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Issoria Hübner, [1819]
Issoria Hübner, [1819]; Verz. bek. Schmett. (2): 31, TS: Papilio lathonia Linnaeus
Rathora Moore, 1900; Lepidoptera Indica, 4 : 241, TS: Papilio lathonia Linnaeus
Yramea Reuss, 1920; Ent. Mitt. 9 : 192, TS: Papilio cytheris Drury
Pseudorathora Reuss, 1926; Dt. ent. Z. 1926 (1) : 68, TS: Rathora isaeae[sic] f.geogr. isaeoides Reuss
Prokükenthaliella Reuss, [1927]; Deuts. ent. Z. 1926 (5): 435, TS: Argynnis excelsior Butler
Prokuekenthaliella Reuss, [1927]; (corrected)
Chilargynnis Bryk, 1944; Ark. Zool. 36A (1): 8, TS: Papilio cytheris Drury
= Yramea ; [NL4A, 261]
Issoria, NSG Voucher Specimen [Wahlberg]
Issoria (Nymphalidae) ; [BRU2, 40]
Yramea (Argynnini) ; [Wahlberg]; [NL4A, 261]
Issoria (Argynnini) ; [Wahlberg]
Prokuekenthaliella (Argynnina) ; [AfrL]
Issoria (Argynnina) ; [AfrL]

fiIssoria lathonia (Linnaeus, 1758)
EU, NAF, Canary Islands, tr, C.Asia, Himalaya, Baluchistan, Safed Koh, Chitral - Sikkim, Chin Hills, W.China. See [About maps] Helmihopeatäplä Queen of Spain Fritillary Storfläckig pärlemorfjäril Spegelpärlefjäril Kleiner Perlmutterfalter Le Petit Nacré Sofia Dostojka latonia
Papilio lathonia Linnaeus, 1758; Syst. Nat. (Edn 10) 1 : 481, TL: [Sweden]
Papilio valdensis Esper, [1804]; Die Schmett., Suppl. Th 1 (10): 112, pl. 115, f. 4
saturata (Röber, 1896)
hungarica (Aigner-Abafi, 1906)
florens Verity, 1916
sheljuzhkoi (Stauder, 1923)
Issoria lathonia, NSG Voucher Specimen [Wahlberg]
Issoria lathonia, Larvae of North-European Lepidoptera [Kimmo Silvonen]
Issoria lathonia, Svenska fjärilar [SVF]
Issoria lathonia, Norges Sommerfugler [NORLEP]
Issoria lathonia, Moths and Butterflies of Europe [MBE]
Issoria lathonia, Europaische Schmetterlinge [Mario Maier]
Issoria lathonia, Captain's European Butterfly Guide [Simon Coombes]
Issoria lathonia ; [BOW: pl. 2, f. 19-20]; [EBW]; [MRS, 479]; [BMAT: 51, pl. 16, f. 17-21]; [BRU2: 40, pl. 28, f. 7-9]; [Otakar Kudrna]
[lathonia-1v.jpg] 500x368 (~85Kb) underside Morbihan, France 12/08/91, Photo © Simon Coombes [lathonia-2u.jpg] 672x556 (~115Kb) upperside A road on a field surrounded by birch forests at the village Klyuchi, Iskitim District, Novosibirsk Province, West Siberia, Russia. 1st October 1988, Photo © Oleg Kosterin [lathonia-3mfv.jpg] 1128x684 (~103Kb) underside male female A meadow in the Koyon River valley at the village Morozovo, Iskitim District, Novosibirsk Province, West Siberia, Russia. 25th July 1998, Photo © Oleg Kosterin
[lathonia-4l.jpg] 596x400 (~42Kb) larva Central Europe 1990-2000, Photo © Michael Borsch [lathonia-4v.jpg] 591x400 (~42Kb) underside Central Europe 1990-2000, Photo © Michael Borsch [lathonia-5u.jpg] 600x435 (~58Kb) upperside Morocco, Djebel Siroua, Anti-Atlas, Photo © Michel Tarrier
[lathonia-6.jpg] 600x450 (~50Kb) Sweden, Ekebo, Julita, 17.9.2003, Photo © Leif Wahlberg [lathonia-6u.jpg] 559x745 (~67Kb) upperside Sweden, Ekebo, Julita, 17.9.2003, Photo © Leif Wahlberg [lathonia-7v.jpg] 689x919 (~94Kb) underside Sweden, Ekebo, Julita, 8.8.2004, Photo © Leif Wahlberg
[lathonia-8v.jpg] 800x905 (~106Kb) underside Russia, Tatarstan, Varkljed-Bodja (56°13'N 52°49'E), about 20km S of Agriz, 8.8.2004, Photo © Markku Savela

Larva on Viola, V. canina [SPRK], V. odorata [BMAT], V. tricolor, V. arvensis, Anchusa, Rubus, Onobrychis [BRU2, 40]


I. l. isaaea (Gray, 1846)
Himalaya, Yunnan Argynnis isaeea Gray, 1846; Descr. lep. Ins. Nepal : 11 (nom. nud.)
Argynnis sp. ; Doubleday, [1845], List. lepid. Ins. Brit. Mus. 1 : 68
Argynnis issaea[sic, recte isaaea] ; Doubleday, [1848], Gen. diurn. Lep. (1): 174; Moore, 1857, in Horsfield & Moore, Cat. lep. Ins. Mus. East India Coy 1 : 156
Issoria lathonia isaea[sic, recte isaaea] ; [MRS, 479]; [BRU2, 40]
Issoria lathonia issaea[sic, recte isaaea] ; Huang, 2001; Huang, 2003, Neue Ent. Nachr. 55: 80 (note)
I. l. messora (Fruhstorfer, 1909)
lathonia messora (Fruhstorfer, 1909); , TL: Ta-Tsien-Lou
Issoria lathonia messora ; [BRU2, 40]
I. l. isaeoides Reuss, 1925
Szetschuan Rathora isaeae[sic] f.geogr. isaeoides Reuss, 1925; Iris 39 (4): 218
Issoria lathonia isaeoides ; [BRU2, 40]

Issoria smaragdifera (Butler, 1895)
Rhodesia, Nyasaland. See [About maps] Argynnis smaragdifera Butler, 1895; Proc. zool. Soc. Lond. 1895 : 629
Issoria smaragdifera, NSG Voucher Specimen [Wahlberg]
Issoria smaragdifera ; [BOW: pl. 104, f. 15]; [AfrL]

Larva on Viola abyssinia [PBSA]

highlands (Malawi, E.Rhodesia) Issoria smaragdifera smaragdifera ; [BAFR, 211]; [AfrL]
I. s. reducta Carcasson, 1961
highlands (S.Tanzania) Issoria smaragdifera reducta ; [BAFR, 211 (text)]; [AfrL]

Issoria hanningtoni Elwes, 1889
E.Africa. See [About maps] Hannington's Fritillary
Issoria hanningtoni Elwes, 1889; Trans. ent. Soc. Lond. 1889 : 558
Issoria hanningtoni, NSG Voucher Specimen [Wahlberg]
Issoria hanningtoni ; [EBW]; [BOW: pl. 104, f. 7]
Prokuekenthaliella hanningtoni ; [AfrL]

Larva on Viola [BK]

N.Tanzania (Mt Kilimanjaro, Mt Meru, Ngorongoro Crater) Issoria hanningtoni hanningtoni ; [BAFR, 211]
Prokuekenthaliella hanningtoni hanningtoni ; [AfrL]
I. h. jeanneli Bernardi, 1968²
Kenya highlands (Mt Kenya, Mt Elgon, Aberdare range) Issoria hanningtoni jeanneli ; [BAFR, 211 (text)]; [BK, 356, pl. 51]
Prokuekenthaliella hanningtoni jeanneli ; [AfrL]
I. h. imatonga Riley, 1932²
S.Sudan (Imatong Mts.) Issoria hanningtoni imatonga ; [BAFR, 211]
Prokuekenthaliella hanningtoni imatonga ; [AfrL]

Issoria baumanni Rebel & Rogenhofer, 1894²
-. See [About maps] Prokuekenthaliella baumanni ; [AfrL]
Burundi, SW.Uganda (Kigezi), Rwanda? Issoria baumanni baumanni ; [BAFR, 211 (text)]
Prokuekenthaliella baumanni baumanni ; [AfrL]
I. b. excelsior (Butler, [1896])
Uganda-Zaire border (Ruwenzori Mts) Argynnis excelsior Butler, [1896]; Proc. zool. Soc. Lond. 1895 : 729, pl. 42, f. 4, TL: Ruwenzori
Issoria excelsior ; [BOW: pl. 104, f. 6]
Issoria baumanni excelsior ; [BAFR, 211]
Prokuekenthaliella baumanni excelsior ; [AfrL]
I. b. katangae Neave, 1910²
SW.Tanzania, S.Zaire (Shaba), N.Zambia Issoria baumanni katangae ; [BAFR, 211 (text)]
Prokuekenthaliella baumanni katangae ; [AfrL]
I. b. orientalis Kielland, 1990²
baumanni orientalis Kielland, 1990²
Prokuekenthaliella baumanni orientalis ; [AfrL]

Issoria uganda Rob.²
Issoria uganda ; [EBW, (name only)]

Subgenus Yramea Reuss, 1920
Yramea, NSG Voucher Specimen [Wahlberg]

Issoria cytheris (Drury, [1773])
-. See [About maps] Papilio cytheris Drury, [1773]; Illust. Nat. Hist. Exot. Insects 2: index, : 7, pl. 4, f. 3-4, TL: Falkland Is.
Yramea cytheris, NSG Voucher Specimen [Wahlberg]
Argynnis cytheris ; Staudinger, 1899, Naturhist. Mus. Hamburg 2 (6) : 28
Issoria cytheris ; [EBW]; [BOW: pl. 38, f. 14]
Yramea cytheris ; ; [NL4A, #2290]
Falkland Is. Argynnis cytheris falklandica Watkins, 1924; , TL: Falkland Is.
= ; [NL4A, #2290a]
Yramea cytheris cytheris ; [NL4A, #2290a]
I. c. siga (Geyer, 1832)
Chile, Argentina cytheris siga (Geyer, 1832) (Argynnis); , TL: "Java"
Argynnis anna Blanchard, 1852; , TL: Chile
= ; [NL4A, #2290b]
Argynnis chilensis Reed, 1877; (nom. nud.), TL: Chile
= ; [NL4A, #2290b]
Chilargynnis cytheris subtusviola Bryk, 1944; , TL: Argentina
= ; [NL4A, #2290b]
Yramea cytheris siga ; [NL4A, #2290b]

Issoria inca (Staudinger, 1894)
-. See [About maps] Argynnis inca Staudinger, 1894; Dt. Ent. Z. Iris 7 (1) : 68, pl. 2, f. 1-2, TL: Bolivia
Yramea inca, NSG Voucher Specimen [Wahlberg]
Issoria inca ; [BOW: pl. 38, f. 13]
Yramea inca ; [NL4A, #2291]
Bolivia Yramea inca inca ; [NL4A, #2291a]
I. i. cora (Lehmann, 1913)
Peru (Brenthis) inca cora (Lehmann, 1913) (Argynnis); , TL: Peru
Issoria cora ; [EBW, (name only)]
Yramea inca cora ; [NL4A, #2291b]
Argentina Yramea inca ssp.
Yramea inca ssp. ; Lamas, MS, [NL4A, #2291c]

Issoria lathonioides (Blanchard, 1852)
Chile, Argentina. See [About maps] Argynnis lathonioides Blanchard, 1852; , TL: Chile
Argynnis dexamene Boisduval, 1859; , TL: Argentina
= ; [NL4A, #2292]
Argynnis dioides Boisduval, 1859; (nom. nud.)
= ; [NL4A, #2292]
Argynnis montana Reed, 1877; , TL: Chile
= ; [NL4A, #2292]
Argynnis euterpe Reed, 1877; (nom. nud.), TL: Chile
= ; [NL4A, #2292]
Argynnis darwini Staudinger, 1899; Naturhist. Mus. Hamburg 2 (6) : 32, TL: Chile
= ; [NL4A, #2292]
Issoria darwini ; [EBW, (name only)]
Yramea lathonioides ; [NL4A, #2292]

Issoria modesta (Blanchard, 1852)
Chile. See [About maps] Argynnis modesta Blanchard, 1852; , TL: Chile
Issoria modesta ; [EBW]
Yramea modesta ; [NL4A, #2293]

Issoria sobrina (Weymer, 1890)
Bolivia, Peru. See [About maps] Argynnis sobrina Weymer, 1890; in Weymer & Maassen, Reisen Süd-America : 118, pl. 3, f. 15, TL: Bolivia
Boloria tessellata Rosenberg & Talbot, 1914; Trans. ent. Soc. Lond. 1913 (4) : 673, TL: Peru
= ; [NL4A, #2294]
Yramea blancasi Hayward, 1960; , TL: Peru
= ; [NL4A, #2294]
Yramea sobrina ; [NL4A, #2294]

Issoria sp.
Peru. See [About maps] Yramea sp.
Yramea sp. ; Lamas & Grados, MS, [NL4A, #2295]


17.2.2013 (10)

References:
[About maps]
Warning! The maps are automatically generated from the textual information, and the process does not always produce acceptable result; See about maps for more info.
[AfrL]
Mark C. Williams; Checklist of Afrotropical Papilionoidea and Hesperoidea; Compiled by Mark C. Williams, 7th ed. (2008) (April 2007); Afrototropical Butterflies; e-mail: Mark C. Williams
[BAFR]
D'Abrera, 1980; Butterflies of the Afrotropical Region
[BHL]
The; Biodiversity Heritage Library; and on; Internet Archive; Note: The linking to the content of Internet Archive is somewhat delicate -- notify me when links break!
[BK]
Larsen, 1996; The Butterflies of Kenya and their natural history
[BMAT]
Tennent, 1996; The butterflies of Morocco, Algeria and Tunisia; ISBN 0 906802 05 9
[BOW]
Lewis, 1974; Butterflies of the World; ISBN 0 245 52097 X
[BRU2]
Tuzov (ed.), Bogdanov, Churkin, Devyatkin, Dantchenko, Murzin, Samodurov, Zhdanko, 2000; Guide to the Butterflies of Russia and adjacent territories; Libytheidae, Danaidae, Nymphalidae, Riodinidae, Lycaenidae; Volume 2
[EBW]
Smart, Paul, 1976; The Illustrated Encyclopedia of the Butterfly World; ISBN 0 600 31381 6
[ESP]
Heppner, 1981; The dates of E. J. C. Esper's Die Schmetterlinge in Abblidungen... 1776-[1830]; Archives of Natural History (1981) 10 (2): 251-254
[HBN]
Francis Hemming, 1937; HÜBNER, A bibliographical and systematic account of the entomological works of Jacob Hübner and of the supplements thereto by Carl Geyer, Gottfried Franz von Frölich and Gottlieb August Wilhelm Herrich-Schäffer, Vol 1 & 2
[Kimmo Silvonen]
Espoo, Finland; Kimmo's Lep Site
[LAMAS-NEO]
Gerardo Lamas; Bibliography of butterflies; An Annocated Bibliography of the Neotropical Butterflies and Skipper (Lepididoptera: Papilionoidea and Hesperioidea); Revised Electronic Edition (2007)
[Leif Wahlberg]
Julita, Sweden; e-mail: Leif Wahlberg
[MBE]
Moths and Butteflies of Europe; Paolo Mazzei, Diego Reggianti and Ilaria Pimpinelli; http://www.leps.it/
[MRS]
Chou Io (Ed.); Monographia Rhopalocerum Sinensium, 1-2
[Mario Maier]
Europäische Schmetterlinge; e-mail: Mario Maier
[Michael Borsch]
Geesthacht, Germany; contact through; e-mail: Francisco Welter-Schultes
[Michel Tarrier]
Michel R. Tarrier; Malaga, Spain; http://www.tarrier.org
[NL4A]
Gerardo Lamas, 2004; Atlas of Neotropical Lepidoptera; Checklist: Part 4A; Hesperioidea - Papilionoidea
[NORLEP]
Norges Sommerfugler
[Oleg Kosterin]
Institue of Cytology & Genetics; Siberian Division of Russian Academy of Sciences; Novosibirsk; e-mail: Oleg Kosterin; http://pisum.bionet.nsc.ru/kosterin/
[PBSA]
Dickson, C.G.C (ed.), Kroon, D.M.; Pennington's Butterflies of Southern Africa; AD. DONKER 1978
[SPRK]
Seppänen, E. J, 1970; Suomen suurperhostoukkien ravintokasvit, Animalia Fennica 14
[SVF]
Svenska fjärilar; Naturhistoriska riksmuseet; (Sweden)
[Simon Coombes]
1 Park Street, Plymouth, PL3 4BL, UK; 01752 607854; e-mail: Simon Coombes; http://www.butterfly-guide.co.uk
[Wahlberg]
Niklas Wahlberg; The NSG's voucher specimen database; Nymphalidae Systematics Group
[²]
This may require parentheses or not. I don't have the necessary information for this taxon.


Some related literature:
Butler, 1895
On a small Collection of Butterflies sent by Mr. Richard Crawshay from the country west of Lake Nyasa Proc. zool. Soc. Lond. 1895 : 627-634, pl. 35
Butler, [1896]
On Lepidoptera recently collected in British East Africa by Mr. G. F. Scott Elliot Proc. zool. Soc. Lond. 1895 : 722-742, pl. 42-43 (1896)
Doubleday, [1845]
List of the Specimens of Lepidopterous insects in The Collection of the British Museum List. lepid. Ins. Brit. Mus. 1 : 1-150
Doubleday, [1848]
The genera of diurnal Lepidoptera, comprising their generic characters, a notice of their habitats and transformations, and a catalogue of the species of each genus; illustrated with 86 plates by W. C. Hewitson Gen. diurn. Lep. (1): 1-18, pl. A, 1-4 (1-2) (1846), (1): 19-132, pl. 4*, 5-25, 28 (3-14) (1847), (1): 133-200, pl. 1*, 26, 27, 29, 31-44 (15-23) (1848), (1): 201-242, pl. 30, 45-52, 56-58, 60-62, 64 (24-31) (1849), (1): 243-250 [?], (2): 243-326, pl. 53-55, 63, 65, 66 (32-38) (1850), (2): 327-466, pl. 59, 67-77 (39-50) (1851), (2): 467-534, 143, 144, pl. 78-80 (Index) (51-54) (1852)
Drury, [1773]
Illustrations of natural history; wherein are exhibited ... Illust. Nat. Hist. Exot. Insects 2 : 1-90, pl. 1-50
Elwes, 1889
A revision of he genus Argynnis Trans. ent. Soc. Lond. 1889 : 535-575
Esper, [1804]
Die Schmetterlinge in Abbildungen nach der Natur mit Beschreibungen. Theil I. Die Tagschmetterlinge. Supplement Theil 1. Abschnitt 1 Die Schmett., Suppl. Th 1 (1-2): 1-8, pl. 94 (1789), (3-4): 9-60, pl. 95-101 (1789), (5-6): 61-68, pl. 102-106 (1794), ([7]): [69-88], pl. [107-109] (1798), (8-9): 89-104, pl. 110-112 (1800), (10): 105-120, pl. 113-116 ([1803-1804])
Gray, 1846
Descriptions and Figures of some new Lepidopterous Insects chiefly from Nepal Descr. lep. Ins. Nepal : 1-16, pl. 1-14
Huang, 2003
A list of butterflies collected from Nujiang (Lou Tse Kiang) and Dulongiang, China with descriptions of new species, and revisional notes Neue Ent. Nachr. 55: 3-114
Hübner, [1819]
Verzeichniss bekannter Schmettlinge, 1816-[1826] Verz. bek. Schmett. (1): [1-3], 4-16 (1816), (2): 17-32 (1819), (3): 33-48 (1819), (4): 49-64 (1819), (5): 65-80 (1819), (6): 81-96 (1819), (7): 97-112 (1819), (8): 113-128 (1819), (9): 129-144 (1819), (10): 145-160 (1819), (11): 161-176 (1819), (12): 177-192 (1820), (13): 193-208 (1820), (14): 209-224 (1821), (15): 225-240 (1821), (16): 241-256 (1821), (17): 257-272 (1823), (18): 273-288 (1823), (19): 289-304 (1823), (20): 305-320 (1825), (21): 321-336 (1825), (22): 337-352 (1825), (23-27): 353-431 ([1825])
Linnaeus, 1758
Systema Naturae per Regna Tria Naturae, Secundum Clases, Ordines, Genera, Species, cum Characteribus, Differentiis, Symonymis, Locis. Tomis I. 10th Edition Syst. Nat. (Edn 10) 1 : 1-338, : 339-824
Moore, 1857
A Catalogue of the Lepidopterous Insects in the Museum of the Hon. East-India Company in Horsfield & Moore, Cat. lep. Ins. Mus. East India Coy 1: : 1-278, pl. 1-12,1a,2a,3a,4a,5a,6a
Reuss, 1920
Die Androconien von Yramea cytheris Drury und nächtststehenden analogen Schuppenbildungen bei Dione Hbn. und Brenthis Hbn. (Lep.) Ent. Mitt. 9 : 192
Reuss, 1926
Systemischer uberlich der Dryadinae T. Rss mit einigen Neubeschreibungen Dt. ent. Z. 1926 (1) : 65-70
Rosenberg & Talbot, 1914
New South American butterflies Trans. ent. Soc. Lond. 1913 (4) : 671-682
Staudinger, 1894
Hochlandine Lepidoptera Dt. Ent. Z. Iris 7 (1) : 43-100, pl. 1-2
Staudinger, 1899
Lepidopteren. Hamburger Magalhaensische Sammelreise Naturhist. Mus. Hamburg 2 (6) : 1-117, pl., f. 1-27
Weymer in Weymer & Maassen, 1890
Lepidoptera gesammelt auf einer Reise durch Colombia, Ecuador, Peru, Brasilien, Argentinien und Bolivien in den Jahren 1868-1877 von Alphons Stübel Reisen Süd-America : 1-182, pl. 1-9


If you have corrections, comments or information to add into these pages, just send mail to Markku Savela
Keep in mind that the taxonomic information is copied from various sources, and may include many inaccuracies. Expert help is welcome.