis] ; McDunnough, 1938
Lemonias nubigena ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 17
Occidryas editha nubigena ; [NACL, #4520s]
Euphydryas nubigena ; [BOW: pl. 13, f. 27]
E. e. remingtoni Burdick, 1959
ab. thielsenensis Gunder, 1931²
thielsonensis ; McDunnough, 1938, (missp.)
Occidryas editha remingtoni ; [NACL, #4520q]
E. e. rubicunda (H. Edwards, 1881)
California Melitaea quino var. rubicunda H. Edwards, 1881; Papilio 1 (4) : 52
Lemonias rubicunda ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 18
Occidryas editha rubicunda ; [NACL, #4520k]
Euphydryas editha rubicunda ; [BOW: pl. 13, f. 20]
E. e. taylori (Edwards, 1888)
British Columbia Melitaea taylori Edwards, 1888; Can. Ent. 20 (5) : 82
victoriae Gunder, 1926²
barnesi Gunder, 1928²
Euphydryas editha taylori, Butterflies of America [BOA]
Lemonias taylori ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 18
Occidryas editha taylori ; [NACL, #4520d]
Euphydryas taylori ; [BOW: pl. 2, f. 6 (text only)]
[taylori-1fu.jpg] 1822x1234 (~544Kb) upperside female Canada: British Columbia, Victoria, 6 May 1917 (topotype, W. Downes leg.), Photo 2007 © Norbert Kondla [taylori-1fv.jpg] 1772x1184 (~504Kb) underside female Canada: British Columbia, Victoria, 6 May 1917 (topotype, W. Downes leg.), Photo 2007 © Norbert Kondla

E. e. wrighti Gunder, 1929
fieldi Gunder, 1934²
thornei Gunder, 1934²
rubyae Hower, 1935²
Occidryas editha wrighti ; [NACL, #4520a]

The "Eurodryas" species-group
Euphydrini eurodryas ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3) : 165

fiEuphydryas aurinia (Rottemburg, 1775)
Yakutia, NW.China, Mongolia. See [About maps] Punakeltaverkkoperhonen Marsh Fritillary Ärenprisnätfjäril Guldnätfjäril Goldener Scheckenfalter Ondas rojas Przeplatka aurinia
Papilio aurinia Rottemburg, 1775; Der Naturforscher 6 : 5, TL: Paris
Papilio artemis Schiffermüller, 1775; (preocc.)
Papilio maturna Esper, 1777; Die Schmett. Th. I, Bd. 1 (8): 209, (3): pl. 16, f. 2 (preocc.)
?Papilio maturna Esper, 1778; Die Schmett. Th. I, Bd. 1 (8): 209, (7): pl. 37
Papilio lye Bergstrasser, 1780
Papilio koloswarensis Piller & Mitterpacher, 1783
Papilio matutina Thunberg, 1791
Melitaea aurinia ab. dubia Krulikovsky, 1890; Bull. Soc. imp. Nat. Moscou (n.s.) 4 (2) : 234, pl. 8, f. f
Melitaea aurinia ab. obscurata Krulikovsky, 1890; Bull. Soc. imp. Nat. Moscou (n.s.) 4 (2) : 235, pl. 8, f. c
Euphydryas aurinia, NSG Voucher Specimen [Wahlberg]
Euphydryas aurinia, Larvae of North-European Lepidoptera [Kimmo Silvonen]
Euphydryas aurinia, Svenska fjärilar [SVF]
Euphydryas aurinia, Moths and Butterflies of Europe [MBE]
Euphydryas aurinia, Europaische Schmetterlinge [Mario Maier]
Euphydryas aurinia, Captain's European Butterfly Guide [Simon Coombes]
Eurodryas aurinia ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3) : 165 (name); [BMAT]
Euphydryas aurinia ; [BOW: pl. 2, f. 6, 7]; [EBW]; [BARU, #155]; [MRS, 585]; [Wahlberg]; [Otakar Kudrna]
Euphydryas (aurinia) aurinia ; [BRU2: 55, pl. 36, f. 19-21, 25-27]
[aurinia-1u.jpg] 500x420 (~58Kb) upperside Criptor, Devon, UK 25/05/92, Photo © Simon Coombes [aurinia-2l.jpg] 550x371 (~40Kb) larva Central Europe 1990-2000, Photo © Michael Borsch [aurinia-2p.jpg] 371x550 (~48Kb) pupa Central Europe 1990-2000, Photo © Michael Borsch
[aurinia-2u.jpg] 595x400 (~57Kb) upperside Central Europe 1990-2000, Photo © Michael Borsch [aurinia-3.jpg] 1024x560 (~109Kb) Russia, Moscow area, 26.05.2007, Photo © D. Smirnov [aurinia-4l.jpg] 758x488 (~92Kb) larva Russia, Moscow area, 18.5.2008, Photo © D. Smirnov
[aurinia-4u.jpg] 748x596 (~109Kb) upperside Russia, Moscow area (36°25'E, 56°00'N), 14.6.2009, Photo © D. Smirnov [aurinia-4v.jpg] 809x652 (~98Kb) underside Russia, Moscow area, 14.6.2008, Photo © D. Smirnov

Larva on Plantago lanceolata, Succisa pratensis [SPRK], Knautia arvense [BARU]

Larva on Scabiosa succisa, Digitalis, Plantago, Veronica, Geranium, Sambucus, Gentiana, Valeriana, Lonicera, Spiraea, Viburnum, Succisa [BRU2, 55]


CEU, SEU, W.Siberia estonia (Petersen, 1902)
Euphydryas aurinia aurinia ; [BRU2, 55]; [Wahlberg]
E. a. bulgarica (Fruhstorfer, 1916)
Carphathians Euphydryas aurinia bulgarica ; [BRU2, 55]
E. a. laeta (Christoph, 1893)
C.Siberia, Altai, Sayan, Transbaikalia Melitaea aurinia var. laeta Christoph, 1893; Dt. Ent. Zeit. Iris, 6 (1) : 86
Euphydryas aurinia laeta ; [BRU2, 55]

Larva on Succisa pratensis [BRU2, 55]

E. a. beckeri (Lederer, 1853)
Morocco (Middle Atlas, Rif Mts) Melitaea beckeri Lederer, 1853; Verh. zool.-bot. Ver. Wien 2 (Abh.) : 39, TL: S.Spain
Melitaea aurinia f. iberica Oberthür, 1881; Étud. d'Ent. 6: ?, TL: Spain
Melitaea aurinia ellisoni Rungs, 1950; Bull. Soc. Sci. nat. Maroc 28: 145, TL: Ifrane (Morocco)
Melitaea aurinia kandarensis Rungs, 1950; Bull. Soc. Sci. nat. Maroc 28: 146 (= ellisoni)
Eurodryas aurinia beckeri ; [BMAT: 56, pl. 18, f. 18-25]

Larva on Lonicera implexa, L. etrusca?, L. biflora?, L. periclymenum?, (in Spain) Knautia arvensis, Succisa pratensis, Digitalis sp., Plantago sp., Veronica sp., Geranium sp. [BMAT]


ellisoni
[ellisoni-1u.jpg] 600x435 (~36Kb) upperside Morocco, Imouzzèr-Kandar, Moyen Atlas, Photo © Michel Tarrier

E. a. barraguei (Betz, 1956)
Algeria Melitaea aurinia barraguei Betz, 1956; Revue fr. Lépidopt. 15: 144, TL: Berrouaghia (Algeria)
Eurodryas aurinia barraguei ; [BMAT: 56, pl. 18, f. 26-29]
E. a. banghaasi (Seitz, 1908)
Euphydryas aurinia f. banghaasi ;
Euphydryas banghaasi ; Lukhtanov, 1994
[banghaasi-1fu.jpg] 684x616 (~64Kb) upperside female An alpine meadow in the valley of the left headwater of the Chikty Rivulet, 2600 m above sea level, the southern principle slope of the Yuzhno-Chuiskii mountain range, the upper Dzhazator River basin, Kosh-Agach District, Altai Republic, West Siberia, Russia. 10th July 1998 (on Viola altaiaca), Photo © Oleg Kosterin [banghaasi-1mu.jpg] 527x440 (~21Kb) upperside male An alpine meadow in the valley of the left headwater of the Chikty Rivulet, 2600 m above sea level, the southern principle slope of the Yuzhno-Chuiskii mountain range, the upper Dzhazator River basin, Kosh-Agach District, Altai Republic, West SIberia, Russia. 10th July 1998, Photo © Oleg Kosterin [banghaasi-2u.jpg] 762x503 (~54Kb) upperside An alpine meadow in the valley of the left headwater of the Chikty Rivulet, 2600 m above sea level, the southern principle slope of the Yuzhno-Chuiskii mountain range, the upper Dzhazator River basin, Kosh-Agach District, Altai Republic, West SIberia, Russia. 10th July 1998, Photo © Oleg Kosterin

E. a. debilis Oberthür, 1909

Euphydryas glaciegenita (Verity, 1928)
Melitaea glaciegenita Verity, 1928; Ent. Rec. 40: 43
Euphydryas glaciegenita ; [Otakar Kudrna]

Euphydryas provincialis (Boisduval, 1828)
S.Europe, N.Africa, Asia Minor. See [About maps] provincialis (Boisduval, 1828); Eur. Lepid. Index Meth. 1: 17, TL: Provence [France]
Euphydryas (aurinia) provincialis ; [BRU2: 55, pl. 36, f. 22-24]

Larva on Scabiosa pratensis, S. columbaria, (Turkey) [BRU2, 56]

E. p. volhynica (Ksienschopolsky, 1912)
Caucasus Major, SEEU Euphydryas provincialis volhynica ; [BRU2, 56]

Larva on Cephalaria gigantea [BRU2, 56]


Euphydryas merope (Prunner, 1798)
-. See [About maps] merope (Prunner, 1798); Lepidopt. Pedemont. Suppl.: 73, TL: Valle Varaitana circa Lyaiol [Alps]
Euphydryas (aurinia) merope ; [BRU2: 56, pl. 36, f. 28-30]
Alps Euphydryas merope merope ; [BRU2, 56]
E. m. pellucida (Christoph, 1893)
Caucasus Major merope pellucida (Christoph, 1893); Dt. Ent. Zeit. Iris, 6 (1) : 86
Melitaea aurinia var. pellucida Christoph, 1893; Dt. Ent. Zeit. Iris, 6 (1) : 86
Euphydryas merope pellucida ; [BRU2, 56]
E. m. altivolans Tuzov, 2000
Altai, Sayan, S.Siberia Euphydryas merope altivolans ; [BRU2, 56]

Euphydryas orientalis (Herrich-Schäffer, [1851])
-. See [About maps] Melitaea orientalis Herrich-Schäffer, [1851]; Syst. Bearb. Schmett. Europ. 1 (10): (ii) pl. 56, f. 265-266, TL: [Amasya, Istanbul, Turkey]
Eurodryas orientalis ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3) : 165 (name)
Euphydryas orientalis ; [Wahlberg]
Euphydryas (aurinia) orientalis ; [BRU2: 56, pl. 36, f. 31-36]
Turkey, Transcaucasia Euphydryas orientalis orientalis ; [BRU2, 56]
E. o. sareptensis (Staudinger, 1878)
S.Ural Melitaea sareptensis Staudinger, 1878; Horae Soc. Ent. Ross. 14 : 265, TL: Sarepta
Melitaea sareptana Staudinger, 1861; in Staudinger & Wocke, Cat. Lep. (ed. 1): 8 (nom. nud.?)
Melitaea sareptana Staudinger, 1871; in Staudinger & Wocke, Cat. Lepid. eur. Faunegeb. (Edn. 2) : 17 (nom. nud.?)
Euphydryas sareptana ; [BOW: pl. 194, f. 20]
Euphydryas orientalis sareptensis ; [BRU2, 56]
E. o. emba (Fruhstorfer, 1917)
N.Kazakhstan Euphydryas orientalis emba ; [BRU2, 56]

Euphydryas asiatica (Staudinger, 1881)
Turkestan. See [About maps] Melitaea aurinia var. asiatica Staudinger, 1881; Stettin ent. Ztg 42 (7-9): 287, TL: "Ala Tau" [Dzhungarsky Alatau Mts., Kazakhstan]
Euphydryas asiatica ; [Oleg Kosterin]
Euphydryas aurinia asiatica ; [Wahlberg]
Euphydryas (aurinia) asiatica ; [BRU2: 57, pl. 37, f. 1-6]
[asiatica-1mu.jpg] 533x400 (~60Kb) upperside male A meadow in the lower valley of the Kora River about 10 km upstream of the Tekeli City (1300 m above sea level), the North Dzhungarskiy Alatau mountain chain (NE part of Tien Shan), Kopal District, Taldy-Kurgan Province, SE Kazakhstan. 13th June 1993, Photo © Oleg Kosterin

Tarbagatai, Saur, Dzhungarsky Alatau Euphydryas asiatica asiatica ; [BRU2, 57]
E. a. alexandrina (Staudinger, 1887)
Tian-Shan Melitaea aurinia var. alexandrina Staudinger, 1887; Stettin Ent. Ztg 48 (1-3) : 52
Euphydryas alexandrina, NSG Voucher Specimen [Wahlberg]
Eurodryas alexandrina ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3) : 165 (name)
Euphydryas alexandrina ; [Wahlberg]
Euphydryas asiatica alexandrina ; [BRU2, 57]
E. a. narina (Oberthür, 1909)
Inner Tian-Shan Euphydryas asiatica narina ; [BRU2, 57]

Euphydryas sibirica (Staudinger, 1871)
-. See [About maps] sibirica (Staudinger, 1871); in Staudinger & Wocke, Cat. Lepid. eur. Faunegeb. (Edn. 2) : 17, TL: Sib.[eria] or.[ientalis] Daür.[ia]
Euphydryas (aurinia) sibirica ; [BRU2: 57, pl. 37, f. 7-15]
Transbaikalia Euphydryas sibirica sibirica ; [BRU2, 57]
E. s. eothena (Röber, 1926)
Amur, Ussuri mandschurica (Staudinger, 1892); (preocc.)
= Euphydryas aurinia davidi (Oberthür, 1881); [Wahlberg]
Euphydryas tjutjujensis Higgins, 1950
Euphydryas aurinia mandschurica ; [MRS, 585]
Euphydrya davidi tjutjujensis ; [BARU, (note)]
Euphydryas sibirica eothena ; [BRU2, 57]

Larva on Scabiosa lachnophylla [BRU2, 57]

E. s. davidi (Oberthür, 1881)
N.China, Tuva?, Mongolia? Melitaea artemis var. davidi Oberthür, 1881; , TL: N.China, Peking
Euphydryas davidi ; [BARU, #156]
Euphydryas aurinia davidi ; [Wahlberg]
Euphydryas sibirica davidi ; [BRU2, 57]
[davidi-1u.jpg] 800x600 (~151Kb) upperside Wet ground at the pool in the pine forest called Tsasucheiskii Bor at the village Nizhnii Tsasuchei, Onon District, SE Chita Province, Transbaikalia, Siberia, Russia. 22nd June 1995, Photo © Oleg Kosterin [davidi-2v.jpg] 450x600 (~56Kb) underside A meadow on the bank of Lake Butyvken, surrounded by a magnificent pine forest within a steppe zone called Tsasucheiskii Bor, 10 km SW of the village Nizhnii Tsasuchei, Onon District, SE Chita Province, Transbaikalia, Siberia, Russia. 25th June 1995 (a congregation), Photo © Oleg Kosterin

E. s. tenebricosa (Bang-Haas, 1927)
Gansu Euphydryas sibirica tenebricosa ; [BRU2, 57]
E. s. phyllis Hemming, 1941
N.Korea koreana (Collier, 1933); (preocc.)
discalis (Bryk, 1946)
Euphydryas sibirica phyllis ; [BRU2, 57]

Euphydryas desfontainii (Godart, 1819)
NAF, Spain, Pyrenees. See [About maps] Spanish Fritillary Dientes gualdos
Argynnis desfontainii Godart, 1819; Encyclopédie Méthodique. 9 (1): 278, TL: N.Africa, Spain
Euphydryas desfontainii, NSG Voucher Specimen [Wahlberg]
Euphydryas desfontainii, Moths and Butterflies of Europe [MBE]
Euphydryas desfontainii, Europaische Schmetterlinge [Mario Maier]
Eurodryas desfontainii ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3) : 165 (name)
Euphydryas desfontainii ; [BOW: pl. 2, f. 9]; [Wahlberg]; [Otakar Kudrna]

Larva on Knautia [H&R]

Morocco (Rif Mts, Middle Atlas), W.Algeria Eurodryas desfontainii desfontainii
Melitaea desfontainii var. gibrati Oberthür, 1922; Étud. Lépid. Comp. 19 (1): 46, (2): pl. 532, f. 4418-4422, TL: Morocco
Eurodryas desfontainii desfontainii ; [BMAT: 56, pl. 18, f. 30-37]

Larva on Knautia arvensis [BMAT]


gibrati
[gibrati-1mf.jpg] 600x435 (~77Kb) male female Morocco, Tizi-n-Tretten, Moyen Atlas, Photo © Jean Delacre [gibrati-1mfv.jpg] 600x435 (~61Kb) underside male female Morocco, Tizi-n-Tretten, Moyen Atlas, Photo © Jean Delacre [gibrati-2mfu.jpg] 600x435 (~58Kb) upperside male female Morocco, Ito, Moyen Atlas, Photo © Michel Tarrier

[martae-1u.jpg] 800x580 (~68Kb) upperside Morocco, Djebel Azourki, Haut Atlas, Photo © Jean Delacre


21.2.2013 (17)

References:
[ALBERTA]
University of Alberta, E.H. Strickland Entomological Museum; Browse Entomology Collection
[About maps]
Warning! The maps are automatically generated from the textual information, and the process does not always produce acceptable result; See about maps for more info.
[BAMONA]
Opler, Pavulaan, Stanford & Pogue; Butterflies and Moths of North America; Mountain Prairie Information Node
[BARU]
Korshunov & Gorbunov; Butterflies of the Asian part of Russia; English translation by Oleg Kosterin; [Dnevnye babochki aziatskoi chasti Rossii. Spravochnik]
[BHL]
The; Biodiversity Heritage Library; and on; Internet Archive; Note: The linking to the content of Internet Archive is somewhat delicate -- notify me when links break!
[BMAT]
Tennent, 1996; The butterflies of Morocco, Algeria and Tunisia; ISBN 0 906802 05 9
[BOA]
Interactive Listing of American Butterflies; Listado Interactivo de la Mariposas Americanas; Butterflies of America
[BOW]
Lewis, 1974; Butterflies of the World; ISBN 0 245 52097 X
[BRU2]
Tuzov (ed.), Bogdanov, Churkin, Devyatkin, Dantchenko, Murzin, Samodurov, Zhdanko, 2000; Guide to the Butterflies of Russia and adjacent territories; Libytheidae, Danaidae, Nymphalidae, Riodinidae, Lycaenidae; Volume 2
[Bruce Walsh]
Bruce Walsh; e-mail: Bruce Walsh; The Moths of Southeastern Arizona; Butterflies of Southeastern Arizona
[Christopher A. Rickards]
Ann Arbor, Mi 48105, USA.; e-mail: Christopher A. Rickards
[CiNii]
http://ci.nii.ac.jp/
[D. Smirnov]
Russia, Moscow; e-mail: D. Smirnov
[EBW]
Smart, Paul, 1976; The Illustrated Encyclopedia of the Butterfly World; ISBN 0 600 31381 6
[ESP]
Heppner, 1981; The dates of E. J. C. Esper's Die Schmetterlinge in Abblidungen... 1776-[1830]; Archives of Natural History (1981) 10 (2): 251-254
[GALEPS]
Moths and Butterflies of Georgia and the Southeastern United States; James K. Adams; Dept. of Natural Science, Dalton State College, Dalton, GA ; http://www.daltonstate.edu/galeps/index.htm
[H&R]
Higgins, L.G, Riley, N.D, 1970; A Field Guide to the Butterflies of Britain and Europe
[HBN]
Francis Hemming, 1937; HÜBNER, A bibliographical and systematic account of the entomological works of Jacob Hübner and of the supplements thereto by Carl Geyer, Gottfried Franz von Frölich and Gottlieb August Wilhelm Herrich-Schäffer, Vol 1 & 2
[ILMB]
Interactive Listing of Mexican Butterflies
[Jean Delacre]
http://homepage.mac.com/jdelacre
[Kimmo Silvonen]
Espoo, Finland; Kimmo's Lep Site
[MBE]
Moths and Butteflies of Europe; Paolo Mazzei, Diego Reggianti and Ilaria Pimpinelli; http://www.leps.it/
[MRS]
Chou Io (Ed.); Monographia Rhopalocerum Sinensium, 1-2
[Mario Maier]
Europäische Schmetterlinge; e-mail: Mario Maier
[Michael Borsch]
Geesthacht, Germany; contact through; e-mail: Francisco Welter-Schultes
[Michel Tarrier]
Michel R. Tarrier; Malaga, Spain; http://www.tarrier.org
[NAB]
Pyle, 1981; Field Guide to North American Butterflies, National Audubon Society; ISBN 0-394-51914-0
[NACL]
Hodges, 1983; Check List of the Lepidoptera of America North of Mexico
[NL4A]
Gerardo Lamas, 2004; Atlas of Neotropical Lepidoptera; Checklist: Part 4A; Hesperioidea - Papilionoidea
[NORLEP]
Norges Sommerfugler
[Norbert Kondla]
Canada, BC; e-mail: Norbert Kondla
[Oleg Kosterin]
Institue of Cytology & Genetics; Siberian Division of Russian Academy of Sciences; Novosibirsk; e-mail: Oleg Kosterin; http://pisum.bionet.nsc.ru/kosterin/
[Opler]
Opler & Warren, 2003 (April); Butterflies of North America. 2. Scientific Names List for Butterfly Species of North America, north of Mexico.
[Pertti Pakkanen]
e-mail: Pertti Pakkanen; Butterfly and moth observations from Finland
[SPRK]
Seppänen, E. J, 1970; Suomen suurperhostoukkien ravintokasvit, Animalia Fennica 14
[SVF]
Svenska fjärilar; Naturhistoriska riksmuseet; (Sweden)
[SZMN]
Lepidoptera; Siberian Zoological Museum
[Simon Coombes]
1 Park Street, Plymouth, PL3 4BL, UK; 01752 607854; e-mail: Simon Coombes; http://www.butterfly-guide.co.uk
[Vitaly Charny]
Birmingham, Alabama, USA
[Wahlberg]
Niklas Wahlberg; The NSG's voucher specimen database; Nymphalidae Systematics Group
[²]
This may require parentheses or not. I don't have the necessary information for this taxon.


Some related literature:
Barnes, 1897
Some new species and varieties of Lepidoptera from the Western U. S. Can. Ent. 29 (2) : 39-42
Barnes, 1900
New species and varietes of North American Lepidoptera Can. Ent. 32 (2) : 42-48
Behr, 1863
On Californian Lepidoptera - No. III Proc. Calif. Acad. Sci. 3 (1) : 84-93
Boisduval, [1833]
Icones historique des Lépidoptères nouveaux ou peu connus. Collection, avec Figures coloritées, des Papillons d'Europe nouvellement découverts; ouvrage formant le complément de tous les auteurs iconographes Icon. hist. Lépid. Europe 1 (1-2): 1-24, pl. 1-4 (1832), (3-4): 25-40, pl. 5-8 (1832), (5-6): 41-56, pl. 9-12 (1832), (7-8): 57-72, pl. 13-16 (1832), (9-10): 73-104, pl. 17-20 (1833), (11-12): 105-120, pl. 21-24 (1833), (13-14): 121-144, pl. 25-28 (1833), (15-16): 145-160, pl. 29-32 (1833), (17-18): 161-184, pl. 33-36 (1833), (19-20): 185-208, pl. 37-40 (1834), (21-22): 209-224, pl. 41-44 (1834), (23-24): 225-248, pl. 45-47 (1834), (25-28): 249-252 (1834)
Boisduval, 1852
Lépidoptères de la Californie Ann. Soc. ent. Fr. (2) 10 : 275-324
Christoph, 1893
Lepidoptera Nova Faunae Palaearcticae Dt. Ent. Zeit. Iris, 6 (1) : 86-96
Clark, 1927
Notes on the Melitaeid Butterfly Euphydryas phaëton (Drury) with Descriptions of a New Subspecies and a New Variety Proc. U.S. Nat. Mus. 71 (2683) : 1-22, pl. 1-5
Dalman, 1816
Försök till systematiks Uppställing af Sveriges Fjärilar K. VetenskAcad. Handl. 1816 (1): 48-101, (2): 199-225
dos Passos, 1938
A new race of Euphydryas chalcedonica Dbldy. & Hew. from Arizona Can. Ent. 70: 199-200
Doubleday, [1847]
The genera of diurnal Lepidoptera, comprising their generic characters, a notice of their habitats and transformations, and a catalogue of the species of each genus; illustrated with 86 plates by W. C. Hewitson Gen. diurn. Lep. (1): 1-18, pl. A, 1-4 (1-2) (1846), (1): 19-132, pl. 4*, 5-25, 28 (3-14) (1847), (1): 133-200, pl. 1*, 26, 27, 29, 31-44 (15-23) (1848), (1): 201-242, pl. 30, 45-52, 56-58, 60-62, 64 (24-31) (1849), (1): 243-250 [?], (2): 243-326, pl. 53-55, 63, 65, 66 (32-38) (1850), (2): 327-466, pl. 59, 67-77 (39-50) (1851), (2): 467-534, 143, 144, pl. 78-80 (Index) (51-54) (1852)
Drury, [1773]
Illustrations of natural history; wherein are exhibited ... Illust. Nat. Hist. Exot. Insects 1 : 1-130, pl. 1-50 (1770), (1773)
Dyar, 1903
A List of North American Lepidoptera and Key to the Literature of this Order of Insects Bull. U.S. nat. Mus. 52 : 1-723
Ellsworth, 1902
Notes on Butterflies and Description of an Aberration Ent. News. 13 : 103-104
Edwards, 1881
Description of new species of butterflies Papilio 1 (4) : 43-48
Edwards, 1888
Descriptions of two new species of Melitaea belonging to North America Can. Ent. 20 (5) : 81-84
Edwards, 1890
Description of a new species of Melitaea from southern California Can. Ent. 22 (2) : 21-23
H. Edwards, 1881
On some apparently new forms of diurnal lepidoptera Papilio 1 (4) : 50-55
Esper, 1777; Esper, 1778
Die Schmetterlinge in Abbildungen nach der Natur mit Beschreibungen. Theil I. Die Tagschmetterlinge. Band 1 Die Schmett. Th. I, Bd. 1 (1): 1-40, pl. 1-6 (1776), (2): 41-72, pl. 7-12 (1776), (3): 73-104, pl. 13-18 (1777), (4): 105-120, pl. 19-24 (1777), (5): 121-144, pl. 25-30 (1777), (6): 145-176, pl. 31-36 (1777), (7): 177-200, pl. 37-46 (1778), (8): 201-216, pl. 43-48 (1778), (9): 217-388, pl. 49-50 (1779)
Godart, 1819
in Latreille & Godart, Encyclopédie Méthodique. Histoire naturelle Entomologie, ou histoire naturelle des crustacés, des arachnides et des insectes Encyclopédie Méthodique. 9 (1): 3-328 (1819), (2): 329-828 ([1824])
Gunder, 1925
Several New Aberrant Lepidoptera (Rhopalocera) Ent. News 36 (7) : 193-200, pl. 5
Gunder, 1929
The genus Euphydryas Scud. of boreal America Pan-Pacific Entomol. 6: 1-8
Herrich-Schäffer, [1851]
Systematische Bearbeitung der Schmetterlinge von Europa, zugleich als Text, Revision und Supplement zu J. Hübner's Sammlung europäischer Schmetterlinge, 1843 (-1855), Die Tagfalter Syst. Bearb. Schmett. Europ. 1 (9): Title ("1843"), (1): [1-3], 4-16, (ii) pl. 1-5, f. 1-29 (1843), (2): 17-40, (ii) pl. 29-34, f. 131-156 (1843), (3): 41-56, (ii) pl. 6-12, f. 30-60, pl. 35-36, f. 157-164 (1844), (4): 57-80, (ii) pl. 13-22, f. 61-107 (1844), (5): 81-90, (ii) pl. 23-28, f. 108-130, pl. 37-41, f. 165-189 (1844), (6): 91-98, (ii) pl. 42-44, f. 190-203 (1844), (7): 99-114, (ii) pl. 45-52, f. 204-248 (1844), (9): 115-122 (1844), (10): 123-146, (ii) pl. 53-57, f. 249-274 (1845), (11): 147-162 (1845), (12): (iii) pl. 1-3, f. 1-17 (1845), (13): 163-164, (ii) pl. 58-65, f. 275-315 (1846), (14): (ii) pl. 66-70, f. 316-338, (iii) pl. 4-5, f. 18-30 (1846), (26): (ii) pl. 71-77, f. 339-376 (1847), (32): (ii) pl. 78-82, f. 377-400 (1848), (36): (ii) pl. 83-89, f. 401-426, (iii) pl. 6, f. 31-36 (1848), (37): (ii) pl. 90-91, f. 427-437 (1849), (43): (ii) pl. 92-94, f. 438-450, 452-453 (1850), (45): (ii) pl. 95-102, f. 453-489 (1850), (47): (ii) pl. 103, f. 490-495 (1851), (49): (ii) pl. 104-109, f. 496-528 (1851), (51): (ii) pl. 110-118, f. 529-570 (1851), (54): (ii) pl. 119-123, f. 571-596 (1852), (56): (ii) pl. 124-127, f. 597-618, pl. 130-132, f. 628-639, pl. 134, f. 646-651 (1852), (59): (ii) pl. 128-129, f. 619-627, pl. 133, f. 640-645 (1853), (65): (iii) pl. 7, f. 37-42, (index) [1], 2-20 (1854), (66): (index) 21-24 (1855)
Higgins, 1981
A revision of Phyciodes Hübner and related genera, with a review of the classification of the Melitaeinae Bull. Br. Mus. nat. Hist. (Ent.) 43 (3) : 77-243, f. 1-490,124a,125a
Hübner, [1806]
Tentamen determinationis digestionis alque denominationis singlarum stripium Lepidopterorum, peritisad inspiciendum et dijudicandum communicatum, a Jacob Hübner Tentamen determinationis digestionis ...: -
Krulikovsky, 1890
[russian: ? "experimental catalog of lepidoptera of Kazakh province"] Bull. Soc. imp. Nat. Moscou (n.s.) 4 (2) : 200-251, pl. 8
Lederer, 1853
Versuch die europäischen Lepidopteren (einschlissig der ihrem Habitus nach noch zure europäischen Fauna gehörigen Arten Labradors, der asiatischen Türkei un des asiatischen Russlands) in möglichst natürliche Reihenfolge zu stellen, nebst Bemerkungen zu eineg Familien und Arten. Rhopaloceren & Heteroceren Verh. zool.-bot. Ver. Wien 2 (Abh.) : 14-54 (Rhopaloceren), : 65-126 (Heteroceren)
Linnaeus, 1758
Systema Naturae per Regna Tria Naturae, Secundum Clases, Ordines, Genera, Species, cum Characteribus, Differentiis, Symonymis, Locis. Tomis I. 10th Edition Syst. Nat. (Edn 10) 1 : 1-338, : 339-824
Matsumura, 1927
Some new Butterflies Ins. Matsumurana 2 (2) : 114-118
Ménétriés, 1859
Lépidoptères de la Sibérie orientale et en particulier des rives de l'Amour in Schrenck, Reise Forschungen Amur-Lande 2 (1) : 1-75, pl. 1-5
Murayama, 1955
New or Little Known Rhopalocera from China and Korea (with 10 text figures) Tyô to Ga 6 (1) : 1-4
Oberthür, 1922
Les Lépidoptères du Maroc Étud. Lépid. Comp. 19 (1): 13-402, (2): pl. 530-546
Rottemburg, 1775
Unmertungen zu den Hufnagelifchen Tabellen der Schmetterlinge Der Naturforscher 6 : 1-34
Scudder, 1869
Report upon a Collection of Diurnal Lepidoptera made in Alaska by the Scientific Corps of the Russo-American Telegraph Expedition under the Direction of Lieut. W. H. Dall Proc. Boston Soc. nat. Hist. 12 : 404-408
Scudder, 1872
A systematic Revision of some of the American Butterflies; with brief notes on those known to occur in Essex County, Mass. 4th Ann. Rep. Peabody Acad. Sci. (1871): 24-83
Skinner, 1897
Notes on Rhopalocera, with descriptions of new species and varieties Can. Ent. 29 (6) : 154-156
Skinner, 1899
A new butterfly from Utah Can. Ent. 31 (1) : 12
Skinner, 1921
A New Species of Melitaea from Montana (Lepid., Rhop.) Ent. News 32 (3) : 89
Staudinger, 1861
Catalog der Lepidopteren Europas und der angrendzenden Länder in Staudinger & Wocke, Cat. Lep. (ed. 1): 1-192
Staudinger, 1878
Lepidopteren-Fauna Kleinasiens Horae Soc. Ent. Ross. 14 : 176-482
Staudinger, 1881
Beitrag zur Lepidopterenfauna Central-Asiens Stettin ent. Ztg 42 (7-9): 253-300, (10-12): 393-424
Staudinger, 1887
Centralasiatische Lepidopteren Stettin Ent. Ztg 48 (1-3) : 49-102
Staudinger in Staudinger & Wocke, 1871
Catalog der Lepidopteren des europaeischen Faunegebiets (Edn. 2): - Cat. Lepid. eur. Faunegeb. (Edn. 2) : 1-421, : 422-426 (Corrigenda et Addenda)


If you have corrections, comments or information to add into these pages, just send mail to Markku Savela
Keep in mind that the taxonomic information is copied from various sources, and may include many inaccuracies. Expert help is welcome.