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Chlosyne Butler, 1870
Chlosyne Butler, 1870; Cistula ent. 1: 38 (repl. Synchloe Doubleday, 1844), TS: Papilio janais Drury
Morpheis Geyer, [1833]; in Hübner, Samml. exot. Schmett. 3 : pl. [5] (preocc. Morpheis Hübner, [1820]), TS: Morpheis ehrenbergii Geyer
= ; [NL4A, 252]
Synchloe Doubleday, 1844; List. lepid. Ins. Brit. Mus. 1 : 76 (preocc. Synchloa Hübner, 1818), TS: Papilio janais Drury
= ; [NL4A, 252]
Coatlantona Kirby, 1871; Syn. Cat. diurn. Lep. : 178 (repl. Synchloe Doubleday, 1844), TS: Papilio janais Drury
= ; [NL4A, 252]
Anemeca Kirby, 1871; Syn. Cat. diurn. Lep. : 179 (repl. Morpheis Geyer, [1833]), TS: Morpheus ehrenbergii Geyer
= ; [NL4A, 252]
Charidryas Scudder, 1872; 4th Ann. Rep. Peabody Acad. Sci. (1871): 47, TS: Melitaea nycteis Doubleday
= ; [Wahlberg]; [NL4A, 252]
Limnaecia Scudder, 1872; 4th Ann. Rep. Peabody Acad. Sci. (1871): 47 (preocc. Limnaecia Stainton, 1851), TS: Melitaea harrisii Scudder
= Charidryas ;
= ; [Wahlberg]; [NL4A, 252]
Thessalia Scudder, 1875; Bull. Buffalo Soc. nat. Sci. 2 : 238, 265, TS: Melitaea leanira C. & R. Felder
= ; [Wahlberg]; [NL4A, 252]
Chlosyne, NSG Voucher Specimen [Wahlberg]
Synchloe ; Godman & Salvin, [1882], Biol. centr.-amer., Lep. Rhop. 1: 176
Morpheis ; Godman & Salvin, [1882], Biol. centr.-amer., Lep. Rhop. 1: 211
Thessalia (Melitaeini) ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3) : 167
Chlosyne (Melitaeini) ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3) : 166 (included Charidryas also)
Chlosyne (Chlosynina) ; [Wahlberg]
Chlosyne ([I]) ; [NL4A, 252]


[sp-1u.jpg] 700x605 (~70Kb) upperside USA: Bridge Creek Trailhead, North Cascades Highway, (48°30'30N 120°43'00W), Chelan Co., WA, 28.7.1999, Photo © Markku Savela [sp-1v.jpg] 700x679 (~62Kb) underside USA: Bridge Creek Trailhead, North Cascades Highway, (48°30'30N 120°43'00W), Chelan Co., WA, 28.7.1999, Photo © Markku Savela [sp-2v.jpg] 550x619 (~38Kb) underside USA: Table Mtn, Kittitas Co., WA, 30.7.1999, Photo © Markku Savela
[sp-3u.jpg] 600x829 (~63Kb) upperside USA: nr Crystal Mountain / FR9726 (47°14'37N 120°38'39W), Kittitas Co., WA, 11.7.2000, Photo © Markku Savela [sp-3v.jpg] 600x722 (~55Kb) underside USA: nr Crystal Mountain / FR9726 (47°14'37N 120°38'39W), Kittitas Co., WA, 11.7.2000, Photo © Markku Savela

The exact identification of these species is still unknown, but tentatively assumed to belong into this group.


The "lacinia" species-group

Chlosyne janais (Drury, [1782])
Texas, Mexico - Colombia. See [About maps] Janais Patch
Papilio janais Drury, [1782]; Illust. Nat. Hist. Exot. Insects 3: index, : 22, pl. 17, f. 5-6, TL: ["Brazil, Rio de Janeiro"]
Chlosyne janais janais, Listing of Mexican Butterflies [ILMB]
Chlosyne janais janais, Butterflies of America [BOA]
Chlosyne janais, NSG Voucher Specimen [Wahlberg]
Chlosyne janais, Butterflies and Moths of North America [BAMONA]
Synchloe janais ; Godman & Salvin, [1882], Biol. centr.-amer., Lep. Rhop. 1: 178; Godman & Salvin, [1901], Biol. centr.-amer., Lep. Rhop. 2: 671
Chlosyne janais ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 21; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3) : 167 (name); [NACL, #4503]; [BCR, 199]; [EBW]; [BOW: pl. 13, f. 13]; [Opler]; [NL4A, #2083]

Larva on Anisacanthus wrightii, Odontonema callistachus [NAB], Odontonema [BCR]

Chlosyne janais janais ; [NL4A, #2083a]
C. j. gloriosa Bauer, [1960]
Mexico Chlosyne gloriosa Bauer, [1960]; J. Lep. Soc. 13: 165, TL: Tepic, Nayarit, Mexico
Chlosyne janais gloriosa, Listing of Mexican Butterflies [ILMB]
Chlosyne janais gloriosa, Butterflies of America [BOA]
Chlosyne gloriosa ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3) : 167 (name)
Chlosyne janais gloriosa ; [NL4A, #2083b]
C. j. marianna Röber, [1914]
Mexico Chlosyne marianna Röber, [1914]; , TL: Mexico
Chlosyne hyperia var. irrubescens Hall, 1917; , TL: Mexico, Morelos
= ; [NL4A, #2083c]
Chlosyne janais marianna, Listing of Mexican Butterflies [ILMB]
Chlosyne janais marianna, Butterflies of America [BOA]
Chlosyne marianna, NSG Voucher Specimen [Wahlberg]
Chlosyne marianna ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3) : 167 (name)
Chlosyne janais marianna ; [NL4A, #2083c]

Chlosyne rosita Hall, 1924
-. See [About maps] Rosita Patch
Chlosyne rosita Hall, 1924; , TL: Guatemala
Chlosyne rosita rosita, Listing of Mexican Butterflies [ILMB]
Chlosyne rosita rosita, Butterflies of America [BOA]
Chlosyne rosita, NSG Voucher Specimen [Wahlberg]
Chlosyne rosita, Butterflies and Moths of North America [BAMONA]
Chlosyne rosita ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3) : 167 (name); [NACL, #4504]; [Opler]; [NL4A, #2091]
Guatemala Chlosyne rosita rosita ; [NL4A, #2091a]
C. r. montana Hall, 1924
Mexico Chlosyne lacinia montana Hall, 1924; Entomologist 57: (241-242)
Chlosyne rosita montana, Listing of Mexican Butterflies [ILMB]
Chlosyne rosita montana, Butterflies of America [BOA]
Chlosyne rosita montana, Butterflies of Southeastern Arizon [Bruce Walsh]
Chlosyne montana ;
Chlosyne rosita montana ; [NL4A, #2091d]
C. r. browni Bauer, [1961]
Mexico Chlosyne rosita browni Bauer, [1961]; J. Lep. Soc. 14 (2): 150, TL: El Salto, 1600ft, San Luis Potosí, Mexico
Chlosyne rosita browni, Listing of Mexican Butterflies [ILMB]
Chlosyne rosita browni, Butterflies of America [BOA]
Chlosyne rosita browni ; [NACL, #4504a]; [NL4A, #2091b]
C. r. riobalsensis Bauer, [1961]
Mexico Chlosyne riobalsensis Bauer, [1961]; J. Lep. Soc. 14 (2): 152, TL: Mexcala, 2000ft, Guerrero, Mexico
Chlosyne rosita riobalsensis, Listing of Mexican Butterflies [ILMB]
Chlosyne rosita riobalsensis, Butterflies of America [BOA]
Chlosyne riobalsensis ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3) : 167 (name)
Chlosyne rosita riobalsensis ; [NL4A, #2091e]
C. r. mazarum Miller & Rotger, 1979
Mexico Chlosyne rosita mazarum Miller & Rotger, 1979; , TL: Mexico, Morelos
Chlosyne rosita mazarum, Listing of Mexican Butterflies [ILMB]
Chlosyne rosita mazarum, Butterflies of America [BOA]
Chlosyne rosita mazarum ; [NL4A, #2091c]

Chlosyne hippodrome (Geyer, 1837)
Mexico - Colombia. See [About maps] Araschnia hippodrome Geyer, 1837; in Hübner, Zuträge. Samml. exot. Schmett. 5 : f. 863-864, TL: Mexico
Chlosyne hippodrome hippodrome, Listing of Mexican Butterflies [ILMB]
Chlosyne hippodrome hippodrome, Butterflies of America [BOA]
Chlosyne hippodrome, NSG Voucher Specimen [Wahlberg]
Chlosyne hippodrome, Butterflies and Moths of North America [BAMONA]
Chlosyne hippodrome ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3) : 167 (name); [BCR, 199]; [NL4A, #2082]

Larva on Melanthera aspera [BCR]

Mexico Chlosyne hippodrome hippodrome ; [NL4A, #2082a]
C. h. fabricii Higgins, 1960
hippodrome fabricii Higgins, 1960; (repl. name), TL: "Indiis"
Papilio hyperia Fabricius, 1793; Ent. Syst. 3 (1): 119 (preocc. Cramer, 1779), TL: "Indiis"
= ; [NL4A, #2082b]
Synchloe hyperia ; Godman & Salvin, [1882], Biol. centr.-amer., Lep. Rhop. 1: 179; Godman & Salvin, [1901], Biol. centr.-amer., Lep. Rhop. 2: 672
Chlosyne hippodrome fabricii ; [NL4A, #2082b]

Chlosyne gaudialis (Bates, 1864)
Panama, .... See [About maps] Synchloe gaudialis Bates, 1864; Ent. mon. Mag. 1 (4): 84, n. 40, TL: Guatemala
Chlosyne gaudialis gaudialis, Listing of Mexican Butterflies [ILMB]
Chlosyne gaudialis gaudialis, Butterflies of America [BOA]
Chlosyne gaudialis, NSG Voucher Specimen [Wahlberg]
Synchloe gaudialis ; Godman & Salvin, [1882], Biol. centr.-amer., Lep. Rhop. 1: 182; Godman & Salvin, [1901], Biol. centr.-amer., Lep. Rhop. 2: 673
?Coatlantona gaudialis ;
Chlosyne gaudealis[sic?] ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3) : 167 (name)
Chlosyne gaudealis[sic] ; [BCR, 200 (missp.)]
Chlosyne gaudialis ; [EBW]; [NL4A, #2080]
[gaudialis-1.jpg] 681x619 (~46Kb) Belize, Central America, Photo © John Shuey

Larva on Justicia [BCR]


Guatemala, Nicaragua Chlosyne gaudialis laeta Röber, [1914]; , TL: Nicaragua
= ; [NL4A, #2080a]
Chlosyne gaudialis gaudialis ; [NL4A, #2080a]
C. g. wellingi Miller & Rotger, 1979
Mexico gaudialis wellingi Miller & Rotger, 1979; , TL: Mexico, Oaxaca
Chlosyne gaudialis wellingi, Listing of Mexican Butterflies [ILMB]
Chlosyne gaudialis wellingi, Butterflies of America [BOA]
Chlosyne gaudialis wellingi ; [NL4A, #2080b]

Chlosyne narva (Fabricius, 1793)
Nicaragua - Venezuela. See [About maps] Papilio narva Fabricius, 1793; , TL: "Africa"
Chlosyne narva, NSG Voucher Specimen [Wahlberg]
Chlosyne narva ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3) : 167 (name); [BCR, 200]; [BOW: pl. 34, f. 7]; [NL4A, #2089]

Larva on Amaranthus? [BCR]

Chlosyne narva narva ; [NL4A, #2089a]
C. n. bonpland (Latreille, [1809])
Panama Cethosia bonpland Latreille, [1809]; in Humboldt & Bonpland, Voy. rég. équinoxiales..., 1 (4): 199, pl. 18, f. 5, TL: "[Ecuador]"
Chlosyne narva var. brunhilda Staudinger, 1885; , TL: Panama
= ; [NL4A, #2089b]
Synchloe bonplandi[sic, recte bonpland] ; Godman & Salvin, [1882], Biol. centr.-amer., Lep. Rhop. 1: 182; Godman & Salvin, [1901], Biol. centr.-amer., Lep. Rhop. 2: 673
Chlosyne narva bonpland ; [NL4A, #2089b]
Misidentified/misapplied:
?Synchloe narva; Godman & Salvin, [1882], Biol. centr.-amer., Lep. Rhop. 3: pl. 20, f. 7-8

Chlosyne ehrenbergii (Geyer, [1833])
Mexico. See [About maps] Morpheis ehrenbergii Geyer, [1833]; in Hübner, Samml. exot. Schmett. 3 : pl. [5], TL: [Mexico]
Chlosyne ehrenbergii, Listing of Mexican Butterflies [ILMB]
Chlosyne ehrenbergii, Butterflies of America [BOA]
Chlosyne ehrenbergii, NSG Voucher Specimen [Wahlberg]
Chlosyne ehrenbergi, Butterflies and Moths of North America [BAMONA]
Anemeca ehrenbergi ;
Morpheis ehrenbergi ; Godman & Salvin, [1882], Biol. centr.-amer., Lep. Rhop. 1: 211; Godman & Salvin, [1901], Biol. centr.-amer., Lep. Rhop. 2: 682
Chlosyne ehrenbergi ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3) : 167 (name); [BOW: pl. 34, f. 18]
Chlosyne ehrenbergii ; [NL4A, #2075]

Chlosyne lacinia (Geyer, 1837)
-. See [About maps] Bordered Patch
Araschnia lacinia Geyer, 1837; in Hübner, Zuträge. Samml. exot. Schmett. 5 : 25, : f. 899-900, TL: Mexico
Chlosyne lacinia lacinia, Listing of Mexican Butterflies [ILMB]
Chlosyne lacinia lacinia, Butterflies of America [BOA]
Chlosyne lacinia, NSG Voucher Specimen [Wahlberg]
Chlosyne lacinia, Butterflies and Moths of North America [BAMONA]
Synchloe lacinia ; Godman & Salvin, [1882], Biol. centr.-amer., Lep. Rhop. 1: 176, 3: pl. 19, f. 6-17; Godman & Salvin, [1901], Biol. centr.-amer., Lep. Rhop. 2: 671
Chlosyne lacinia ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 21; [NACL, #4499]; [BCR, 201]; [BOW: pl. 13, f. 14, pl. 34, f. 19]; [Opler]; [NL4A, #2085]
Chlosyne lacina[sic] ; [EBW]

Larva on Helianthus annuus, Ambrosia trifida, Verbesina encelioides [NAB], Helianthus, Verbesina, Ambrosia [BCR]

Mexico, Guatemala, Panama, Costa Rica Synchloe tellias Bates, 1864; Ent. mon. Mag. 1 (4): 82, TL: Guatemala
= ; [NL4A, #2085a]
Synchloe quehtala Reakirt, 1866; Proc. Acad. nat. Sci. Philad. 18 (3) : 248, TL: Mexico
= Chlosyne lacinia ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 21
= ; [NL4A, #2085a]
Synchloe ardema Reakirt, [1867]; Proc. Acad. nat. Sci. Philad. 18 (4): 336, TL: Mexico, Veracruz
= Chlosyne lacinia ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 21
= ; [NL4A, #2085a]
Synchloe misera R. Felder, 1869; Verh. zool.-bot. Ges. Wien. 19 : 471, TL: Mexico, Oaxaca; Morelos
= Chlosyne lacinia ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 21
= ; [NL4A, #2085a]
Synchloe pretona Boisduval, 1870; Considérations Lépid. Guatemala: 37, TL: Guatemala; Mexico
= Chlosyne lacinia ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 21
= ; [NL4A, #2085a]
Synchloe adelina Staudinger, 1876
Synchloë adelina Staudinger, 1876; Verh. zool.-bot. Ges. Wien, 25 (1) : 102, TL: Panama
= ; [NL4A, #2085a]
f. indigens Higgins, 1960; , TL: Costa Rica
= ; [NL4A, #2085a]
pretoria[sic] ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 21 (missp.)
Chlosyne lacinia adelina ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 21
Chlosyne lacinia lacinia ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3) : 167 (name); [NL4A, #2085a]
C. l. saundersi (Doubleday, [1847])
Venezuela, Colombia, Ecuador, Bolivia, Argentina Synchloe saundersi Doubleday, [1847]; Gen. diurn. Lep. (1): 186, (1): pl. 24, f. 2, TL: Venezuela
Synchloë mediatrix C. & R. Felder, [1867]; Reise Fregatte Novara, Bd 2 (Abth. 2) (3): 395, TL: Colombia
= ; [NL4A, #2085d]
Synchloë paupera C. & R. Felder, [1867]; Reise Fregatte Novara, Bd 2 (Abth. 2) (3): 395, TL: Colombia
= Chlosyne lacinia ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 21
= ; [NL4A, #2085d]
Chlosyne mediatrix f. fassli Röber, [1914]; , TL: Colombia
= ; [NL4A, #2085d]
Chlosyne mediatrix f. felderi Röber, [1914]; , TL: Colombia
= ; [NL4A, #2085d]
Chlosyne mediatrix f. fruhtorferi Röber, [1914]; , TL: Bolivia
= ; [NL4A, #2085d]
Chlosyne saundersi f. splendida Hayward, 1931; , TL: Argentina
= ; [NL4A, #2085d]
Chlosyne saundersi f. splendida ab. reducta Hayward, 1931; , TL: Argentina
= ; [NL4A, #2085d]
Chlosyne saundersi f. infuscata Hayward, 1935; , TL: Argentina
= ; [NL4A, #2085d]
Synchloë saundersii ; Doubleday, [1847], (missp.)
Chlosyne lacinia saundersi ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 21; Brown & Mielke, 1967, J. Lep. Soc. 21 (2): 98; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3) : 167 (name); [EcuL]; [NL4A, #2085d]
C. l. crocale (Edwards, 1874)
Arizona, New Mexico, Mexico Synchloe crocale Edwards, 1874; Trans. amer. ent. Soc. 5 (1): 17, TL: Arizona
Synchloe lacinia f. nigra Cockerell, 1893; , TL: New Mexico
= ; [NL4A, #2085c (part.)]
Synchloe lacinia f. bicolor Cockerell, 1893; , TL: New Mexico
= ; [NL4A, #2085c (part.)]
Synchloe lacinia f. rufa Cockerell, 1893; , TL: New Mexico
= ; [NL4A, #2085c (part.)]
Synchloe lacinia ab. nigrescens Edwards, 1893; , TL: New Mexico
= ; [NL4A, #2085c]
Synchloe lacinia ab. rufescens Edwards, 1893; , TL: Mexico, Chihuahua
= ; [NL4A, #2085c]
Synchloe lacinia ab. rufescens Cockerell, 1894; , TL: Mexico, Chihuahua
= ; [NL4A, #2085c]
Chlosyne lacinia f. flavida Higgins, 1960; , TL: Mexico, Chihuahua
= ; [NL4A, #2085c]
Chlosyne lacinia crocale, Listing of Mexican Butterflies [ILMB]
Chlosyne lacinia crocale, Butterflies of America [BOA]
Chlosyne lacinia crocale, Butterflies of Southeastern Arizon [Bruce Walsh]
Chlosyne lacinia crocale ; [NL4A, #2085c]; [NACL, #4499a]
C. l. adjutrix (Scudder, 1875)
Texas, New Mexico, Arizona Chlosyne adjutrix Scudder, 1875; Bull. Buffalo Soc. nat. Sci. 2 : 269, TL: Texas
Synchloe lacinia f. nigra Cockerell, 1893; , TL: New Mexico
= ; [NL4A, #2085b (part.)]
Synchloe lacinia f. bicolor Cockerell, 1893; , TL: New Mexico
= ; [NL4A, #2085b (part.)]
Synchloe lacinia f. rufa Cockerell, 1893; , TL: New Mexico
= ; [NL4A, #2085b (part.)]
Chlosyne lacinia f. rufescens tr.f. inghami Gunder, 1928; , TL: Arizona
= ; [NL4A, #2085b]
Chlosyne lacinia adjutrix, Listing of Mexican Butterflies [ILMB]
Chlosyne lacinia adjutrix, Butterflies of America [BOA]
Chlosyne lacinia adjutrix ; [NACL, #4499b]; [NL4A, #2085b]

Chlosyne californica (Wright, 1905)
California. See [About maps] California Patch
Synchloe californica Wright, 1905; Butts. West Coast: 170, TL: California
Chlosyne califonia[sic] ab. chinoi Gunder, 1924; , TL: California
= ; [NL4A, #2072]
Chlosyne californica, Listing of Mexican Butterflies [ILMB]
Chlosyne californica, Butterflies of America [BOA]
Chlosyne californica, NSG Voucher Specimen [Wahlberg]
Chlosyne californica, Butterflies and Moths of North America [BAMONA]
Chlosyne californica ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3) : 167 (name); [NACL, #4498]; [Opler]; [NL4A, #2072]

Larva on Viguiera deltoidea f. parishii, Helianthus annuus [NAB]


Chlosyne melanarge (Bates, 1864)
Mexico - Costa Rica, Guatemala. See [About maps] Synchloe melanarge Bates, 1864; Ent. mon. Mag. 1 (4): 85, TL: Guatemala
Chlosyne paupera Boisduval, 1870; Considérations Lépid. Guatemala: 38 (preocc. C. & R. Felder, 1867), TL: Guatemala
= ; [NL4A, #2088]
Chlosyne melanarge, Listing of Mexican Butterflies [ILMB]
Chlosyne melanarge, Butterflies of America [BOA]
Chlosyne melanarge, NSG Voucher Specimen [Wahlberg]
Synchloe melanarge ; Godman & Salvin, [1882], Biol. centr.-amer., Lep. Rhop. 1: 179, 3: pl. 20, f. 9-10; Godman & Salvin, [1901], Biol. centr.-amer., Lep. Rhop. 2: 672
Chlosyne melanarge ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3) : 167 (name); [BCR, 200]; [EBW]; [BOW: pl. 34, f. 22]; [NL4A, #2088]

Larva on Aphelandra deppiana [BCR]


Chlosyne erodyle (Bates, 1864)
Mexico - Costa Rica. See [About maps] Synchloe erodyle Bates, 1864; Ent. mon. Mag. 1 (4): 84, TL: Guatemala
Chlosyne erodyle, Listing of Mexican Butterflies [ILMB]
Chlosyne erodyle erodyle, Butterflies of America [BOA]
Chlosyne erodyle, NSG Voucher Specimen [Wahlberg]
Synchloe erodyle ; Godman & Salvin, [1882], Biol. centr.-amer., Lep. Rhop. 1: 180, 3: pl. 20, f. 3-4; Godman & Salvin, [1901], Biol. centr.-amer., Lep. Rhop. 2: 672
Chlosyne erodyle ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 21; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3) : 167 (name); [NACL, #4502]; [BCR, 201]; [BOW: pl. 34, f. 16]; [NL4A, #2077]
Guatemala Synchloe erodyle Doubleday, [1845]; (nom. nud.)
= ; [NL4A, #2077a]
Chlosyne erodyle erodyle ; [NL4A, #2077a]
C. e. poecile (C. & R. Felder, 1867)
Costa Rica - Colombia, Venezuela Synchloë poecile C. & R. Felder, [1867]; Reise Fregatte Novara, Bd 2 (Abth. 2) (3): 396, TL: Colombia
Synchloe poecile C. & R. Felder, 1867
Chlosyne poecile, NSG Voucher Specimen [Wahlberg]
Synchloe poecile ; Godman & Salvin, [1882], Biol. centr.-amer., Lep. Rhop. 1: 180, 3: pl. 20, f. 1-2; Godman & Salvin, [1901], Biol. centr.-amer., Lep. Rhop. 2: 672
Chlosyne poecile ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3) : 167 (name); [BCR, 201]
Chlosyne erodyle poecile ; [NL4A, #2077b]
C. e. rubrigutta Röber, [1914]
Colombia erodyle rubrigutta Röber, [1914]; , TL: Colombia
Chlosyne erodyle rubrigutta ; [NL4A, #2077c]
Venezuela Chlosyne erodyle ssp. ; Lamas, MS, [NL4A, #2077d]

Chlosyne marina (Geyer, 1837)
-. See [About maps] Araschnia marina Geyer, 1837; in Hübner, Zuträge. Samml. exot. Schmett. 5 : 20, : f. 877-878, TL: Mexico
Chlosyne marina, Listing of Mexican Butterflies [ILMB]
Chlosyne marina, Butterflies of America [BOA]
Chlosyne marina, NSG Voucher Specimen [Wahlberg]
Chlosyne marina, Butterflies and Moths of North America [BAMONA]
Synchloe marina ; Godman & Salvin, [1882], Biol. centr.-amer., Lep. Rhop. 1: 181; Godman & Salvin, [1901], Biol. centr.-amer., Lep. Rhop. 2: 673
Chlosyne marina ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3) : 167 (name); [BOW: pl. 34, f. 21]; [NL4A, #2087]
Mexico Synchloe hylaeus Godman & Salvin, 1894
Synchloë hylaeus Godman & Salvin, 1894; Ann. Mag. nat. Hist. (6) 14 (80) : 97, TL: Mexico, Durango
= ; [NL4A, #2087a]
Synchloe hylaeus ; Godman & Salvin, [1901], Biol. centr.-amer., Lep. Rhop. 2: 673
Chlosyne marina marina ; [NL4A, #2087a]
C. m. melitaeoides (C. & R. Felder, [1867])
Mexico Synchloe melitaeoides C. & R. Felder, [1867]; Reise Fregatte Novara, Bd 2 (Abth. 2) (3): 396, TL: Mexico
Chlosyne melitaeoides, Listing of Mexican Butterflies [ILMB]
Chlosyne melitaeoides, Butterflies of America [BOA]
Chlosyne melitaeoides, NSG Voucher Specimen [Wahlberg]
Synchloe melitaeoides ; Godman & Salvin, [1882], Biol. centr.-amer., Lep. Rhop. 1: 181
Chlosyne melitaeoides ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3) : 167 (name); [Opler]
Chlosyne marina melitaeoides ; [NL4A, #2087c]
C. m. eumeda (Godman & Salvin, 1894)
Mexico Synchloë eumeda Godman & Salvin, 1894; Ann. Mag. nat. Hist. (6) 14 (80) : 96, TL: Mexico, Guerrero
Synchloë dryope Godman & Salvin, 1894; Ann. Mag. nat. Hist. (6) 14 (80) : 96, TL: Mexico, Jalisco
= ; [NL4A, #2087b]
Chlosyne fasciata Röber, [1914]; , TL: Mexico, Guerrero
= ; [NL4A, #2087b]
Chlosyne eumeda, Listing of Mexican Butterflies [ILMB]
Chlosyne eumeda, Butterflies of America [BOA]
Chlosyne eumeda, NSG Voucher Specimen [Wahlberg]
Synchloe eumeda ; Godman & Salvin, [1901], Biol. centr.-amer., Lep. Rhop. 2: 672, 3: pl. 108, f. 1-2
Synchloe dryope ; Godman & Salvin, [1901], Biol. centr.-amer., Lep. Rhop. 2: 672, 3: pl. 108, f. 3-4
Chlosyne eumeda ; [EBW]; [BOW: pl. 34, f. 20]; [Opler]
Chlosyne marina eumeda ; [NL4A, #2087b]

Chlosyne endeis (Godman & Salvin, 1894)
-. See [About maps] Synchloe endeis Godman & Salvin, 1894
Synchloë endeis Godman & Salvin, 1894; Ann. Mag. nat. Hist. (6) 14 (80) : 97, TL: Mexico, Nayarit
Chlosyne endeis, NSG Voucher Specimen [Wahlberg]
Chlosyne endeis, Butterflies and Moths of North America [BAMONA]
Synchloe endeis ; Godman & Salvin, [1901], Biol. centr.-amer., Lep. Rhop. 2: 673, 3: pl. 108, f. 5-6
Chlosyne endeis ; [NACL, #4501]; [Opler]; [NL4A, #2076]
Mexico Chlosyne endeis endeis ; [NL4A, #2076a]
C. e. pardelina Higgins, 1960
Mexico endeis pardelina Higgins, 1960; , TL: Mexico, Coahuila
Chlosyne endeis pardelina, Listing of Mexican Butterflies [ILMB]
Chlosyne endeis pardelina, Butterflies of America [BOA]
Chlosyne endeis pardelina ; [NL4A, #2076b]

Chlosyne definita (Aaron, [1885])
Texas. See [About maps] Definite Patch
Melitaea definita Aaron, [1885]; Papilio 4 (9/10) : 176, TL: Texas
Chlosyne definita definita, Listing of Mexican Butterflies [ILMB]
Chlosyne definita definita, Butterflies of America [BOA]
Chlosyne definita, NSG Voucher Specimen [Wahlberg]
Chlosyne definita, Butterflies and Moths of North America [BAMONA]
Thessalia definita ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 19
Chlosyne definita ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3) : 167 (name); [NACL, #4500]; [Opler]; [NL4A, #2074]

Larva on Stenardrium barbatum [NAB]

Texas, Mexico Melitaea albiplaga Aaron, [1885]; Papilio 4 (9/10) : 175, TL: Texas
= ; [NL4A, #2074a]
Melitaea schausi Godman, [1901]; in Godman & Salvin, Biol. centr.-amer., Lep. Rhop. 2: 676, 3: pl. 108, f. 11-12, TL: Mexico, Paso de San Juan, Veracruz
= ; [NL4A, #2074a]
Cinclidia albiplaga ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 19
Chlosyne definita definita ; [NL4A, #2074a]
C. d. anastasia (Hemming, 1934)
Mexico definita anastasia (Hemming, 1934) (Melitaea); (repl. name), TL: Mexico, Durango
Melitaea beckeri Godman, [1901]; in Godman & Salvin, Biol. centr.-amer., Lep. Rhop. 2: 676, 3: pl. 108, f. 9-10 (preocc. Herrich-Schäffer, 1851), TL: Mexico, Durango
= ; [NL4A, #2074b]
Chlosyne definita anastasia, Listing of Mexican Butterflies [ILMB]
Chlosyne definita anastasia, Butterflies of America [BOA]
Chlosyne definita anastasia ; [NL4A, #2074b]

The ? species-group

Chlosyne kendallorum Opler, 1999
Mexico. See [About maps] Chlosyne kendallorum Opler, 1999; , TL: Mexico, Nuevo León
Chlosyne kendallorum, Listing of Mexican Butterflies [ILMB]
Chlosyne kendallorum, Butterflies of America [BOA]
Chlosyne kendallorum, NSG Voucher Specimen [Wahlberg]
Chlosyne kendallorum ; [NL4A, #2084]

The "Charidryas" species-group

Chlosyne palla (Boisduval, 1852)
British Columbia - California. See [About maps] Northern Checkerspot
Melitaea palla Boisduval, 1852; Ann. Soc. ent. Fr. (2) 10 : 305
eremita Wright, 1905²; (female)
wardi Oberthür, 1913²
blackmorei Gunder, 1926²
stygiana Comstock, 1926²
hemifusa Gunder, 1930²
Chlosyne palla, E.H. Strickland Ent. Mus, U of [ALBERTA]
Chlosyne palla, NSG Voucher Specimen [Wahlberg]
Chlosyne palla, Butterflies and Moths of North America [BAMONA]
Lemonias palla ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 18
Charidryas palla ; [NACL, #4492]
Chlosyne palla ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3) : 167 (name); [EBW]; [Opler]

Larva on Aster conspicuus, A. occidentalis, Chrysothamnus nauseosus, C. viscidiflorus, Chrysothamnus paniculatus, Erigeron speciosus, Solidago californica [NAB]

C. p. vallismortis F. Johnson, 1938²
palla vallismortis F. Johnson, 1938²
Charidryas palla vallismortis ; [NACL, #4492b]
C. p. flavula Barnes & McDunnough, 1918²
palla flavula Barnes & McDunnough, 1918²
Charidryas palla flavula ; [NACL, #4492c]
C. p. calydon Strecker, 1878²
palla calydon Strecker, 1878²
Charidryas palla calydon ; [NACL, #4492d]
C. p. sterope (Edwards, 1870)
Oregon Melitaea sterope Edwards, 1870; Trans. amer. ent. Soc. 3 : 190
hewesi Leussler, 1931²
hopfingeri Gunder, 1934²
dorothyi Bauer, 1975²
Lemonias sterope ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 18
Charidryas palla sterope ; [NACL, #4492e]
C. p. australomontana Emmel & Mattoon, 1998²
California Chlosyne palla australomontana, Butterflies of America [BOA]

Chlosyne whitneyi (Behr, 1863)
California - Nevada. See [About maps] whitneyi (Behr, 1863); Proc. Calif. Acad. Sci. 3 (1) : 88
vanduzeei Gunder, 1928²
Chlosyne whitneyi, Butterflies of America [BOA]
Chlosyne whitneyi, NSG Voucher Specimen [Wahlberg]
Chlosyne whitneyi, Butterflies and Moths of North America [BAMONA]
Lemonias whitneyi ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 18
Charidryas palla whitneyi ; [NACL, #4492a]
Chlosyne whitneyi ; [Opler]
C. w. damoetas (Skinner, 1902)
Rockslide Checkerspot
Melitaea damoetas Skinner, 1902; Ent. News 13 (10) : 304
damoetella McDunnough, 1927²
Chlosyne damaetas[sic, recte damoetas] ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3) : 167 (name)
Charidryas damoetas ; [NACL, #4496]

Larva on Erigeron leiomeris, Solidago multiradiata [NAB]

C. w. malcolmi Comstock, 1926²
whitneyi malcolmi Comstock, 1926²
malcolmi Comstock, 1926²
Chlosyne malcolmi ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3) : 167 (name)
Charidryas damoetas malcolmi ; [NACL, #4496a]

Chlosyne acastus (Edwards, 1874)
Utah, Nevada, Montana. See [About maps] Sagebrush Checkerspot
Melitaea acastus Edwards, 1874; Trans. amer. ent. Soc. 5 (1): 16
pearlae Gunder, 1926²
Chlosyne acastus, E.H. Strickland Ent. Mus, U of [ALBERTA]
Chlosyne acastus, NSG Voucher Specimen [Wahlberg]
Chlosyne acastus, Butterflies and Moths of North America [BAMONA]
Lemonias acastus ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 18
Charidryas acastus ; [NACL, #4493]
Chlosyne acastus ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3) : 167 (name); [BOW: pl. 13, f. 9]; [Opler]; [NL4A, #2071]

Larva on Chrysothamnus viscidiflorus, Machaeranthera canescens?, Machaeranthera viscosa? [NAB]

C. a. neumoegeni (Skinner, 1895)
Utah, California Desert Checkerspot
acastus neumoegeni (Skinner, 1895) (Melitaea); Entomol. News 6 (4) : 113, TL: Utah
Melitaea neumoegeni tr.f. friday Gunder, 1932; , TL: California
= ; [NL4A, #2071a]
boharti Gunder, 1933²
Chlosyne acastus neumoegeni; #
Chlosyne acastus neumoegeni, Butterflies of America [BOA]
Chlosyne acastus neumoegeni ; [NL4A, #2071a]
Lemonias neumoegeni ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 18
Chlosyne neumeogeni[sic] ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3) : 167 (name, missp.)
Charidryas neumoegeni ; [NACL, #4494]

Larva on Acamptopappus sphaerocephalus, Machaeranthera tortifolia [NAB]

C. a. sabina (Wright, 1905)
Arizona acastus sabina (Wright, 1905) (Melitaea); , TL: Arizona
Chlosyne acastus sabina, Listing of Mexican Butterflies [ILMB]
Chlosyne acastus sabina, Butterflies of America [BOA]
Charidryas neumoegeni sabina ; [NACL, #4494a]
Chlosyne acastus sabina ; [NL4A, #2071b]

Chlosyne gabbii (Behr, 1863)
W.USA. See [About maps] Gabb's Checkerspot
Melitaea gabbii Behr, 1863; Proc. Calif. Acad. Sci. 3 (1) : 89, TL: California
Chlosyne gabbii gabbii, Listing of Mexican Butterflies [ILMB]
Chlosyne gabbii gabbii, Butterflies of America [BOA]
Chlosyne gabbii, NSG Voucher Specimen [Wahlberg]
Chlosyne gabbii, Butterflies and Moths of North America [BAMONA]
Lemonias gabbii ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 18
Charidryas gabbii ; [NACL, #4495]
Chlosyne gabbii ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3) : 167 (name); [EBW]; [BOW: pl. 13, f. 10]; [Opler]; [NL4A, #2079]

Larva on Heterotheca grandiflora, Haplopappus squarrosus [NAB]

California Melitaea pola Boisduval, 1869; , TL: [California]
= ; [NL4A, #2079a]
Melitaea sonorae Boisduval, 1869; , TL: [California]
= ; [NL4A, #2079a]
ab. pasadenae (Gunder, 1924) (Euphydryas); , TL: California
= ; [NL4A, #2079a]
ab. gunderi (Comstock, 1926) (Melitaea); , TL: California
= ; [NL4A, #2079a]
ab. newcombi (Comstock, 1926) (Melitaea); , TL: California
= ; [NL4A, #2079a]
Chlosyne gabbii gabbii ; [NL4A, #2079a]
C. g. atrifasciata Emmel & Mattoon, 1998²
gabbii atrifasciata Emmel & Mattoon, 1998²; , TL: Santa Cruz, California

Chlosyne hoffmanni (Behr, 1863)
W.USA. See [About maps] Aster Checkerspot
Melitaea hoffmanni Behr, 1863; Proc. Calif. Acad. Sci. 3 (1) : 89
Melitaea helicta Boisduval, 1869; Ann. Soc. ent. Belg. 12 : 55
abnorma Wright, 1905²
hollandae Gunder, 1928²
Chlosyne hoffmanni hoffmanni, Butterflies of America [BOA]
Chlosyne hoffmanni, NSG Voucher Specimen [Wahlberg]
Chlosyne hoffmanni, Butterflies and Moths of North America [BAMONA]
Lemonias hoffmanni ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 18
Lemonias hoffmanni helcita[sic] ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 18
Charidryas hoffmanni ; [NACL, #4497]
Chlosyne hoffmanni ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3) : 167 (name); [BOW: pl. 13, f. 12]; [Opler]

Larva on Aster conspicuus, Chrysoptis breweri [NAB]

C. h. segregata Barnes & McDunnough, 1918²
hoffmanni segregata Barnes & McDunnough, 1918²
bridgei Comstock, 1924²
Charidryas hoffmanni segregata ; [NACL, #4497a]
C. h. manchada Bauer, [1960]
Washington Chlosyne hoffmanni manchada Bauer, [1960]; JLS#b: 208, TL: 1600ft, Tumwater Canyon, Drury, Chelan Co., Washington
Charidryas hoffmanni manchada ; [NACL, #4497b]

The "harrisii" species-group

Chlosyne harrisii (Scudder, 1863)
NE.USA, Illinois. See [About maps] Harris' Checkerspot
Melitaea harrisii Scudder, 1863; Proc. Essex Inst. 3: 167
albimontana Avinoff, 1930²
Chlosyne harrisii harrisii, Butterflies of America [BOA]
Chlosyne harrisii, NSG Voucher Specimen [Wahlberg]
Chlosyne harrisii, Butterflies and Moths of North America [BAMONA]
Charidryas harrisii ; [NACL, 61]
Cinclidia harrisii ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 18
Chlosyne harrisii ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3) : 167 (name); [BOW: pl. 13, f. 11]; [Opler]

Larva on Aster umbellatus, less Verbesina helianthoides [NAB]

C. h. hanhami (Fletcher, 1904)
Hanham's Crescent
Phyciodes hanhami Fletcher, 1904; Can. Ent. 36 (5) : 122
Charidryas harrisii hanhami ; [NACL, 61]
C. h. liggetti Avinoff, 1930²
harrisii liggetti Avinoff, 1930²
Charidryas harrisii liggetti ; [NACL, 61]

Chlosyne gorgone (Hübner, [1810])
-. See [About maps] Gorgone Crescentspot
Dryas gorgone Hübner, [1810]; Samml. exot. Schmett. 1 : pl. 41
Chlosyne gorgone, E.H. Strickland Ent. Mus, U of [ALBERTA]
Chlosyne gorgone gorgone, Butterflies of America [BOA]
Chlosyne gorgone, NSG Voucher Specimen [Wahlberg]
Chlosyne gorgone, Butterflies and Moths of North America [BAMONA]
Chlosyne gorgone, Georgia Lepidoptera (USA) [GALEPS]
Chlosyne gorgone ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3) : 166 (name); [Opler]
Charidryas gorgone ; [NACL, #4489]
Chlosyne gorgone ; [NL4A, #2081]

Larva on Helianthus, Ambrosia trifida, Iva xanthifolia, Viguiera multiflora [NAB]

South Carolina, NA.Georgia, ... Melitaea ismeria Boisduval & Le Conte, [1835]; Hist. Lép. Am. Sept. (11-22): 168, pl. 46, f. 1, 7, 9, 12, 16, TL: South Carolina; Georgia
= Chlosyne gorgone ; Calhoun, 2003, J. Lep. Soc. 57 (3): 204-209
= ; [NL4A, #2081a]
Charidryas ismeria ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 20
Chlosyne gorgone gorgone ; [NL4A, #2081a]
C. g. carlota (Reakirt, 1866)
Eresia carlota Reakirt, 1866
nigra Cary, 1901²
nox Gunder, 1928²
Charidryas gorgone carlota ; [NACL, #4489a]

Larva on Lippia lanceolata, L. nodiflora, Lysimachia sp., Eriogonum sp., Helianthus annuus, Linaria vulgaris, Ambrosia trifida Iftner, 1983, J. Lep. Soc. 37 (1): 80-81


Chlosyne nycteis (Doubleday, [1847])
E.USA. See [About maps] Silvery Crescentspot
Melitaea nycteis Doubleday, [1847]; Gen. diurn. Lep. (1): 181, (1): pl. 23, f. 3, TL: USA
?Charidryas harrisii Skinner, 1904
Chlosyne nycteis; [BOW: pl. 13, f. 15-16]
Chlosyne nycteis nycteis, Butterflies of America [BOA]
Chlosyne nycteis, NSG Voucher Specimen [Wahlberg]
Chlosyne nycteis, Butterflies and Moths of North America [BAMONA]
Charidryas nycteis ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 20; [NACL, #4490]
Chlosyne nycteis ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3) : 166 (name); [Opler]; [NL4A, #2090]
[nycteis-1u.jpg] 1025x700 (~167Kb) upperside USA: Alabama, 12.5.2001, Photo © Vitaly Charny [nycteis-1v.jpg] 719x906 (~125Kb) underside USA: Alabama, 12.5.2001, Photo © Vitaly Charny [nycteis-2u.jpg] 1332x919 (~183Kb) upperside USA: Alabama, 26.5.2001, Photo © Vitaly Charny
[nycteis-2v.jpg] 742x919 (~86Kb) underside USA: Alabama, 26.5.2001, Photo © Vitaly Charny [nycteis-3u.jpg] 1200x782 (~120Kb) upperside USA: Alabama, 12.6.2001, Photo © Vitaly Charny [nycteis-4a.jpg] 876x1158 (~90Kb) USA: Alabama, 29.4.2007, Photo © Vitaly Charny
[nycteis-4b.jpg] 1267x878 (~127Kb) USA: Alabama, 29.4.2007, Photo © Vitaly Charny

Larva on Rudbeckia laciniata, Actinomeris alternifolia, Aster, Helianthus, Verbesina helianthoides, V. virginica [NAB]


Massachusetts, Maine, New Jersey, Missouri, Kansas, ... Melitaea oenone Scudder, 1863; , TL: (Massachusetts; Maine
= ; [NL4A, #2090a]
ab. millburni (Rummel, 1926) (Phyciodes); , TL: New Jersey
= ; [NL4A, #2090a]
tr.f. lacteus (Gunder, 1928) (Phyciodes); , TL: Missouri
= ; [NL4A, #2090a]
tr.f. greyi (Field, 1934) (Phyciodes); , TL: Kansas
= ; [NL4A, #2090a]
Chlosyne nycteis nycteis ; [NL4A, #2090a]
C. n. drusius Edwards, 1884²
nycteis drusius Edwards, 1884²; Papilio 4 (3) : 57
hewitsoni Field, 1936²
Chlosyne nycteis drusius, Butterflies of America [BOA]
Charidryas nycteis drusius ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 20; [NACL, #4490a]
C. n. reversa F. & R. Chermock, 1940²
nycteis reversa F. & R. Chermock, 1940²
Charidryas nycteis reversa ; [NACL, #4491b]

The "Thessalia" species-group

Chlosyne leanira (C. & R. Felder, 1860)
Mexico, California. See [About maps] Leanira Checkerspot
Melitaea leanira C. & R. Felder, 1860; Wien. ent. Monats. 4 (4): 106, TL: California
Thessalia daviesi Wind, 1947
Chlosyne leanira leanira, Butterflies of America [BOA]
Chlosyne leanira, NSG Voucher Specimen [Wahlberg]
Chlosyne leanira, Butterflies and Moths of North America [BAMONA]
Thessalia leanira ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 19; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3) : 167 (name); [NACL, #4505]
Phyciodes leanira ; [BOW: pl. 15, f. 1]
Chlosyne leanira ; [Opler]; [NL4A, #2086]

Larva on Castilleja, Cordylanthus pilosus [NAB]

C. l. oregonensis (Bauer, 1975)
Thessalia leanira oregonensis Bauer, 1975
Melitaea leanira var. obsoleta H. Edwards, 1877; Proc. Calif. Acad. Sci. 7 : 171
obliterata Strecker, 1878²; (misspell.)
Chlosyne leanira oregonensis, Butterflies of America [BOA]
Thessalia leanira obsoleta ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 19
Thessalia leanira oregonensis ; [NACL, #4505a]
C. l. wrightii (Edwards, 1886)
California leanira wrightii (Edwards, 1886) (Melitaea); Can. Ent. 18 (4) : 64, TL: California
Melitaea cerrita Wright, 1905; , TL: California
= ; [NL4A, #2086b]
Melitaea wrightii ab. carlynae Gunder, 1926; , TL: California
= ; [NL4A, #2086b]
Melitaea wrightii f. pelona Gunder, 1930; , TL: California
= ; [NL4A, #2086b]
leona Wright, 1905²
Thessalia wrighti; [BOW: pl. 16, f. 13, 15]
Chlosyne leanira wrightii, Listing of Mexican Butterflies [ILMB]
Thessalia wrighti ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 19
Thessalia leanira wrighti ; [NACL, #4505b]
Chlosyne leanira wrightii ; [NL4A, #2086b]
C. l. cerrita Wright, 1905²
leanira cerrita Wright, 1905²
Thessalia leanira cerrita ; [NACL, #4505c]
C. l. alma Strecker, [1878]²
Utah, Arizona leanira alma Strecker, [1878]²; Lep. Rhopal. Het. (15): 135
koebelei Gunder, 1927²
Chlosyne leanira alma, Butterflies of America [BOA]
Thessalia alma ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 19
Thessalia leanira alma ; [NACL, #4505d]
Thessalia alma ; [BOW: pl. 16, f. 18]
C. l. austrima (Austin & Smith, 1998)
Mexico(Baja California) Thessalia leanira austrima Austin & Smith, 1998; , TL: Mexico, Baja California (Norte)
Chlosyne leanira austrima, Listing of Mexican Butterflies [ILMB]
Chlosyne leanira austrima, Butterflies of America [BOA]
Chlosyne leanira austrima ; [NL4A, #2086a]
C. l. elegans Preistaf & Emmel, 1998²
California Chlosyne leanira elegans, Butterflies of America [BOA]

Chlosyne cyneas (Godman & Salvin, 1878)
-. See [About maps] Cyneas Checkerspot
Phyciodes cyneas Godman & Salvin, 1878; Proc. zool. Soc. Lond. 1878 (1) : 269, TL: Mexico, Oaxaca
Chlosyne cyneas, Listing of Mexican Butterflies [ILMB]
Chlosyne cyneas, Butterflies of America [BOA]
Chlosyne cyneas, NSG Voucher Specimen [Wahlberg]
Chlosyne cyneas, Butterflies and Moths of North America [BAMONA]
Thessalia cyneas, Butterflies of Southeastern Arizon [Bruce Walsh]
Phyciodes cyneas ; Godman & Salvin, [1882], Biol. centr.-amer., Lep. Rhop. 1: 191, 3: pl. 21, f. 10-11; Godman & Salvin, [1901], Biol. centr.-amer., Lep. Rhop. 2: 677
Thessalia cyneas ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 19; [NACL, #4507]; [BOW: pl. 16, f. 14]
Chlosyne cyneas ; [Opler]; [NL4A, #2073]

Larva on Seymeria tenuisecta [NAB]

Arizona, Mexico tr.f. infrequens (Gunder, 1928) (Melitaea); , TL: Arizona
= ; [NL4A, #2073a]
Chlosyne cyneas cyneas ; [NL4A, #2073a]
C. c. cynisca (Godman & Salvin, 1882)
Mexico Phyciodes cynisca Godman & Salvin, [1882]; Biol. centr.-amer., Lep. Rhop. 1: 191, 3: pl. 21, f. 12, TL: Mexico, Orizaba; Jalapa
Thessalia cynisca ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3) : 167 (name)
Chlosyne cyneas cynisca ; [NL4A, #2073b]

Chlosyne theona (Ménétriés, 1855)
-. See [About maps] Theona Checkerspot
Melitaea theona Ménétriés, 1855; Cat. lep. Petersb. 2: 86, TL: Nicaragua
Chlosyne theona, NSG Voucher Specimen [Wahlberg]
Chlosyne theona, Butterflies and Moths of North America [BAMONA]
Phyciodes theona ; Godman & Salvin, [1882], Biol. centr.-amer., Lep. Rhop. 1: 192; Godman & Salvin, [1901], Biol. centr.-amer., Lep. Rhop. 2: 677
Thessalia theona ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 19; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3) : 167 (name); [NACL, #4508]; [BCR, 202]
Chlosyne theona ; [Opler]; [NL4A, #2092]
[theona-1.jpg] 800x877 (~98Kb) USA: on road to Miller Canyon, Sierra Vista, Cochise Co., Arizona, 8.8.2005, Photo © Markku Savela

Larva on Castilleja lanata, Leucophyllum frutescens, Verbena [NAB], Castilleja [BCR]


Nicaragua Chlosyne theona theona ; [NL4A, #2092a]
C. t. perlula (C. & R. Felder, 1861)
Venezuela theona perlula (C. & R. Felder, 1861) (Eresia); , TL: Venezuela
Chlosyne theona perlula ; [NL4A, #2092j]
C. t. ezra (Hewitson, 1864)
Costa Rica, Panama, Honduras Eresia ezra Hewitson, 1864; Ill. exot. Butts [2] (Eresia III-VI): [66], pl. [34], f. 29
Eresia yorita Reakirt, [1866]; Proc. Ent. Soc. Philad. 5 (2) : 224, TL: Honduras
= ; [NL4A, #2092f]
Phyciodes ezra ; Godman & Salvin, [1882], Biol. centr.-amer., Lep. Rhop. 1: 192; Godman & Salvin, [1901], Biol. centr.-amer., Lep. Rhop. 2: 677
Thessalia ezra ; [BCR, 202]
Chlosyne theona ezra ; [NL4A, #2092f]
C. t. thekla (Edwards, 1870)
California - Texas, Arizona Melitaea thekla Edwards, 1870; Trans. amer. ent. Soc. 3 : 191, TL: Arizona
tr.f. benajmini (Gunder, 1928) (Melitaea); , TL: Arizona
= ; [NL4A, #2092k]
Chlosyne theona thekla, Listing of Mexican Butterflies [ILMB]
Chlosyne theona thekla, Butterflies of America [BOA]
Thessalia theona thekla, Butterflies of Southeastern Arizon [Bruce Walsh]
Thessalia thekla ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 19
Thessalia theona thekla ; [NACL, #4508a]
Chlosyne theona thekla ; [NL4A, #2092k]
C. t. bollii (Edwards, [1878])
Texas, Arizona theona bollii (Edwards, [1878]) (Melitaea); , TL: Texas
Chlosyne theona bollii, Listing of Mexican Butterflies [ILMB]
Chlosyne theona bollii, Butterflies of America [BOA]
Thessalia bollii ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 19
Thessalia theona bollii ; [NACL, #4508b]
Chlosyne theona bollii ; [NL4A, #2092b]
C. t. hondana (Weymer, 1890)
Colombia Phyciodes hondana Weymer, 1890; in Weymer & Maassen, Reisen Süd-America : 119, pl. 3, f. 5, TL: Colombia
Chlosyne theona hondana ; [NL4A, #2092g]
C. t. chinatiensis (Tinkham, 1944)
Texas Chinati Checkerspot
Melitaea chinatiensis Tinkham, 1944; Can. Ent. 76: 12, TL: Texas
Chlosyne theona chinatiensis, Listing of Mexican Butterflies [ILMB]
Chlosyne theona chinatiensis, Butterflies of America [BOA]
Thessalia chinatiensis ; [NACL, #4509]
Chlosyne theona chinatiensis ; [NL4A, #2092d]

Larva on Leucophyllum minus [NAB]

C. t. brocki (Austin & Smith, 1998)
Mexico Thessalia theona brocki Austin & Smith, 1998; , TL: Mexico, Sonora
Chlosyne theona brocki, Listing of Mexican Butterflies [ILMB]
Chlosyne theona brocki, Butterflies of America [BOA]
Chlosyne theona brocki ; [NL4A, #2092c]
C. t. costaricensis (Austin & Miller, 1998)
Costa Rica Thessalia theona costaricensis Austin & Miller, 1998; , TL: Costa Rica
Chlosyne theona costaricensis ; [NL4A, #2092e]
C. t. minimus (Austin & Smith, 1998)
Guatemala Thessalia theona minimus Austin & Smith, 1998; , TL: Guaetemala
Chlosyne theona minimus ; [NL4A, #2092h]
C. t. mullinsi (Austin & Smith, 1998)
Mexico Thessalia theona mullinsi Austin & Smith, 1998; , TL: Mexico, Oaxaca
Chlosyne theona mullinsi, Listing of Mexican Butterflies [ILMB]
Chlosyne theona mullinsi, Butterflies of America [BOA]
Chlosyne theona mullinsi ; [NL4A, #2092i]

The ? species-group

Chlosyne fulvia (Edwards, 1879)
-. See [About maps] Fulvia Checkerspot
Melitaea fulvia Edwards, 1879; Can. Ent. 11 (6) : 117, TL: Texas
sinefascia R. C. Williams, 1914²
Chlosyne fulvia fulvia, Listing of Mexican Butterflies [ILMB]
Chlosyne fulvia fulvia, Butterflies of America [BOA]
Chlosyne fulvia, NSG Voucher Specimen [Wahlberg]
Thessalia fulvia ; [NACL, #4506]
Chlosyne fulvia ; [Opler]; [NL4A, #2078]

Larva on Castilleja integra, C. lanata [NAB]

Texas Chlosyne fulvia fulvia ; [NL4A, #2078a]
C. f. coronado (Smith & Brock, 1988)
Arizona fulvia coronado (Smith & Brock, 1988) (Thessalia); , TL: Arizona
Chlosyne fulvia coronado, Listing of Mexican Butterflies [ILMB]
Chlosyne fulvia coronado, Butterflies of America [BOA]
Chlosyne fulvia coronado ; [NL4A, #2078b]


Unknown or unplaced taxa

Synchloe analis Herrich-Schäffer, 1865; (nom. nud.)
analis ; [NL4A, 253]

Synchloe dromippe Herrich-Schäffer, 1865; (nom. nud.)
dromippe ; [NL4A, 254]

Synchloe najas Herrich-Schäffer, 1865; (nom. nud.)
najas ; [NL4A, 254]

Synchloe tegea Boisduval, 1870; (nom. nud.)
tegea ; [NL4A, 254]

Synchloe tyrinthe Doubleday, [1845]; (nom. nud.), TL: Mexico
tyrinthe ; [NL4A, 254]


22.2.2013 (26)

References:
[ALBERTA]
University of Alberta, E.H. Strickland Entomological Museum; Browse Entomology Collection
[About maps]
Warning! The maps are automatically generated from the textual information, and the process does not always produce acceptable result; See about maps for more info.
[BAMONA]
Opler, Pavulaan, Stanford & Pogue; Butterflies and Moths of North America; Mountain Prairie Information Node
[BCA]
electronic; BIOLOGIA CENTRALI-AMERICANA
[BCR]
DeVries; The Butterflies of Costa Rica 1; Papilionidae, Pieridae, Nymphalidae
[BHL]
The; Biodiversity Heritage Library; and on; Internet Archive; Note: The linking to the content of Internet Archive is somewhat delicate -- notify me when links break!
[BOA]
Interactive Listing of American Butterflies; Listado Interactivo de la Mariposas Americanas; Butterflies of America
[BOW]
Lewis, 1974; Butterflies of the World; ISBN 0 245 52097 X
[Bruce Walsh]
Bruce Walsh; e-mail: Bruce Walsh; The Moths of Southeastern Arizona; Butterflies of Southeastern Arizona
[EBW]
Smart, Paul, 1976; The Illustrated Encyclopedia of the Butterfly World; ISBN 0 600 31381 6
[EcuL]
Tommaso & Racheli, 2001; Fragmenta entomologica 33 (2): 213-380; An annotated list of Ecuadorian Butterflies (Lepidoptera: Papilionidae, Pieridae, Nymphalidae)
[GALEPS]
Moths and Butterflies of Georgia and the Southeastern United States; James K. Adams; Dept. of Natural Science, Dalton State College, Dalton, GA ; http://www.daltonstate.edu/galeps/index.htm
[HBN]
Francis Hemming, 1937; HÜBNER, A bibliographical and systematic account of the entomological works of Jacob Hübner and of the supplements thereto by Carl Geyer, Gottfried Franz von Frölich and Gottlieb August Wilhelm Herrich-Schäffer, Vol 1 & 2
[ILMB]
Interactive Listing of Mexican Butterflies
[JLS]
Journal of the Lepidopterists' Society; the original J. Lep. Soc. articles
[John Shuey]
e-mail: jshuey at tnc.org; Director of Conservation Science; Indiana Office of The Nature Conservancy; http://www.nature.org
[LAMAS-NEO]
Gerardo Lamas; Bibliography of butterflies; An Annocated Bibliography of the Neotropical Butterflies and Skipper (Lepididoptera: Papilionoidea and Hesperioidea); Revised Electronic Edition (2007)
[NAB]
Pyle, 1981; Field Guide to North American Butterflies, National Audubon Society; ISBN 0-394-51914-0
[NACL]
Hodges, 1983; Check List of the Lepidoptera of America North of Mexico
[NL4A]
Gerardo Lamas, 2004; Atlas of Neotropical Lepidoptera; Checklist: Part 4A; Hesperioidea - Papilionoidea
[Opler]
Opler & Warren, 2003 (April); Butterflies of North America. 2. Scientific Names List for Butterfly Species of North America, north of Mexico.
[Vitaly Charny]
Birmingham, Alabama, USA
[Wahlberg]
Niklas Wahlberg; The NSG's voucher specimen database; Nymphalidae Systematics Group
[²]
This may require parentheses or not. I don't have the necessary information for this taxon.


Some related literature:
Aaron, [1885]
List of a collection of diurnal Lepidoptera from southern Texas Papilio 4 (9/10) : 172-182
Bates, 1864
New species of butterflies from Guatemala and Panama, collected by Osbert Salvin and F. du Cane Godman, Esqs. Ent. mon. Mag. 1 (1): 1-6 (1864), (2): 31-35 (1864), (3): 55-59 (1864), (4): 81-85 (1864), (5): 113-116 (1864), (6): 126-131 (1864), (7): 161-164 (1864), (8): 178-180 (1865), (9): 202-205 (1865)
Bauer, [1960]
A new species of Chlosyne (Nymphalidae) from western Mexico J. Lep. Soc. 13: 165-169
Bauer, [1961]
Descriptions of two new Chlosyne (Nymphalidae) from Mexico, with a discussion of related forms J. Lep. Soc. 14 (2): 148-154
Behr, 1863
On Californian Lepidoptera - No. III Proc. Calif. Acad. Sci. 3 (1) : 84-93
Boisduval, 1852
Lépidoptères de la Californie Ann. Soc. ent. Fr. (2) 10 : 275-324
Boisduval, 1869
Lépidoptères de la Californie Ann. Soc. ent. Belg. 12 : 5-28,37-94
Boisduval, 1870
Considerations sur des Lépidoptères envoyés du Guatemala à M. de l'Orza Considérations Lépid. Guatemala: 100pp
Boisduval & Le Conte, [1835]
Histoire générale et iconographie des lépidoptères et des chenilles de l'Amérique septentrionale Hist. Lép. Am. Sept. (1): 1-16. pl. 1-3 (1829), (2): 17-24, pl. 4-6 (1829), (3/4): 25-40, pl. 7-12 ([1830]), (5/6): 41-56, pl. 13-18 ([1830]), (7/8): 57-80, pl. 19-24 ([1830]), (9/10): 81-100, pl. 25-30 (1833), (11-22): 101-196, pl. 31-65 ([1835]), (23-26): 197-228, pl. 66-78 ([1837])
Brown & Mielke, 1967
Lepidoptera of the Central Brazil Plateau. I. Preliminary list of Rhopalocera J. Lep. Soc. 21 (2): 77-106
Calhoun, 2003
The history and true identity of Melitaea ismeria (Nymphalidae): a remarkable tale of duplication , misinterpretation, and presumption J. Lep. Soc. 57 (3): 204-219
Doubleday, 1844
List of the Specimens of Lepidopterous insects in The Collection of the British Museum List. lepid. Ins. Brit. Mus. 1 : 1-150
Doubleday, [1847]
The genera of diurnal Lepidoptera, comprising their generic characters, a notice of their habitats and transformations, and a catalogue of the species of each genus; illustrated with 86 plates by W. C. Hewitson Gen. diurn. Lep. (1): 1-18, pl. A, 1-4 (1-2) (1846), (1): 19-132, pl. 4*, 5-25, 28 (3-14) (1847), (1): 133-200, pl. 1*, 26, 27, 29, 31-44 (15-23) (1848), (1): 201-242, pl. 30, 45-52, 56-58, 60-62, 64 (24-31) (1849), (1): 243-250 [?], (2): 243-326, pl. 53-55, 63, 65, 66 (32-38) (1850), (2): 327-466, pl. 59, 67-77 (39-50) (1851), (2): 467-534, 143, 144, pl. 78-80 (Index) (51-54) (1852)
Drury, [1782]
Illustrations of natural history; wherein are exhibited ... Illust. Nat. Hist. Exot. Insects 3 : 1-76, pl. 1-50
Dyar, 1903
A List of North American Lepidoptera and Key to the Literature of this Order of Insects Bull. U.S. nat. Mus. 52 : 1-723
Edwards, 1870
Descriptions of new North American diurnal Lepidoptera Trans. amer. ent. Soc. 3 : 189-196
Edwards, 1874
Descriptions of new species of diurnal Lepidoptera found in North America Trans. amer. ent. Soc. 5 (1): 13-19, (2): 103-111
Edwards, 1879
Description of a new species of Melitaea from Texas Can. Ent. 11 (6) : 117-118
Edwards, 1884
Description. of new species of butterflies mostly from Arizona Papilio 4 (3) : 53-58
Edwards, 1886
Description of new species of butterflies found in the United States Can. Ent. 18 (4) : 61-65
H. Edwards, 1877
Pacific Coast Lepidoptera. - No. 22. Notes on some Diurnal Lepidoptera, with description of new varieties Proc. Calif. Acad. Sci. 7 : 163-173
R. Felder, 1869
Diagnosen neuer von dem K. K. Oberlieutenant H. v. Hedemann in Mexico in den Jahren 1865-1867 gesammelter Lepidopteren. Erste Folge Verh. zool.-bot. Ges. Wien. 19 : 465-480
C. & R. Felder, 1860
Lepidopterologische Fragmente. V-VI Wien. ent. Monats. 4 (4): 97-112, pl. 1-2, (8): 225-251, pl. 3-4
C. & R. Felder, [1867]
Reise der österreichischen Fregatte Novara um die Erde in den Jahren 1857, 1858, 1859 unter den Behilfen des Commodore B. von Wüllerstorf-Urbair. Zoologischer Theil. Band 2. Abtheilung 2. Lepidoptera. Rhopalocera Reise Fregatte Novara, Bd 2 (Abth. 2) (1): 1-136, pl. 1-21 (Rhop.) (C. & R. Felder, [1865]), (2): 137-378, pl. 22-47 (Rhop.) (C. & R. Felder, 1865), (3): 379-536, pl. 48-74 (Rhop.) (C. & R. Felder, 1867), (4): 537-548, pl. 75-120, (Inhalts-Verz.) 1-9 (pl. 1-74), (Felder & Rogenhofer, 1874) , (5): pl. 121-140, (erklärung) 1-10 (pl. 75-107), (Inhalts-Verz.) 1-20 (pl. 108-140), (Felder & Rogenhofer, 1875)
Fletcher, 1904
Descriptions of some new species and varieties of Canadian butterflies Can. Ent. 36 (5) : 121-129
Godman & Salvin, 1878
Descriptions of nineteen new species of Diurnal Lepidoptera from Central America Proc. zool. Soc. Lond. 1878 (1) : 264-271
Godman & Salvin, [1882]
Biologia Centrali-Americana. Rhopalocera. Vol. 1. (1879-1886) Biol. centr.-amer., Lep. Rhop. 1: 1-487, 3: pl. 1-24,24a,25-112
Godman & Salvin, [1901]; Godman in Godman & Salvin, [1901]
Biologia Centrali-Americana. Rhopalocera. (1887-1901) Biol. centr.-amer., Lep. Rhop. 2: 1-782, 3: pl. 1-112
Godman & Salvin, 1894
Descriptions of new Species of Rhopalocera from Mexico and Central America Ann. Mag. nat. Hist. (6) 14 (80) : 95-98
Hall, 1924
Descriptions of two new forms of Chlosyne (Lep. Nymphalidae) Entomologist 57: 241-242
Hewitson, 1864
Illustrations of new species of exotic Butterflies selected chiefly from the collections of W. Wilson Saunders and William C. Hewitson Ill. exot. Butts [2] : (Systematic Index) II, (Ithomia XIV): [1-2], pl. [1] (1855), (Ithomia XV-XIX): [3-12], pl. [2-6] (1856), (Ithomia XX): 13x,14x, [13-14] (1861), pl. [7] (1857), (Ithomia XXI): 15x,16x, [15-16] (1861), pl. [8] (1858), (Ithomia XXII): [17-18], pl. [9] (1859), (Ithomia XXIII): [19-20], pl. [10] (1861), (Ithomia XXIV): [21-22], pl. [11] (1863), (Ithomia XXV): [23-24], pl. [12] (1867), (Ithomia XXVI): [25-26], pl. [13] (1868), (Ithomia XXVII): [27-28], pl. [14] (1869), (Ithomia XXVIII-XXIX): [29-32], pl. [15-16] (1871), (Ithomia XXX-XXXI): [33-36], pl. [17-18] (1872), (Ithomia XXXII-XXXIII): [37-40], pl. [19-20], (1874) , (Ithomia XXXIV-XXXV): [41-44], pl. [21-22] (1876), (Acraea I): [45-46], pl. [23] (1852), (Acraea II): [47-48], pl. [24] (1861), (Acraea III): [49-50], pl. [25] (1865), (Acraea IV-V): [51-52], pl. [26-27] (1867), (Acraea VI): [53-54], pl. [28] (1868), (Acraea VII): [55-56], pl. [29] (1875), (Messaras): [57-58], pl. [30] (1859), (Eresia I): [59-60], pl. [31] (1852), (Eresia II): [61-62], pl. [32] (1857), (Eresia III-VI): [63-70], pl. [33-36] (1864), (Eresia VII): [71-72], pl. [37] (1868), (Eresia VIII-IX): [73-76], pl. [38-39] (1870), (Eresia X): [77-78], pl. [40] (1872), (Junonia): [79-80], pl. [41] (1864), (Cybdelis I-II): [81-84], pl. [42-43] (1852), (Cybdelis III): [85-86], pl. [44] (1857), (Cybdelis IV): [87-88], pl. [45] (1869), (Epiphile I): [89-90], pl. [46] (1857), (Epiphile II & Paromia): [91-92], pl. [47] (1861), (Eubagis I): [93-94], pl. [48] (1852), (Eubagis II): [95-96], pl. [49] (1857), (Eubagis III): [97-98], pl. [50] (1859), (Catagramma I): [99-100], pl. [51] (1851), (Catagramma II): [101-102], pl. [52] (1853), (Catagramma III-V): [103-108], pl. [53-55] (1855), (Catagramma VI): [109-110], pl. [56] (1857), (Catagramma VII-VIII): [111-114], pl. [57-58] (1858), (Catagramma IX and Callithea): [115-116], pl. [59] (1858), (Catagramma X): [117-118], pl. [60] (1864), (Catagramma XI-XII): [119-122], pl. [61-62] (1868), (Catagramma XIII): [123-124], pl. [63] (1876), : iv
Higgins, 1981
A revision of Phyciodes Hübner and related genera, with a review of the classification of the Melitaeinae Bull. Br. Mus. nat. Hist. (Ent.) 43 (3) : 77-243, f. 1-490,124a,125a
Hübner, [1810]
Sammlung exotischer Schmetterlinge, Vol. 1 (1806 - [1819]) Samml. exot. Schmett. 1 : [1-4] (1807), : [5-12] (1807-1809), : [13-36] (1823-1824), : pl. [1-2], [5-8], [12], [15], [29], [47] (1806), : pl. [9], [11], [17], [21], [25-27], [32], [35], [43-44], [56], [60], [79], [102], [104], [116], [141], [148], [150-151], [155], [159], [200] (1807), : pl. [19], [39], [57], [73], [82], [114], [157], [167], [170], [173], [187] (1807), : pl. [3], [31], [37], [46], [54], [58], [62], [76], [84], [92], [95], [105], [112-113], [117], [137-138], [146], [154], [158], [180], [209] (1808), : pl. [16], [28], [65], [89], [142], [160], [162], [166], [182], [188], [193] (1809), : pl. [4], [22], [42], [53], [61], [88], [90], [97], [100], [107], [110], [118], [120], [140], [143], [149], [161], [171], [178], [184], [196], [202] (1809), : pl. [10], [18], [30], [41], [51], [80], [144], [172], [189] (1810), : pl. [36], [70], [87], [91], [101], [124], [129], [177], [183], [207], [210] (1811), : pl. [20], [38], [45], [55], [85], [98], [103], [108], [136], [139], [152-153], [156], [163], [179], [186], [190], [205-206] (1812), : pl. [13-14], [40], [48], [50], [67], [74], [86], [119], [121], [130], [135], [175-176], [197-199], [204], [212] (1813), : pl. [72], [75], [77-78], [83], [93], [109], [111], [115], [128], [131], [165], [174], [194] (1813), : pl. [24], [49], [52], [63-64], [66], [68-69], [71], [81], [123], [145], [181], [192] (1814), : pl. [126], [132-134], [168] (1815), : pl. [23], [96], [122], [125], [169], [195] (1816), : pl. [33], [99], [127], [187], [185], [213] (1817), : pl. [59], [106], [147], [164], [208], [211] (1818), : pl. [34], [94], [191], [201] (1819), : (Index Systematicus) [1-4] (1819 - April 1823)
Geyer, [1833]
Sammlung exotischer Schmetterlinge, Vol. 3 ([1827] - [1838]) in Hübner, Samml. exot. Schmett. 3 : pl. [9], [11], [21], [26-27], [32], [43], [45] (1827), : pl. [7], [13], [35], [42] (1828), : pl. [20], [22], [37-38] (1829), : pl. [17], [24], [41], [49] (1830), : pl. [30] (1831), : pl. [25], [28], [31], [44], [50-51] (1832), : pl. [5-6], [14-16], [29], [48], [52], [54] (1833), : pl. [1], [12], [18] (1834), : pl. [2-4], [10], [19], [23], [39], [46] (1835), : pl. [40] (1836-1837), : pl. [8], [33-34], [36], [47], [53] (1838)
Geyer, 1837
Zuträge zur Sammlung exotischer Schmettlinge, Vol. 5 [1833]-1837 in Hübner, Zuträge. Samml. exot. Schmett. 5 : [1-3], 4, [5], 6-52 (1837), : pl. [138-151], f. 801-880 (1833), : pl. [152-172], f. 881-1000 (1834 - 1836)
Iftner, 1983
A new food plant record for Chlosyne gorgone carlota J. Lep. Soc. 37 (1): 80-81
Kirby, 1871
A synonymic catalogue of the diurnal Lepidoptera Syn. Cat. diurn. Lep.: vii + 690pp : 1-690
Latreille, [1809]
Insectes de l'Amérique équinoxiale, recueillis pendant de voyage de MM. de Humboldt et Bonpland. In Humbolt & Bonpland, aux régions équinoxiales du Nouveau Continent. 2. Recueil d'observations de zoologie et d'anatomie comparée; faites dans l'Océan Atlantique, dans l'intérieur du Nouveau Continent et dans la Mer du Sund, pendant les années 1799, 1800, 1801, 1802 et 1803 in Humboldt & Bonpland, Voy. rég. équinoxiales..., 1 (4): 197-283, pl. 15-18 (1809), (5/6): 344-397, pl. 22-25 (1809), (7): 127-252, pl. 15-18,22-25 (1811)
Ménétriés, 1855
Enumeratio corporum animalium Musei Imperialis Academiae Scientiarum Petropilitanae. Classis Insectorum, Ordo Lepidopterorum Cat. lep. Petersb. 1: 1-66, pl. 1-6, (supplement) 67-112 (1855), 2: 67-97,99-144, pl. 7-14 (1857), 3: 145-161, pl. 15-18 (1863)
Reakirt, [1866]
Descriptions of some new species of Eresia Proc. Ent. Soc. Philad. 5 (2) : 224-227
Reakirt, 1866
Descriptions of some new species of Diurnal Lepidoptera Proc. Acad. nat. Sci. Philad. 18 (3) : 238-249
Reakirt, [1867]
Descriptions of some new species of Diurnal Lepidoptera. Series II Proc. Acad. nat. Sci. Philad. 18 (4): 331-336 (1867), (5): 337-342 (1867)
Scudder, 1872
A systematic Revision of some of the American Butterflies; with brief notes on those known to occur in Essex County, Mass. 4th Ann. Rep. Peabody Acad. Sci. (1871): 24-83
Scudder, 1875
Synonymic list of the butterflies of North Amercia, north of Mexico. (1) Nymphales Bull. Buffalo Soc. nat. Sci. 2 : 233-269
Skinner, 1895
Notes on Rhopalocera, with descriptions of new species Entomol. News 6 (4) : 112-114
Skinner, 1902
A New Species of Melitaea Ent. News 13 (10) : 304-305
Staudinger, 1876
Neue Lepidopteren des südamerikanischen Faunengebietes Verh. zool.-bot. Ges. Wien, 25 (1) : 89-124
Strecker, [1878]
Lepidoptera, Rhopaloceres and Heteroceres, indigenous and exotic, with descriptions and colored illustrations Lep. Rhopal. Het. (1): 1-7, pl. 1 (1872), (2): 9-15, pl. 2 (1873), (3): 17-24, pl. 3 (1873), (4): 25-32, pl. 4 (1873), (5): 33-43, pl. 5 (1873), (6): 45-50, pl. 6 (1873), (7): 51-60, pl. 7 (1873), (8): 61-70, pl. 8 (1874), (9): 71-80, pl. 9 (1874), (10): 81-94, pl. 10 (1874), (11): 95-100, pl. 11 (1874), (12): 101-108, pl. 12 (1875), (13): 109-123, pl. 13 (1876), (14): 125-134, pl. 14 (1878), (15): 135-143, pl. 15 (1878)
Weymer in Weymer & Maassen, 1890
Lepidoptera gesammelt auf einer Reise durch Colombia, Ecuador, Peru, Brasilien, Argentinien und Bolivien in den Jahren 1868-1877 von Alphons Stübel Reisen Süd-America : 1-182, pl. 1-9
Wright, 1905
The Butterflies of the West Coast of the United States Butts. West Coast: 257pp


If you have corrections, comments or information to add into these pages, just send mail to Markku Savela
Keep in mind that the taxonomic information is copied from various sources, and may include many inaccuracies. Expert help is welcome.