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Heliconius Kluk, 1780
Heliconius Kluk, 1780; Hist. nat. pocz. gospod. 4: 82, TS: Papilio charithonia Linnaeus
Heliconius Latreille, 1804; Nouv. Dict. Hist. nat. 24 (6): 185, 199, TS: Papilio charithonia Linnaeus
= ; [NL4A, 266]
Migonitis Hübner, 1816; Verz. bek. Schmett. (1): 12 (preocc. Migonitis Rafinesque, 1815), TS: Papilio erato Linnaeus
= ; [NL4A, 266]
Sunias Hübner, 1816; Verz. bek. Schmett. (1): 12, TS: Papilio melpomene Linnaeus
= ; [NL4A, 266]
Sicyonia Hübner, 1816; Verz. bek. Schmett. (1): 13 (suppr.), TS: Papilio rhea Cramer
= ; [NL4A, 266]
Ajantis Hübner, 1816; Verz. bek. Schmett. (1): 13, TS: Papilio sappho Drury
= ; [NL4A, 266]
Apostraphia Hübner, 1816; Verz. bek. Schmett. (1): 13, TS: Papilio charithonia Linnaeus
= ; [NL4A, 266]
Heliconia Latreille, 1818; (emend.)
= ; [NL4A, 266]
Phlogris Hübner, [1825]; Samml. exot. Schmett. 2 : pl. [5], TS: Papilio melpomene Linnaeus
= ; [NL4A, 266]
Podalirius Gistel, 1848; (preocc. Latreille, 1802, repl. name)
= ; [NL4A, 266]
Blanchardia Buchecker, 1880; Syst. Ent. 2: pl. 51 (preocc. Blanchardia Castelnau, 1875), TS: Heliconia choarina Hewitson
= ; [NL4A, 266]
Heliconius, NSG Voucher Specimen [Wahlberg]
Sycionia ; Hübner, [1826], Verz. bek. Schmett. (Anz.) (1-9): 7 (missp.)
Heliconia ; Godart, 1819, Encyclopédie Méthodique. 9 (1): 203 (missp.)
Helicona ; Guérin-Méneville, [1844], Icon. Règne anim. Cuvier 3 (Insectes) : 472 (missp.)
Heliconius ; Godman & Salvin, [1881], Biol. centr.-amer., Lep. Rhop. 1: 143
Heliconius (Heliconiini) ; [Wahlberg]; [NL4A, 266]

Heliconius antiochus (Linnaeus, 1767)
-. See [About maps] Papilio antiochus Linnaeus, 1767; , TL: "Indiis" [South America]
Heliconius antiochus, NSG Voucher Specimen [Wahlberg]
Heliconius antiochus ; [EBW]; [BOW: pl. 43, f. 37]; [NL4A, #2394]
Brazil(Pará), Peru, Surinam Heliconius zobeide Butler, 1869; Ann. Mag. nat. Hist. (4) 3 (13) : 18, pl. 9, f. 3, TL: Brazil, Pará; Peru
= ; [NL4A, #2394a]
Heliconius antiochus ab. divisus Staudinger, [1897]; Dt. Ent. Z. Iris 9 (2) : 294, TL: Brazil, Pará
= ; [NL4A, #2394a]
Heliconius antiochus ab. alba Riffarth, 1900; Berl. ent. Zs. 45 (3/4) : 208, TL: Peru,"Sarayacu, Amazon",Surinam
= ; [NL4A, #2394a]
Heliconius antiochus alba ab. trimaculata Krüger, 1933; , TL: Brazil, Pará
= ; [NL4A, #2394a]
Heliconius antiochus alba f. zobeide ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 209, pl. 37, f. 382
Heliconius antiochus alba ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 177, pl. 37, f. 381; [EBW, (name only)]
Heliconius antiochus antiochus ; [NL4A, #2394a]
H. a. aranea (Fabricius, 1793)
Papilio aranea Fabricius, 1793; Ent. Syst. 3 (1): 168
Heliconius antiochus aranea ; [EBW]; [EcuL]; [NL4A, #2394b]
H. a. salvinii Dewitz, 1877
Venezuela Heliconius salvinii Dewitz, 1877; Mitt. Münch. Ent. Ver. 1 : 86, TL: [Venezuela]
Heliconius antiochus salvinii ; [EBW]; [NL4A, #2394d]
H. a. araneides Staudinger, 1897
Venezuela Heliconius antiochus var. (ab?) araneides Staudinger, [1897]; Dt. Ent. Z. Iris 9 (2) : 294, TL: Venezuela
Heliconius antiochus araneides ; [NL4A, #2394c]
Peru Heliconius antiochus ssp. ; Lamas, MS, [NL4A, #2394e]

Heliconius astraea Staudinger, 1897
-. See [About maps] Heliconius astraea Staudinger, 1897; , TL: Brazil, Amazonas
Heliconius astraea ; [EBW, (name only)]; [NL4A, #2395]
Brazil(Amazonas) Heliconius astraea astraea ; [NL4A, #2395a]
H. a. rondonia Brown, 1973
Brazil(Rondônia) Heliconius astraea rondonia Brown, 1973; Bull. Allyn Mus. 13: 2, TL: Brazil, Rondônia
Heliconius astraea rondonia ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 202, pl. 9, f. 92; [NL4A, #2395b]

Heliconius atthis Doubleday, [1847]
Ecuador. See [About maps] Heliconia atthis Doubleday, [1847]; Gen. diurn. Lep. (1): pl. 14, f. 3, TL: Ecuador
Heliconia bourcieri Becker, 1851; (nom. nud.)
= ; [NL4A, #2396]
f. gerstneriana Descimon & Mast, 1971; , TL: Ecuador
= ; [NL4A, #2396]
Heliconius atthis, NSG Voucher Specimen [Wahlberg]
Heliconius atthis ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 179, pl. 17, f. 174; [EBW]; [BOW: pl. 44, f. 2]; [EcuL]; [NL4A, #2396]

Mimics Tithorea pavoni. [BOW]


Heliconius besckei Ménétriés, 1857
Brazil(Rio de Janeiro, Minas Gerais, Santa Catarina, São Paulo). See [About maps] Heliconia besckei Ménétriés, 1857; Cat. lep. Petersb. 2: 114, TL: Brazil, Rio de Janeiro; Minas Gerais
Heliconia besckii [= besckei] Ménétriés, 1857; Cat. lep. Petersb. 2: pl. 8, f. 3
Heliconius epiphyllis Riffarth, 1901; Berl. ent. Zs. 46 (1/2) : 106 (nom. nud.)
= ; [NL4A, #2397]
Heliconius besckei principalis Röber, 1921; , TL: Brazil, Santa Catarina; São Paulo
= ; [NL4A, #2397]
f. intermixta Neustetter, 1931; , TL: Brazil, Rio de Janeiro
= ; [NL4A, #2397]
Heliconius besckei, NSG Voucher Specimen [Wahlberg]
Heliconius beschkei[sic, recte besckei] ; [BOW: pl. 44, f. 20 (text only)]
Heliconius (Heliconius) besckei ; Brown & Mielke, 1967, J. Lep. Soc. 21 (2): 96
Heliconius besckei ; [EBW]; [NL4A, #2397]

Heliconius burneyi (Hübner, [1831])
Guianas - Peru, Bolivia. See [About maps] Migonitis burneyi Hübner, [1831]; Zuträge Samml. exot. Schmett. 3 : 7, : pl. [70], f. 401-402, TL: Brazil
Heliconius burneyi, NSG Voucher Specimen [Wahlberg]
Heliconius burneyi ; [BOW: pl. 44, f. 3]; [NL4A, #2398]
French Guiana, .... var. jeanneae Boullet & Le Cerf, 1909; , TL: French Guiana
= ; [NL4A, #2398a]
f. oberthüri Boullet & Le Cerf, 1909; (preocc. Riffarth, 1903), TL: ["Peru"]
= ; [NL4A, #2398a]
f. roseni Krüger, 1928; , TL: "Surinam"
= ; [NL4A, #2398a]
Heliconius burneyi burneyi ; [NL4A, #2398a]
H. b. catharinae Staudinger, 1885
Brazil(Pará, Amazonas) Heliconius catharinae Staudinger, 1885; in Staudinger & Schatz, Exot. Schmett. 1 (9): 79, TL: [Brazil, Pará]
Heliconius burneyi serpensis Kaye, 1919; Ann. Mag. nat. Hist. (9) 3 (14) : 218, TL: Brazil, Amazonas
= Heliconius burneyi burneyi × burneyi catherinae ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 204, pl. 8, f. 90
= ; [NL4A, #2398e (hyb.)]
Heliconius burneyi f. catharinae ab. vitiosa Krüger, 1933; , TL: Brazil, Pará
= ; [NL4A, #2398e]
Heliconius burneyi f. miraculosa Kotzsch, 1936; , TL: Brazil, Pará
= ; [NL4A, #2398e]
Heliconius burneyi serpensis ; [EBW, (name only)]
Heliconius burneyi catharinae ; [EBW]; [NL4A, #2398e]
H. b. huebneri Staudinger, 1897
Peru Heliconius burneyi var. huebneri Staudinger, [1897]; Dt. Ent. Z. Iris 9 (2) : 312, TL: "Ob. Amazonas (u. Rio Negro) Gebiet" [Peru]
Heliconius burneyi huebneri ; [EBW]; [EcuL]; [NL4A, #2398f]
H. b. lindigii C. & R. Felder, 1865
Colombia Heliconius lindigii C. & R. Felder, [1865]; Reise Fregatte Novara, Bd 2 (Abth. 2) (2): 377, pl. 47, f. 1, TL: Colombia
Heliconius burneyi lindigii ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 195, pl. 8, f. 89; [EBW, (name only)]; [NL4A, #2398i]
H. b. ada Neustetter, 1925
Brazil(Rondônia) burneyi ada Neustetter, 1925; , TL: Brazil, Rondônia
Heliconius burneyi ada ; [NL4A, #2398b]
H. b. skinneri Brown & Fernández, 1985
Venezuela burneyi skinneri Brown & Fernández, 1985; , TL: Venezuela
Heliconius burneyi skinneri ; [NL4A, #2398j]
H. b. anjae Neukirchen, 1995
Brazil(Amazonas) Heliconius burneyi anjae Neukirchen, 1995; Atalanta 26 (1/2): 202, pl. VII, f. 1, TL: Brazil, Amazonas, Barcelos
Heliconius burneyi anjae ; [NL4A, #2398c]
H. b. jamesi Neukirchen, 1995
Peru Heliconius burneyi jamesi Neukirchen, 1995; Atalanta 26 (1/2): 203, pl. VII, f. 3, TL: Peru, Pt. Alegre, Rio Pachitea
Heliconius burneyi jamesi ; [NL4A, #2398g]
H. b. koenigi Neukirchen, 1995
Peru Heliconius burneyi koenigi Neukirchen, 1995; Atalanta 26 (1/2): 204, pl. VIII, f. 1, TL: Peru, Madre de Dios, Puerto Maldonado, 200m
Heliconius burneyi koenigi ; [NL4A, #2398h]
H. b. boliviensis Neukirchen, 1995
Bolivia Heliconius burneyi boliviensis Neukirchen, 1995; Atalanta 26 (1/2): 205, pl. VIII, f. 3, TL: Bolivia, Cochabamba, Rio Chapare, 500-800m
Heliconius burneyi jeanneae f. reducta Boullet & Le Cerf, 1909; , TL: Bolivia
= ; [NL4A, #2398d]
Heliconius burneyi boliviensis ; [NL4A, #2398d]

Heliconius charithonia (Linnaeus, 1767)
S.USA, CA, SA, West Indies. See [About maps] Zebra Longwing
Papilio charithonia Linnaeus, 1767; Syst. Nat. (Edn 12) 1 (2): 757, TL: "America" [?Virgin Islands]
Heliconius charithonia, NSG Voucher Specimen [Wahlberg]
Heliconius charithonia, Butterflies and Moths of North America [BAMONA]
Apostraphia charithonia ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 32
Heliconius charitonius[sic, recte charithonia] ; [NACL, #4418]; [BCR, 193]
Heliconius charitonia[sic, recte charithonia] ; [BOW: pl. 44, f. 4]
Heliconius charithonia ; Godman & Salvin, [1881], Biol. centr.-amer., Lep. Rhop. 1: 151; Godman & Salvin, [1901], Biol. centr.-amer., Lep. Rhop. 2: 667; [EBW]; [Opler]; [NL4A, #2399]
[charithonia-1.jpg] 655x469 (~42Kb) Belize, Central America, Photo © John Shuey

Larva on Passiflora [NAB], Tetrastylis lobata, less Passiflora adenopoda [BCR]


Ecuador Papilio charitonia[sic, recte charithonia] ; Linnaeus, 1767; Cramer, [1777], Uitl. Kapellen 2 (9-16): pl. 191, f. F
Heliconius charithonia charithonia ; [EcuL]; [NL4A, #2399a]
H. c. simulator Röber, 1921
Jamaica charithonia simulator Röber, 1921; , TL: Jamaica
Heliconius charithonia simulator ; [EBW, (name only)]; [NL4A, #2399f]
H. c. bassleri Comstock & Brown, 1950
Colombia charithonia bassleri Comstock & Brown, 1950; , TL: Colombia
Heliconius charithonia bassleri ; [NL4A, #2399c]
H. c. churchi Comstock & Brown, 1950
Haiti charithonia churchi Comstock & Brown, 1950; , TL: Haiti
Heliconius charithonia churchi ; [NL4A, #2399d]
H. c. tuckeri Comstock & Brown, 1950
Florida charithonia tuckeri Comstock & Brown, 1950; , TL: Florida
Heliconius charitonia[sic] tuckerorum Lamas, 1988; (emend.)
= ; [NL4A, #2399g]
Heliconius charithonia tuckeri, Butterflies of America [BOA]
Heliconius charitonius tuckeri ; [NACL, #4418a]
Heliconius charithonia tuckeri ; [NL4A, #2399g]
H. c. vazquezae Comstock & Brown, 1950
Mexico charithonia vazquezae Comstock & Brown, 1950; , TL: Mexico, Campeche
Heliconius charithonia vazquezae, Listing of Mexican Butterflies [ILMB]
Heliconius charithonia vazquezae, Butterflies of America [BOA]
Heliconius charitonius vasquezae[sic] ; [NACL, #4418b]
Heliconius charithonia vazquezae ; [NL4A, #2399h]
H. c. ramsdeni Comstock & Brown, 1950
Cuba charithonia ramsdeni Comstock & Brown, 1950; , TL: Cuba
Heliconius charithonia ramsdeni ; [NL4A, #2399e]
H. c. antiquus Lamas, 1988
St. Kitts, Antigua Heliconius charithonia antiquus Lamas, 1988; (repl. name), TL: St. Kitts, Antigua
Heliconius charithonia punctata Hall, 1936; Entomologist 69 : 276 (preocc. Neustetter, 1907), TL: St. Kitts, Antigua
= ; [NL4A, #2399b]
Heliconius charithonia punctata ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 201, pl. 36, f. 366; [EBW, (name only)]
Heliconius charithonia antiquus ; [NL4A, #2399b]

Heliconius clysonymus Latreille, [1817]
Honduras - Colombia, Ecuador. See [About maps] Heliconius clysonymus Latreille, [1817]; , TL: [Colombia]
Heliconius clysonymus, NSG Voucher Specimen [Wahlberg]
Heliconius clysonimus[sic, recte clysonymus] ; [BOW: pl. 44, f. 15 (text only)]; [EBW, [spelling?]]
Heliconius clysonymus ; [NL4A, #2400]
Colombia, Venezuela Heliconius clysonimus[sic] fischeri Fassl, 1912; Ent. Rdsch. 29 (8) : 55, TL: Colombia
= Heliconius clysonymus clysonymus × clysonymus hygiana ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 189, pl. 36, f. 364
= ; [NL4A, #2400a (hyb.)]
Heliconius clysonimus[sic] flavopunctatus Fassl, 1912; Ent. Rdsch. 29 (8) : 55, TL: Colombia
= Heliconius clysonymus clysonymus × clysonymus hygiana ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 189, pl. 35, f. 362
= ; [NL4A, #2400a (hyb.)]
Heliconius micrus Seitz, 1912; , TL: [Venezuela]
= ; [NL4A, #2400a]
Heliconius clysonymus apicalis Joicey & Kaye, 1917; (nom. nud.)
= ; [NL4A, #2400a]
Heliconius clysonymus apicalis ab. semirubra Joicey & Kaye, 1917; Ann. Mag. nat. Hist. (8) 20 (115) : 90, TL: Colombia
= Heliconius clysonymus clysonymus × clysonymus hygiana ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 204, pl. 36, f. 363
= ; [NL4A, #2400a (hyb.)]
Heliconius clysonymus perbellus Stichel, 1923; , TL: Colombia
= ; [NL4A, #2400a]
Heliconius hygianus fischeri f. leonis H. & R. Holzinger, 1970; , TL: Colombia
= ; [NL4A, #2400a (hyb.)]
Heliconius hygianus fischeri f. leoncitonis H. & R. Holzinger, 1970; , TL: Colombia
= ; [NL4A, #2400a (hyb.)]
Heliconius clysonymus semirubra ; [EBW, (name only)]
Heliconius clysonymus fischeri ; [EBW, (name only)]
Heliconius clysonymus clysonymus ; [EcuL]; [NL4A, #2400a]
H. c. hygiana Hewitson, 1867
Ecuador Heliconia hygiana Hewitson, 1867; Ill. exot. Butts [1] (Heliconia V): [78], pl. [40], f. 15, TL: Ecuador, Quito
Heliconius hygiana ab. albescens Kaye, 1916; Ent. Rec. J. Var. 28 : 194, TL: Ecuador
= ; [NL4A, #2400b]
Heliconius clysonymus hygiana ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 193, pl. 36, f. 365; [EBW]; [EcuL]; [NL4A, #2400b]
H. c. montanus Salvin, 1871
Costa Rica, Panama Heliconius montanus Salvin, 1871; Ann. Mag. nat. Hist. (4) 4 (21) 7 (42): 414, TL: Costa Rica, ?Orosi
Heliconius montanus ; Godman & Salvin, [1881], Biol. centr.-amer., Lep. Rhop. 1: 152, 3: pl. 17, f. 11; Godman & Salvin, [1901], Biol. centr.-amer., Lep. Rhop. 2: 668
Heliconius clysonymus montanus ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 197, pl. 35, f. 361; [NL4A, #2400c]; [BCR, 196]

Larva on Passiflora apetala, P. biflora [BCR]

H. c. tabaconas Brown, 1976
Peru Heliconius clysonymus tabaconas Brown, 1976; 221, TL: N.Peru, Charapi, Cajamarca
Heliconius clysonymus tabaconas ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 206, pl. 38, f. 391; [EcuL, (?)]; [NL4A, #2400d]

Heliconius congener Weymer, 1890
-. See [About maps] Heliconius congener Weymer, 1890; in Weymer & Maassen, Reisen Süd-America : 117, pl. 2, f. 5, TL: "Baños un Jivaria del Píntuc [=Puyo]"
Heliconius congener, NSG Voucher Specimen [Wahlberg]
Heliconius congener ; [EcuL]; [NL4A, #2401]
Ecuador, Peru Heliconius apseudes var. paranapurae Staudinger, [1897]; Dt. Ent. Z. Iris 9 (2) : 292, TL: Peru
= ; [NL4A, #2401a]
Heliconius congener congener ; [NL4A, #2401a]
H. c. ocannensis Stichel, 1905
Colombia congener ocannensis Stichel, 1905; (repl. name), TL: Colombia
Heliconius ocannus Staudinger, 1885; in Staudinger & Schatz, Exot. Schmett. 1 (9): 75 (preocc. Buchecker, [1880]), TL: Colombia
= ; [NL4A, #2401c]
Heliconius congener ocannensis ; [NL4A, #2401c]
H. c. aquilionaris Brown, 1976
Colombia Heliconius congener aquilionaris Brown, 1976; 222, TL: Colombia
Heliconius congener aquilionaris ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 179, pl. 37, f. 384; [NL4A, #2401b]

Heliconius cydno Doubleday, 1847
Mexico - Colombia, Ecuador. See [About maps] Heliconius cydno Doubleday, 1847; Gen. diurn. Lep. (1): pl. 15, f. 3, TL: Santa Fé de Bogotá
Heliconius cydno, NSG Voucher Specimen [Wahlberg]
Heliconius cydno ; Godman & Salvin, [1901], Biol. centr.-amer., Lep. Rhop. 2: 668; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 185, pl. 26, f. 264; [EBW]; [BOW: pl. 44, f. 5]; [NL4A, #2402]

Larva on Passiflora vitifolia, P. biflora, Passiflora spp. [BCR]

Colombia ab. mediocydno Neustetter, 1907; , TL: Colombia
= ; [NL4A, #2402a]
f. punctana Neustetter, 1907; , TL: Colombia
= ; [NL4A, #2402a]
Heliconius cydno cydno ; [NL4A, #2402a]
H. c. hermogenes Hewitson, 1858
Colombia Heliconia hermogenes Hewitson, 1858; Ill. exot. Butts [1] (Heliconia IV): [76], pl. [39], f. 12, TL: [Colombia]
Heliconia temerinda Hewitson, 1873; Ill. exot. Butts [1] (Heliconia and Tithorea VII): [81], pl. [42], f. 23, TL: Colombia
= ; [NL4A, #2402i]
Heliconius hermogenes ab. lutescens Kaye, 1916; Ent. Rec. J. Var. 28 : 194, TL: Colombia, Muzo
= ; [NL4A, #2402i]
f. anthelea Neustetter, 1925; , TL: Colombia
= ; [NL4A, #2402i]
Heliconius cydno dolores Apolinar, 1926; , TL: Colombia
= ; [NL4A, #2402i]
Heliconius cydno f. temerinda ; [BOW: pl. 44. f. 5 (text only)]
Heliconius cydno hermogenes f. temerinda ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 206, pl. 26, f. 268
Heliconius cydno hermogenes ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 192, pl. 27, f. 269; [NL4A, #2402i]
H. c. chioneus Bates, 1864
Panama, Costa Rica - Colombia Heliconius chioneus Bates, 1864; Ent. mon. Mag. 1 (3): 58, TL: Panama
Heliconius cydno galanthus f. exornata Riffarth, 1907; Dt. ent. Zs. 1907 (5) : 505, TL: Costa Rica
= ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 189; [NL4A, #2402d (hyb.)]
Heliconius chioneus ; Godman & Salvin, [1881], Biol. centr.-amer., Lep. Rhop. 1: 156, 3: pl. 18, f. 7-8
Heliconius cydno chioneus ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 183, pl. 26, f. 263; [BCR, 194]; [NL4A, #2402d]
H. c. galanthus Bates, 1864
Mexico - Costa Rica, Guatemala, Nicaragua Heliconius galanthus Bates, 1864; Ent. mon. Mag. 1 (3): 58, TL: Guatemala
Heliconia diotrephes Hewitson, 1869; Trans. ent. Soc. Lond. 1869 (1) : 33, TL: Nicaragua
= ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 186, pl. 26, f. 261; [NL4A, #2402h]
Heliconius cydno ab. stübeli Riffarth, 1900; Berl. ent. Zs. 45 (3/4) : 199, TL: Costa Rica
= ; [NL4A, #2402h]
Heliconius piera Riffarth, 1901; Berl. ent. Zs. 46 (1/2) : 80 (nom. nud.)
= ; [NL4A, #2402h]
Heliconius galanthus ab. subrufescens Schaus, 1913; Proc. zool. Soc. Lond. 1913 (3) : 343, pl. 53, f. 7, TL: Costa Rica
= ; [NL4A, #2402h (hyb.)]
Heliconius cydno galanthus, Listing of Mexican Butterflies [ILMB]
Heliconius cydno galanthus, Butterflies of America [BOA]
Heliconius galanthus ; Godman & Salvin, [1881], Biol. centr.-amer., Lep. Rhop. 1: 155, 3: pl. 18, f. 1-2; Godman & Salvin, [1901], Biol. centr.-amer., Lep. Rhop. 2: 668
Heliconius cydno galanthus ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 190, pl. 26, f. 260; [BCR, 194]; [EBW]; [NL4A, #2402h]
H. c. zelinde Butler, 1869
Heliconius zelinde Butler, 1869; Ann. Mag. nat. Hist. (4) 3 (13) : 17, pl. 9, f. 1, TL: "W coast of America"
f. inca Neustetter, 1928
= ; [NL4A, #2402n]
Heliconius cydno cydnides f. flavidior Neustetter, 1928; Int. ent. Zs. 22 : 258, TL: Colombia
= ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 189, pl. 27, f. 271; [NL4A, #2402n (hyb.)]
Heliconius cydno cydnides f. confluens Neustetter, 1928; Int. ent. Zs. 22 : 259, TL: Colombia
= Heliconius cydno zelinde × cydno cydnides ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 184, pl. 26, f. 266
= ; [NL4A, #2402n (hyb.)]
Heliconius cydno cydnides f. albidior Neustetter, 1928; Int. ent. Zs. 22 : 259, TL: Colombia
= Heliconius cydno cydnides × cydno × zelinde ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 177, pl. 27, f. 272
= ; [NL4A, #2402n (hyb.)]
Heliconius cydno zelinde ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 209, pl. 27, f. 270; [EBW]; [EcuL]; [NL4A, #2402n]
H. c. alithea Hewitson, 1869
Ecuador Heliconia alithea Hewitson, 1869; Equat. Lepid. Buckley (1): 10, TL: Ecuador, [Jorge]
Heliconius cydno ab. haenschi Riffarth, 1900; Berl. ent. Zs. 45 (3/4) : 200, TL: Ecuador, Balzapamba
= ; [EcuL]; [NL4A, #2402b]
Heliconius cydno broncus Stichel, 1906; Genera Insectorum 37 : 21, pl. 2, f. 6a, TL: "Peru?"
= Heliconius cydno alithea × cydno zelinde ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 181, pl. 28, f. 280
= ; [NL4A, #2402b (hyb.)]
Heliconius cydno alithea f. egregia Riffarth, 1907; Dt. ent. Zs. 1907 (5) : 505, TL: Ecuador, Balzapamba
= ; [EcuL]; [NL4A, #2402b]
Heliconius alithea (?var.) ab. minor Neustetter, 1907; , TL: Ecuador
= ; [EcuL]; [NL4A, #2402b]
Heliconius alithea f. neustetteri Riffarth, 1908; Dt. ent. Zs. 1908 (1) : 114, TL: Ecuador, Balzapamba und Riobamba
= ; [EcuL]; [NL4A, #2402b]
Heliconia alithea ; Hewitson, 1871, Ill. exot. Butts [1] (Heliconia VI): [79], pl. [41], f. 18
Heliconius cydno alithea f. egregia ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 187, pl. 28, f. 284
Heliconius cydno alithea f. haenschi ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 191, pl. 28, f. 281
Heliconius cydno alithea f. neustetteri ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 198, pl. 28, f. 283
Heliconius cydno alithea ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 177, pl. 28, f. 282; [EBW, (name only)]; [EcuL]; [NL4A, #2402b]
H. c. pachinus Salvin, 1871
Costa Rica, Panama Heliconius pachinus Salvin, 1871; Ann. Mag. nat. Hist. (4) 4 (21) 7 (42): 414, TL: Panama
Heliconius pachinus, NSG Voucher Specimen [Wahlberg]
Heliconia pachinus ; Hewitson, 1875, Ill. exot. Butts [1] (Heliconia VIII): [83], pl. [43], f. 26
Heliconius pachinus ; Godman & Salvin, [1881], Biol. centr.-amer., Lep. Rhop. 1: 158, 3: pl. 18, f. 11; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 199, pl. 28, f. 287; [BCR, 195]; [EBW]
Heliconius cydno pachinus ; [NL4A, #2402k]

Larva on Passiflora costaricensis, P. vitifolia, Passiflora spp. [BCR]

H. c. cydnides Staudinger, 1885
Colombia cydno cydnides Staudinger, 1885; in Staudinger & Schatz, Exot. Schmett. 1 (9): 77, TL: Colombia
Heliconius cydno var. epicydnides Staudinger, [1897]; Dt. Ent. Z. Iris 9 (2) : 289, pl. 6, f. 5, TL: Colombia
= ; [NL4A, #2402f]
Heliconius cydno var. subcydnides Staudinger, [1897]; Dt. Ent. Z. Iris 9 (2) : 289, TL: Colombia
= ; [NL4A, #2402f]
Heliconius interrupta Riffarth, 1901; Berl. ent. Zs. 46 (1/2) : 83 (nom. nud.)
= ; [NL4A, #2402f]
f. werneri Neustetter, 1928; , TL: Colombia
= ; [NL4A, #2402f]
Heliconius cydno cydnides f. azteka Neustetter, 1928; Int. ent. Zs. 22 : 259, TL: Colombia
= Heliconius cydno cydnides × cydno zelinde ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 180, pl. 26, f. 265
= ; [NL4A, #2402f]
f. semicydnides H. & R. Holzinger, 1968; , TL: Colombia
= ; [NL4A, #2402f]
f. tenebrosa H. & R. Holzinger, 1968; , TL: Colombia
= ; [NL4A, #2402f]
Heliconius cydno f. epicydnides ; [BOW: pl. 44, f. 5 (text only)]
Heliconius cydno cydnides ; [NL4A, #2402f]
H. c. weymeri Staudinger, [1897]
Colombia Heliconius weymeri Staudinger, [1897]; Dt. Ent. Z. Iris 9 (2) : 287, pl. 6, f. 5, TL: Colombia
Heliconius weymeri ab. gustavi Staudinger, [1897]; Dt. Ent. Z. Iris 9 (2) : 287, pl. 6, f. 1, TL: Colombia
= ; [NL4A, #2402m]
Heliconius weymeri submarginatus Fassl, 1912; Ent. Rdsch. 29 (8) : 56, TL: Colombia
= ; [NL4A, #2402m]
Heliconius weymeri ab. sulphureomaculata Fassl, 1914; Ent. Rdsch. 31 (8): 43, TL: Colombia
= ; [NL4A, #2402m]
Heliconius cydno gustavi ab. flavomaculata Seitz, 1916; , TL: Colombia
= ; [NL4A, #2402m]
Heliconius hermogenes ab. leucosticta Apolinar, 1926; , TL: Colombia
= ; [NL4A, #2402m]
Heliconius hermogenes ab. xanthosticta Apolinar, 1926; , TL: Colombia
= ; [NL4A, #2402m]
Heliconius cydno gerstneri H. & R. Holzinger, 1968; , TL: Colombia
= ; [NL4A, #2402m (hyb.)]
Heliconius cydno gerstneri f. pseudweymeri H. & R. Holzinger, 1968; , TL: Colombia
= ; [NL4A, #2402m (hyb.)]
Heliconius cydno gerstneri f. denhezi H. & R. Holzinger, 1968; , TL: Colombia
= ; [NL4A, #2402m (hyb.)]
Heliconius cydno gerstneri f. flavissima H. & R. Holzinger, 1968; , TL: Colombia
= ; [NL4A, #2402m (hyb.)]
Heliconius cydno weymeri f. gustavi ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 191, pl. 27, f. 276
Heliconius cydno weymeri f. submarginatus ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 205, pl. 27, f. 275
Heliconius cydno weymeri f. sulphureomaculata ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 205, pl. 27, f. 274
Heliconius cydno weymeri ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 209, pl. 27, f. 273; [EBW]; [NL4A, #2402m]
H. c. cordula Neustetter, 1913
Venezuela, Colombia cydno cordula Neustetter, 1913; , TL: Venezuela
Heliconius cydno ab. larseni Niepelt, 1916; in Strand, Lepidoptera Niepeltiana (2) : 13, pl. 14, f. 1, TL: Colombia
= ; [NL4A, #2402e]
Venezuela cydno flaveola Joicey & Kaye, 1917; Ann. Mag. nat. Hist. (8) 20 (115) : 93, pl. 6, f. 3, TL: Mocotone, Venezuela
= Heliconius cydno perijaensis × cydno barinasensis ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 189, pl. 38, f. 389
= ; [NL4A, #2402e (hyb.)]
Heliconius cydno perijaensis Masters, 1973; , TL: Venezuela
= ; [NL4A, #2402e]
Heliconius cydno cordula ; [NL4A, #2402e]
H. c. barinasensis Masters, 1973
Venezuela cydno barinasensis Masters, 1973; , TL: Venezuela
Heliconius cydno barinasensis ; [NL4A, #2402c]
H. c. gadouae Brown & Fernández, 1985
Venezuela cydno gadouae Brown & Fernández, 1985; , TL: Venezuela
Heliconius hahneli Staudinger, 1885; in Staudinger & Schatz, Exot. Schmett. 1 (9): 77, (7): pl. 31, TL: Venezuela
= ; [NL4A, #2402g (hyb.)]
Heliconius cydno gadouae ; [NL4A, #2402g]
H. c. wanningeri Neukirchen, 1991
Colombia cydno wanningeri Neukirchen, 1991; , TL: Colombia
Heliconius cydno wanningeri ; [NL4A, #2402l]
H. c. lisethae Neukirchen, 1995
Colombia cydno lisethae Neukirchen, 1995; , TL: Colombia
Heliconius cydno lisethae ; [NL4A, #2402j]

Heliconius demeter Staudinger, [1897]
-. See [About maps] Heliconius demeter Staudinger, [1897]; Dt. Ent. Z. Iris 9 (2) : 310, pl. 7, f. 6, TL: [Peru] Iquitos
Heliconius demeter, NSG Voucher Specimen [Wahlberg]
Heliconius demeter ; [EBW, (name only)]; [NL4A, #2403]
Peru, Brazil(Amazonas) f. similis Neustetter, 1931; , TL: Brazil, Amazonas
= ; [NL4A, #2403a]
Heliconius demeter demeter ; [NL4A, #2403a]
H. d. bouqueti Nöldner, 1901
French Guiana Heliconius demeter var. bouqueti Nöldner, 1901; Berl. ent. Z. 46 : 7, TL: French Guiana
Heliconia xanthoceras Oberthür, 1902; , TL: French Guiana
= ; [NL4A, #2403d]
Heliconia eueidina Oberthür, 1916; 37, pl. 406, f. 3487, TL: [French Guiana]
= ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 188, pl. 36, f. 370; [NL4A, #2403d]
Eueides egeriformis Joicey & Kaye, 1917; Trans. ent. Soc. Lond. 1916 : 430, pl. 107, f. 8, TL: French Guiana
= ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 187, pl. 36, f. 369; [NL4A, #2403d]
Heliconia automatia Oberthür, 1925; Etud. Lépid. Comp. 22 : 81, f. 5, TL: French Guiana
= ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 180; [NL4A, #2403d]
Heliconius eratoformis Neustetter, 1931; , TL: French Guiana
= ; [NL4A, #2403d]
Heliconius demeter bouqueti ; [EBW, (name only)]; [NL4A, #2403d]
H. d. beebei Turner, 1966
Guyana Heliconius demeter beebei Turner, 1966; Proc. R. ent. Soc. Lond. (B) 35 : 128, TL: Guyana
Heliconius demeter beebei ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 181, pl. 36, f. 371; [NL4A, #2403c]; [EBW, (name only)]
H. d. eratosignis (Joicey & Talbot, 1925)
Brazil(Mato Grosso) Eueides eratosignis Joicey & Talbot, 1925; Ann. Mag. Nat. Hist. (9) 16 (96) : 649, TL: Brazil, Mato Grosso
Heliconius demeter eratosignis ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 188, pl. 36, f. 372; [EBW, (name only)]; [NL4A, #2403e]
H. d. ucayalensis H. & R. Holzinger, 1975
Peru demeter ucayalensis H. & R. Holzinger, 1975; , TL: Peru
Heliconius demeter ucayalensis ; [NL4A, #2403l]
H. d. ulysses Brown & Benson, 1975
Bolivia Heliconius demeter ulysses Brown & Benson, 1975; Bull. Allyn Mus. 26: (1-19), TL: Bolivia
Heliconius demeter ulysses ; [NL4A, #2403m]
H. d. terrasanta Brown & Benson, 1975
Brazil(Pará) Heliconius demeter terrasanta Brown & Benson, 1975; Bull. Allyn Mus. 26: (1-19), TL: Brazil, Pará
Heliconius demeter terrasanta ; [NL4A, #2403i]
H. d. turneri Brown & Benson, 1975
Brazil(Amazonas) Heliconius demeter turneri Brown & Benson, 1975; Bull. Allyn Mus. 26: 2, TL: Brazil, Amazonas
Heliconius demeter turneri ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 207, pl. 36, f. 373; [EBW, (name only)]; [NL4A, #2403k]
H. d. zikani Brown & Benson, 1975
Brazil(Amazonas) Heliconius demeter zikani Brown & Benson, 1975; Bull. Allyn Mus. 26: (1-19), TL: Brazil, Amazonas
Heliconius demeter zikani ; [NL4A, #2403n]
H. d. tambopata Lamas, 1985
Peru demeter tambopata Lamas, 1985; , TL: Peru
Heliconius demeter tambopata ; [NL4A, #2403h]
H. d. karinae Neukirchen, 1990
Brazil(Pará) demeter karinae Neukirchen, 1990; , TL: Brazil, Pará
Heliconius demeter karinae ; [NL4A, #2403f]
H. d. titan Neukirchen, 1995
Brazil(Amazonas) demeter titan Neukirchen, 1995; , TL: Brazil, Amazonas
Heliconius demeter titan ; [NL4A, #2403j]
H. d. neildi Neukirchen, 1997
Ecuador demeter neildi Neukirchen, 1997; , TL: Ecuador, Napo, El Capricho 850m
Heliconius demeter neildi ; [EcuL]; [NL4A, #2403g]
H. d. angeli Neukirchen, 1997
Peru demeter angeli Neukirchen, 1997; , TL: Peru
Heliconius demeter angeli ; [NL4A, #2403b]
Peru Heliconius demeter ssp. ; Lamas, MS, [NL4A, #2403o]

Heliconius egeria (Cramer, 1775)
-. See [About maps] Papilio egeria Cramer, 1775; , TL: Surinam
Heliconius egeria, NSG Voucher Specimen [Wahlberg]
Heliconius egeria ; [NL4A, #2404]
Surinam, French Guiana Migonitis isaea Hübner, 1816; (repl. name), TL: Surinam
= ; [NL4A, #2404a]
Heliconia ergatis Godart, 1819; (repl. name), TL: Surinam
= ; [NL4A, #2404a]
Heliconia egeria ab. clearista Oberthür, 1923; Etud. Lépid. Comp. 20: 304, pl. 365, f. 4863, TL: French Guiana
= ; [NL4A, #2404a]
Heliconius egeria egeria ab. clearista ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 183, pl. 8, f. 91
Heliconius egeria egeria ; [NL4A, #2404a]
H. e. hyas Weymer, 1883
Brazil (Amazonas) Heliconius hyas Weymer, 1883; Stettin ent. Ztg 45 (1-3) : 26, pl. 1, f. 5, TL: [Brazil, Amazonas]
Heliconius egeria hylas[sic] ; [EBW, (name only, missp.?)]
Heliconius egeria hyas ; Neukirchen, 1991, Atalanta 22 (2/4): 87 (note), pl. XXI, f. 1-8; [NL4A, #2404e]
H. e. egerides Staudinger, 1897
Brazil(Amazonas) Heliconius egeria var. egerides Staudinger, [1897]; Dt. Ent. Z. Iris 9 (2) : 311, TL: Brazil, Amazonas
Heliconius egeria egerides ; [EBW, (name only)]; [NL4A, #2404c]
H. e. homogena Bryk, 1953
Brazil(Amazonas) egeria homogena Bryk, 1953; , TL: Brazil, Amazonas
Heliconius egeria-hyas f. asterope Zikán, 1937; Ent. Rdsch. 54: 329, TL: Brazil, Amazonas
= ; [NL4A, #2404d]
Heliconius egeria homogena ; [EBW, (name only)]; [NL4A, #2404d]
H. e. mariasibyllae Neukirchen, 1991
Brazil (Para) Heliconius egeria mariasibyllae Neukirchen, 1991; Atalanta 22 (2/4): 89, pl. XX, f. 1-8, TL: Brazil, Pará, Santarém
Heliconius egeria mariasibyllae ; [NL4A, #2404g]
H. e. keithbrowni Neukirchen, 1992
Venezuela egeria keithbrowni Neukirchen, 1992; , TL: Venezuela
Heliconius egeria keithbrowni ; [NL4A, #2404f]
H. e. christiani Neukirchen, 1997
Brazil(Pará) egeria christiani Neukirchen, 1997; , TL: Brazil, Pará
Heliconius egeria christiani ; [NL4A, #2404b]

Heliconius eleuchia Hewitson, 1854
Costa Rica - Ecuador. See [About maps] Heliconia eleuchia Hewitson, 1854; Ill. exot. Butts [1] (Heliconia II-III): [73], pl. [38], f. 8, TL: New Granada
Heliconius eleuchia, NSG Voucher Specimen [Wahlberg]
Helicounis eleuchia ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 187, pl. 37, f. 385
Heliconius eleuchia ; [EBW]; [NL4A, #2405]
Costa Rica - Colombia Heliconius eleuchia eleuchia ; [BCR, 198]; [NL4A, #2405a]

Larva on Passiflora tica? [BCR]

H. e. primularis Butler, 1869
Ecuador Heliconius primularis Butler, 1869; Ann. Mag. nat. Hist. (4) 3 (13) : 18, pl. 9, f. 2, TL: Ecuador, "Guayaquil and Rio Napo"
Heliconius eleuchia primularis ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 201, pl. 38, f. 387; [EBW]; [EcuL]; [NL4A, #2405c]
H. e. eleusinus Staudinger, 1885
Colombia, Ecuador eleuchia eleusinus Staudinger, 1885; in Staudinger & Schatz, Exot. Schmett. 1 (9): 76, (7): pl. 31, TL: "westlichen Columbien Rio San Juan"
Heliconius sapho primularis ab. deflava Joicey & Kaye, 1917; Ann. Mag. nat. Hist. (8) 20 (115) : 93, pl. 6, f. 6, TL: Paramba, Ecuador
= ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 185, pl. 38, f. 386; [EcuL]; [NL4A, #2405b (hyb.)]
Heliconius ceres Oberthür, 1920; 30, pl. 519, f. 4324, TL: Colombia
= ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 183, pl. 38, f. 388; [NL4A, #2405b (hyb.)]
Heliconius primularis ceres Stichel, 1923; (preocc. Oberthür, 1920), TL: Colombia
= ; [NL4A, #2405b (hyb.)]
Heliconius eleuchia eleusinus ; [EBW, (name only)]; [EcuL]; [NL4A, #2405b]

Heliconius elevatus Nöldner, 1901
Amazon. See [About maps] Heliconius elevatus Nöldner, 1901; Berl. ent. Z. 46 : 5, TL: "Amazona"
Heliconius elevatus, NSG Voucher Specimen [Wahlberg]
Heliconius elevatus ; [EBW, (name only)]; [NL4A, #2406]
Peru f. griseoviridis Neustetter, 1938; , TL: Peru
= ; [NL4A, #2406a]
Heliconius elevatus elevatus ; [NL4A, #2406a]
H. e. bari Oberthür, 1902
French Guiana Heliconia bari Oberthür, 1902; Étud. d'Ent. 21 : 23, pl. 11, f. 129-130, TL: French Guiana
?Heliconius melpomene f. bari ; [BOW: pl. 44, f. 18 (text only)]
Heliconius elevatus bari ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 180, pl. 17, f. 176; [EBW, (name only)]; [NL4A, #2406b]
H. e. tumatumari Kaye, 1906
Guyana Heliconius elevatus tumatumari Kaye, 1906; Entomologist 39 : 53, pl. 2, f. 2, TL: British Guyana
Heliconius elevatus tumaturnari[sic] ; [EBW, (name only, missp.)]
Heliconius elevatus tumatumari ; [NL4A, #2406j]
H. e. perchlora Joicey & Kaye, 1917
Bolivia Heliconius elevatus perchlora Joicey & Kaye, 1917; Ann. Mag. nat. Hist. (8) 20 (115) : 94, TL: Bolivia
Heliconius elevatus perchlora ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 200, pl. 17, f. 178; [EBW, (name only)]; [NL4A, #2406d]
H. e. schmassmanni Joicey & Talbot, 1925
Brazil(Mato Grosso, Rondônia) Heliconius elevatus schmassmanni Joicey & Talbot, 1925; Ann. Mag. Nat. Hist. (9) 16 (96) : 647, TL: Brazil, Mato Grosso
Heliconius melpomene thelxiope f. aquilina Neustetter, 1925; , TL: Brazil, Rondônia
= ; [NL4A, #2406g]
Heliconius elevatus schmassmanni ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 203, pl. 17, f. 177; [EBW, (name only)]; [NL4A, #2406g]
H. e. pseudocupidineus Neustetter, 1931
Peru elevatus pseudocupidineus Neustetter, 1931; , TL: Peru
Heliconius elevatus f. nigromacula Neustetter, 1932; , TL: Peru
= ; [NL4A, #2406e]
Heliconius elevatus f. nöldneri Neustetter, 1938; , TL: Peru
= ; [NL4A, #2406e]
Heliconius elevatus pseudocupidineus ; [NL4A, #2406e]; [EBW, (name only)]
H. e. taracuanus Bryk, 1953
Brazil(Amazonas) elevatus taracuanus Bryk, 1953; , TL: Brazil, Amazonas
Heliconius elevatus taracuanus ; [NL4A, #2406i]; [EBW, (name only)]
H. e. roraima Turner, 1966
Guyana Heliconius elevatus roraima Turner, 1966; J. Res. Lepid. 5 : 109, TL: Guyana
Heliconius elevatus roraima ; [EBW, (name only)]; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 202, pl. 17, f. 175; [NL4A, #2406f]
[roraima-1.jpg] 500x352 (~41Kb) Venezuela, Parai Tepui near Mount Roraima in the Gran Sabana, July 2002, Photo © Simon Coombes

H. e. lapis Lamas, 1976
Peru elevatus lapis Lamas, 1976; , TL: Peru
Heliconius elevatus lapis ; [NL4A, #2406c]
H. e. schmidt-mummi Takahashi, 1977
Heliconius (Heliconius) elevatus schmidt-mummi Takahashi, 1977; Tyô to Ga 28 (1): 13, TL: nr Leticia, Brazil
H. e. zoelleri Neukirchen, 1990
Venezuela elevatus zoelleri Neukirchen, 1990; , TL: Venezuela
Heliconius elevatus zoelleri ; [NL4A, #2406l]
H. e. sonjae Neukirchen, 1997
Brazil(Pará) Heliconius elevatus sonjae Neukirchen, 1997; , TL: Brazil, Pará
Heliconius elevatus sonjae ; [NL4A, #2406h]
H. e. willmotti Neukirchen, 1997
Ecuador Heliconius elevatus willmotti Neukirchen, 1997; , TL: Ecuador, Napo, Zarzayacu, 520m
Heliconius elevatus willmotti ; [EcuL]; [NL4A, #2406k]

Heliconius erato (Linnaeus, 1764)
Mexico - Amazon Basin. See [About maps] Crimson-patched Longwing
Papilio erato Linnaeus, 1764; Mus. Lud. Ulr.: 467, TL: "Indiis" [South America, Surinam]
Heliconius erato, NSG Voucher Specimen [Wahlberg]
Heliconius erato, Butterflies and Moths of North America [BAMONA]
Heliconius erato ; Godman & Salvin, [1881], Biol. centr.-amer., Lep. Rhop. 1: 160; Godman & Salvin, [1901], Biol. centr.-amer., Lep. Rhop. 2: 669; [NACL, #4419]; [EBW]; [BOW: pl. 44, f. 10]; [Opler]; [NL4A, #2407]

Larva on Passiflora [NAB]

Surinam, French Guiana Papilio vesta Cramer, 1777; , TL: Surinam
= ; [NL4A, #2407a (hyb.)]
Papilio erythrea Cramer, [1777]; Uitl. Kapellen 2 (9-16): 140, pl. 189, f. A, TL: Surinam
= Heliconius erato erato × erato hydara ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 188, pl. 39, f. 395
= ; [NL4A, #2407a (hyb.)]
Papilio andremona Cramer, [1780]; Uitl. Kapellen 4 (25-26a): 16, pl. 297, f. A, TL: Surinam
= Heliconius erato erato × erato hydara ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 178, pl. 39, f. 393
= ; [NL4A, #2407a (hyb.)]
Papilio udalrica Cramer, 1780; , TL: Surinam
= ; [NL4A, #2407a (hyb.)]
Heliconia cynisca Godart, 1819; , TL: French Guiana
= ; [NL4A, #2407a]
Heliconia vesta tellus Oberthür, 1902; pl. 2, f. 26, TL: French Guiana
= Heliconius erato erato × erato hydara ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 206, pl. 32, f. 330
= ; [NL4A, #2407a (hyb.)]
Heliconius erato oberthüri Riffarth, 1903; Berl. ent. Zs. 47 (3/4) : 162, TL: Surinam
= Heliconius erato erato × erato hydara ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 199, pl. 32, f. 326
= ; [NL4A, #2407a (hyb.)]
Heliconius erato erato f. fuliginosa Riffarth, 1907; Dt. ent. Zs. 1907 (5) : 511, TL: French Guiana,Surinam
= ; [NL4A, #2407a]
Heliconius erato ab. hemicycla Joicey & Kaye, 1917; Trans. ent. Soc. Lond. 1916 : 427, TL: French Guiana
= ; [NL4A, #2407a (hyb.)]
Heliconius erato ab. protea Joicey & Kaye, 1917; Trans. ent. Soc. Lond. 1916 : 427, TL: French Guiana
= Heliconius erato erato × erato hydara ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 201, pl. 33, f. 332
= ; [NL4A, #2407a (hyb.)]
Heliconius erato ab. constricta Joicey & Kaye, 1917; Trans. ent. Soc. Lond. 1916 : 427, TL: French Guiana
= Heliconius erato erato × erato hydara ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 184, pl. 32, f. 329
= ; [NL4A, #2407a (hyb.)]
f. albida Joicey & Kaye, 1919; , TL: French Guiana
= ; [NL4A, #2407a]
Heliconius erato ab. telloides Joicey & Talbot, 1925; Ann. Mag. Nat. Hist. (9) 16 (96) : 648, TL: French Guiana
= Heliconius erato erato × erato hydara ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 206, pl. 33, f. 331
= ; [NL4A, #2407a (hyb.)]
Heliconius erato erato f. roseoflava Neustetter, 1926; Int. ent. Z. 20 : 292, TL: French Guiana
= Heliconius erato erato × erato hydara ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 203, pl. 33, f. 333
= ; [NL4A, #2407a (hyb.)]
f. fumata Neustetter, 1926; , TL: French Guiana
= ; [NL4A, #2407a (hyb.)]
f. latiflava Neustetter, 1931; , TL: French Guiana
= ; [NL4A, #2407a (hyb.)]
f. nigrobasalis Neustetter, 1931; , TL: French Guiana
= ; [NL4A, #2407a (hyb.)]
Migonitis ulrica ; Hübner, 1816, (missp.)
Heliconius erato erato f. fuliginosa ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 190, pl. 32, f. 328
Heliconius erato erato f. hemicycla ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 192, pl. 32, f. 327
Heliconius erato erato ; [NL4A, #2407a]
H. e. phyllis (Fabricius, 1775)
Guianas, Peru, Bolivia, Brazil(Espírito Santo, Rio de Janeiro), Paraguay, Uruguay, Argentina Papilio phyllis Fabricius, 1775; Syst. Ent.: 463, TL: Brazil
Papilio roxane Cramer, 1775; , TL: "Surinam"
= ; [NL4A, #2407y]
Heliconius phyllidis Grose-Smith & Kirby, 1892; Rhop. Exot. [2] 1: (Heliconius) 2, pl. 1, f. 4, TL: Bolivia
= ; [NL4A, #2407y]
Heliconius anacreon Grose-Smith & Kirby, 1892; Rhop. Exot. [2] 1: (Heliconius) 3, pl. 1, f. 5, TL: Bolivia
= ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 178, pl. 34, f. 347; [NL4A, #2407y (hyb.)]
Heliconius amatus Staudinger, [1897]; Dt. Ent. Z. Iris 9 (2) : 301, TL: Bolivia
= ; [NL4A, #2407y (hyb.)]
Heliconius phyllis ab. artifex Stichel, 1899; Ent. Nachr. 25 (2) : 29, TL: Paraguay,Brazil, Espírito Santo
= ; [NL4A, #2407y]
Heliconius erato phyllis f. diffluens Riffarth, 1907; Dt. ent. Zs. 1907 (5) : 513, TL: "Peru",Bolivia
= ; [NL4A, #2407y]
Heliconius erato anacreon f. anaitis Riffarth, 1907; Dt. ent. Zs. 1907 (5) : 513, TL: Bolivia
= ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 178, pl. 34, f. 351; [NL4A, #2407y (hyb.)]
f. athene Neustetter, 1909; , TL: Bolivia
= ; [NL4A, #2407y]
Heliconius phyllis ab. miletus d'Almeida, 1928; , TL: Brazil, Rio de Janeiro
= ; [NL4A, #2407y]
Heliconius phyllis ab. cohaerens Hayward, 1931; , TL: Argentina
= ; [NL4A, #2407y]
Heliconius phyllis ab. alicia Schweizer & Kay, 1941; , TL: Uruguay
= ; [NL4A, #2407y]
Heliconius (Heliconius) erato phyllis ; Brown & Mielke, 1967, J. Lep. Soc. 21 (2): 96
Heliconius erato phyllis f. artifex ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 179, pl. 35, f. 353
Heliconius erato phyllis f. diffluens ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 186, pl. 34, f. 348
Heliconius erato phyllis f. phyllidis ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 201, pl. 34, f. 352
Heliconius erato f. phyllis ; [BOW: pl. 44, f. 26]
Heliconius erato phyllis ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 201, pl. 34, f. 349; [EBW]; [NL4A, #2407y]
H. e. cyrbia Godart, 1819
Ecuador, Colombia cyrbia (Godart, 1819) (Heliconia); Encyclopédie Méthodique. 9 (1): 203, TL: America [West Ecuador]
Heliconius cyrbia ab. diformata Riffarth, 1900; Berl. ent. Zs. 45 (3/4) : 209, TL: Ecuador, "Quito?"
= Heliconius erato cyrbia × erato venus ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 186, pl. 29, f. 297
= ; [NL4A, #2407i (hyb.)]
Heliconius cyrbia cyrbia f. bella Riffarth, 1907; Dt. ent. Zs. 1907 (5) : 511, TL: Ecuador
= ; [NL4A, #2407i]
Heliconius erato cyrbia f. bella ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 181, pl. 29, f. 298
Heliconius cyrbia ; [BOW: pl. 44, f. 6]
Heliconius erato cyrbia ; [EBW]; [EcuL]; [NL4A, #2407i]
H. e. demophoon Ménétriés, 1855
Nicaragua Heliconius demophoon Ménétriés, 1855; Cat. lep. Petersb. 2: 86, 1: pl. 2, f. 4, TL: Nicaragua
Heliconius chiriquensis Riffarth, 1900; Berl. ent. Zs. 45 (3/4) : 209 (nom. nud.)
= ; [NL4A, #2407j]
Heliconius erato demophon[sic] ; [EBW, (name only, missp.)]
Heliconius erato demophoon ; [NL4A, #2407j]
H. e. chestertonii Hewitson, 1872
Colombia Heliconia chestertonii Hewitson, 1872; Ill. exot. Butts [1] (Athesis): [86], pl. [44], f. 22, TL: Colombia
Heliconius damysus Hopffer, 1874; Stettin ent. Ztg 35 (10-12) : 349, TL: "Bolivia"
= ; [NL4A, #2407f]
Heliconius molina Grose-Smith, 1898; Ann. Mag. nat. Hist. (7) 2 (7) : 70, TL: Colombia
= Heliconius erato chestertonii × erato hydara ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 197, pl. 29, f. 295
= ; [NL4A, #2407f]
Heliconius hydara ab. nocturna Riffarth, 1900; Berl. ent. Zs. 45 (3/4) : 210, TL: "Venezuela"
= Heliconius erato chestertonii × erato hydara ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 198, pl. 29, f. 296
= ; [NL4A, #2407f (hyb.)]
Heliconius erato f. extrema Kaye, 1919; (nom. nud.), TL: Colombia
= ; [NL4A, #2407f]
Heliconius erato extrema Kaye, 1919; , TL: Colombia
= ; [NL4A, #2407f (hyb.)]
Heliconius erato chestertonii ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 183, pl. 29, f. 294; [EBW]; [NL4A, #2407f]
H. e. petiverana Doubleday, 1847
Mexico - Panama, Honduras Heliconia petiverana Doubleday, 1847; Gen. diurn. Lep. (1): 103, TL: Mexico
Heliconia mexicana Boisduval, 1870; Considérations Lépid. Guatemala: 28, TL: Mexico, Veracruz; Honduras
= ; [NL4A, #2407x]
Heliconius petiverea Riffarth, 1901; (nom. nud.)
= ; [NL4A, #2407x]
Heliconius erato petiverana, Listing of Mexican Butterflies [ILMB]
Heliconius erato petiverana, Butterflies of America [BOA]
Heliconius petiveranus ; Godman & Salvin, [1881], Biol. centr.-amer., Lep. Rhop. 1: 153; Godman & Salvin, [1901], Biol. centr.-amer., Lep. Rhop. 2: 668
Heliconius erato petiveranus ; [NACL, #4419a]; [BCR, 195]
Heliconius erato petiverana ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 200, pl. 29, f. 292; [EBW]
Heliconius petiverana ; [BOW: pl. 44, f. 25]
Heliconius erato petiverana ; [NL4A, #2407x]

Larva on Passiflora talamancensis, P. coreacea, P. biflora [BCR]

H. e. estrella Bates, 1862
Brazil(Pará) Heliconius estrella Bates, 1862; Trans. linn. Soc. Lond. 23 (3) : 560, TL: Brazil, Pará
Heliconius erato estrella ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 188, pl. 33, f. 334; [EBW, (name only)]; [NL4A, #2407m]
H. e. hydara Hewitson, 1867
Panama, Colombia, Surinam, Brazil(Amazonas, Pará) Heliconia hydara Hewitson, 1867; Ill. exot. Butts [1] (Heliconia V): [77], pl. [40], f. 14, TL: Colombia
Papilio callycopis Cramer, [1777]; Uitl. Kapellen 2 (9-16): 143, pl. 190, f. E, F, TL: Surinam
= Heliconius erato hydara × erato amalfreda ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 182, pl. 39, f. 394
= ; [NL4A, #2407q (hyb.)]
Papilio carolina Herbst, 1790
= ; [NL4A, #2407q (artifact)]
Heliconius coralii Butler, 1877; Cistula Ent. 2 : 151, TL: Brazil, Amazonas
= ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 184, pl. 32, f. 324; [NL4A, #2407q (hyb.)]
Heliconius erato f. palantia Möschler, 1883; , TL: Surinam
= ; [NL4A, #2407q (hyb.)]
Heliconius cyrbia var. juno Riffarth, 1900; Berl. ent. Zs. 45 (3/4) : 209, TL: ?
= Heliconius erato hydara f. guarica ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 194, pl. 31, f. 317
= ; [NL4A, #2407q (hyb.)]
Heliconius petiverana ab. tristis Riffarth, 1900; Berl. ent. Zs. 45 (3/4) : 210, TL: Panama, Chiriqui
= Heliconius erato hydara × erato petiverana ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 207, pl. 31, f. 315
= ; [NL4A, #2407q (hyb.)]
Heliconius phyllis var. viculata Riffarth, 1900; Berl. ent. Zs. 45 (3/4) : 211, TL: Surinam, "Paricatuba, Santarem, Obidos, Porto Cabello"
= ; [NL4A, #2407q (hyb.)]
Heliconius phyllis ab. callista Riffarth, 1900; Berl. ent. Zs. 45 (3/4) : 212, TL: Surinam
= Heliconius erato hydara f. callycopis ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 182, pl. 31, f. 320
= ; [NL4A, #2407q (hyb.)]
Heliconius phyllis ab. dryope Riffarth, 1900; Berl. ent. Zs. 45 (3/4) : 212, TL: Surinam
= ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 186, pl. 32, f. 323; [NL4A, #2407q (hyb.)]
Heliconius palantes Riffarth, 1900; (nom. nud.)
= ; [NL4A, #2407q]
Heliconius erato callycopis f. belticopis Joicey & Kaye, 1917; , TL: Surinam
= ; [NL4A, #2407q (hyb.)]
Heliconius erato rubrizona Joicey & Kaye, 1917; Ann. Mag. nat. Hist. (8) 20 (115) : 93, pl. 6, f. 7, TL: Brazil, Pará
= ; [NL4A, #2407q (hyb.)]
?Heliconius petiverana f. viculata ; [BOW: pl. 44, f. 25 (text only)]
Heliconius erato hydara f. rubrizona ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 203, pl. 31, f. 319
Heliconius erato hydara f. viculata ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 208, pl. 31, f. 318
Heliconius erato hydara ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 192, pl. 31, f. 316; [EBW]; [NL4A, #2407q]
H. e. guarica Reakirt, 1868
Colombia, Venezuela Heliconius guarica Reakirt, 1868; Proc. Acad. nat. Sci. Philad. 20 (1/2) : 91, TL: Colombia
Heliconius guayana Herrich-Schäffer, 1865; (nom. nud.)
= ; [NL4A, #2407p]
Heliconia euryas Boisduval, 1870; Considérations Lépid. Guatemala: 29, TL: [Venezuela]
= ; [NL4A, #2407p]
Heliconius erato guarica ; [NL4A, #2407p]
H. e. notabilis Salvin & Godman, 1868
Ecuador Heliconius notabilis Salvin & Godman, 1868; 145, TL: Ecuador, Canelos
Heliconius erato estrella f. beata Riffarth, 1907; Dt. ent. Zs. 1907 (5) : 512, pl. 5, f. 14, TL: Ecuador
= Heliconius erato notabilis × erato lativitta ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 180, pl. 30, f. 304
= ; [NL4A, #2407w (hyb.)]
Heliconius erato simplex f. rosacea Riffarth, 1907; Dt. ent. Zs. 1907 (5) : 512, pl. 5, f. 14, TL: Ecuador
= ; [NL4A, #2407w (hyb.)]
Heliconius erato estrella f. ilia Niepelt, 1908; Dt. ent. Zs. 1908 : 506, TL: Ecuador
= Heliconius erato notabilis × erato lativitta ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 193, pl. 30, f. 307
= ; [NL4A, #2407w (hyb.)]
Heliconius erato estrella f. rothschildi Niepelt, 1909; Berl. ent. Zs. 54 : 107, TL: Ecuador
= Heliconius erato × notabilis × erato lativitta ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 203, pl. 30, f. 305
= ; [NL4A, #2407w (hyb.)]
Heliconia microclea notabilis radiata Oberthür, 1916; Étud. Lépid. Comp. 12 (2) : 33, pl. 404, f. 3479, TL: Ecuador, Guayaquil
= Heliconius erato notabilis × erato lativitta ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 202, pl. 30, f. 303
= Heliconius erato lativitta ; [EcuL]
= ; [NL4A, #2407w (hyb.)]
Heliconius notabilis rubrescens Röber, 1921; , TL: Ecuador
= ; [NL4A, #2407w (hyb.)]
Heliconius microclea notabilis f. flavopunctatus Neustetter, 1925; , TL: Ecuador
= ; [NL4A, #2407w (hyb.)]
Heliconius erato notabilis f. rosacea ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 202, pl. 30, f. 302
Heliconius erato notabilis ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 198, pl. 30, f. 301; [EBW]; [EcuL]; [NL4A, #2407w]
H. e. etylus Salvin, 1871
Ecuador Heliconius etylus Salvin, 1871; Ann. Mag. nat. Hist. (4) 4 (21) 7 (42): 414, TL: Ecuador, Guadalquiza
= Heliconius erato lativitta × erato notabilis ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 188, pl. 30, f. 308
Heliconius erato etylus ab. insignis Kaye, 1916; Ent. Rec. J. Var. 28 : 194, TL: Ecuador
= ; [NL4A, #2407n (hyb.)]
Heliconius erato etylus ; [EcuL]; [NL4A, #2407n]
H. e. venustus Salvin, 1871
Bolivia Heliconius venustus Salvin, 1871; Ann. Mag. nat. Hist. (4) 4 (21) 7 (42): 413, TL: Bolivia
Heliconia anactorie Doubleday, 1847; 103, pl. 15, f. 4, TL: Bolivia
= Heliconius erato venustus × erato phyllis ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 178, pl. 33, f. 341
= ; [NL4A, #2407ac (hyb.)]
Heliconius anactorie ab. sanguineus Staudinger, 1894; , TL: Bolivia
= ; [NL4A, #2407ac (hyb.)]
Heliconius phyllis ab. ottonis Riffarth, 1900; Berl. ent. Zs. 45 (3/4) : 212, TL: Bolivia, Chiquitos, Bueyes
= ; [NL4A, #2407ac (hyb.)]
Heliconius locris Riffarth, 1901; (nom. nud.)
= ; [NL4A, #2407ac]
Heliconius erato diva Stichel, 1906; Genera Insectorum 37 : 47, TL: Bolivia
= Heliconius erato venustus × erato lativitta ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 186, pl. 33, f. 340
= ; [NL4A, #2407ac (hyb.)]
Heliconius phyllis ab. confluens Seitz, 1913; , TL: Bolivia
= ; [NL4A, #2407ac (hyb.)]
Heliconius phyllis kruegeri f. anacreonides Neustetter, 1925; Int. ent. Z. 19 : 60, TL: Bolivia
= Heliconius erato venustus × erato phyllis ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 178, pl. 34, f. 346
= ; [NL4A, #2407ac (hyb.)]
Heliconius phyllis f. krügeri Neustetter, 1925; Int. ent. Z. 19 : 60, TL: Bolivia
= Heliconius erato venustus × erato phyllis ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 194, pl. 34, f. 344
= ; [NL4A, #2407ac (hyb.)]
Heliconius phyllis f. leonora Krüger, 1927; Int. ent. Z. 21: 86, TL: Bolivia
= Heliconius erato venustus × erato phyllis ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 195, pl. 34, f. 343
= ; [NL4A, #2407ac (hyb.)]
Heliconius phyllis anacreon f. clelia Neustetter, 1927; , TL: Bolivia
= Heliconius erato venustus × erato phyllis ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 183, pl. 34, f. 345
= ; [NL4A, #2407ac (hyb.)]
Heliconius phyllis anactorie f. henrici Krüger, 1929; Int. ent. Z. 22 : 376, TL: Bolivia
= Heliconius erato venustus × erato phyllis ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 192, pl. 34, f. 342
= ; [NL4A, #2407ac (hyb.)]
Heliconius phyllis venustus f. pseudoleda Neustetter, 1932; , TL: Bolivia
= ; [NL4A, #2407ac (hyb.)]
Heliconius xanthocles f. denieri Hayward, 1939; , TL: Bolivia
= ; [NL4A, #2407ac]
Heliconius erato venustus ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 208, pl. 33, f. 339; [NL4A, #2407ac]; [EBW, (name only)]
H. e. favorinus (Hopffer, 1874)
Peru Heliconius favorinus Hopffer, 1874; Stettin ent. Ztg 35 (10-12) : 348, TL: "Bolivia"
Heliconius amaryllis var.? pseudamaryllis Staudinger, [1897]; Dt. Ent. Z. Iris 9 (2) : 297, TL: Peru
= ; [NL4A, #2407o]
Heliconius erato eratophylla Joicey & Kaye, 1917; Ann. Mag. nat. Hist. (8) 20 (115) : 91, pl. 6, f. 2, TL: Peru
= Heliconius erato favorinus × erato lativitta ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 187, pl. 29, f. 299
= ; [NL4A, #2407o (hyb.)]
Heliconius favorinus pseudoanacreon Neustetter, 1932; , TL: Peru
= ; [NL4A, #2407o (hyb.)]
Heliconius erato favorinus ; [EBW, (name only)]; [NL4A, #2407o]
H. e. lativitta Butler, 1877
Brazil(Amazonas), Ecuador, Colombia Heliconius lativitta Butler, 1877; Cistula Ent. 2 : 150, TL: "Ega, Guayaquil, Rio Madeira"
Heliconius vesta var. vestalis Staudinger, 1885; , TL: [Brazil, Amazonas]
= ; [NL4A, #2407r]
Heliconius erato estrella f. ochracea Riffarth, 1907; Dt. ent. Zs. 1907 (5) : 511, pl. 5, f. 12, TL: Ecuador
= ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 199, pl. 30, f. 306; [NL4A, #2407r (hyb.)]
Heliconius erato estrella f. feyeri Niepelt, 1908; Dt. ent. Zs. 1908 : 506, TL: Ecuador
= ; [NL4A, #2407r (hyb.)]
Heliconius erato estrella f. anactorina Stichel, 1923; , TL: Colombia
= ; [NL4A, #2407r (hyb.)]
Heliconius erato estrella f. sanguinella Stichel, 1923; , TL: Colombia
= ; [NL4A, #2407r (hyb.)]
Heliconius erato estrella f. perplexa Stichel, 1923; , TL: Colombia
= ; [NL4A, #2407r (hyb.)]
Heliconius microclea beata f. zoraida Neustetter, 1925; , TL: Ecuador
= ; [NL4A, #2407r (hyb.)]
Heliconius erato estrella f. aurivillii Bryk, 1953; , TL: Brazil, Amazonas
= ; [NL4A, #2407r (hyb.)]
Heliconius erato lativitta ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 194, pl. 31, f. 311; [EBW, (name only)]; [EcuL]; [NL4A, #2407r]
H. e. venus Staudinger, 1882
Colombia Heliconius venus Staudinger, 1882; Proc. zool. Soc. Lond. 1882 (3) : 396, pl. 24, f. 2, TL: Colombia
Heliconius erato venus ; [EBW]; [NL4A, #2407ab]
H. e. amazona Staudinger, 1897
Brazil(Pará) Heliconius vesta var. amazona Staudinger, [1897]; Dt. Ent. Z. Iris 9 (2) : 306, TL: Brazil, Pará
Heliconius philadelphus Kirby, [1900]; , TL: Brazil
= ; [NL4A, #2407d]
Heliconius phyllis andremona ab. androdaixa Seitz, 1912; 393, TL: [Brazil, Pará]
= Heliconius erato amazona × erato hydara ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 178, pl. 33, f. 335
= ; [NL4A, #2407d (hyb.)]
Heliconius erato amazona ; [NL4A, #2407d]
H. e. colombina Staudinger, 1897
Colombia Heliconius petiveranus var.? colombina Staudinger, [1897]; Dt. Ent. Z. Iris 9 (2) : 295, TL: Colombia
Heliconius hydara var. antigona Riffarth, 1900; Berl. ent. Zs. 45 (3/4) : 210, TL: Colombia, Medellin, Rio Dagua
= ; [NL4A, #2407g]
Heliconius erato demophoon f. antigona ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 178, pl. 29, f. 293
Heliconius erato colombina ; [NL4A, #2407g]
H. e. magnifica Riffarth, 1900
Guyana Heliconius phyllis var. magnifica Riffarth, 1900; Berl. ent. Zs. 45 (3/4) : 211, TL: Guyana
Heliconius erato magnifica ; [NL4A, #2407u]
H. e. amalfreda Riffarth, 1900
Guyana, French Guiana, Surinam, Brazil(Pará, Amazonas, Pará, Roraima) Heliconius phyllis ab. amalfreda Riffarth, 1900; Berl. ent. Zs. 45 (3/4) : 212, TL: Brazil, Pará
Heliconia elimaea Erichson, [1849]; , TL: Guyana
= ; [NL4A, #2407c (hyb.)]
Heliconius cybele ab. cybelina Staudinger, [1897]; Dt. Ent. Z. Iris 9 (2) : 304, pl. 7, f. 2, TL: Brazil, Amazonas
= ; [NL4A, #2407c]
Heliconius vesta ab. leda Staudinger, [1897]; Dt. Ent. Z. Iris 9 (2) : 306, TL: French Guiana
= ; [NL4A, #2407c (hyb.)]
Heliconius phyllis ab. amalfreda Riffarth, 1900; Berl. ent. Zs. 45 (3/4) : 212, TL: Manicoré, Faro, Manaos, Obidos, Berg en Dal, Surin
= ; [NL4A, #2407c]
Heliconius erato cybelinus f. helena Riffarth, 1907; Dt. ent. Zs. 1907 (5) : 511, TL: Brazil, Pará,Surinam
= Heliconius erato amalfreda × erato hydara ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 191, pl. 32, f. 321
= ; [NL4A, #2407c (hyb.)]
Heliconius erato erato ab. cybelellus Joicey & Kaye, 1917; Ann. Mag. nat. Hist. (8) 20 (115) : 92, pl. 6, f. 4, TL: Brazil, Amazonas
= Heliconius erato amalfreda × erato hydara ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 185, pl. 32, f. 325
= ; [NL4A, #2407c (hyb.)]
Heliconius melpomene f. pyritosa Zikán, 1937; , TL: Brazil, Pará
= ; [NL4A, #2407c]
Heliconius erato andremona f. juanita Neustetter, 1938; , TL: Brazil, Roraima
= ; [NL4A, #2407c (hyb.)]
Heliconius f. heydei Stammeshaus, 1962; , TL: Surinam
= ; [NL4A, #2407c]
Heliconius erato amalfreda ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 177, pl. 32, f. 322; [EBW]; [NL4A, #2407c]
H. e. amphitrite Riffarth, 1901
Peru Heliconius phyllis amphitrite Riffarth, 1901; 157, TL: Peru
Heliconius erato estrella f. simplex Riffarth, 1906; Insekten-Börse 23 (14): 56, TL: Peru
= ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 204, pl. 33, f. 336; [NL4A, #2407e (hyb.)]
Heliconius erato phyllis f. sperata Riffarth, 1907; Dt. ent. Zs. 1907 (5) : 513, TL: Peru,"Bolivia"
= Heliconius erato phyllis × erato venustus ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 204, pl. 34, f. 350
= ; [NL4A, #2407e (hyb.)]
Heliconius melpomene hyperplea Dyar, 1913; Proc. U.S. natn. Mus. 45 (2006) : 631, TL: Peru, Paltaymbamba, 5000ft
= ; [NL4A, #2407e]
Heliconius erato amphitrite ab. unipuncta Joicey & Kaye, 1917; Ann. Mag. nat. Hist. (8) 20 (115) : 93, pl. 6, f. 8, TL: Peru
= ; [NL4A, #2407e]
Heliconius erato amphitrite f. unipuncta ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 208, pl. 33, f. 338
Heliconius erato amphitrite ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 178, pl. 33, f. 337; [EBW, (name only)]; [NL4A, #2407e]
H. e. dignus Stichel, 1923
Colombia erato dignus Stichel, 1923; , TL: Colombia
Heliconius dignus f. discerpta Stichel, 1923; , TL: Colombia
= ; [NL4A, #2407k (hyb.)]
Heliconius erato estrella f. problemata Neustetter, 1928; Int. ent. Zs. 22 : 78, TL: Colombia
= Heliconius erato dignus × erato lativitta ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 201, pl. 31, f. 314
= ; [NL4A, #2407k (hyb.)]
Heliconius estrella f. meliorina Neustetter, 1928; Int. ent. Zs. 22 : 79, TL: Colombia
= Heliconius erato dignus × erato lativitta ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 196, pl. 31, f. 313
= ; [NL4A, #2407k (hyb.)]
Heliconius estrella f. glaucina Neustetter, 1928; Int. ent. Zs. 22 : 80, TL: Colombia
= Heliconius erato dignus × erato lativitta ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 191, pl. 31, f. 312
= ; [NL4A, #2407k (hyb.)]
Heliconius dignus f. elvira Niepelt, 1928; , TL: Columbia
= ; [NL4A, #2407k (hyb.)]
Heliconius erato meliorina ; [EBW, (name only)]
Heliconius erato dignus ; [EBW, (name only)]; [NL4A, #2407k]
H. e. emma Riffarth, 1901
Peru Heliconius phyllis emma Riffarth, 1901; Berl. ent. Zs. 46 (1/2) : 164, TL: "Sarayacu, Pachitea, Jurimaguas Peru, Archidona"
Heliconius augusta Riffarth, 1901; Berl. ent. Zs. 46 (1/2) : 164 (nom. nud.)
= ; [NL4A, #2407l]
Heliconius erato estrella f. palmata Stichel, 1906; Genera Insectorum 37 : 46, TL: Peru
= ; [NL4A, #2407l (hyb.)]
Heliconius erato estrella f. agnata Stichel, 1906; Genera Insectorum 37 : 46, TL: Peru
= ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 176, pl. 29, f. 300; [NL4A, #2407l (hyb.)]
Heliconius erato lativitta f. palmata ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 199, pl. 30, f. 310
Heliconius erato emma ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 187, pl. 30, f. 309; [EcuL]; [NL4A, #2407l]
H. e. microclea Kaye, 1907
Peru erato microclea Kaye, 1907; , TL: Peru
f. microfluens Descimon & Mast, 1971; , TL: Peru
= ; [NL4A, #2407v (hyb.)]
Heliconius erato microlea[sic] ; [EBW, (name only, missp.)]
Heliconius erato microclea ; [NL4A, #2407v]
Brazil(Amazonas) erato reductimacula Bryk, 1953; , TL: Brazil, Amazonas
f. venusta Bryk, 1953; , TL: Brazil, Amazonas
= ; [NL4A, #2407z]
Heliconius erato reductimacula ; [EBW, (name only)]
?Heliconius erato f. venusta ; [BOW: pl. 44, f. 10 (text only)]
Heliconius erato reductimacula ; [NL4A, #2407z]
H. e. adana Turner, 1967
Trinidad erato adana Turner, 1967; , TL: [Trinidad]
Heliconius hydara ab. adana Seitz, 1913; , TL: [Trinidad]
= ; [NL4A, #2407b]
Heliconius hydarus hydarus f. vitellina Stichel, 1919; , TL: Trinidad
= ; [NL4A, #2407b]
Heliconius erato adana ; [NL4A, #2407b]
H. e. fuscombei Lamas, 1976
Peru erato fuscombei Lamas, 1976; , TL: Peru
Heliconius erato fuscombei ; [NL4A, #2407t]
H. e. tobagoensis Barcant, 1982
Tobago erato tobagoensis Barcant, 1982; , TL: Tobago
Heliconius erato tobagoensis ; [NL4A, #2407aa]
H. e. lichyi Brown & Fernández, 1985
Venezuela erato lichyi Brown & Fernández, 1985; , TL: Venezuela
Heliconius erato lichyi ; [NL4A, #2407s]
H. e. cruentus Lamas, 1998
Mexico Heliconius erato cruentus Lamas, 1998; , TL: Mexico, Chiapas
Heliconius erato punctata Beutelspacher, 1992; (preocc. Neustetter, 1907), TL: [Mexico, Jalisco]
= ; [NL4A, #2407h]
Heliconius erato cruentus, Listing of Mexican Butterflies [ILMB]
Heliconius erato cruentus, Butterflies of America [BOA]
Heliconius erato cruentus ; [NL4A, #2407h]

Heliconius ethilla Godart, 1819
-. See [About maps] Heliconia ethilla Godart, 1819; , TL: "Antilles"
Heliconius ethilla, NSG Voucher Specimen [Wahlberg]
Heliconius ethilla ; [NL4A, #2408]
Venezuela, Trinidad Heliconius metalilis var. flavidus Weymer, 1894; Dt. ent. Z. Iris 6 (2) : 302, TL: Venezuela
= ; [NL4A, #2408a]
Heliconius ethilla metalilis f. depuncta Boullet & Le Cerf, 1909; , TL: Trinidad
= ; [NL4A, #2408a]
Heliconius flavidus ; [EBW]
Heliconius ethilla ethilla ; [NL4A, #2408a]
H. e. narcaea Godart, 1819
Brazil(Rio de Janeiro, ...?) narcaea (Godart, 1819) (Heliconia); Encyclopédie Méthodique. 9 (1): 217, TL: "Antilles"
Mechanites[sic] ecrate Hübner, [1823]; Samml. exot. Schmett. 2 : pl. 1
= ; [NL4A, #2408p]
Heliconius satis Weymer, 1875; Stettin ent. Ztg 36 (7/9) : 380, pl. 1, f. 6, TL: Brazil
= ; [NL4A, #2408p]
Heliconius eucrate var. infuscata Staudinger, 1885; , TL: Brazil
= ; [NL4A, #2408p]
Heliconius narcaea ab. brunnescens Neustetter, 1907; , TL: Brazil, Rio de Janeiro
= ; [NL4A, #2408p]
Heliconius connexa Seitz, 1912; , TL: Brazil, Rio de Janeiro
= ; [NL4A, #2408p]
Heliconius (Heliconius) ethillus narceus[sic, recte narcaea] ; Brown & Mielke, 1967, J. Lep. Soc. 21 (3): 96
Heliconius ethilla narcaea ; [EBW]
Heliconius narcaea ; [BOW: pl. 44, f. 21]
Heliconius ethilla narcaea ; [NL4A, #2408p]
H. e. eucoma (Hübner, [1831])
Peru - Panama Eueides eucoma Hübner, [1831]; Zuträge Samml. exot. Schmett. 3 : 38, : f. 577-578, TL: Brazil
Eueides eucoma Hübner, 1816; (nom. nud.)
= ; [NL4A, #2408g]
Heliconius ethilla eucoma ; [NL4A, #2408g]; [EBW, (name only)]
Heliconius eucoma ; [BOW: pl. 44, f. 16 (text only)]
H. e. aerotome C. & R. Felder, 1862
Heliconius aerotome C. & R. Felder, 1862; Wien. ent. Monats. 6 (3): 79, TL: "Brazil, Amazonas"
Heliconius ethilla aerotome ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 176, pl. 17, f. 172; [EBW, (name only)]; [NL4A, #2408c]
H. e. cephallenia C. & R. Felder, [1865]
Surinam Heliconius cephallenia C. & R. Felder, [1865]; Reise Fregatte Novara, Bd 2 (Abth. 2) (2): 373, TL: Surinam
Heliconius ethilla cephallenia ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 183, pl. 17, f. 170; [NL4A, #2408d]
H. e. polychrous C. & R. Felder, [1865]
Brazil(São Paulo) Heliconius polychrous C. & R. Felder, [1865]; Reise Fregatte Novara, Bd 2 (Abth. 2) (2): 375, pl. 47, f. 7, TL: Brazil, São Paulo
Heliconius physcoa Seitz, 1912; , TL: [Brazil, São Paulo]
= ; [NL4A, #2408u]
Heliconius ethilla polychrous ; [NL4A, #2408u]
H. e. metalilis Butler, 1873
Venezuela Heliconius metalilis Butler, 1873; Cist. Ent. 1 (7) : 167, TL: Venezuela
Heliconius ethilla metalilis ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 197, pl. 17, f. 169; [EBW, (name only)]; [NL4A, #2408n]
H. e. claudia Godman & Salvin, 1881
Panama Heliconius claudia Godman & Salvin, [1881]; Biol. centr.-amer., Lep. Rhop. 1: 145, TL: Panama, Calobre
Heliconius assimilis Röber, 1921; , TL: Panama
= ; [NL4A, #2408f]
Heliconius ethilla claudia ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 183, pl. 16, f. 168; [EBW, (name only)]; [NL4A, #2408f]
H. e. mentor Weymer, 1883
Colombia Heliconius mentor Weymer, 1883; Stettin ent. Ztg 45 (1-3) : 22, pl. 1, f. 3, TL: Colombia
Heliconius orchamus Weymer, 1912; Ent. Rdsch. 29 (11) : 73, TL: Colombia
= ; [NL4A, #2408m]
Heliconius ethilla mentor ; [NL4A, #2408m]
Brazil(Pará) Heliconius eucoma var. flavofasciatus Weymer, 1894; Dt. ent. Z. Iris 6 (2) : 302, TL: Brazil, Pará
Heliconius ethilla flavofasciatus ; [NL4A, #2408h]
Brazil(Pernambuco) Heliconius narcaea var. flavomaculatus Weymer, 1894; Dt. ent. Z. Iris 6 (2) : 341, TL: Brazil, Pernambuco
Heliconius ethilla flavomaculatus ; [EBW, (name only)]; [NL4A, #2408i]
H. e. numismaticus Weymer, 1894
Brazil(Pará) Heliconius eucoma var. numismaticus Weymer, 1894; Dt. ent. Z. Iris 6 (2) : 303, TL: Brazil, Pará
Heliconius ethilla numismaticus ; [NL4A, #2408s]
H. e. semiflavidus Weymer, 1894
Colombia Heliconius metalilis var. semiflavidus Weymer, 1894; Dt. ent. Z. Iris 6 (2) : 302, TL: Colombia
Heliconius eucoma var. daguanus Staudinger, [1897]; Dt. Ent. Z. Iris 9 (2) : 286, TL: Colombia
= ; [NL4A, #2408v]
Heliconius eucoma var. juntana Riffarth, 1900; Berl. ent. Zs. 45 (3/4) : 196, TL: Rio Juntas, Colombia
= ; [NL4A, #2408v]
Heliconius ethilla semiflavidus ; [EBW, (name only)]; [NL4A, #2408v]
H. e. tyndarus Weymer, 1897
Bolivia Heliconius tyndarus Weymer, 1897; Dt. ent. Z. Iris 9 (2) : 317, pl. 6, f. 6, TL: Bolivia
Heliconius ethilla tyndarus ; [EBW, (name only)]; [NL4A, #2408x]
H. e. thielei Riffarth, 1900
French Guiana Heliconius gradatus thielei Riffarth, 1900; Berl. ent. Zs. 45 (3/4) : 195, TL: French Guiana
Heliconius numatus isabellinus ab. fusca Boullet & Le Cerf, 1909; , TL: French Guiana
= ; [NL4A, #2408w]
Heliconius ethilla thielei ; [EBW, (name only)]; [NL4A, #2408w]
H. e. adela Neustetter, 1912
Peru ethilla adela Neustetter, 1912; , TL: Peru
Heliconius ethilla adela ; [EBW, (name only)]; [NL4A, #2408b]
H. e. nebulosa Kaye, 1916
Peru Heliconius quitalinus nebulosa Kaye, 1916; Ent. Rec. J. Var. 28 : 194, TL: SE.Peru, Yahuarmayao, 1200ft
Heliconius ethilla nebulosa ; [EBW, (name only)]; [NL4A, #2408q]
H. e. hyalina Neustetter, 1928
Brazil(Roraima) ethilla hyalina Neustetter, 1928; , TL: Brazil, Roraima
Heliconius ethilla hyalina ; [NL4A, #2408j]
H. e. chapadensis Brown, 1973
Brazil(Mato Grosso) Heliconius ethilla chapadensis Brown, 1973; Bull. Allyn Mus. 13: 6, TL: Brazil, Mato Grosso
Heliconius ethilla chapadensis ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 183, pl. 17, f. 171; [NL4A, #2408e]
H. e. jaruensis Brown, 1976
Brazil(Rondônia) ethilla jaruensis Brown, 1976; , TL: Brazil, Rondônia
Heliconius ethilla jaruensis ; [NL4A, #2408k]
H. e. yuruani Brown & Fernández, 1985
Venezuela ethilla yuruani Brown & Fernández, 1985; , TL: Venezuela
Heliconius ethilla yuruani ; [NL4A, #2408y]
Peru Heliconius ethilla michaelianius Lamas, 1988; (repl. name), TL: Peru
Heliconius aurotome[sic] f. clarus Michael, 1926; (preocc. Fabricius, 1793), TL: Peru
= ; [NL4A, #2408o]
Heliconius ethilla michaelianius ; [NL4A, #2408o]
H. e. penthesilea Neukirchen, 1994
Brazil(Amazonas) ethilla penthesilea Neukirchen, 1994; , TL: Brazil, Amazonas
Heliconius ethilla penthesilea ; [NL4A, #2408t]
H. e. latona Neukirchen, 1998
Colombia ethilla latona Neukirchen, 1998; , TL: Colombia
Heliconius ethilla latona ; [NL4A, #2408l]
H. e. neukircheni Lamas, 1998
Peru ethilla neukircheni Lamas, 1998; , TL: Peru
Heliconius ethilla neukircheni ; [NL4A, #2408r]

Heliconius hecale (Fabricius, 1775)
Mexico - Peruvian Amazon. See [About maps] Papilio hecale Fabricius, 1775; , TL: Surinam
?Papilio hecale Fabricius, 1777; Genera Insectorum : 254
Papilio urania Müller, 1774; Natursystem 5 (1): pl. 18, f. 2 (preocc. Papilio urania Linnaeus, 1758)
Heliconius hecale, NSG Voucher Specimen [Wahlberg]
Heliconius hecale ; [BOW: pl. 44, f. 12]; [EBW]; [NL4A, #2409]
Heliconius urania ; [EBW]

Copy of Tithocea hecalesia. [BOW]

Surinam, Guyana Papilio pasithoe Cramer, [1775]; Uitl. Kapellen 1 (1-7): 25, pl. 17, f. A-B (preocc. Linnaeus, 1767), TL: Surinam
= ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 200, pl. 39, f. 400-402; [NL4A, #2409a]
Heliconius pasithoë var. fulvescens Lathy, 1906; Proc. zool. Soc. Lond. 1906 : 452, pl. 34, f. 1, TL: Guyana
= ; [NL4A, #2409a]
Heliconius hecale hecale f. fulvescens ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 190, pl. 16, f. 159
Heliconius hecale hecale ; [NL4A, #2409a]
H. h. anderida Hewitson, 1853
Venezuela Heliconia anderida Hewitson, 1853; Ill. exot. Butts [1] (Heliconia I): [69], pl. [36], f. 2, TL: "Honduras" [Venezuela]
Heliconius zagora Riffarth, 1901; Berl. ent. Zs. 46 (1/2) : 74 (nom. nud.)
= ; [NL4A, #2409b]
Heliconius rebeli Neustetter, 1907; , TL: "Colombia"
= ; [NL4A, #2409b]
Heliconius anderida estebana Kaye, 1914; , TL: Venezuela
= ; [NL4A, #2409b]
Heliconius anderida ; [BOW: pl. 43, f. 36]
Heliconius hecale anderida ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 178, pl. 15, f. 151; [EBW, (name only)]; [NL4A, #2409b]
H. h. quitalena Hewitson, 1853
Bolivia, Peru, Ecuador Heliconia quitalena Hewitson, 1853; Ill. exot. Butts [1] (Heliconia I): [70], pl. [36], f. 3, TL: Ecuador, Quito
Heliconius quitalena ; [BOW: pl. 44, f. 27]
Heliconius hecale quitalenus ; [EcuL]
Heliconius hecale quitalena ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 202, pl. 15, f. 155; [EBW]; [NL4A, #2409v]
H. h. fornarina Hewitson, 1854
Guatemala Heliconia fornarina Hewitson, 1854; Ill. exot. Butts [1] (Heliconia II-III): [73], pl. [38], f. 9, TL: ["S. Amer."]
Heliconius anderida fornarina f. bouvieri Boullet & Le Cerf, 1909; , TL: Guatemala
= ; [NL4A, #2409j]
Heliconius anderida f. styx Niepelt, 1921; Int. Ent. Zs. 15 : 19, TL: Guatemala
= ; [NL4A, #2409j]
Heliconius hecale fornarina, Listing of Mexican Butterflies [ILMB]
Heliconius hecale fornarina, Butterflies of America [BOA]
Heliconius hecale styx ; [EBW, (name only)]
Heliconius fornarina ; Godman & Salvin, [1881], Biol. centr.-amer., Lep. Rhop. 1: 145; Godman & Salvin, [1901], Biol. centr.-amer., Lep. Rhop. 2: 667
Heliconius hecale fornarina ab. styx ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 205, pl. 13, f. 138
Heliconius hecale fornarina ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 190, pl. 13, f. 139; [EBW]
Heliconius anderida fornarina ; [BOW: pl. 43, f. 35 (text only)]
Heliconius hecale fornarina ; [NL4A, #2409j]
H. h. zuleika Hewitson, 1854
Nicaragua - Panama Heliconia zuleika Hewitson, 1854; Ill. exot. Butts [1] (Heliconia II-III): [74], pl. [38], f. 10, TL: Nicaragua
Heliconius jucundus Bates, 1864; Ent. mon. Mag. 1 (3): 56, TL: Panama
= ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 194, pl. 14, f. 146; [NL4A, #2409ac]
Heliconius xanthicus Bates, 1864; Ent. mon. Mag. 1 (3): 57, TL: Panama
= Heliconius jucundus ; Godman & Salvin, [1881], Biol. centr.-amer., Lep. Rhop. 1: 146
= ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 209, pl. 14, f. 147; [NL4A, #2409ac]
Heliconius chrysantis Godman & Salvin, [1881]; Biol. centr.-amer., Lep. Rhop. 1: 146, TL: Nicaragua
= Heliconius hecale zuleika × hecale fornarina ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 183, pl. 14, f. 141
= ; [NL4A, #2409ac]
Heliconius zuleika ab. albipunctata Riffarth, 1900; Berl. ent. Zs. 45 (3/4) : 199, TL: Panama, Chiriqui
= ; [NL4A, #2409ac]
Heliconius zuleika ab. dentata Neustetter, 1907; , TL: Panama
= ; [NL4A, #2409ac]
Heliconius hecale zuleika, Listing of Mexican Butterflies [ILMB]
Heliconius hecale zuleika, Butterflies of America [BOA]

?Heliconius hecale xanthicles[sic] ; [EBW, (name only, missp.?)]
Heliconius hecale zuleica f. albipunctata ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 177, pl. 14, f. 142
Heliconius zuleika ; Godman & Salvin, [1881], Biol. centr.-amer., Lep. Rhop. 1: 147; Godman & Salvin, [1901], Biol. centr.-amer., Lep. Rhop. 2: 667
Heliconius xanthicus ; Godman & Salvin, [1881], Biol. centr.-amer., Lep. Rhop. 3: pl. 16, f. 10-11
Heliconius jucundus ; Godman & Salvin, [1881], Biol. centr.-amer., Lep. Rhop. 1: 146, 3: pl. 17, f. 1-2
Heliconius hecale jucundus ; [EBW, (name only)]
Heliconius hecale zuleika ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 209, pl. 13, f. 140; [BCR, 196]; [NL4A, #2409ac]

Larva on Passiflora oerstedii, P. vitifolia, P. auriculata, P. platyloba [BCR]

H. h. ithaca C. & R. Felder, 1862
Colombia Heliconius ithaca C. & R. Felder, 1862; Wien. ent. Monats. 6 (12) : 418, TL: Colombia
Heliconius vittatus Butler, 1873; Cist. Ent. 1 (7) : 166, TL: Bogota, Colombia
= ; [NL4A, #2409m]
Heliconius marius Weymer, 1890; in Weymer & Maassen, Reisen Süd-America : 116, pl. 2, f. 1, TL: Colombia
= ; [NL4A, #2409m]
Heliconius ithaka[sic] f. cajetani Neustetter, 1909; , TL: [Colombia]
= ; [NL4A, #2409m]
Heliconius ithaca f. hero Weymer, 1912; Ent. Rdsch. 29 (11) : 74, TL: Colombia
= ; [NL4A, #2409m]
Heliconius aristiona indecisa Joicey & Kaye, 1917; Ann. Mag. nat. Hist. (8) 20 (115) : 91, TL: [Colombia]
= ; [NL4A, #2409m]
Heliconius ithaka ithaka[sic] f. sulphureofasciata Neustetter, 1925; , TL: Colombia
= ; [NL4A, #2409m]
Heliconius ithaka[sic] vittatus f. nigroapicalis Neustetter, 1925
= ; [NL4A, #2409m]
Heliconius hecale vittatus ; [EBW, (name only)]
Heliconius hecale ithaca f. indecisa ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 193, pl. 15, f. 153
Heliconius hecale ithaca f. vittatus ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 208, pl. 15, f. 154
Heliconius hecale ithaca ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 194, pl. 15, f. 152; [EBW, (name only)]; [NL4A, #2409m]
H. h. melicerta Bates, 1866
Panama, Colombia Heliconius melicerta Bates, 1866; Ent. mon. Mag. 3 (28): 87, TL: Panama; Colombia
Papilio clara Fabricius, 1793; Ent. Syst. 3 (1): 161 (preocc. Cramer, 1775), TL: "Surinam"
= ; [NL4A, #2409o]
Heliconius albucilla Bates, 1866; Ent. mon. Mag. 3 (28): 88, TL: Panama
= Heliconius hecale melicerta × hecale zuleika ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 177
= ; [NL4A, #2409o]
Heliconius etholea Riffarth, 1901; Berl. ent. Zs. 46 (1/2) : 72 (nom. nud.)
= ; [NL4A, #2409o]
Heliconius anderida melicerta f. zygia Riffarth, 1907; Dt. ent. Zs. 1907 (5) : 504, pl. 5, f. 5, TL: Colombia
= ; [NL4A, #2409o]
Heliconius anderida melicerta f. muzoënsis Neustetter, 1909; , TL: Colombia
= ; [NL4A, #2409o]
Heliconius albucilla ; Godman & Salvin, [1881], Biol. centr.-amer., Lep. Rhop. 1: 144, 3: pl. 16, f. 7-8, 9 (var.)
Heliconius melicerta ; Godman & Salvin, [1881], Biol. centr.-amer., Lep. Rhop. 1: 144, 3: pl. 16, f. 12-13
Heliconius hecale melicerta f. zygia ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 209, pl. 14, f. 144
Heliconius hecale melicerta ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 196, pl. 14, f. 143; [EBW, (name only)]; [NL4A, #2409o]
H. h. novatus Bates, 1867
Bolivia, Peru, Brazil(Pará, Maranhão, Pará) Heliconius novatus Bates, 1867; Trans. ent. Soc. Lond. (3) 5 : 539, TL: Brazil, Pará; Maranhão
Heliconius schulzi Riffarth, 1899; Berl. ent. Zs. 43 (3/4) : 405, TL: Brazil, Pará
= ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 203, pl. 16, f. 164; [NL4A, #2409s]
Heliconius schultzi Seitz, 1924; (emend.)
= ; [NL4A, #2409s]
Heliconius novartus[sic, recte novatus] ; [BOW: pl. 44, f. 22]
Heliconius hecale novatus ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 198, pl. 16, f. 163; [EBW]; [NL4A, #2409s]
H. h. sisyphus Salvin, 1871
Peru, Brazil(Amazonas) Heliconius sisyphus Salvin, 1871; Ann. Mag. nat. Hist. (4) 4 (21) 7 (42): 413, TL: Peru
Heliconius jonas Weymer, 1894; Dt. ent. Z. Iris 6 (2) : 307, TL: Brazil, Amazonas
= ; [NL4A, #2409y]
Heliconius hecale sisyphus ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 204, pl. 15, f. 158; [EBW, (name only)]; [NL4A, #2409y]
H. h. vetustus Butler, 1873
Guyanas, Lower Amazon Heliconius vetustus Butler, 1873; Cist. Ent. 1 (7) : 165, TL: Guyana
Heliconius clarissa Riffarth, 1901; (nom. nud.)
= ; [NL4A, #2409aa]
Heliconius vetustus ; [BOW: pl. 45, f. 1]
Heliconius hecale vetustus ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 208, pl. 16, f. 162; [EBW]; [NL4A, #2409aa]
Honduras Heliconius discomaculatus Weymer, 1891; Stettin ent. Ztg 51 (7-12) : 289, TL: Honduras
Heliconius hecale discomaculatus ; [NL4A, #2409g]
H. h. ennius Weymer, 1891
Brazil(Amazonas) Heliconius ennius Weymer, 1891; Stettin ent. Ztg 51 (7-12) : 283, TL: Brazil, Amazonas
Heliconius ennius ; [BOW: pl. 44, f. 27 (text only)]
Heliconius hecale ennuis[sic] ; [EBW, (name only, missp.)]
Heliconius hecale ennius ; [NL4A, #2409h]
H. h. felix Weymer, 1894
Peru Heliconius felix Weymer, 1894; Dt. ent. Z. Iris 6 (2) : 315, pl. 4, f. 10, TL: Peru
Heliconius felix var. concors Weymer, 1894; Dt. ent. Z. Iris 6 (2) : 317, pl. 4, f. 11, TL: Peru
= ; [NL4A, #2409i]
Heliconius versicolor Weymer, 1894; Dt. ent. Z. Iris 6 (2) : 317, pl. 4, f. 12, TL: "Brazil, Amazonas"
= ; [NL4A, #2409i]
Heliconius quitalena felix f. umbrina Neustetter, 1931; , TL: Peru
= ; [NL4A, #2409i (hyb.)]
Heliconius hecale felix ; [NL4A, #2409i]
H. h. metellus Weymer, 1894
Brazil(Pará) Heliconius metellus Weymer, 1894; Dt. ent. Z. Iris 6 (2) : 290, pl. 4, f. 1, TL: Brazil, Pará
Heliconius vetustus boyi Röber, 1923; , TL: Brazil, Pará
= ; [NL4A, #2409p]
Heliconius hecale metellus ; [NL4A, #2409p]
H. h. sulphureus Weymer, 1894
Brazil(Amazonas) Heliconius sulphureus Weymer, 1894; Dt. ent. Z. Iris 6 (2) : 311, pl. 4, f. 8, TL: Brazil, Amazonas
Heliconius quitalenus denticulatus Riffarth, 1907; , TL: Brazil, Amazonas
= ; [NL4A, #2409z]
Heliconius ?sulphureus subsulphureus Röber, 1919; (nom. dub.)
= ; [NL4A, #2409z]
Heliconius quitalenus denticulatus ; Fruhstorfer, 1907, Stettin ent. Ztg 68 (2): 219
Heliconius hecale sulphureus ; [NL4A, #2409z]
H. h. holcophorus Staudinger, [1897]
Colombia Heliconius semiphorus ab.? (var.?) holcophorus Staudinger, [1897]; Dt. Ent. Z. Iris 9 (2) : 285, pl. 6, f. 3, TL: Colombia
Heliconius metaphorus var. semiphorus Staudinger, [1897]; Dt. Ent. Z. Iris 9 (2) : 284, TL: Colombia
= ; [NL4A, #2409k]
Heliconius eucherius Weymer, 1906; Dt. ent. Z. Iris 19 (2) : 68, TL: Colombia
= ; [NL4A, #2409k]
Heliconius hecale holcophorus f. eucherius ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 188, pl. 14, f. 149
Heliconius hecale melicerta f. semiphorus ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 204, pl. 14, f. 145
Heliconius hecale semiphorus ; [EBW, (name only)]
Heliconius hecale holcophorus ; [NL4A, #2409k]
Brazil(Amazonas) Heliconius ennius var. nigrofasciatus Weymer, 1894; Dt. ent. Z. Iris 6 (2) : 306, pl. 4, f. 6, TL: Brazil, Amazonas
Heliconius hecale nigrofasciatus ; [EBW, (name only)]; [NL4A, #2409r]
H. h. annetta Riffarth, 1900
Colombia Heliconius clara var. annetta Riffarth, 1900; Berl. ent. Zs. 45 (3/4) : 198, TL: Colombia
Heliconius hecale annetta ; [NL4A, #2409c]
H. h. latus Riffarth, 1900
Brazil(Pará) Heliconius paraensis var. latus Riffarth, 1900; Berl. ent. Zs. 45 (3/4) : 197, TL: Itaituba
Heliconius gradatus xinguensis Neustetter, 1925; , TL: Brazil, Pará
= ; [NL4A, #2409n]
Heliconius hecale latus ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 194, pl. 16, f. 166; [EBW, (name only)]; [NL4A, #2409n]
H. h. paraensis Riffarth, 1900
Brazil(Pará) Heliconius paraensis Riffarth, 1900; Berl. ent. Zs. 45 (3/4) : 197, TL: Brazil, Pará
Heliconius hecale paraensis ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 200, pl. 16, f. 167; [NL4A, #2409t]
H. h. humboldti Neustetter, 1928
Peru Heliconius humboldti Neustetter, 1928; Int. ent. Zs. 21 : 442
Heliconius humboldti f. alexander Neustetter, 1928; Int. ent. Zs. 21 : 443, TL: Peru
= ; [NL4A, #2409l]
Heliconius hecale humboldti f. alexander ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 177, pl. 15, f. 157
Heliconius hecale humboldti ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 192, pl. 15, f. 156; [NL4A, #2409l]
H. h. clearei Hall, 1930
Guyana Heliconius hecale clearei Hall, 1930; Entomologist 63 : 278, TL: Guyana
Heliconius hecale clearei ; [EBW, (name only)]; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 183, pl. 16, f. 161; [NL4A, #2409f]
H. h. australis Brown, 1976
Ecuador Heliconius hecale australis Brown, 1976; 218, TL: West Ecuador: Manabi, Palmar 200m
Heliconius hecale australis ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 180, pl. 15, f. 150; [EcuL]; [NL4A, #2409d]
H. h. barcanti Brown, 1976
Venezuela Heliconius hecale barcanti Brown, 1976; 219, TL: Venezuela
Heliconius hecale barcanti ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 180, pl. 16, f. 160; [EBW, (name only)]; [NL4A, #2409e]
H. h. rosalesi Brown & Fernández, 1976
Venezuela hecale rosalesi Brown & Fernández, 1976; , TL: Venezuela
Heliconius hecale rosalesi ; [NL4A, #2409w]
H. h. shanki Lamas & Brown, 1976
Peru hecale shanki Lamas & Brown, 1976; , TL: Peru
Heliconius hecale shanki ; [NL4A, #2409x]
H. h. zeus Neukirchen, 1995
Bolivia hecale zeus Neukirchen, 1995; , TL: Bolivia
Heliconius felix var. concors Weymer, 1894
= ; [NL4A, #2409ab (part)]
Heliconius hecale zeus ; [NL4A, #2409ab]
H. h. naxos Neukirchen, 1998
Brazil(Pará) hecale naxos Neukirchen, 1998; , TL: Brazil, Pará
Heliconius hecale naxos ; [NL4A, #2409q]
H. h. paulus Neukirchen, 1998
Brazil(Pará) hecale paulus Neukirchen, 1998; , TL: Brazil, Pará
Heliconius hecale paulus ; [NL4A, #2409u]

Heliconius hecalesia Hewitson, 1854
Mexico - Colombia, Venezuela. See [About maps] Heliconia hecalesia Hewitson, 1854; Ill. exot. Butts [1] (Heliconia II-III): [71], pl. [37], f. 6, TL: Colombia
Heliconius hecalesia, NSG Voucher Specimen [Wahlberg]
Heliconius hecalesia ; [EBW]; [NL4A, #2410]
Colombia Heliconius hecalesia hecalesia ; [NL4A, #2410a]
H. h. formosus Bates, 1866
Costa Rica, Panama, Nicaragua Heliconius formosus Bates, 1866; Ent. mon. Mag. 3 (28): 87, TL: Panama
Heliconius formosus ; Godman & Salvin, [1881], Biol. centr.-amer., Lep. Rhop. 1: 148, 3: pl. 17, f. 7-8
Heliconius hecalesia formosus ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 190, pl. 35, f. 355; [BCR, 195]; [EBW]; [NL4A, #2410c]

Larva on Passiflora biflora, Passiflora lancearea [BCR]

H. h. octavia Bates, 1866
Guatemala Heliconius octavia Bates, 1866; Ent. mon. Mag. 3 (28): 86, TL: Guatemala
Heliconius hecalesia octavia, Listing of Mexican Butterflies [ILMB]
Heliconius hecalesia octavia, Butterflies of America [BOA]
Heliconius octavia ; Godman & Salvin, [1881], Biol. centr.-amer., Lep. Rhop. 1: 148, 3: pl. 17, f. 9-10
Heliconius hecalesia octavia ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 199, pl. 35, f. 354; [NL4A, #2410f]
H. h. gynaesia Hewitson, 1875
Heliconia gynaesia Hewitson, 1875; Ent. mon. Mag. 11 : 182
Heliconia gynaesia ; Hewitson, 1875, Ill. exot. Butts [1] (Heliconia VIII): [84], pl. [43], f. 28
Heliconius hecalesia gynaesia ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 191, pl. 35, f. 356; [NL4A, #2410d]
H. h. longarena Hewitson, 1875
Colombia Heliconia longarena Hewitson, 1875; Ent. mon. Mag. 11 : 182, TL: Colombia
Heliconius hecalesia ernestus Brown & Benson, 1975; , TL: Colombia
= ; [NL4A, #2410e (hyb.)]
Heliconia longarena ; Hewitson, 1875, Ill. exot. Butts [1] (Heliconia VIII): [84], pl. [43], f. 29
Heliconius hecalesia longarena ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 195, pl. 35, f. 357; [NL4A, #2410e]
H. h. eximius Stichel, 1923
Colombia hecalesia eximius Stichel, 1923; , TL: Colombia, Rio Jacula
Heliconius hecalesia eximus[sic] ; [EBW, (name only, missp.)]
Heliconius hecalesia eximius ; [NL4A, #2410b]
H. h. romeroi Brown & Fernández, 1985
Venezuela hecalesia romeroi Brown & Fernández, 1985; , TL: Venezuela
Heliconius hecalesia romeroi ; [NL4A, #2410g]

Heliconius hecuba Hewitson, 1858
Ecuador. See [About maps] Heliconia hecuba Hewitson, 1858; Ill. exot. Butts [1] (Heliconia IV): [75], pl. [39], f. 11, TL: New Granada
Heliconius hecuba, NSG Voucher Specimen [Wahlberg]
Heliconius hecuba ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 191, pl. 7, f. 72; [EBW]; [NL4A, #2411]
Colombia Heliconius hecuba intermedius Riffarth, 1907; Dt. ent. Zs. 1907 (5) : 509, pl. 5, f. 11, TL: Colombia
= Heliconius hecuba hecuba × hecuba cassandra ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 194, pl. 7, f. 75
= ; [NL4A, #2411a (hyb.)]
Heliconius hecuba hecuba ; [NL4A, #2411a]
H. h. cassandra C. & R. Felder, 1862
Colombia Heliconius cassandra C. & R. Felder, 1862; Wien. ent. Monats. 6 (12) : 419, TL: Colombia
Heliconius hecuba cassandra ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 182, pl. 7, f. 74; [NL4A, #2411c]
H. h. choarina Hewitson, 1872
Ecuador Heliconia choarina Hewitson, 1872; Ent. mon. Mag. 9 : 83, TL: Ecuador
Blanchardia dismorphia Buchecker, [1880]; (nom. nud.), TL: "Colombia"
= ; [NL4A, #2411d]
f. cacica Neustetter, 1928; Int. ent. Zs. 22 : 247, TL: Ecuador (Macas)
= ; [NL4A, #2411d]
Heliconia choarina ; Hewitson, 1873, Ill. exot. Butts [1] (Heliconia and Tithorea VII): [82], pl. [42], f. 24-25
Heliconius hecuba cacicus ; Brown, 1979; [EcuL]
Heliconius hecuba choarina ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 183, pl. 7, f. 77; [EcuL]; [NL4A, #2411d]
H. h. crispus Staudinger, 1885
Colombia hecuba crispus Staudinger, 1885; , TL: Colombia
Heliconius hecuba crispus ; [NL4A, #2411f]
H. h. tolima Fassl, 1912
Colombia Heliconius tolima Fassl, 1912; Ent. Rdsch. 29 (8) : 55, TL: Colombia
Heliconius hecuba tolima ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 207, pl. 7, f. 73; [NL4A, #2411j]
H. h. flava Brown, 1979
Ecuador hecuba flava Brown, 1979; , TL: Ecuador (Macas)
Heliconius hecuba choarinus f. flava Neustetter, 1928; Int. ent. Zs. 22 : 247, TL: Ecuador
= ; [NL4A, #2411g]
Heliconius hecuba choarina f. flava ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 189, pl. 7, f. 78
Heliconius hecuba flavus ; [EcuL]
Heliconius hecuba flava ; [NL4A, #2411g]
H. h. creusa H. & R. Holzinger, 1989
Colombia hecuba creusa H. & R. Holzinger, 1989; , TL: Colombia
Heliconius crispus var. crespinus Krüger, 1925; , TL: Colombia
= ; [NL4A, #2411e (hyb.)]
?Heliconius crispinus[sic] ; [EBW, (name only, missp.?)]
Heliconius hecuba crispus f. crespinus ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 185, pl. 7, f. 76
Heliconius hecuba creusa ; [NL4A, #2411e]
H. h. bonplandi Neukirchen, 1991
Ecuador hecuba bonplandi Neukirchen, 1991; , TL: Ecuador, Morona Santiago, Macas
= Heliconius hecuba cacicus ; [EcuL]
Heliconius hecuba bonplandi ; [NL4A, #2411b]
H. h. lamasi Neukirchen, 1991
Ecuador hecuba lamasi Neukirchen, 1991; , TL: Ecuador, Zamorea, Chinchipe, Rio Cristal 1000m
Heliconius hecuba lamasi ; [EcuL]; [NL4A, #2411h]
H. h. salazari Neukirchen, 1993
Colombia hecuba salazari Neukirchen, 1993; , TL: Colombia
Heliconius hecuba salazari ; [NL4A, #2411i]
H. h. walteri Salazar, 1998
Colombia hecuba walteri Salazar, 1998; , TL: Colombia
Heliconius hecuba walteri ; [NL4A, #2411k]

Heliconius hermathena Hewitson, 1854
-. See [About maps] Heliconia hermathena Hewitson, 1854; Ill. exot. Butts [1] (Heliconia II-III): [71], pl. [37], f. 5, TL: [Brazil]
Heliconius hermathena, NSG Voucher Specimen [Wahlberg]
Heliconius hermathena ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 192, pl. 29, f. 290; [NL4A, #2412]; [BOW: pl. 44, f. 13]
Heliconius hermathena hermathena ; [NL4A, #2412a]
H. h. vereatta Stichel, 1912
Brazil(Pará) Heliconius hermathena f. vereatta Stichel, 1912; Int. ent. Z. 6 (1) : 1, TL: Brazil, Pará
Heliconius hermathena f. hydarina Stichel, 1912; Int. ent. Z. 6 (1) : 1, TL: Brazil, Pará
= ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 193, pl. 29, f. 289; [NL4A, #2412f]
Heliconius hermathena vereatta ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 208, pl. 28, f. 288; [EBW, (name only)]; [NL4A, #2412f]
H. h. duckei Brown & Benson, 1977
Brazil(Pará, ...) hermathena duckei Brown & Benson, 1977; , TL: Brazil, Pará
Heliconius hermathena f. rubropunctata D'Abrera, 1984; (nom. nud.), TL: Brazil
= ; [NL4A, #2412b]
Heliconius hermathena duckei ; [NL4A, #2412b]
H. h. sheppardi Brown & Benson, 1977
Brazil(Amazonas) hermathena sheppardi Brown & Benson, 1977; , TL: Brazil, Amazonas
Heliconius hermathena sheppardorum Lamas, 1988; (emend.)
= ; [NL4A, #2412e]
Heliconius hermathena sheppardi ; [NL4A, #2412e]
H. h. renatae Brown & Fernández, 1984
Venezuela hermathena renatae Brown & Fernández, 1984; , TL: Venezuela
Heliconius hermathena renatae ; [NL4A, #2412c]
H. h. sabinae Neukirchen, 1992
Brazil(Amazonas) hermathena sabinae Neukirchen, 1992; , TL: Brazil, Amazonas
Heliconius hermathena sabinae ; [NL4A, #2412d]

Heliconius heurippa Hewitson, 1854
Colombia. See [About maps] Heliconia heurippa Hewitson, 1854; Ill. exot. Butts [1] (Heliconia II-III): [72], pl. [37], f. 7, TL: Colombia
Heliconius heurippa, NSG Voucher Specimen [Wahlberg]
?Heliconius heurippa f. wernickei (Staudinger); [BOW: pl. 44, f. 14 (text only)]
Heliconius heurippa ; [EBW]; [BOW: pl. 44, f. 14]; [NL4A, #2413]

Heliconius hierax Hewitson, 1869
Ecuador. See [About maps] Heliconia hierax Hewitson, 1869; Equat. Lepid. Buckley (1): 11, TL: Ecuador [Rio Topo]
Heliconius hierax, NSG Voucher Specimen [Wahlberg]
Heliconia hierax ; Hewitson, 1871, Ill. exot. Butts [1] (Heliconia VI): [80], pl. [41], f. 20
Heliconius hierax ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 192, pl. 7, f. 71; [EBW, (text only)]; [EcuL]; [NL4A, #2414]
Ecuador f. semibrunnea Niepelt, 1923; , TL: Ecuador
= Heliconius hierax ; [EcuL]
= ; [NL4A, #2414a]
Heliconius hierax hierax ; [NL4A, #2414a]
H. h. leveri Attal, 1999
Ecuador Heliconius hierax leveri Attal, 1999; Bull. Soc. ent. Fr. 104 (4): (369-374), TL: Ecuador
Heliconius hierax leveri ; [NL4A, #2414b]

Heliconius himera Hewitson, 1867
Ecuador. See [About maps] Heliconia himera Hewitson, 1867; Ill. exot. Butts [1] (Heliconia V): [78], pl. [40], f. 16, TL: Ecuador
Heliconius himera, NSG Voucher Specimen [Wahlberg]
Heliconius himera ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 192, pl. 29, f. 291; [EBW, (text only)]; [EcuL]; [NL4A, #2415]

Heliconius hortense Guérin, [1844]
Mexico, Honduras - Ecuador. See [About maps] Heliconia hortense Guérin-Méneville, [1844]; Icon. Règne anim. Cuvier 3 (Insectes) : 469, TL: Mexico
Heliconius hortense, Listing of Mexican Butterflies [ILMB]
Heliconius hortense, Butterflies of America [BOA]
Heliconius hortense, NSG Voucher Specimen [Wahlberg]
Heliconia hortensia[sic, recte hortense] ; Doubleday, [1847], Gen. diurn. Lep. (1): 103, pl. 15, f. 1; Boisduval, 1870, Considérations Lépid. Guatemala: 29
Heliconius hortensia[sic, recte hortense] ; Godman & Salvin, [1881], Biol. centr.-amer., Lep. Rhop. 1: 151; Godman & Salvin, [1901], Biol. centr.-amer., Lep. Rhop. 2: 668
Heliconius hortense ; [EBW]; [BOW: pl. 44, f. 15]; [NL4A, #2416]

Heliconius ismenius Latreille, [1817]
-. See [About maps] Heliconius ismenius Latreille, [1817]; in Humboldt & Bonpland, Voy. rég. équinoxiales..., 2 (10): 125, pl. 41, f. 5-6, TL: [S. Amer.]
Heliconius ismenius, NSG Voucher Specimen [Wahlberg]
Heliconius ismenius ; [EBW]; [BOW: pl. 44, f. 16]; [NL4A, #2417]

Copy of Melinaea messatis. [BOW]

Larva on Passiflora alata, Passiflora pedata, P. ambigua, P. platyloba [BCR]

Colombia Heliconia fritscheï Möschler, 1872; Stettin Ent. Ztg 33 (7-9) : 336, TL: "Peru or Colombia"
= ; [NL4A, #2417a]
Heliconius ocanna Buchecker, [1880]; , TL: Colombia
= ; [NL4A, #2417a]
Heliconius faunus Staudinger, 1885; in Staudinger & Schatz, Exot. Schmett. 1 (9): 74, (7): pl. 31, TL: Colombia
= ; [NL4A, #2417a]
Heliconius faunus var. antioquensis Staudinger, 1885; in Staudinger & Schatz, Exot. Schmett. 1 (9): 74, TL: Colombia
= ; [NL4A, #2417a]
Heliconius ismenius var. (ab.?) hermanni Riffarth, 1899; Berl. ent. Zs. 43 (3/4) : 407, TL: [Colombia]
= ; [NL4A, #2417a]
Heliconius distincta Riffarth, 1901; Berl. ent. Zs. 46 (1/2) : 31 (nom. nud.)
= ; [NL4A, #2417a]
Heliconius ismenius ismenius f. immoderata Stichel, 1906; Genera Insectorum 37 : 9, TL: Colombia
= ; [NL4A, #2417a]
ab. albofasciatus Neustetter, 1907; , TL: Colombia
= ; [NL4A, #2417a]
f. defasciatus Neustetter, 1909; , TL: Colombia
= ; [NL4A, #2417a]
Heliconius ismenius abadiae Apolinar, 1926; , TL: Colombia
= ; [NL4A, #2417a]
Heliconius ismenius ismenius f. hermanni ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 192, pl. 13, f. 134
Heliconius ismenius ismenius f. immoderata ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 193, pl. 13, f. 133
Heliconius ismenius immoderata ; [EBW, (name only)]
Heliconius fritschei ; [EBW]
Heliconius ismenius ismenius ; [NL4A, #2417a]
H. i. telchinia Doubleday, 1847
Mexico - Panama, Honduras Heliconia telchinia Doubleday, 1847; Gen. diurn. Lep. (1): 104, pl. 14, f. 4, TL: [Honduras]
Heliconius ismenius telchinia, Listing of Mexican Butterflies [ILMB]
Heliconius ismenius telchinia, Butterflies of America [BOA]
Heliconius telchinia ; Godman & Salvin, [1881], Biol. centr.-amer., Lep. Rhop. 1: 149; Godman & Salvin, [1901], Biol. centr.-amer., Lep. Rhop. 2: 667
Heliconius ismenius telchinia ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 206, pl. 12, f. 129; [BCR, 196]; [EBW]; [NL4A, #2417g]
H. i. clarescens Butler, 1875
Costa Rica - Panama Heliconius clarescens Butler, 1875; Ann. Mag. Nat. Hist. (4) 15 (87) : 223, TL: Panama
Heliconius clarescens ; Godman & Salvin, [1881], Biol. centr.-amer., Lep. Rhop. 1: 150, 3: pl. 17, f. 5-6
Heliconius ismenius clarescens ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 183, pl. 12, f. 131; [BCR, 196]; [NL4A, #2417c]
H. i. fasciatus Godman & Salvin, 1877
Panama Heliconius fasciatus Godman & Salvin, 1877; Proc. zool. Soc. Lond. 1877 (1) : 62, TL: Panama
Heliconius fasciatus ; Godman & Salvin, [1881], Biol. centr.-amer., Lep. Rhop. 1: 150, 3: pl. 17, f. 3-4
Heliconius ismenius ismenius f. fasciatus ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 189, pl. 13, f. 135
Heliconius ismenius fasciatus ; [EBW, (name only)]; [NL4A, #2417d]
H. i. metaphorus Weymer, 1883
Ecuador, Colombia Heliconius metaphorus Weymer, 1883; Stettin ent. Ztg 45 (1-3) : 24, pl. 2, f. 1, TL: Chonana, Ecuador
Heliconius catilina Riffarth, 1901; Berl. ent. Zs. 46 (1/2) : 39 (nom. nud.)
= ; [NL4A, #2417e]
Heliconius ismenius faunus f. hoppi Neustetter, 1928; , TL: Colombia
= ; [NL4A, #2417e (hyb.)]
Heliconius metaphorus ; [EBW]
Heliconius ismenius metaphorus ; [EcuL]; [NL4A, #2417e]
H. i. boulleti Neustetter, 1928
ismenius boulleti Neustetter, 1928; , TL: "French Guiana"
Heliconius sylvana sylvana[sic] f. sticheli Boullet & Le Cerf, 1909; , TL: "French Guiana"
= ; [NL4A, #2417b]
Heliconius ismenius boulleti ; [NL4A, #2417b]
H. i. occidentalis Neustetter, 1928
Colombia Heliconius ismenius occidentalis Neustetter, 1928; Int. ent. Zs. 22 : 258, TL: Colombia
Heliconius ismenius f. occidentalis ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 199
Heliconius ismenius occidentalis ; [EBW, (name only)]; [NL4A, #2417f]
H. i. tilletti Brown & Fernández, 1976
Venezuela ismenius tilletti Brown & Fernández, 1976; , TL: Venezuela
Heliconius ismenius tilletti ; [NL4A, #2417h]

Heliconius lalitae Brévignon, 1996
French Guiana. See [About maps] lalitae Brévignon, 1996; , TL: French Guiana
Heliconius lalitae ; [NL4A, #2418]

Heliconius leucadia Bates, 1862
Ecuador. See [About maps] Heliconius leucadia Bates, 1862; Trans. linn. Soc. Lond. 23 (3) : 556, TL: St. Paulo, Upper Amazons
Heliconius leucadia, NSG Voucher Specimen [Wahlberg]
Heliconius leucadia ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 195, pl. 37, f. 380; [BOW: pl. 30, f. 30 (text only)]; [NL4A, #2419]
Brazil(Amazonas) Heliconius leucadia leucadia ; [NL4A, #2419a]
H. l. pseudorhea Staudinger, 1897
Brazil(Amazonas) Heliconius pseudorhea Staudinger, [1897]; Dt. Ent. Z. Iris 9 (2) : 291, TL: Brazil, Amazonas
Heliconius leucadia pseudorhea ; [NL4A, #2419d]
H. l. andromeda Neukirchen, 1996
Ecuador leucadia andromeda Neukirchen, 1996; , TL: Ecuador, Pastaza, Puyo-Tena km 25
Heliconius leucadia andromeda ; [EcuL]; [NL4A, #2419b]
H. l. birgitae Neukirchen, 1996
Peru leucadia birgitae Neukirchen, 1996; , TL: Peru
Heliconius leucadia birgitae ; [NL4A, #2419c]

Heliconius luciana Lichy, 1960
-. See [About maps] luciana Lichy, 1960; , TL: Venezuela
Heliconius luciana ; [NL4A, #2420]
Venezuela Heliconius luciana luciana ; [NL4A, #2420a]
Heliconius elevatus luciana ; [EBW, (name only)]
H. l. watunna Lichy, 1960
Venezuela luciana watunna Lichy, 1960; , TL: Venezuela
Heliconius luciana watunna ; [NL4A, #2420b]

Heliconius melpomene (Linnaeus, 1758)
-. See [About maps] Papilio melpomene Linnaeus, 1758; Syst. Nat. (Edn 10) 1 : 467, TL: "America" [Sout America, ?Guianas]
Heliconius melpomene, NSG Voucher Specimen [Wahlberg]
Heliconius melpomene ; Godman & Salvin, [1881], Biol. centr.-amer., Lep. Rhop. 1: 154; [NL4A, #2421]; [EBW]; [BOW: pl. 44, f. 18]
Surinam, French Guiana, Guyana, Brazil(Amazonas, Pará) Papilio lucia Stoll, [1781]; in Cramer, Uitl. Kapellen 4 (29-31): 117, pl. 350 f. E-F, TL: Surinam
= ; [NL4A, #2421a (hyb.)]
Heliconius melpomene var. tyche Bates, 1862; Trans. linn. Soc. Lond. 23 (3) : 559, TL: Brazil, Amazonas
= Heliconius melpomene melpomene × melpomene madeira ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 207, pl. 24, f. 243
= ; [NL4A, #2421a (hyb.)]
Heliconius hippolyte Bates, 1862; Trans. linn. Soc. Lond. 23 (3) : 559, TL: Brazil, Amazonas; Pará
= Heliconius melpomene melpomene × melpomene madeira ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 192, pl. 24, f. 241
= ; [NL4A, #2421a (hyb.)]
Heliconius rufolimbatus Butler, 1873; Cist. Ent. 1 (7) : 169, TL: Brazil, Pará
= Heliconius melpomene melpomene × melpomene madeira ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 203, pl. 24, f. 242
= ; [NL4A, #2421a (hyb.)]
Heliconius mutabilis Butler, 1877; Cistula Ent. 2 : 151, TL: Brazil, Amazonas
= Heliconius melpomene melpomene × melpomene meriana ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 197, pl. 23, f. 230
= ; [NL4A, #2421a (hyb.)]
Heliconius funebris Möschler, 1877; Verh. zool.-bot. Ges. Wien 26 : 314, pl. 3, f. 8, TL: Surinam
= ; [NL4A, #2421a (hyb.)]
Heliconius amor Staudinger, 1885; in Staudinger & Schatz, Exot. Schmett. 1 (9): 78, (7): pl. 32, TL: Brazil, Amazonas
= ; [NL4A, #2421a (hyb.)]
Heliconius timareta ab.? erebia Riffarth, 1900; Berl. ent. Zs. 45 (3/4) : 201, TL: French Guiana
= ; [NL4A, #2421a (hyb.)]
Heliconius melpomene ab. atrosecta Riffarth, 1900; Berl. ent. Zs. 45 (3/4) : 202, TL: Brazil, Pará, Obidos
= ; [NL4A, #2421a]
Heliconius melpomene ab. lucinda Riffarth, 1900; Berl. ent. Zs. 45 (3/4) : 203, TL: Surinam, Massauary
= ; [NL4A, #2421a (hyb.)]
Heliconius melpomene ab. melpomenides Riffarth, 1900; Berl. ent. Zs. 45 (3/4) : 203, TL: Surinam, Villa Bella
= ; [NL4A, #2421a (hyb.)]
Heliconius melpomene ab. karschi Riffarth, 1900; Berl. ent. Zs. 45 (3/4) : 203, TL: Brazil, Pará; Surinam
= ; [NL4A, #2421a (hyb.)]
Heliconius melpomene var. melanippe Riffarth, 1900; Berl. ent. Zs. 45 (3/4) : 204, TL: Surinam
= ; [NL4A, #2421a (hyb.)]
Heliconius melpomene ab. diana Riffarth, 1900; Berl. ent. Zs. 45 (3/4) : 204, TL: Surinam?
= Heliconius melpomene melpomene × melpomene meriana ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 185, pl. 23, f. 229
= ; [NL4A, #2421a (hyb.)]
Heliconius gaea Riffarth, 1901; Berl. ent. Zs. 46 (1/2) : 91 (nom. nud.)
= ; [NL4A, #2421a]
Heliconius jussa Riffarth, 1901; Berl. ent. Zs. 46 (1/2) : 92 (nom. nud.)
= ; [NL4A, #2421a]
Heliconius justina Riffarth, 1901; Berl. ent. Zs. 46 (1/2) : 92 (nom. nud.)
= ; [NL4A, #2421a]
Heliconius melpomene melpomene ab. collis Joicey & Kaye, 1917; Trans. ent. Soc. Lond. 1916 : 413, pl. 107, f. 3, TL: French Guiana
= Heliconius melpomene melpomene × melpomene meriana ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 184, pl. 21, f. 215
= ; [NL4A, #2421a (hyb.)]
Heliconius melpomene melpomene ab. primus Joicey & Kaye, 1917; Trans. ent. Soc. Lond. 1916 : 414, pl. 107, f. 1, TL: French Guiana
= Heliconius melpomene melpomene × melpomene meriana ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 201, pl. 21, f. 214
= ; [NL4A, #2421a (hyb.)]
Heliconius melpomene melpomene ab. melpina Joicey & Kaye, 1917; Trans. ent. Soc. Lond. 1916 : 414, TL: French Guiana
= Heliconius melpomene melpomene × melpomene meriana ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 197, pl. 21, f. 217
= ; [NL4A, #2421a (hyb.)]
Heliconius melpomene cybele ab. dianides Joicey & Kaye, 1917; Trans. ent. Soc. Lond. 1916 : 417, pl. 107, f. 7, TL: French Guiana
= Heliconius melpomene melpomene × melpomene meriana ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 186, pl. 22, f. 225
= ; [NL4A, #2421a (hyb.)]
Heliconius melpomene cybele ab. elegantula Joicey & Kaye, 1917; Trans. ent. Soc. Lond. 1916 : 417, TL: French Guiana
= Heliconius melpomene melpomene × melpomene meriana ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 187, pl. 22, f. 227
= ; [NL4A, #2421a (hyb.)]
Heliconius melpomene cybele ab. maris Joicey & Kaye, 1917; Trans. ent. Soc. Lond. 1916 : 418, pl. 107, f. 4, TL: French Guiana
= Heliconius melpomene melpomene × melpomene meriana ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 196, pl. 22, f. 228
= ; [NL4A, #2421a (hyb.)]
Heliconius melpomene eltringhami Joicey & Kaye, 1917; Trans. ent. Soc. Lond. 1916 : 418, pl. 107, f. 6, TL: French Guiana
= Heliconius melpomene melpomene × melpomene × meriana ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 187, pl. 21, f. 211
= ; [NL4A, #2421a (hyb.)]
Heliconius melpomene melponene ab. compacta Joicey & Kaye, 1919; Trans. ent. Soc. Lond. 1918 : 349, TL: French Guiana
= Heliconius melpomene melponene × melpomene meriana ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 184, pl. 21, f. 216
= ; [NL4A, #2421a (hyb.)]
Heliconius melpomene melpomene ab. faivrei Joicey & Kaye, 1919; Trans. ent. Soc. Lond. 1918 : 349, TL: French Guiana
= Heliconius melpomene melpomene × melpomene meriana ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 189, pl. 21, f. 218
= ; [NL4A, #2421a (hyb.)]
Heliconius melpomene f. flavorubra Neustetter, 1926; Int. ent. Z. 20 : 279, TL: French Guiana
= Heliconius melpomene melpomene × melpomene meriana ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 190, pl. 21, f. 213
= ; [NL4A, #2421a]
f. lydia Neustetter, 1926; , TL: French Guiana
= ; [NL4A, #2421a (hyb.)]
Heliconius melpomene f. luteipicta Neustetter, 1926; Int. ent. Z. 20 : 281, TL: French Guiana,Guyana
= Heliconius melpomene melpomene × melpomene meriana ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 196, pl. 21, f. 212
= ; [NL4A, #2421a (hyb.)]
Heliconius melpomene f. aurelia Neustetter, 1926; Int. ent. Z. 20 : 289, TL: French Guiana
= Heliconius melpomene melpomene × melpomene meriana ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 180, pl. 22, f. 224
= ; [NL4A, #2421a (hyb.)]
f. bang-haasi Neustetter, 1926; , TL: French Guiana
= ; [NL4A, #2421a (hyb.)]
f. rubroflammea Neustetter, 1927; , TL: French Guiana
= ; [NL4A, #2421a (hyb.)]
Heliconius melpomene f. laurentina Neustetter, 1927; Int. ent. Z. 21 : 229, TL: French Guiana
= Heliconius melpomene melpomene × melpomene thelxiope ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 195, pl. 23, f. 235
= ; [NL4A, #2421a (hyb.)]
f. rufolinea Neustetter, 1928; , TL: French Guiana
= ; [NL4A, #2421a (hyb.)]
f. trimacula Neustetter, 1931; , TL: French Guiana
= ; [NL4A, #2421a (hyb.)]
Heliconius melpomene melpomene f. atrosecta ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 179, pl. 22, f. 219
Heliconius melpomene f. lucia ; [BOW: pl. 44, f. 18 (text only)]
Heliconius melpomene melpomene ; [NL4A, #2421a]
H. m. thelxiope (Hübner, [1806])
Amazon Nerëis melpomene thelxiope Hübner, [1806]; , TL: [Brazil]
Heliconius melpomene thelxiope ; [EBW]; [NL4A, #2421x]
H. m. vicina Ménétriés, 1857
Rio Negro Heliconia vicina Ménétriés, 1857; Cat. lep. Petersb. 2: 114, TL: Brazil
Heliconius melpomene vicinus ; [EBW]
Heliconius melpomene vicina ; [NL4A, #2421z]
H. m. xenoclea Hewitson, 1853
Ecuador, Peru Heliconia xenoclea Hewitson, 1853; Ill. exot. Butts [1] (Heliconia I): [69], pl. [36], f. 1
Heliconius batesi Riffarth, 1900; Berl. ent. Zs. 45 (3/4) : 207, TL: "Ecuador, Chanchamago",Peru
= ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 180, pl. 18, f. 186; [NL4A, #2421ab]
Heliconius xenoclea var. superba Lathy, 1906; Proc. zool. Soc. Lond. 1906 : 453, pl. 34, f. 2, TL: Peru
= Heliconius melpomene xenoclea × melpomene aglaope ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 205, pl. 18, f. 188
= ; [NL4A, #2421ab (hyb.)]
Heliconius xenoclea var. confluens Lathy, 1906; Proc. zool. Soc. Lond. 1906 : 453, pl. 34, f. 3, TL: Peru
= Heliconius melpomene xenoclea × melpomene euryades ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 184, pl. 18, f. 185
= ; [NL4A, #2421ab (hyb.)]
Heliconius xenoclea ab. zio Röber, 1906; , TL: Peru
= ; [NL4A, #2421ab (hyb.)]
Heliconius f. integra Descimon & Mast, 1971; , TL: Peru
= ; [NL4A, #2421ab (hyb.)]
Heliconius xenoclea ; [BOW: pl. 45, f. 6]
Heliconius melpomene xenoclea ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 209, pl. 18, f. 187; [EBW]; [NL4A, #2421ab]
H. m. aglaope C. & R. Felder, 1862
Peru (Ucayali) Heliconius aglaope C. & R. Felder, 1862; Wien. ent. Monats. 6 (3): 79, TL: "Rio Negro" [Peru, San Martin]
Heliconius melpomene ab. mirabilis Riffarth, 1900; Berl. ent. Zs. 45 (3/4) : 205, TL: Peru, Yurimaguas
= ; [NL4A, #2421b (hyb.)]
Heliconius melpomene riffarthi Stichel, 1906; Genera Insectorum 37 : 26, pl. 2, f. 8a, TL: Peru
= ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 202, pl. 19, f. 189; [NL4A, #2421b (hyb.)]
Heliconius melpomene riffarthi f. rubescens Stichel, 1906; , TL: Peru
= ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 203, pl. 19, f. 190; [NL4A, #2421b (hyb.)]
Heliconius melpomene aglaope f. cognata Riffarth, 1907; Dt. ent. Zs. 1907 (5) : 506, TL: Peru
= ; [NL4A, #2421b]
Heliconius aglaope f. flavotenuiata Neustetter, 1931; , TL: Peru
= ; [NL4A, #2421b]
Heliconius aglaope f. magnimacula Neustetter, 1932; , TL: Peru
= ; [NL4A, #2421b (hyb.)]
Heliconius aglaope f. daira Neustetter, 1932; , TL: Peru
= ; [NL4A, #2421b (hyb.)]
Heliconius melpomene aglaope f. cognata ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 184, pl. 20, f. 204
Heliconius melpomene aglaope ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 176, pl. 20, f. 206; [EBW]; [EcuL]; [NL4A, #2421b]
H. m. amaryllis C. & R. Felder, 1862
Costa Rica, Panama, Colombia, Peru Heliconius amaryllis C. & R. Felder, 1862; Wien. ent. Monats. 6 (3): 80, TL: "Brazil, Amazonas"
Heliconius amaryllis ab. pseudo-penelamanda Michael, 1912; , TL: Peru
= ; [NL4A, #2421d (hyb.)]
Heliconius melpomene melpophylla Joicey & Kaye, 1917; Ann. Mag. nat. Hist. (8) 20 (115) : 92, TL: Peru?
= Heliconius melpomene amaryllis × melpomene aglaope ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 197, pl. 19, f. 192
= ; [NL4A, #2421d (hyb.)]
Heliconius amaryllis ; [BOW: pl. 43, f. 35]
Heliconius melpomene amaryllis ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 177, pl. 19, f. 191; [EBW]; [NL4A, #2421d]
H. m. vulcanus Butler, 1865
Colombia, Panama Heliconius vulcanus Butler, 1865; Proc. zool. Soc. Lond. 1865 (2) : 433, TL: "Guyana",Panama
Heliconius amaryllis rosina f. fumosa Neustetter, 1925; , TL: Colombia
= ; [NL4A, #2421aa (hyb.)]
Heliconius vulcanus ; [BOW: pl. 45, f. 3]
Heliconius melpomene vulcanus ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 209, pl. 18, f. 180; [EBW]; [NL4A, #2421aa]
H. m. cythera Hewitson, 1869
Ecuador Heliconia cythera Hewitson, 1869; Equat. Lepid. Buckley (1): 9, TL: Ecuador [Angus = rio Angas, Cañar]
Heliconius vulcanus ab. modesta Riffarth, 1900; Berl. ent. Zs. 45 (3/4) : 206, TL: Ecuador
= Heliconius melpomene cythera × melpomene vulcanus ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 197, pl. 18, f. 181
= ; [NL4A, #2421h (hyb.)]
Heliconius hypna Riffarth, 1901; Berl. ent. Zs. 46 (1/2) : 105 (nom. nud.)
= ; [NL4A, #2421h]
Heliconius vulcanus sticheli Riffarth, 1907; Dt. ent. Zs. 1907 (5) : 509, pl. 5, f. 10, TL: Ecuador
= Heliconius melpomene cythera × melpomene vulcanus ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 205, pl. 18, f. 182
= ; [NL4A, #2421h (hyb.)]
Heliconius vulcanus tenuistriga Kaye, 1920; , TL: Ecuador
= ; [NL4A, #2421h]
Heliconia cythera ; Hewitson, 1871, Ill. exot. Butts [1] (Heliconia VI): [79], pl. [41], f. 17
Heliconius melpomene sticheli ; [EBW]
Heliconius melpomene modesta ; [EBW, (name only)]
Heliconius melpomene cythera ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 185, pl. 18, f. 183; [EBW, (name only)]; [EcuL]; [NL4A, #2421h]
H. m. rosina Boisduval, 1870
Mexico - Panama, Costa Rica Heliconia rosina Boisduval, 1870; Considérations Lépid. Guatemala: 29, TL: Costa Rica,"Mexico"
Heliconius melpomene rosina ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 203, pl. 18, f. 179; [BCR, 194]; [NL4A, #2421u]

Larva on Passiflora oerstedii, P. menispermifolia [BCR]

H. m. amandus Grose-Smith & Kirby, 1892
Bolivia Heliconius amandus Grose-Smith & Kirby, 1892; Rhop. Exot. [2] 1: (Heliconius) 1, pl. 1, f. 3, TL: Bolivia
Heliconius penelope ab. pelopeia Staudinger, 1894; Dt. Ent. Z. Iris 7 (1) : 67, TL: Bolivia
= ; [NL4A, #2421c (hyb.)]
Heliconius penelope ab. penelamanda Staudinger, 1894; Dt. Ent. Z. Iris 7 (1) : 67, TL: Bolivia
= ; [NL4A, #2421c (hyb.)]
Heliconius aphrodyte Staudinger, [1897]; Dt. Ent. Z. Iris 9 (2) : 299, pl. 6, f. 4, TL: Bolivia
= ; [NL4A, #2421c (hyb.)]
Heliconius penelope ab. penelopeia Staudinger, [1897]; Dt. Ent. Z. Iris 9 (2) : 303 (emend.)
= ; [NL4A, #2421c]
Heliconius phyllis f. flammea Niepelt, 1925; Int. ent. Zs. 18 : 252, TL: Bolivia
= Heliconius melpomene amandus × melpomene penelope ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 189, pl. 25, f. 253
= ; [NL4A, #2421c (hyb.)]
Heliconius penelope f. biedermanni Niepelt, 1926; Int. ent. Zs. 19 : 337, TL: Bolivia
= Heliconius melpomene amandus × melpomene penelope ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 181, pl. 25, f. 254
= ; [NL4A, #2421c (hyb.)]
Heliconius penelope f. amandoides Neustetter, 1926; Int. ent. Z. 20 : 39, TL: Bolivia
= ; [NL4A, #2421c (hyb.)]
Heliconius penelope f. carnea Neustetter, 1926; Int. ent. Z. 20 : 38, TL: Bolivia
= ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 182, pl. 24, f. 248; [NL4A, #2421c (hyb.)]
Heliconius melpomene amandus f. amandoides ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 177, pl. 25, f. 255
Heliconius melpomene amandus ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 177, pl. 25, f. 252; [EBW, (name only)]; [NL4A, #2421c]
H. m. penelope Staudinger, 1894
Bolivia, Brazil(Mato Grosso) Heliconius penelope Staudinger, 1894; Dt. Ent. Z. Iris 7 (1) : 67, TL: Bolivia
Heliconius penelope ab. pluto Staudinger, [1897]; Dt. Ent. Z. Iris 9 (2) : 303, pl. 7, f. 4, TL: Bolivia
= ; [NL4A, #2421r (hyb.)]
Heliconius melpomene ab. margarita Riffarth, 1900; Berl. ent. Zs. 45 (3/4) : 205, TL: Bolivia
= Heliconius melpomene penelope × melpomene amandus ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 196, pl. 25, f. 251
= ; [NL4A, #2421r (hyb.)]
f. praxedis Neustetter, 1925; , TL: Bolivia
= ; [NL4A, #2421r (hyb.)]
Heliconius penelope f. noctis Neustetter, 1926; Int. ent. Z. 20 : 36, TL: Bolivia
= Heliconius melpomene penelope × melpomene amandus ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 198, pl. 25, f. 249
= ; [NL4A, #2421r (hyb.)]
Heliconius penelope f. excellens Neustetter, 1926; Int. ent. Z. 20 : 37, TL: Bolivia
= Heliconius melpomene penelope × melpomene amandus ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 188, pl. 24, f. 247
= ; [NL4A, #2421r (hyb.)]
Heliconius penelope f. aida Neustetter, 1926; Int. ent. Z. 20 : 37, TL: Bolivia
= Heliconius melpomene penelope × melpomene amandus ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 176, pl. 4, f. 246
= ; [NL4A, #2421r (hyb.)]
Heliconius penelope f. amneris Neustetter, 1926; Int. ent. Z. 20 : 37, TL: Bolivia
= Heliconius melpomene penelope × melpomene amandus ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 177, pl. 25, f. 250
= ; [NL4A, #2421r (hyb.)]
Heliconius penelope f. rufofascia Neustetter, 1926; , TL: Bolivia
= ; [NL4A, #2421r (hyb.)]
Heliconius melpomene burchelli f. obscurifascia Talbot, 1928; Bull. Hill Mus. Witley 2 : 220, TL: Brazil, Mato Grosso
= Heliconius melponeme penelope × melpomene burchelli ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 199, pl. 25, f. 258
= ; [NL4A, #2421r (hyb.)]
Heliconius penelope f. flavodiscalis Neustetter, 1931; , TL: Bolivia
= ; [NL4A, #2421r (hyb.)]
?Heliconius melpomene penelopeia[sic] ; [BOW: pl. 44, f. 18 (text only, missp.?)]
Heliconius melpomene penelope ; [EBW]; [NL4A, #2421r]
H. m. thelxiopeia Staudinger, 1897
French Guiana, Surinam Heliconius thelxiope ab. (var.) thelxiopeia Staudinger, [1897]; Dt. Ent. Z. Iris 9 (2) : 305, TL: French Guiana
Heliconius thelxiope var. aglaopeia Staudinger, [1897]; Dt. Ent. Z. Iris 9 (2) : 305, TL: French Guiana
= ; [NL4A, #2421y]
Heliconius melpomene var. augusta Riffarth, 1900; Berl. ent. Zs. 45 (3/4) : 204, TL: Surinam
= Heliconius melpomene melpomene × melpomene thelxiope ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 180, pl. 23, f. 237
= ; [NL4A, #2421y (hyb.)]
Heliconius judith Riffarth, 1901; Berl. ent. Zs. 46 (1/2) : 93 (nom. nud.)
= ; [NL4A, #2421y]
Heliconius milesia Riffarth, 1901; Berl. ent. Zs. 46 (1/2) : 93 (nom. nud.)
= ; [NL4A, #2421y]
Heliconius melpomene thelxiope ab. punctarius Joicey & Kaye, 1917; Trans. ent. Soc. Lond. 1916 : 422, pl. 107, f. 5, TL: French Guiana
= Heliconius melpomene melpomene × melpomene thelxiope ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 201, pl. 24, f. 240
= ; [NL4A, #2421y (hyb.)]
Heliconius melpomene thelxiope ab. lucindella Joicey & Kaye, 1917; Trans. ent. Soc. Lond. 1916 : 423, TL: French Guiana
= Heliconius melpomene melpomene × melpomene thelxiope ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 195, pl. 24, f. 239
= ; [NL4A, #2421y (hyb.)]
Heliconius melpomene thelxiope ab. majestica Joicey & Kaye, 1917; Trans. ent. Soc. Lond. 1916 : 423, TL: French Guiana
= Heliconius melpomene melpomene × melpomene thelxiope ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 196, pl. 23, f. 238
= ; [NL4A, #2421y (hyb.)]
Heliconius melpomene thelxiope ab. stygianus Joicey & Kaye, 1917; Trans. ent. Soc. Lond. 1916 : 423, TL: French Guiana
= Heliconius melpomene thelxiope × melpomene melpomene ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 205, pl. 23, f. 231
= ; [NL4A, #2421y (hyb.)]
Heliconius melpomene thelxiope ab. negroides Joicey & Kaye, 1917; Trans. ent. Soc. Lond. 1916 : 424, TL: French Guiana
= Heliconius melpomene melpomene × melpomene thelxiope ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 198, pl. 23, f. 234
= ; [NL4A, #2421y (hyb.)]
Heliconius melpomene f. athalia Neustetter, 1927; Int. ent. Z. 21 : 228, TL: French Guiana
= ; [NL4A, #2421y (hyb.)]
Heliconius melpomene f. penelopides Neustetter, 1927; , TL: French Guiana
= Heliconius melpomene thelxiope × melpomene melpomene ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 200, pl. 23, f. 233
= ; [NL4A, #2421y (hyb.)]
Heliconius melpomene f. nigrointerrupta Neustetter, 1931; , TL: French Guiana
= ; [NL4A, #2421y (hyb.)]
Heliconius melpomene thelxiopeia f. aglaopeia ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 176, pl. 23, f. 232
Heliconius melpomene thelxiopeia f. athalia ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 179, pl. 23, f. 236
Heliconius melpomene thelxiopeia ; [NL4A, #2421y]
H. m. nanna Stichel, 1899
Brazil(Espírito Santo, Minas Gerais, Rio de Janeiro) Heliconius nanna Stichel, 1899; Ent. Z. 12 (19): 143, TL: Brazil, Espírito Santo; Minas Gerais
Heliconius bidentatus Riffarth, 1901; Berl. ent. Zs. 46 (1/2) : 106 (nom. nud.)
= ; [NL4A, #2421q]
Heliconius mayi d'Almeida, 1928; , TL: Brazil, Rio de Janeiro
= ; [NL4A, #2421q]
Heliconius (Heliconius) melpomene nannus ; Brown & Mielke, 1967, J. Lep. Soc. 21 (2): 96
Heliconius nanna ; [BOW: pl. 44, f. 20]
Heliconius melpomene nanna ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 198, pl. 26, f. 259; [EBW, (name only)]; [NL4A, #2421q]
H. m. euryades Riffarth, 1900
Peru Heliconius amaryllis var. euryades Riffarth, 1900; Berl. ent. Zs. 45 (3/4) : 205, TL: Peru, (Garlepp), 3000'
amaryllis euryades f. rubrica Stichel, 1919; , TL: Peru
= ; [NL4A, #2421j (hyb.)]
Heliconius melpomene euryades ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 188, pl. 24, f. 245; [EBW, (name only)]; [NL4A, #2421j]
H. m. plesseni Riffarth, 1907
Ecuador Heliconius batesi f. plesseni Riffarth, 1907; Dt. ent. Zs. 1907 (3) : 333, TL: Baraneas[sic], ober Pastazza, Ecuador
Heliconia unimaculata Hewitson, 1869; Equat. Lepid. Buckley (1): 10, TL: Canelos
= Heliconius melpomene plesseni × melpomene aglaope ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 208, pl. 20, f. 200
= Heliconius erato notabilis ; [EcuL]
= ; [NL4A, #2421s (hyb.)]
Heliconius radiatus Riffarth, 1901; Berl. ent. Zs. 46 (1/2) : 96 (nom. nud.)
= ; [NL4A, #2421s]
Heliconius batesi plesseni f. rubicunda Niepelt, 1907; Soc. ent. 22 (6) : 42, TL: Ecuador
= ; [NL4A, #2421s]
Heliconius batesi plesseni f. pura Niepelt, 1907; Soc. ent. 22 (6) : 42, TL: Ecuador
= ; [NL4A, #2421s]
Heliconius melpomene aglaope f. niepelti Riffarth, 1907; Dt. ent. Zs. 1907 (5) : 507, pl. 5, f. 7, TL: Ecuador
= ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 198, pl. 20, f. 201; [NL4A, #2421s (hyb.)]
Heliconius batesi plesseni f. adonis Riffarth, 1907; Dt. ent. Zs. 1907 (5) : 507, pl. 5, f. 9, TL: Ecuador
= Heliconius melpomene plesseni × melpomene aglaope ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 176, pl. 19, f. 197
= ; [NL4A, #2421s (hyb.)]
Heliconius batesi plesseni f. corona Niepelt, 1908; Berl. ent. Zs. 52 : 213, TL: Ecuador
= ; [NL4A, #2421s]
Heliconius batesi plesseni f. diadema Niepelt, 1908; Berl. ent. Zs. 52 : 213, TL: Ecuador
= ; [NL4A, #2421s (hyb.)]
Heliconius melpomene aglaope f. fraterna Niepelt, 1909; Berl. ent. Zs. 54 : 106, TL: Ecuador
= ; [NL4A, #2421s]
Heliconia microclea virgo Oberthür, 1916; , TL: Ecuador
= ; [NL4A, #2421s]
Heliconius xenoclea plesseni f. dido Neustetter, 1925; , TL: Ecuador
= ; [NL4A, #2421s (hyb.)]
Heliconius xenoclea plesseni f. clytie Neustetter, 1927; Int. ent. Z. 21 : 229, TL: Ecuador
= Heliconius melpomene plesseni × melpomene aglaope ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 184, pl. 20, f. 199
= ; [NL4A, #2421s (hyb.)]
Heliconius xenoclea psesseni f. mimetica Neustetter, 1928; , TL: Ecuador
= ; [NL4A, #2421s (hyb.)]

Heliconia unimaculata ; Hewitson, 1871, Ill. exot. Butts [1] (Heliconia VI): [79], pl. [41], f. 19
?Heliconius xanthocles f. plesseni ; [BOW: pl. 45, f. 6 (text only)]
Heliconius melpomene plesseni f. corona ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 185, pl. 19, f. 195
Heliconius melpomene plesseni f. fraterna ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 190, pl. 19, f. 198
Heliconius melpomene plesseni f. pura ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 202, pl. 19, f. 196
Heliconius melpomene plesseni f. rubicunda ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 203, pl. 19, f. 194
Heliconius melpomene unimaculata ; [EBW, (name only)]
Heliconius melpomene plesseni ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 201, pl. 19, f. 193; [EBW]; [EcuL]; [NL4A, #2421s]
H. m. pyrforus Kaye, 1907
Guyana, Brazil(Roraima) melpomene pyrforus Kaye, 1907; , TL: Guyana
Heliconius vulcanus f. immarginata Neustetter, 1938; , TL: Brazil, Roraima
= ; [NL4A, #2421t]
Heliconius melpomene pyrforus ; [NL4A, #2421t]
H. m. burchelli Poulton, 1910
Brazil(Goiás, Mato Grosso) Heliconius nanna burchelli Poulton, 1910; in Joseph, Ann. Mat. nat. Hist. (8) 5 (28) : 331, TL: Villa Nivac, S.E. of Matto Grosso
Heliconius melpomene burchelli f. curvifascia Talbot, 1928; Bull. Hill Mus. Witley 2 : 220, TL: Brazil, Mato Grosso
= Heliconius melpomene penelope × melpomene burhcelli ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 185, pl. 25, f. 257
= ; [NL4A, #2421g (hyb.)]
Heliconius melpomene burchelli ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 181, pl. 25, f. 256; [NL4A, #2421g]
H. m. flagrans Stichel, 1919
Trinidad Heliconius amaryllis flagrans Stichel, 1919; Neue Beitr. syst. Ins. 1 (15) : 120, TL: Trinidad
Heliconius melpomene melpomene f. flagrans ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 189, pl. 21, f. 210
Heliconius melpomene flagrans ; [NL4A, #2421k]
H. m. madeira Riley, 1919
Brazil(Amazonas) Heliconius melpomene madeira Riley, 1919; Entomologist 52 : 185, TL: Brazil, Amazonas
Heliconius melpomene madeira ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 196, pl. 24, f. 244; [NL4A, #2421m]
H. m. intersectus Neustetter, 1928
Brazil(Pará) melpomene intersectus Neustetter, 1928; , TL: Brazil, Pará
Heliconius melpomene intersectus ; [NL4A, #2421l]
H. m. ecuadorensis Emsley, 1964
Ecuador melpomene ecuadorensis Emsley, 1964; , TL: Ecuador, Macas
Heliconius melpomene aglaope f. ecuadorensis Neustetter, 1909; , TL: Ecuador
= ; [NL4A, #2421i]
Heliconius equatorensis ; Seitz, 1912, (missp.)
Heliconius melpomene equadoriensis ; Seitz, 1913, (missp.)
Heliconius melpomene ecuadorensis ; [EcuL]; [NL4A, #2421i]
H. m. meriana Turner, 1967
Surinam, French Guiana melpomene meriana Turner, 1967; , TL: Surinam
Papilio cybele Cramer, [1777]; Uitl. Kapellen 2 (9-16): 139, pl. 188, f. A (preocc. Fabricius, 1775), TL: Surinam
= ; [NL4A, #2421o]
Heliconius deinia Möschler, 1877; Verh. zool.-bot. Ges. Wien 26 : 315, TL: [Surinam]
= ; [NL4A, #2421o (hyb.)]
Heliconius faustina Staudinger, 1885; in Staudinger & Schatz, Exot. Schmett. 1 (9): 78, TL: French Guiana
= ; [NL4A, #2421o (hyb.)]
Heliconius melpomene ab. eulalia Riffarth, 1900; Berl. ent. Zs. 45 (3/4) : 204, TL: Surinam
= ; [NL4A, #2421o (hyb.)]
Heliconius melpomene funebris f. obscurata Riffarth, 1907; Dt. ent. Zs. 1907 (5) : 505, TL: Surinam
= ; [NL4A, #2421o]
Heliconius fascinatrix Seitz, 1912; , TL: French Guiana
= ; [NL4A, #2421o (hyb.)]
Heliconius melpomene cybele ab. faustalia Joicey & Kaye, 1917; Trans. ent. Soc. Lond. 1916 : 417, TL: French Guiana
= ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 189, pl. 22, f. 223; [NL4A, #2421o (hyb.)]
Heliconius melpomene cybele ab. negroida Joicey & Kaye, 1917; Trans. ent. Soc. Lond. 1916 : 418, pl. 107, f. 2, TL: French Guiana
= Heliconius melpomene meriana f. faustina ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 198, pl. 22, f. 222
= ; [NL4A, #2421o (hyb.)]
Heliconius melpomene cybele ab. cybeleia Joicey & Kaye, 1919; Trans. ent. Soc. Lond. 1918 : 349, TL: French Guiana
= Heliconius melpomene meriana f. obscurata ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 185, pl. 22, f. 221
= ; [NL4A, #2421o (hyb.)]
Heliconius melpomene f. lavinia Neustetter, 1926; Int. ent. Z. 20 : 290, TL: French Guiana
= Heliconius melpomene meriana × melpomene melpomene ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 195, pl. 22, f. 226
= ; [NL4A, #2421o (hyb.)]
f. alba Stammeshaus, 1982; , TL: Surinam
= ; [NL4A, #2421o]
Heliconius melpomene meriana f. obscurata ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 199, pl. 22, f. 220
Heliconius melpomene meriana ; [EBW, (name only)]; [NL4A, #2421o]
H. m. bellula Turner, 1971
Colombia melpomene bellula Turner, 1971; , TL: Colombia
Heliconius amaryllis amaryllis f. bellula Stichel, 1923; , TL: Colombia
= ; [NL4A, #2421f]
Heliconius amaryllis amaryllis f. permira Stichel, 1923; , TL: Colombia
= ; [NL4A, #2421f (hyb.)]
Heliconius amaryllis amaryllis f. degener Stichel, 1923; , TL: Colombia
= ; [NL4A, #2421f (hyb.)]
Heliconius amaryllis amaryllis anacreontica Stichel, 1923; Dt. ent. Z. 1923 (3) : 263, TL: Colombia
= ; [NL4A, #2421f (hyb.)]
Heliconius amaryllis amaryllis f. perrara Stichel, 1923; , TL: Colombia
= ; [NL4A, #2421f (hyb.)]
Heliconius melpomene aglaope f. parva Neustetter, 1928; Int. ent. Zs. 22 : 77, TL: Colombia
= ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 200, pl. 20, f. 207; [NL4A, #2421f (hyb.)]
Heliconius melpomene mocoa Brower, 1996; , TL: Colombia
= ; [NL4A, #2421f]
Heliconius melpomene bellula ; [NL4A, #2421f]
H. m. schunkei Lamas, 1976
Peru (Madre de Dios) melpomene schunkei Lamas, 1976; , TL: Peru
Heliconius melpomene aglaope ab. rubra Stichel, 1906; Genera Insectorum 37 : 26, TL: Peru
= Heliconius melpomene aglaope ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 203, pl. 20, f. 205
= ; [NL4A, #2421v]
Heliconius aglaope var. incarnata Stichel, 1906; (nom. nud.)
= ; [NL4A, #2421v]
Heliconius melpomene schunkei ; [NL4A, #2421v]
H. m. michellae Neukirchen, 1982
Peru melpomene michellae Neukirchen, 1982; , TL: Peru
Heliconius melpomene michellae ; [NL4A, #2421p]
H. m. tessa Barcant, 1982
Tobago melpomene tessa Barcant, 1982; , TL: Tobago
Heliconius melpomene tessa ; [NL4A, #2421w]
H. m. anduzei Brown & Fernández, 1985
Venezuela melpomene anduzei Brown & Fernández, 1985; , TL: Venezuela
Heliconius melpomene anduzei ; [NL4A, #2421e]
H. m. malleti Lamas, 1988
Peru, Ecuador, Colombia Heliconius melpomene malleti Lamas, 1988; , TL: Ecuador, Napo, Misahualli 500m
Heliconius melpomene aglaope f. iris Riffarth, 1907; Dt. ent. Zs. 1907 (5) : 506, pl. 5, f. 6, TL: Ecuador
= Heliconius melpomene aglaope × melpomene plesseni ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 194, pl. 20, f. 202
= ; [NL4A, #2421n (hyb.)]
Heliconius melpomene aglaope f. isolda Niepelt, 1908; Dt. ent. Zs. 1908 : 505, TL: Ecuador
= Heliconius melpomene aglaope × melpomene plesseni ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 194, pl. 20, f. 203
= ; [NL4A, #2421n (hyb.)]
Heliconius melpomene aglaope f. adonides Niepelt, 1908; Dt. ent. Zs. 1908 : 505, TL: Ecuador
= ; [NL4A, #2421n (hyb.)]
Heliconius melpomene aglaope f. rubripicta Niepelt, 1908; Dt. ent. Zs. 1908 : 505, TL: Ecuador
= ; [NL4A, #2421n (hyb.)]
Heliconius melpomene aglaope f. gisela Niepelt, 1908; Dt. ent. Zs. 1908 : 506, TL: Ecuador
= ; [NL4A, #2421n (hyb.)]
Heliconius melpomene aglaope f. anna Neustetter, 1909; , TL: Ecuador
= ; [NL4A, #2421n (hyb.)]
Heliconius melpomene aglaope f. dione Neustetter, 1909; , TL: Ecuador
= ; [NL4A, #2421n (hyb.)]
Heliconius melpomene aglaope f. gratiosa Niepelt, 1909; Berl. ent. Zs. 54 : 106, TL: Ecuador
= ; [NL4A, #2421n (hyb.)]
Heliconius amaryllis amaryllis f. rufata Stichel, 1923; , TL: Colombia
= ; [NL4A, #2421n (hyb.)]
Heliconius amaryllis amaryllis f. aglaspis Stichel, 1923; , TL: Colombia
= ; [NL4A, #2421n (hyb.)]
Heliconius melpomene aglaope f. aurofasciata Neustetter, 1928; Int. ent. Zs. 22 : 76, TL: Colombia
= Heliconius melpomene aglaope × melpomene bellula ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 180, pl. 21, f. 209
= ; [NL4A, #2421n (hyb.)]
Heliconius melpomene aglaope f. paula Neustetter, 1928; Int. ent. Zs. 22 : 77, TL: Colombia
= Heliconius melpomene aglaope × melpomene bellula ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 200, pl. 20, f. 208
= ; [NL4A, #2421n (hyb.)]
Heliconius aglaope f. paulina Niepelt, 1928; , TL: Colombia
= ; [NL4A, #2421n (hyb.)]
Heliconius aglaope f. carminata Niepelt, 1928; , TL: Colombia
= ; [NL4A, #2421n (hyb.)]
Heliconius melpomene malleti ; [EcuL]; [NL4A, #2421n]
Colombia Heliconius melpomene ssp. ; Neukirchen, MS, [NL4A, #2421ac]

Heliconius nattereri C. & R. Felder, [1865]
Brazil(Bahia, Espírito Santo). See [About maps] Heliconius nattereri C. & R. Felder, [1865]; Reise Fregatte Novara, Bd 2 (Abth. 2) (2): 375, pl. 47, f. 8, TL: Brazil, Bahia
Heliconius fruhstorferi Riffarth, 1899; Berl. ent. Zs. 43 (3/4) : 406, TL: Brazil, Espírito Santo
= ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 190, pl. 9, f. 100; [NL4A, #2422]
Heliconius nattereri, NSG Voucher Specimen [Wahlberg]
Heliconius fruhstorferi ; [EBW, (name only)]
Heliconius nattereri ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 198, pl. 9, f. 99; [NL4A, #2422]

Heliconius numata (Cramer, [1780])
Amazon. See [About maps] Papilio numata Cramer, [1780]; Uitl. Kapellen 4 (25-26a): pl. 297, f. C-D, TL: Suriname
Heliconius numata, NSG Voucher Specimen [Wahlberg]
Heliconius numata ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 199, pl. 39, f. 403; [BOW: pl. 44, f. 23]; [NL4A, #2423]
Surinam, French Guiana, Guyana Eueides pione Hübner, 1816; (repl. name), TL: Surinam
= ; [NL4A, #2423a]
Heliconius numata var. melanops Weymer, 1894; Dt. ent. Z. Iris 6 (2) : 303, TL: French Guiana
= ; [NL4A, #2423a]
Heliconius numata var. guiensis Riffarth, 1900; Berl. ent. Zs. 45 (3/4) : 198, TL: Guyana
= ; [NL4A, #2423a]
Heliconius numata isabellinus f. intermedia Boullet & Le Cerf, 1909; , TL: French Guiana
= ; [NL4A, #2423a]
Heliconius numata f. melanopors Joicey & Kaye, 1917; Trans. ent. Soc. Lond. 1916 : 425, TL: French Guiana
= ; [NL4A, #2423a]
Heliconius numata numata f. guiensis ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 191, pl. 11, f. 122
Heliconius numata numata f. melanopors ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 196, pl. 12, f. 124
Heliconius numata numata ; [NL4A, #2423a]
H. n. silvana (Stoll, 1781)
Surinam, Guyana, Venezuela, Guatemala, Brazil(Pará, Amazonas) Papilio silvana Stoll, [1781]; in Cramer, Uitl. Kapellen 4 (29-31): 143, pl. 364, f. C-D, TL: Surinam
Heliconius diffusus Butler, 1873; Cist. Ent. 1 (7) : 168, TL: Brazil, Pará
= ; [NL4A, #2423w]
Heliconius silvana var. divisus Kaye, 1906; Entomologist 39 : 53, pl. 2, f. 3, TL: Guyana
= ; [NL4A, #2423w]
Heliconius numata silvaniformis Joicey & Kaye, 1917; Ann. Mag. nat. Hist. (8) 20 (115) : 89, pl. 5, f. 7, TL: Brazil, Pará
= ; [NL4A, #2423w]
Heliconius silvana atakama Neustetter, 1931; , TL: Brazil, Amazonas
= ; [NL4A, #2423w]
Heliconius numata silvana f. diffusus ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 186, pl. 12, f. 128
Heliconius numata superioris f. silvaniformis ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 204, pl. 12, f. 127
Heliconius silvana ; [BOW: pl. 44, f. 31]
Heliconius numata silvana ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 204, pl. 39, f. 404; [EBW]; [NL4A, #2423w]
H. n. ethra (Hübner, [1831])
Brazil(Espírito Santo) numata ethra (Hübner, [1831]) (Eueides); Zuträge Samml. exot. Schmett. 3 : 35, : f. 553-554, TL: Brazil
Heliconius dryalus Hopffer, 1869; Stettin Ent. Ztg 30 : 452 (repl. name), TL: Brazil
= ; [NL4A, #2423g]
Heliconius ethra ab. brasiliensis Neustetter, 1907; , TL: Brazil, Espírito Santo
= ; [NL4A, #2423g]
Heliconius ethra ?var. hopfferi Neustetter, 1907; , TL: Brazil
= ; [NL4A, #2423g]
?Heliconius ethra ; [BOW: pl. 44, f. 11]
Heliconius numata ethra ; [NL4A, #2423g]
H. n. aristiona Hewitson, 1853
Bolivia, Peru Heliconia aristiona Hewitson, 1853; Ill. exot. Butts [1] (Heliconia I): [70], pl. [36], f. 4, TL: "Colombia"
Heliconius aristiona var. peruana Hopffer, 1879; Stettin Ent. Ztg 40 (10-12): 434, TL: Peru
= ; [NL4A, #2423c]
Heliconius aristiona var. splendidus Weymer, 1894; Dt. ent. Z. Iris 6 (2) : 334, pl. 5, f. 8, TL: Bolivia
= ; [NL4A, #2423c (hyb.)]
Heliconius aristiona ; [BOW: pl. 44, f. 1]
Heliconius numata aristiona ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 179, pl. 10, f. 106; [EBW]; [NL4A, #2423c]
H. n. aurora Bates, 1862
Brazil(Amazonas), Colombia, Bolivia, Peru Heliconius aurora Bates, 1862; Trans. linn. Soc. Lond. 23 (3) : 555, TL: S.Paulo, Amazonas
Heliconius numata var. isabellinus Bates, 1862; Trans. linn. Soc. Lond. 23 (3) : 554, TL: Brazil, Amazonas
= ; [NL4A, #2423e (hyb.)]
Heliconius euphrasius Weymer, 1890; in Weymer & Maassen, Reisen Süd-America : 116, pl. 2, f. 2, TL: Llanos de San Martin (Colombia) & Ecuador
= Heliconius numata euphone ; [EcuL]
= ; [NL4A, #2423e (hyb.)]
Heliconius gordius Weymer, 1894; Dt. ent. Z. Iris 6 (2) : 312, pl. 4, f. 9, TL: Brazil, Amazonas
= ; [NL4A, #2423e (hyb.)]
Heliconius floridus Weymer, 1894; Dt. ent. Z. Iris 6 (2) : 329, pl. 5, f. 4, TL: Bolivia,Peru
= ; [NL4A, #2423e (hyb.)]
Heliconius seraphion Weymer, 1894; Dt. ent. Z. Iris 6 (2) : 330, pl. 5, f. 5, TL: Peru
= ; [NL4A, #2423e (hyb.)]
Heliconius gradatus Weymer, 1894; Dt. ent. Z. Iris 6 (2) : 335, pl. 5, f. 9, TL: Peru
= ; [NL4A, #2423e (hyb.)]
Heliconius aristiona aurora f. michaeli Neustetter, 1931; , TL: Peru
= ; [NL4A, #2423e]
Heliconius aristiona aurora f. deflavata Neustetter, 1932; , TL: Peru
= ; [NL4A, #2423e (hyb.)]
Heliconius aristiona idalion f. excelsa Neustetter, 1932; , TL: Colombia
= ; [NL4A, #2423e (hyb.)]
Heliconius aristiona euphrasius ; Riffarth, 1901, Berl. ent. Zs. 46 (1/2) : 59
Heliconius numata isabellinus ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 194, pl. 11, f. 113; [EBW]
Heliconius numata aurora ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 180, pl. 10, f. 111; [EcuL]; [NL4A, #2423e]
H. n. euphone C. & R. Felder, 1862
Colombia, Ecuador Heliconius euphone C. & R. Felder, 1862; Wien. ent. Monats. 6 (12) : 418, TL: "Nova Granada, apud flumina Orinoco et Meta. In provincia Bogotá prope Fusagasugá"
Heliconius idalion Weymer, 1894; Dt. ent. Z. Iris 6 (2) : 337, TL: Colombia
= ; [NL4A, #2423h]
Heliconius tleson Riffarth, 1901; (nom. nud.) [?]
= ; [NL4A, #2423h]
Heliconius aganippe Riffarth, 1901; Berl. ent. Zs. 46 (1/2) : 60 (nom. nud.)
= ; [NL4A, #2423h]
Heliconius aristiona lepidus Riffarth, 1907; Dt. ent. Zs. 1907 (5) : 503, pl. 5, f. 2, TL: Ecuador, "Villavicencio"
= ; [EcuL]; [NL4A, #2423h]
Heliconius idalion f. confluens Neustetter, 1912; (preocc. Lathy, 1906), TL: Colombia
= ; [NL4A, #2423h]
Heliconius euphone ab. nephele Seitz, 1916
= ; [NL4A, #2423h]
Heliconius euphone ab. confluxus Seitz, 1916; (nom. nud.)
= ; [NL4A, #2423h]
Heliconius aristiona euphorbius Stichel, 1923; , TL: Colombia
= ; [NL4A, #2423h]
Heliconius mixta Apolinar, 1927; , TL: Colombia
= ; [NL4A, #2423h]
Heliconius numatus laura Neustetter, 1932; , TL: Colombia
= ; [NL4A, #2423h (hyb.)]
Heliconius numata euphone f. lepidus ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 195, pl. 10, f. 103
Heliconius numata euphone ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 188, pl. 10, f. 102; [EBW, (name only)]; [EcuL]; [NL4A, #2423h]
H. n. messene C. & R. Felder, 1862
Colombia Heliconius messene C. & R. Felder, 1862; Wien. ent. Monats. 6 (12) : 418, TL: Colombia
Heliconius sikinos Riffarth, 1901; Berl. ent. Zs. 46 (1/2) : 60 (nom. nud.)
= ; [NL4A, #2423q]
Heliconius aristiona messene f. juncta Neustetter, 1925; , TL: Colombia
= ; [NL4A, #2423q]
Heliconius aristiona colombiana Apolinar, 1927; , TL: Colombia
= ; [NL4A, #2423q]
Heliconius aristiona messene f. euphrasinus Neustetter, 1928; , TL: Colombia
= ; [NL4A, #2423q]
Heliconius numata messene ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 197, pl. 9, f. 101; [EBW, (name only)]; [NL4A, #2423q]
H. n. bicoloratus Butler, 1873
Peru Heliconius bicoloratus Butler, 1873; Cist. Ent. 1 (7) : 167, TL: Peru, Nauta
Heliconius bicoloratus var. phalaris Weymer, 1894; Dt. ent. Z. Iris 6 (2) : 333, pl. 5, f. 7, TL: "Brazil, Amazonas"
= ; [NL4A, #2423f (hyb.)]
Heliconius numata aristiona f. bicoloratus ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 181, pl. 10, f. 108
Heliconius numata bicoloratus ; [EcuL]; [NL4A, #2423f]
H. n. arcuella Druce, 1874
Peru Heliconia arcuella Druce, 1874; Trans. ent. Soc. Lond. 1874 (1) : 156, TL: Peru
Heliconius elegans Weymer, 1894; Dt. ent. Z. Iris 6 (2) : 326, pl. 5, f. 3, TL: Peru
= ; [NL4A, #2423b (hyb.)]
Heliconius numatus praelautus Stichel, 1906; Genera Insectorum 37 : 10, pl. 1, f. 2, TL: Peru
= ; [NL4A, #2423b]
Heliconius numata arcuella ♀-f. praelautus ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 201, pl. 10, f. 109
Heliconius numata arcuella ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 179, pl. 10, f. 110; [NL4A, #2423b]
H. n. nubifer Butler, 1875
Brazil(Amazonas) Heliconius nubifer Butler, 1875; Ann. Mag. Nat. Hist. (4) 15 (87) : 224, TL: Brazil, Amazonas
Heliconius numata nubifer ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 199, pl. 10, f. 112; [NL4A, #2423s]
H. n. superioris Butler, 1875
Brazil(Amazonas, Pará), Venezuela Heliconius superioris Butler, 1875; Ann. Mag. Nat. Hist. (4) 15 (87) : 224, TL: Brazil, Amazonas; Pará
Heliconius numata var. maecenas Weymer, 1894; Dt. ent. Z. Iris 6 (2) : 303, TL: Brazil, Pará
= ; [NL4A, #2423y]
Heliconius numatus sincerus Riffarth, 1907; Dt. ent. Zs. 1907 (5) : 501, TL: Brazil?,Venezuela
= ; [NL4A, #2423y]
Heliconius numata superioris f. sincerus ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 204, pl. 12, f. 125
Heliconius numata superioris ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 205, pl. 12, f. 126; [EBW]; [NL4A, #2423y]
H. n. robigus Weymer, 1875
Heliconius robigus Weymer, 1875; Stettin ent. Ztg 36 (7/9) : 382, pl. 2, f. 5, TL: "Venezuela"
Heliconia numilia Herrich-Schäffer, 1865; (nom. nud.)
= ; [NL4A, #2423v]
Heliconius numata robigus ; [NL4A, #2423v]
H. n. aulicus Weymer, 1883
Venezuela Heliconius aulicus Weymer, 1883; Stettin ent. Ztg 45 (1-3) : 19, pl. 1, f. 1, TL: [Venezuela]
Heliconius numata aulicus ; [NL4A, #2423d]
H. n. lenaeus Weymer, 1891
Ecuador? Heliconius lenaeus Weymer, 1891; Stettin ent. Ztg 51 (7-12) : 284, TL: "Caucathal, Loja, Huayabamba"
Heliconius colepta Riffarth, 1901; Berl. ent. Zs. 46 (1/2) : 58 (nom. nud.)
= ; [NL4A, #2423n]
Heliconius numata lenaeus ; [EcuL]; [NL4A, #2423n]
H. n. lyrcaeus Weymer, 1891
Peru Heliconius lyrcaeus Weymer, 1891; Stettin ent. Ztg 51 (7-12) : 286, TL: Peru?
Heliconius timaeus Weymer, 1894; Dt. ent. Z. Iris 6 (2) : 331, pl. 5, f. 6, TL: Peru
= ; [NL4A, #2423o]
Heliconius numata lyrcaeus ; [NL4A, #2423o]
H. n. geminatus Weymer, 1894
Brazil(Amazonas) Heliconius geminatus Weymer, 1894; Dt. ent. Z. Iris 6 (2) : 299, pl. 4, f. 3, TL: Brazil, Amazonas
Heliconius numata geminatus ; [NL4A, #2423i]
H. n. illustris Weymer, 1894
Peru Heliconius mirus var. illustris Weymer, 1894; Dt. ent. Z. Iris 6 (2) : 298, TL: Peru
Heliconius silvana mirificus Stichel, 1906; Genera Insectorum 37 : 12, pl. 1, f. 3a, TL: Peru
= ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 197, pl. 11, f. 117; [NL4A, #2423l]
Heliconius novatus subnubilus Stichel, 1906; Genera Insectorum 37 : 14, pl. 1, f. 4a, TL: Peru
= ; [NL4A, #2423l]
Heliconius numata illustris f. subnubilis[sic, recte subnubilus] ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 205, pl. 11, f. 116
Heliconius numata illustris ; [NL4A, #2423l]
H. n. mavors Weymer, 1894
Brazil(Amazonas) Heliconius superioris var. mavors Weymer, 1894; Dt. ent. Z. Iris 6 (2) : 305, TL: [Brazil, Amazonas]
Heliconius numata superioris ab. translata Joicey & Kaye, 1917; Ann. Mag. nat. Hist. (8) 20 (115) : 91, TL: Brazil, Amazonas
= ; [NL4A, #2423p]
Heliconius numata mavors f. translata ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 207, pl. 11, f. 121
Heliconius numata mavors ; [NL4A, #2423p]
H. n. mirus Weymer, 1894
Bolivia Heliconius mirus Weymer, 1894; Dt. ent. Z. Iris 6 (2) : 296, pl. 4, f. 2, TL: Bolivia
Heliconius leopardus Weymer, 1894; Dt. ent. Z. Iris 6 (2) : 319, pl. 5, f. 11, TL: Bolivia
= ; [NL4A, #2423r]
Heliconius spadicarius Weeks, 1901; Proc. New England Zool. Club 2 : 72, TL: Bolivia, near Coroice
= ; [NL4A, #2423r]
Heliconius arethusa Riffarth, 1901; Berl. ent. Zs. 46 (1/2) : 51 (nom. nud.)
= ; [NL4A, #2423r]
Heliconius novatus obscurior Stichel, 1906; Genera Insectorum 37 : 15, pl. 1, f. 5, TL: Bolivia
= Heliconius numata leopardus × numata spadicarius ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 199, pl. 11, f. 119
= ; [NL4A, #2423r]
Heliconius novatus artemis Riffarth, 1907; Dt. ent. Zs. 1907 (5) : 502, pl. 5, f. 1, TL: Bolivia
= ; [NL4A, #2423r]
Heliconius insolitus Avinoff, 1926; , TL: Bolivia
= ; [NL4A, #2423r]
Heliconius novatus f. confluens Neustetter, 1931; (preocc. Lathy, 1906), TL: Bolivia
= ; [NL4A, #2423r]
Heliconius numata leopardus f. artemis ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 179, pl. 11, f. 118
Heliconius numata leopardus ; [EBW, (name only)]
Heliconius numata mirus ; [NL4A, #2423r]
H. n. tarapotensis Riffarth, 1901
Peru Heliconius aristiona tarapotensis Riffarth, 1901; Berl. ent. Zs. 46 (1/2) : 59, TL: Peru, Tarapoto
Heliconius staudingeri Weymer, 1894; Dt. ent. Z. Iris 6 (2) : 324, pl. 5, f. 2, TL: Peru
= ; [NL4A, #2423aa (hyb.)]
Heliconius staudingeri var. pretiosus Weymer, 1894; Dt. ent. Z. Iris 6 (2) : 325, TL: Peru
= ; [NL4A, #2423aa (hyb.)]
Heliconius aristiona lepidus f. gracilis Riffarth, 1907; Dt. ent. Zs. 1907 (5) : 504, pl. 5, f. 3, TL: Peru
= ; [NL4A, #2423aa]
Heliconius aristiona timaeus f. aristeus Neustetter, 1931; , TL: Peru
= ; [NL4A, #2423aa (hyb.)]
Heliconius aristiona staudingeri f. lutea Neustetter, 1931; , TL: Peru
= ; [NL4A, #2423aa (hyb.)]
Heliconius numata euphone f. gracilis ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 191, pl. 10, f. 104
Heliconius numata euphone f. tarapotensis ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 206, pl. 10, f. 105
Heliconius numata tarapotensis ; [NL4A, #2423aa]
H. n. zobrysi Fruhstorfer, 1910
Brazil(Mato Grosso) Heliconius aulicus zobrysi Fruhstorfer, 1910; Ent. Zs. 24 (35) : 194, TL: Brazil, Mato Grosso
Heliconius numata zobrysi ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 209, pl. 12, f. 130; [EBW, (name only)]; [NL4A, #2423ab]
H. n. ignotus Joicey & Kaye, 1917
Peru Heliconius ignotus Joicey & Kaye, 1917; Ann. Mag. nat. Hist. (8) 20 (115) : 89, TL: Charape, N.Peru
= Heliconius numata talboti ; [EcuL]
Heliconius numata talbot f. ignotus ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 193, pl. 11, f. 114
Heliconius numata ignotus ; [NL4A, #2423k]
H. n. talboti Joicey & Kaye, 1917
Peru Heliconius numata talboti Joicey & Kaye, 1917; Ann. Mag. nat. Hist. (8) 20 (115) : 88, pl. 5, f. 3, TL: Rentema Falls, N.Peru
Heliconius numata talboti ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 206, pl. 11, f. 115; [EBW, (name only)]; [EcuL]; [NL4A, #2423z]
H. n. pratti Joicey & Kaye, 1917
Peru Heliconius aristiona pratti Joicey & Kaye, 1917; Ann. Mag. nat. Hist. (8) 20 (115) : 90, pl. 5, f. 4, TL: Peru
Heliconius numata aristiona f. pratti ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 201, pl. 10, f. 107
Heliconius numata pratti ; [NL4A, #2423u]
H. n. peeblesi Joicey & Talbot, 1925
Venezuela Heliconius peeblesi Joicey & Talbot, 1925; Ann. Mag. Nat. Hist. (9) 16 (96) : 647, TL: Venezuela
Heliconius numata peeblesi ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 200, pl. 11, f. 120; [EBW, (name only)]; [NL4A, #2423t]
H. n. jiparanaensis Neustetter, 1931
Brazil(Rondônia) numata jiparanaensis Neustetter, 1931; , TL: Brazil, Rondônia
f. mediatrix Neustetter, 1931; , TL: Brazil, Rondônia
= ; [NL4A, #2423m]
Heliconius numata jiparanaensis ; [NL4A, #2423m]
H. n. holzingeri Fernández & Brown, 1976
Venezuela numata holzingeri Fernández & Brown, 1976; , TL: Venezuela
Heliconius numata holzingeri ; [NL4A, #2423j]
H. n. sourensis Brown, 1976
Brazil(Pará) numata sourensis Brown, 1976; , TL: Brazil, Pará
Heliconius numata sourensis ; [NL4A, #2423x]

Heliconius pardalinus Bates, 1862
Ecuador, Peru, Bolivia, Amazon. See [About maps] Heliconius eucoma var. pardalinus Bates, 1862; Trans. linn. Soc. Lond. 23 (3) : 555, TL: S.Paulo, U. Amazonas
Heliconius pardalinus, NSG Voucher Specimen [Wahlberg]
Heliconius pardalinus ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 200, pl. 13, f. 137; [BOW: pl. 44, f. 24]; [NL4A, #2424]
Brazil(Amazonas) Heliconius pardalinus pardalinus ; [NL4A, #2424a]
H. p. radiosus Butler, 1873
Brazil(Pará, Amazonas) Heliconius radiosus Butler, 1873; Cist. Ent. 1 (7) : 166, TL: Brazil, Pará
Heliconius fortunatus Weymer, 1883; Stettin ent. Ztg 45 (1-3) : 21, pl. 1, f. 4, TL: [Brazil, Amazonas]
= ; [NL4A, #2424i]
Heliconius spurius Weymer, 1894; Dt. ent. Z. Iris 6 (2) : 309, pl. 4, f. 7, TL: Brazil, Amazonas
= ; [NL4A, #2424i]
Heliconius hecale radiosus ; [EBW, (name only)]
Heliconius pardalinus radiosus ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 202, pl. 16, f. 165; [NL4A, #2424i]
H. p. maeon Weymer, 1891
Heliconius maeon Weymer, 1891; Stettin ent. Ztg 51 (7-12) : 287
Heliconius pardalinus maeon ; [NL4A, #2424g]
H. p. dilatus Weymer, 1894
Peru Heliconius aerotome var. dilatus Weymer, 1894; Dt. ent. Z. Iris 6 (2) : 323, pl. 5, f. 1, TL: Peru
Heliconius pardalinus radiosus f. colorata Stichel, 1919; , TL: Peru
= ; [NL4A, #2424d]
Heliconius pardalinus dilatus ; [NL4A, #2424d]
H. p. lucescens Weymer, 1894
Brazil(Pará) Heliconius pardalinus var. lucescens Weymer, 1894; Dt. ent. Z. Iris 6 (2) : 321, TL: Brazil, Pará
Heliconius pardalinus lucescens ; [NL4A, #2424f]
H. p. sergestus Weymer, 1894
Peru Heliconius sergestus Weymer, 1894; Dt. ent. Z. Iris 6 (2) : 339, pl. 5, f. 12, TL: Peru
Heliconius sergestus ab. nicacura Michael, 1926; , TL: Peru
= ; [NL4A, #2424j (hyb.)]
Heliconius sergestus ; [BOW: pl. 44, f. 27 (text only)]
Heliconius pardalinus sergestus ; [NL4A, #2424j]
H. p. tithoreides Staudinger, 1900
Peru Heliconius tithoreides Staudinger, 1900; Dt. Ent. Z. Iris 12 (2): 404, TL: Peru
Heliconius tithoreides f. minor Riffarth, 1908; (nom. nud.)
= ; [NL4A, #2424k]
f. garleppi Neustetter, 1928; , TL: Peru
= ; [NL4A, #2424k]
Heliconius pardalinus tithoreides ; [NL4A, #2424k]
H. p. butleri Brown, 1976
Peru Heliconius pardalinus butleri Brown, 1976; 217, TL: Peru
Heliconius pardalinus butleri ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 181, pl. 13, f. 136; [NL4A, #2424c]
H. p. orteguaza Brown, 1976
Colombia pardalinus orteguaza Brown, 1976; , TL: Colombia, Caqueta, Rio Oreguaza
Heliconius pardalinus orteguaza ; [EcuL]; [NL4A, #2424h]
H. p. ariadne Neukirchen, 1995
Bolivia pardalinus ariadne Neukirchen, 1995; , TL: Bolivia
Heliconius pardalinus ariadne ; [NL4A, #2424b]
H. p. julia Neukirchen, 2000
Ecuador pardalinus julia Neukirchen, 2000; , TL: Ecuador
Heliconius pardalinus julia ; [NL4A, #2424e]

Heliconius peruvianus (C. & R. Felder, 1859)
Peru. See [About maps] Heliconia peruvianus C. & R. Felder, 1859; , TL: Peru, [Quito]
Heliconius aganice Riffarth, 1901; (nom. nud.)
= ; [NL4A, #2425]
Heliconius peruvianus, NSG Voucher Specimen [Wahlberg]
Heliconius charithonia peruviana ; [EBW, (name only)]
Heliconius peruvianus ; [NL4A, #2425]

Heliconius ricini (Linnaeus, 1758)
-. See [About maps] Papilio ricini Linnaeus, 1758; , TL: America
Heliconius ricini, NSG Voucher Specimen [Wahlberg]
?Heliconius vicini[sic] ; [BOW: pl. 44, f. 28 (missp.?)]
Heliconius ricini ; [EBW]; [NL4A, #2426]
[ricini.jpg] 500x698 (~63Kb) Plate No. 30 © MERIAN

Surinam Papilio myrti Fabricius, 1775; , TL: Surinam
= ; [NL4A, #2426a]
Papilio polyhymnia Shaw, 1806
= ; [NL4A, #2426a]
Heliconius ricini ricini ; [NL4A, #2426a]
H. r. insulanus (Stichel, 1909)
Trinidad Eueides ricini insulanus Stichel, 1909; Soc. ent. 23 (23) : 179, TL: Trinidad
= Heliconius ricini ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 194, pl. 36, f. 367
insularis[sic, recte insulanus] ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 194
Heliconius insularis[sic, recte insulanus] ; [EBW]
Heliconius ricini insulanus ; [NL4A, #2426b]

Heliconius sapho (Drury, 1782)
Mexico - Colombia, Ecuador. See [About maps] Papilio sapho Drury, 1782; Illust. Nat. Hist. Exot. Insects 3: index, : 54, pl. 38, f. 4, TL: "Jamaica"
Heliconius sapho, NSG Voucher Specimen [Wahlberg]
Heliconius sappho[sic] ; Godman & Salvin, [1881], Biol. centr.-amer., Lep. Rhop. 1: 157, 3: pl. 18, f. 5-6
Heliconius sapho ; [EBW]; [BOW: pl. 44, f. 29]; [NL4A, #2427]
[sapho-1.jpg] 589x449 (~32Kb) Belize, Central America, Photo © John Shuey

Heliconius sapho sapho ; [NL4A, #2427a]
H. s. leuce Doubleday, 1847
Mexico - Costa Rica, Guatemala Heliconia leuce Doubleday, 1847; Gen. diurn. Lep. (1): 102, TL: "Brazil"
Heliconius leuce Bates, 1864; Ent. mon. Mag. 1 (3): 58 (preocc. Doubleday, 1847), TL: Guatemala
= ; [NL4A, #2427e]
Heliconius sapho leuce, Listing of Mexican Butterflies [ILMB]
Heliconius sapho leuce, Butterflies of America [BOA]
Heliconia luce[sic] ; Godman & Salvin, [1881], Biol. centr.-amer., Lep. Rhop. 1: 156 (missp.)
Heliconius leuce ; Godman & Salvin, [1881], Biol. centr.-amer., Lep. Rhop. 1: 156, 3: pl. 18, f. 3-4
Heliconius sapho leuce ; [BCR, 197]; [EBW]; [NL4A, #2427e]

Larva on Passiflora pittieri [BCR]

H. s. hewitsoni Hewitson, 1875
Costa Rica, Panama Heliconia hewitsoni Hewitson, 1875; Ill. exot. Butts [1] (Heliconia VIII): [83], pl. [43], f. 27, TL: Panama
Heliconius hewitsoni Staudinger, 1876; Verh. zool.-bot. Ges. Wien, 25 (1) : 98 (preocc. Hewitson, 1875), TL: Panama
= ; [NL4A, #2427d]
Heliconius hewitsoni, NSG Voucher Specimen [Wahlberg]
Heliconius hewitsoni ; Godman & Salvin, [1881], Biol. centr.-amer., Lep. Rhop. 1: 158; Godman & Salvin, [1901], Biol. centr.-amer., Lep. Rhop. 2: 669; [BCR, 197]
Heliconius sapho hewitsoni ; [NL4A, #2427d]

Larva on Passiflora pittieri [BCR]

H. s. chocoensis Brown & Benson, 1975
Colombia sapho chocoensis Brown & Benson, 1975; , TL: Colombia
Heliconius sapho chocoensis ; [NL4A, #2427c]
H. s. candidus Brown, 1976
Ecuador Heliconius sapho candidus Brown, 1976; 223, TL: Ecuador
Heliconius sapho candidus ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 182, pl. 37, f. 383; [EcuL]; [NL4A, #2427b]

Heliconius sara (Fabricius, 1793)
Mexico - Amazon Basin. See [About maps] Papilio sara Fabricius, 1793; Ent. Syst. 3 (1): 167, TL: Guaiana
Heliconius sara, NSG Voucher Specimen [Wahlberg]
Heliconius sara ; [EBW]; [BOW: pl. 44, f. 30]; [NL4A, #2428]

Larva on Passiflora auriculata [BCR]

Surinam, Brazil(Pará) Papilio rhea Cramer, [1775]; Uitl. Kapellen 1 (1-7): 85, pl. 54, f. C, D (preocc. Poda, 1761), TL: Surinam
= Heliconius sara thamar ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 202, pl. 39, f. 399
= ; [NL4A, #2428a]
Nerëis thamar Hübner, [1806]; Samml. exot. Schmett. 1 : pl. [16]
= ; [NL4A, #2428a]
Heliconius rhea ab. albinea Riffarth, 1899; Berl. ent. Zs. 43 (3/4) : 408, TL: Surinam
= ; [NL4A, #2428a]
Heliconius sara praesignis Stichel, 1919; , TL: Brazil, Pará
= ; [NL4A, #2428a]
Heliconius sara thamar f. nana Stammeshaus, 1982; , TL: Surinam
= ; [NL4A, #2428a]
Heliconius (Heliconius) sarae[sic] thamar ; Brown & Mielke, 1967, J. Lep. Soc. 21 (2): 96
Heliconius sara thamar f. albinea ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 177, pl. 37, f. 379
Heliconius rhea ; [EBW]
Heliconius sara thamar ; [EcuL]
Heliconius sara sara ; [NL4A, #2428a]
H. s. apseudes (Hübner, [1813])
sara apseudes (Hübner, [1813]) (Nerëis); , TL: Brazil
Nerëis apseudes Hübner, [1808]; (nom. nud.)
= ; [NL4A, #2428b]
Heliconius sara apseudes ; [EBW]; [NL4A, #2428b]
H. s. magdalena Bates, 1864
Colombia, Panama, Venezuela Heliconius magdalena Bates, 1864; Ent. mon. Mag. 1 (3): 57, TL: Colombia; Panama
= Heliconius sara sara ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 196, pl. 37, f. 377
Heliconius apseudes var. magdalena ab. albimacula Staudinger, [1897]; Dt. Ent. Z. Iris 9 (2) : 292, TL: Colombia
= ; [NL4A, #2428f]
Heliconius sara ab. albula Riffarth, 1900; Berl. ent. Zs. 45 (3/4) : 208, TL: Venezuela, La Guayra
= ; [NL4A, #2428f]
Heliconius sarae[sic] lilianae Emsley, 1965; (nom. nud.)
= ; [NL4A, #2428f]
Heliconius magdalena ; Godman & Salvin, [1881], Biol. centr.-amer., Lep. Rhop. 1: 159, 3: pl. 18, f. 13; Godman & Salvin, [1901], Biol. centr.-amer., Lep. Rhop. 2: 669
Heliconius sara magdalena ; [NL4A, #2428f]
H. s. sprucei Bates, 1864
Ecuador Heliconius sprucei Bates, 1864; Ent. mon. Mag. 1 (3): 57, TL: Ecuador, Chimborazo
Heliconius sara sprucei ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 205, pl. 37, f. 378; [EBW]; [EcuL]; [NL4A, #2428g]
H. s. veraepacis Bates, 1864
Guatemala Heliconius veraepacis Bates, 1864; Ent. mon. Mag. 1 (3): 57, TL: Guatemala
Heliconius veraepacis ; Godman & Salvin, [1881], Biol. centr.-amer., Lep. Rhop. 1: 159, 3: pl. 18, f. 12; Godman & Salvin, [1901], Biol. centr.-amer., Lep. Rhop. 2: 669
Heliconius sara veraepacis ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 208, pl. 36, f. 374; [EBW, (name only)]; [NL4A, #2428i]
H. s. theudela Hewitson, 1874
Costa Rica, Panama Heliconia theudela Hewitson, 1874; Ent. mon. Mag. 10 (118) : 224, TL: Panama
Heliconius theudela ; Godman & Salvin, [1881], Biol. centr.-amer., Lep. Rhop. 1: 157, 3: pl. 18, f. 9-10; Godman & Salvin, [1901], Biol. centr.-amer., Lep. Rhop. 2: 669
?Heliconius sara theudda[sic] ; [EBW, (name only, missp.?)]
Heliconius sara theudela ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 207, pl. 37, f. 376; [BCR, 197]; [NL4A, #2428h]
H. s. brevimaculata Riffarth, 1901
Colombia sara brevimaculata Riffarth, 1901; , TL: Colombia
Heliconius apseudes var. magdalena ab.? (var.?) brevimaculata Staudinger, [1897]; Dt. Ent. Z. Iris 9 (2) : 292, TL: Colombia
= ; [NL4A, #2428c]
Heliconius sara brevimaculata ; [EBW, (name only)]; [NL4A, #2428c]
H. s. fulgidus Stichel, 1906
Guatemala - Panama, Costa Rica Heliconius sara fulgidus Stichel, 1906; Genera Insectorum 37 : 41, TL: Costa Rica
= Heliconius sara sara ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 190, pl. 37, f. 375
Heliconius sara fulgidus ; [EBW, (name only)]; [BCR, 197]; [NL4A, #2428e]
H. s. williami Neukirchen, 1994
Trinidad sara williami Neukirchen, 1994; , TL: Trinidad
Heliconius sara williami ; [NL4A, #2428j]
H. s. elektra Neukirchen, 1998
Colombia sara elektra Neukirchen, 1998; , TL: Colombia
Heliconius sara elektra ; [NL4A, #2428d]

Heliconius telesiphe (Doubleday, 1847)
-. See [About maps] Heliconia telesiphe Doubleday, 1847; Gen. diurn. Lep. (1): 103, pl. 15, f. 2, TL: Bolivia
Heliconius telesiphe, NSG Voucher Specimen [Wahlberg]
Heliconius telesiphe ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 206, pl. 35, f. 360; [EBW]; [BOW: pl. 44, f. 33]; [NL4A, #2429]
Bolivia Heliconius telesiphe telesiphe ; [NL4A, #2429a]
H. t. sotericus Salvin, 1871
Ecuador, Peru Heliconius sotericus Salvin, 1871; Ann. Mag. nat. Hist. (4) 4 (21) 7 (42): 414, TL: Guaymay, Ecuador
Heliconius telesiphe sotericus ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 204, pl. 35, f. 358; [NL4A, #2429c]; [EBW]; [EcuL]
H. t. cretacea Neustetter, 1916
Peru telesiphe cretacea Neustetter, 1916; , TL: "French Guyana" [= Peru]
= Heliconius telesiphe sotericus ; [EcuL]
Heliconius telesiphe ab. nivea Kaye, 1916; Ent. Rec. J. Var. 28 : 195, TL: Peru
= ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 198, pl. 35, f. 359; [NL4A, #2429b]
Heliconius telesiphe nivea ; [EBW, (name only)]
Heliconius telesiphe cretacea ; [NL4A, #2429b]

Heliconius timareta Hewitson, 1867
-. See [About maps] Heliconia timareta Hewitson, 1867; Trans. ent. Soc. Lond. (3) 5 : 563, TL: Ecuador
Heliconius timareta, NSG Voucher Specimen [Wahlberg]
Heliconia timareta ; Hewitson, 1871, Ill. exot. Butts [1] (Heliconia VI): [80], pl. [41], f. 21
Heliconius timareta ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 207, pl. 28, f. 286; [EBW]; [NL4A, #2430]
Ecuador Heliconius contiguus Weymer, 1890; in Weymer & Maassen, Reisen Süd-America : 118, pl. 2, f. 6, TL: Agoyán, 1600m [Ecuador, Tungurahua]
= ; [NL4A, #2430a]
Heliconius timareta ab. richardi Riffarth, 1900; Berl. ent. Zs. 45 (3/4) : 201, TL: Ecuador, Santa Inéz
= ; [NL4A, #2430a]
Heliconius melpomene timareta ab. virgata Stichel, 1902; Insekten-Börse 19 (45) : 355, Berl. ent. Z. 48 (1/2): (s.b.) 14, TL: Ecuador, Santa Inez
= ; [NL4A, #2430a]
Heliconius melpomene timareta f. insolita Riffarth, 1907; Dt. ent. Zs. 1907 (5) : 508, TL: Ecuador
= ; [NL4A, #2430a]
Heliconius melpomene timareta f. peregrina Stichel, 1909; Soc. ent. 23 (23) : 178, TL: Ecuador
= ; [NL4A, #2430a]
Heliconius timareta f. strandi Neustetter, 1928; , TL: Ecuador (Zarayaquillo)
= ; [NL4A, #2430a]
Heliconius timareta f. insolita ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 193, pl. 28, f. 285
Heliconius timareta f. virgata ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 208, pl. 38, f. 392
Heliconius timareta timareta ; [NL4A, #2430a]; [EcuL]
H. t. timoratus Lamas, 1998
Peru timareta timoratus Lamas, 1998; , TL: Peru, Amazonas, Cordillera del Cóndor, Puesto de Vigilancia 3 (Alfonso Ugarte), 1000-2000m (03°55'S, 78°26'W)
Heliconius timareta timoratus ; [EcuL]; [NL4A, #2430b]

Heliconius tristero Brower, 1996
Colombia. See [About maps] Heliconius tristero Brower, 1996; , TL: Colombia
Heliconius tristero ; [NL4A, #2431]

Heliconius wallacei Reakirt, 1866
Guianas - Peru. See [About maps] Heliconius wallacei Reakirt, 1866; Proc. Acad. nat. Sci. Philad. 18 (3) : 242, TL: [Brazil]
Heliconius wallacei, NSG Voucher Specimen [Wahlberg]
Heliconius wallacei ; [BOW: pl. 45, f. 4]; [NL4A, #2432]
Brazil(Pará) Heliconius clytia ab. parvimaculata Riffarth, 1900; Berl. ent. Zs. 45 (3/4) : 207, TL: Brazil, Pará, "Obidos, Santarem, Faro Yuruty"
= ; [NL4A, #2432a (hyb.)]
Heliconius clytia f. flavescens ab. wucherpfennigi Krüger, 1933; , TL: Brazil, Pará
= ; [NL4A, #2432a (hyb.)]
ab. erichi Krüger, 1933; , TL: Brazil, Pará
= ; [NL4A, #2432a (hyb.)]
Heliconius wallacei wallacei f. parvimaculata ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 200, pl. 8, f. 87
Heliconius wallacei wallacei ; [NL4A, #2432a]
H. w. mimulinus Butler, 1873
Colombia Heliconius mimulinus Butler, 1873; Cist. Ent. 1 (7) : 168, TL: Bogota, Colombia
= Heliconius wallacei wallacei ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 197, pl. 8, f. 84
Heliconius wallacei mimulinus ; [NL4A, #2432f]
H. w. flavescens Weymer, 1891
Ecuador, Peru, Guyana, Bolivia wallacei flavescens Weymer, 1891; , TL: Ecuador
?Heliconius clytia var. colon Weymer, 1891; Stettin ent. Ztg 51 (7-12) : 291 [?], TL: [Brazil]
= ?; [NL4A, #2432d (hyb.)]
Heliconius clytia var. sulphurea Staudinger, [1897]; Dt. Ent. Z. Iris 9 (2) : 295 (preocc. Weymer, 1894), TL: Peru
= ; [NL4A, #2432d]
Heliconius hagar Kirby, [1900]; , TL: Guyana,"Surinam"
= ; [NL4A, #2432d]
f. quadrimaculata Neustetter, 1925; , TL: Bolivia
= ; [NL4A, #2432d]
Heliconius clytia f. flavescens ab. graphitica Krüger, 1933; , TL: Peru
= ; [NL4A, #2432d]
f. inez Stammeshaus, 1982; , TL: Surinam
= ; [NL4A, #2432d]
Heliconius wallacei flavescens ; [EBW]; [EcuL]; [NL4A, #2432d]
H. w. colon Weymer, 1891
Brazil(Amazonas), Surinam Heliconius clytia var. colon Weymer, 1891; Stettin ent. Ztg 51 (7-12) : 291, TL: [Brazil]
Papilio clytia Cramer, 1776; (preocc. Linnaeus, 1758), TL: Surinam
= ; [NL4A, #2432c]
Heliconius clytia var. elsa Riffarth, 1899; Berl. ent. Zs. 43 (3/4) : 407, TL: Surinam
= ; [NL4A, #2432c]
Heliconius wallacei brevimaculata ab. halli Kaye, 1919; Ann. Mag. nat. Hist. (9) 3 (14) : 217, TL: Brazil, Amazonas
= ; [NL4A, #2432c (hyb.)]
Heliconius wallacei wallacei f. elsa ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 187, pl. 8, f. 85
Heliconius wallacei wallacei f. halli ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 191, pl. 8, f. 88
Heliconius wallacei elsa ; [EBW, (name only)]
Heliconius wallacei clytia ; [EBW, (name only)]
Heliconius wallacei colon ; [NL4A, #2432c]
H. w. kayei Neustetter, 1929
Trinidad wallacei kayei Neustetter, 1929; (repl. name), TL: Trinidad
Heliconius wallacei latus Kaye, 1925; (preocc. Riffarth, 1900), TL: Trinidad
= ; [NL4A, #2432e]
Heliconius wallacei kayei ; [NL4A, #2432e]
Heliconius wallacei kayi[sic, recte kayei] ; [EBW, (name only)]
H. w. araguaia Brown, 1976
Brazil(Goiás) Heliconius wallacei araguaia Brown, 1976; 216, TL: Brazil, Goiás
Heliconius wallacei araguaia ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 179, pl. 8, f. 86; [NL4A, #2432b]

Heliconius xanthocles Bates, 1862
Guianas, Ecuador, Peru. See [About maps] Heliconius xanthocles Bates, 1862; Trans. linn. Soc. Lond. 23 (3) : 561, TL: Guyana,"French Guiana"
Heliconius xanthocles, NSG Voucher Specimen [Wahlberg]
Heliconius xanthocles ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 209, pl. 7, f. 79; [NL4A, #2433]; [BOW: pl. 45, f. 5]
Guyana Heliconius xanthocles xanthocles ; [NL4A, #2433a]
H. x. melete C. & R. Felder, 1865
Colombia Heliconius melete C. & R. Felder, [1865]; Reise Fregatte Novara, Bd 2 (Abth. 2) (2): 376, TL: Colombia
Heliconius melinus f. fassli Neustetter, 1912; , TL: Colombia
= ; [NL4A, #2433g]
Heliconius cethosia Seitz, 1912; , TL: Colombia
= ; [NL4A, #2433g]
Heliconius xanthocles flavosia Kaye, 1920; Ent. Rec. J. Var. 32 (10) : 187, TL: E.Colombia, Villavicencia
= ; [NL4A, #2433g]
Heliconia cethosia completa Oberthür, 1920; Étud. Lépid. Comp. 17 : 37, pl. 522, f. 4334, TL: Colombia
= Heliconius xanthocles flavosia ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 184, pl. 8, f. 82
= ; [NL4A, #2433g]
Heliconius xanthocles melete ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 196, pl. 8, f. 83; [EBW]; [NL4A, #2433g]
H. x. paraplesius Bates, 1867
Brazil(Maranhão) Heliconius paraplesius Bates, 1867; Trans. ent. Soc. Lond. (3) 5 : 540, TL: Brazil, Maranhão
Heliconius olede Riffarth, 1901; (nom. nud.)
= ; [NL4A, #2433k]
Heliconius xanthocles paraplesius ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 200, pl. 8, f. 81; [NL4A, #2433k]
H. x. vala Staudinger, 1885
French Guiana xanthocles vala Staudinger, 1885; , TL: French Guiana
Heliconia caternaulti Oberthür, 1902; Étud. d'Ent. 21 : 24, pl. 11, f. 131, TL: French Guiana
= ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 182, pl. 7, f. 80; [NL4A, #2433o]
Heliconius vala ; [EBW]
Heliconius xanthocles caternaulti ; [EBW, (name only)]
?Heliconius xanthocles f. vola[sic] ; [BOW: pl. 45, f. 5 (text only, missp.?)]
Heliconius xanthocles vala ; [NL4A, #2433o]
H. x. melior Staudinger, 1897
Peru Heliconius melior Staudinger, [1897]; Dt. Ent. Z. Iris 9 (2) : 307, TL: Peru
Heliconius xanthocles melior ; [EBW, (name only)]; [NL4A, #2433h]
H. x. melittus Staudinger, 1897
Peru Heliconius (melete var.?) melittus Staudinger, [1897]; Dt. Ent. Z. Iris 9 (2) : 309, TL: Peru
?Heliconius xanthocles mellitus[sic] ; [EBW, (missp.?)]
Heliconius xanthocles melittus ; [NL4A, #2433i]
H. x. donatia Fruhstorfer, 1910
Brazil(Mato Grosso) xanthocles donatia Fruhstorfer, 1910; Ent. Zs. 24 (35) : 194, TL: Brazil, Mato Grosso
Heliconius xanthocles melete f. meridionalis Neustetter, 1925; , TL: Brazil, Mato Grosso
= ; [NL4A, #2433d]
Heliconius xanthocles donatia ; [NL4A, #2433d]
H. x. similatus Zikán, 1937
Brazil(Amazonas) xanthocles similatus Zikán, 1937; , TL: Brazil, Amazonas
Heliconius xanthocles similatus ; [NL4A, #2433n]
H. x. cleoxanthe H. & R. Holzinger, 1972
Venezuela xanthocles cleoxanthe H. & R. Holzinger, 1972; , TL: Venezuela
Heliconius xanthocles cleoxanthe ; [NL4A, #2433c]
H. x. explicata Brown, 1976
Colombia xanthocles explicata Brown, 1976; , TL: Colombia
Heliconius xanthocles melete f. explicata Stichel, 1923; , TL: Colombia
= ; [NL4A, #2433e]
Heliconius xanthocles melete f. paranympha Stichel, 1923; , TL: Colombia
= ; [NL4A, #2433e]
Heliconius xanthocles melete f. latior Neustetter, 1932; , TL: Colombia
= ; [NL4A, #2433e]
Heliconius xanthocles explicata ; [NL4A, #2433e]
H. x. quindecim Lamas, 1976
Peru xanthocles quindecim Lamas, 1976; , TL: Peru
Heliconius xanthocles quindecim ; [NL4A, #2433l]
H. x. hippocrene Holzinger & Brown, 1982
Bolivia xanthocles hippocrene Holzinger & Brown, 1982; , TL: Bolivia
Heliconius hippocrene Stichel, 1906; (nom. nud.)
= ; [NL4A, #2433f]
Heliconius xanthocles hippocrene ; [NL4A, #2433f]
H. x. napoensis Holzinger & Brown, 1982
Ecuador xanthocles napoensis Holzinger & Brown, 1982; , TL: Napo, Oriente, Rio Anzu
Heliconius xanthocles napoensis ; [EcuL]; [NL4A, #2433j]
H. x. rindgei Holzinger & Brown, 1982
Colombia xanthocles rindgei Holzinger & Brown, 1982; , TL: Colombia
Heliconius xanthocles rindgei ; [NL4A, #2433m]
H. x. zamora Holzinger & Brown, 1982
Ecuador xanthocles zamora Holzinger & Brown, 1982; , TL: road to Loja, 4-10km W of Zamora, 1100-1500m
Heliconius xanthocles zamora ; [EcuL]; [NL4A, #2433p]
H. x. buechei Neukirchen, 1992
Venezuela xanthocles buechei Neukirchen, 1992; , TL: Venezuela
Heliconius xanthocles buechei ; [NL4A, #2433b]


Unknown or unplaced taxa

Heliconia hippola Hewitson, 1867; Ill. exot. Butts [1] (Heliconia V): [77], pl. [40], f. 13
= Heliconius melpomene melpomene × ethilla metalilis ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 192, pl. 17, f. 173
Heliconius ethilla latona × melpomene melpomene
= Heliconius xanthocles ; [NL4A, #2433]
hippola ; [NL4A, 273 (hyb.)]

Heliconius rubellius Grose-Smith & Kirby, 1892; Rhop. Exot. [2] 1: (Heliconius) 1, pl. 1, f. 1-2, TL: Colombia
Heliconius melpomene melpomene × cydno cydno ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 203, pl. 28, f. 279
rubellius ; [NL4A, 273 (hyb.)]

Heliconius wernickei Weymer, 1906; Dt. ent. Z. Iris 19 (2) : 68, TL: Colombia
= Heliconius melpomene melpomene × cydno cydno ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 209
Heliconius cydno wernickei ; [EBW, (name only)]
wernickei ; [NL4A, 273 (hyb.)]

Heliconius fassli Seitz, 1912; , TL: [Colombia]
fassli ; [NL4A, 273 (hyb.)]

Heliconius emilius Weymer, 1912; Ent. Rdsch. 29 (11) : 73, TL: Colombia
= Heliconius melpomene melpomene × cydno cydno ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 187, pl. 27, f. 278
emilius ; [NL4A, 273 (hyb.)]

Heliconius seitzi Neustetter, 1916; , TL: [Colombia]
seitzi ; [NL4A, 274 (hyb.)]

Heliconius aventina Oberthür, 1925; Etud. Lépid. Comp. 22 : 81, f. 6, TL: Colombia
= Heliconius cydno hermogenes × melpomene melpomene ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 180, pl. 26, f. 267
aventina ; [NL4A, 274 (hyb.)]

Heliconius cydno chioneus f. eugenius Niepelt, 1928; , TL: Colombia
cydno chioneus f. eugenius ; [NL4A, 274 (hyb.)]

Heliconius heurippa f. tenuifasciata Neustetter, 1928; , TL: ?
melpomene melpomene × cydno cydno
= Heliconius xanthocles ; [NL4A, #2433]
heurippa f. tenuifasciata ; [NL4A, 274 (hyb.)]

Heliconius vulcanus cythera f. concinna Stichel, 1906; Genera Insectorum 37 : 28, TL: Ecuador
melpomene cythera × cydno alithea
= Heliconius xanthocles ; [NL4A, #2433]
vulcanus cythera concinna ; [NL4A, 274 (hyb.)]
Heliconius melpomene cythera f. concinna ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 184, pl. 18, f. 184

Heliconius seraphini Talbot, 1932; Bull. Hill Mus. 4 (3) : 191, TL: French Guiana
= Heliconius numata numata × melpomene thelxiope ; Ackery & Smiles, 1976, Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 204, pl. 12, f. 123
numata numata × melpomene thelxiopeia
= Heliconius xanthocles ; [NL4A, #2433]
seraphini ; [NL4A, 274 (hyb.)]

Heliconia celia Doubleday, 1847; (nom. nud.), TL: Ecuador
celia ; [NL4A, 274]

Heliconia choanita Herrich-Schäffer, 1865; (nom. nud.), TL: ?
choanita ; [NL4A, 274]

Heliconia claudina Doubleday, 1847; (nom. nud.), TL: Ecuador
claudina ; [NL4A, 274]

Heliconia cratena Herrich-Schäffer, 1865; (nom. nud.), TL: ?
cratena ; [NL4A, 274]

Heliconia elvira Doubleday, 1847; (nom. nud.), TL: Ecuador
elvira ; [NL4A, 274]

Heliconia eroxena Doubleday, 1847; (nom. nud.), TL: Bolivia
eroxena ; [NL4A, 274]

Heliconia marinetta Doubleday, 1847; (nom. nud.), TL: French Guiana
marinetta ; [NL4A, 274]

Heliconia matuna Herrich-Schäffer, 1865; (nom. nud.), TL: ?
matuna ; [NL4A, 274]

Heliconia mexicana Herrich-Schäffer, 1865; (nom. nud.), TL: ?
mexicana ; [NL4A, 274]

Heliconia nerina Doubleday, 1847; (nom. nud.), TL: Brazil
nerina ; [NL4A, 274]

Heliconia phaëna Doubleday, 1847; (nom. nud.), TL: Venezuela
phaëna ; [NL4A, 274]

Heliconia siphelia Herrich-Schäffer, 1865; (nom. nud.), TL: ?
siphelia ; [NL4A, 274]

Heliconia tessala Herrich-Schäffer, 1865; (nom. nud.), TL: ?
tessala ; [NL4A, 274]

Heliconia thusania Herrich-Schäffer, 1865; (nom. nud.), TL: ?
thusania ; [NL4A, 274]

Heliconia vilis Herrich-Schäffer, 1865; (nom. nud.), TL: ?
vilis ; [NL4A, 274]

Heliconia xithelia Herrich-Schäffer, 1865; (nom. nud.), TL: ?
xithelia ; [NL4A, 274]

Heliconia zerbinetta Doubleday, 1847; , TL: Bolivia
zerbinetta ; [NL4A, 274]


19.5.2013 (40)

References:
[About maps]
Warning! The maps are automatically generated from the textual information, and the process does not always produce acceptable result; See about maps for more info.
[BAMONA]
Opler, Pavulaan, Stanford & Pogue; Butterflies and Moths of North America; Mountain Prairie Information Node
[BCA]
electronic; BIOLOGIA CENTRALI-AMERICANA
[BCR]
DeVries; The Butterflies of Costa Rica 1; Papilionidae, Pieridae, Nymphalidae
[BHL]
The; Biodiversity Heritage Library; and on; Internet Archive; Note: The linking to the content of Internet Archive is somewhat delicate -- notify me when links break!
[BOA]
Interactive Listing of American Butterflies; Listado Interactivo de la Mariposas Americanas; Butterflies of America
[BOW]
Lewis, 1974; Butterflies of the World; ISBN 0 245 52097 X
[CiNii]
http://ci.nii.ac.jp/
[EBW]
Smart, Paul, 1976; The Illustrated Encyclopedia of the Butterfly World; ISBN 0 600 31381 6
[EcuL]
Tommaso & Racheli, 2001; Fragmenta entomologica 33 (2): 213-380; An annotated list of Ecuadorian Butterflies (Lepidoptera: Papilionidae, Pieridae, Nymphalidae)
[FLNHM]
Florida Museum of Natural History ; Bulletin of the Allyn Museum
[HBN]
Francis Hemming, 1937; HÜBNER, A bibliographical and systematic account of the entomological works of Jacob Hübner and of the supplements thereto by Carl Geyer, Gottfried Franz von Frölich and Gottlieb August Wilhelm Herrich-Schäffer, Vol 1 & 2
[ILMB]
Interactive Listing of Mexican Butterflies
[JLS]
Journal of the Lepidopterists' Society; the original J. Lep. Soc. articles
[John Shuey]
e-mail: jshuey at tnc.org; Director of Conservation Science; Indiana Office of The Nature Conservancy; http://www.nature.org
[LAMAS-2005]
Gerardo Lamas; Entomofauna 26 (6): 57-100; A Bibliography of the Zoological Publications of Hans Fruhstorfer (1886* - 1922+)
[LAMAS-NEO]
Gerardo Lamas; Bibliography of butterflies; An Annocated Bibliography of the Neotropical Butterflies and Skipper (Lepididoptera: Papilionoidea and Hesperioidea); Revised Electronic Edition (2007)
[MERIAN]
Merian, M. S., 1705; Metamorphosis insectorum Surinamensium. Of te verandering der Surinaamsche insecten. Waar in de Surinaamsche rupsen en wormen met alle des zelfs veranderingen, na het leven afgebeeld en beschreeven worden, zynde elk geplaast op die gewassen, bloemen en vruchten, daar sy op gevonden zyn; waar in ook de generatie der kirkvorschen, wonderbaare padden, hagedissen, slangen, spinnen en mieren werden vertoond en beschreeven, alles in America na het leven en levensgroote geschildert en beschreeven; pp. 1-60, pl. 1-60; Image from Francisco Welter-Schultes
[NAB]
Pyle, 1981; Field Guide to North American Butterflies, National Audubon Society; ISBN 0-394-51914-0
[NACL]
Hodges, 1983; Check List of the Lepidoptera of America North of Mexico
[NL4A]
Gerardo Lamas, 2004; Atlas of Neotropical Lepidoptera; Checklist: Part 4A; Hesperioidea - Papilionoidea
[Opler]
Opler & Warren, 2003 (April); Butterflies of North America. 2. Scientific Names List for Butterfly Species of North America, north of Mexico.
[Simon Coombes]
1 Park Street, Plymouth, PL3 4BL, UK; 01752 607854; e-mail: Simon Coombes; http://www.butterfly-guide.co.uk
[Wahlberg]
Niklas Wahlberg; The NSG's voucher specimen database; Nymphalidae Systematics Group


Some related literature:
Ackery & Smiles, 1976
An illustrated list of the type-specimens of the Heliconiidae (Lepidoptera: Nymphalidae) in the British Museum (Natural History) Bull. Brit. Mus. nat. Hist. (Ent.) 32 : 173-214, pl. 1-39
Attal, 1999
Nouveaux Nymphalidae néotropicaux (Lepidoptera, Rhopalocera) Bull. Soc. ent. Fr. 104 (4): 369-374, 3fgs
Bates, 1862
Contributions to an Insect Fauna of the Amazon Valley. Heliconidae Trans. linn. Soc. Lond. 23 (3) : 495-566, pl. 55-56
Bates, 1864
New species of butterflies from Guatemala and Panama, collected by Osbert Salvin and F. du Cane Godman, Esqs. Ent. mon. Mag. 1 (1): 1-6 (1864), (2): 31-35 (1864), (3): 55-59 (1864), (4): 81-85 (1864), (5): 113-116 (1864), (6): 126-131 (1864), (7): 161-164 (1864), (8): 178-180 (1865), (9): 202-205 (1865)
Bates, 1866
New species of butterflies from Guatemala and Panama Ent. mon. Mag. 3 (27): 49-52 (1866), (28): 85-88 (1866), (30): 133-136 (1866), (31): 152-157 (1866)
Bates, 1867
On a Collection of Butterflies formed by Thomas Belt, Esq., in the interior of the Province of Maranham, Brazil Trans. ent. Soc. Lond. (3) 5 : 535-546
Boisduval, 1870
Considerations sur des Lépidoptères envoyés du Guatemala à M. de l'Orza Considérations Lépid. Guatemala: 100pp
Brown, 1973
The Heliconians of Brazil. Part V. Three new subspecies from Mato Grosso and Rondônia Bull. Allyn Mus. 13: 1-19
Brown & Benson, 1975
The Heliconians of Brazil. (Lepidoptera: Nymphalidae). Part VI. Aspects of the biology and ecology of Heliconius demeter, with description of four new subspecies Bull. Allyn Mus. 26: 1-19
Brown & Mielke, 1967
Lepidoptera of the Central Brazil Plateau. I. Preliminary list of Rhopalocera J. Lep. Soc. 21 (2): 77-106
Brown & Mielke, 1967
Lepidoptera of the Central Brazil Plateau. I. Preliminary list of Rhopalocera (continued) J. Lep. Soc. 21 (3): 145-168
Butler, 1865
Description of six new species of diurnal Lepidoptera in the British Museum collection Proc. zool. Soc. Lond. 1865 (2) : 430-434, pl. 25
Butler, 1869
Descriptions of several new Species of Nymphalidian Rhopalocera Ann. Mag. nat. Hist. (4) 3 (13) : 17-21, pl. 9
Butler, 1873
Descriptions of new species of Lepidoptera Cist. Ent. 1 (7) : 151-177
Butler, 1875
On some new Species of Butterflies from Tropical America Ann. Mag. Nat. Hist. (4) 15 (87) : 222-225
Butler, 1877
On some Heliconoid Danainae and Heliconidae in the Collection of the British Museum Cistula Ent. 2 : 149-151
Cramer, [1775]
Uitlandsche Kapellen (Papillons exotiques) Uitl. Kapellen 1 (1-7): 1-132, pl. 1-84 (1775), (8): 133-155, pl. 85-96 (1776)
Cramer, [1777]
Uitlandsche Kapellen (Papillons exotiques) Uitl. Kapellen 2 (9-16): 1-152, pl. 97-192 (1777)
Cramer, [1780]
Uitlandsche Kapellen (Papillons exotiques) Uitl. Kapellen 4 (25-26a): 1-28, pl. 289-304 (1780)
Dewitz, 1877
Neue Schmetterlinge des Berliner museums Mitt. Münch. Ent. Ver. 1 : 85-91, pl. 2
Doubleday, 1847
The genera of diurnal Lepidoptera, comprising their generic characters, a notice of their habitats and transformations, and a catalogue of the species of each genus; illustrated with 86 plates by W. C. Hewitson Gen. diurn. Lep. (1): 1-18, pl. A, 1-4 (1-2) (1846), (1): 19-132, pl. 4*, 5-25, 28 (3-14) (1847), (1): 133-200, pl. 1*, 26, 27, 29, 31-44 (15-23) (1848), (1): 201-242, pl. 30, 45-52, 56-58, 60-62, 64 (24-31) (1849), (1): 243-250 [?], (2): 243-326, pl. 53-55, 63, 65, 66 (32-38) (1850), (2): 327-466, pl. 59, 67-77 (39-50) (1851), (2): 467-534, 143, 144, pl. 78-80 (Index) (51-54) (1852)
Druce, 1874
Descriptions of Fifteen new species of Diurnal Lepidoptera, chiefly from South America Trans. ent. Soc. Lond. 1874 (1) : 155-160
Drury, 1782
Illustrations of natural history; wherein are exhibited ... Illust. Nat. Hist. Exot. Insects 3 : 1-76, pl. 1-50
Dyar, 1903
A List of North American Lepidoptera and Key to the Literature of this Order of Insects Bull. U.S. nat. Mus. 52 : 1-723
Dyar, 1913
Results of the Yale Peruvian expedition of 1911. Lepidoptera Proc. U.S. natn. Mus. 45 (2006) : 627-649
Fabricius, 1775
Systema Entomologiae, sistens Insectorum Classes, Ordines, Genera, Species, Adiectis Synonymis, Locis, Descriptionibus, Observationibus. Syst. Ent.: 832pp
Fabricius, 1777
Genera Insectorum eorumque characters naturales secundum numerum, figuram, situm et proportionem ... Genera Insectorum : 1-310
Fassl, 1912
Neue Heliconiusformen aus Kolumbien Ent. Rdsch. 29 (8) : 55-56
Fassl, 1914
Tropische Reisen. Das obere Caucatal und die Westcordillere Ent. Rdsch. 31 (7): 35-38, (8): 42-46, (9): 50-52, (10): 57-58
C. & R. Felder, 1862
Specimen faunae lepidopterologicae riparum fluminis Negro superioris in Brasilia septentrionali Wien. ent. Monats. 6 (3): 65-80, (4): 109-126, (6): 175-192, (7): 229-235
C. & R. Felder, 1862
Lepidoptera nova Columbiae Wien. ent. Monats. 6 (12) : 409-427
C. & R. Felder, [1865]
Reise der österreichischen Fregatte Novara um die Erde in den Jahren 1857, 1858, 1859 unter den Behilfen des Commodore B. von Wüllerstorf-Urbair. Zoologischer Theil. Band 2. Abtheilung 2. Lepidoptera. Rhopalocera Reise Fregatte Novara, Bd 2 (Abth. 2) (1): 1-136, pl. 1-21 (Rhop.) (C. & R. Felder, [1865]), (2): 137-378, pl. 22-47 (Rhop.) (C. & R. Felder, 1865), (3): 379-536, pl. 48-74 (Rhop.) (C. & R. Felder, 1867), (4): 537-548, pl. 75-120, (Inhalts-Verz.) 1-9 (pl. 1-74), (Felder & Rogenhofer, 1874) , (5): pl. 121-140, (erklärung) 1-10 (pl. 75-107), (Inhalts-Verz.) 1-20 (pl. 108-140), (Felder & Rogenhofer, 1875)
Fruhstorfer, 1907
Verzeichniss der von Herrn Dr. Theodor Koch-Grünberg am oberen Waupes 1903-1905 gesammelten Rhopaloceren mit Besprechung verwandter Arten Stettin ent. Ztg 68 (1): 117-164 (May), (2): 207-309, pl. 1 (July)
Fruhstorfer, 1910
Drei neue Heliconidenformen Ent. Zs. 24 (35) : 194 (3 December)
Godart, 1819
in Latreille & Godart, Encyclopédie Méthodique. Histoire naturelle Entomologie, ou histoire naturelle des crustacés, des arachnides et des insectes Encyclopédie Méthodique. 9 (1): 3-328 (1819), (2): 329-828 ([1824])
Godman & Salvin, 1877
Description of twelwe new species and a new genus of Rhopalocera from Central America Proc. zool. Soc. Lond. 1877 (1) : 60-64
Godman & Salvin, [1881]
Biologia Centrali-Americana. Rhopalocera. Vol. 1. (1879-1886) Biol. centr.-amer., Lep. Rhop. 1: 1-487, 3: pl. 1-24,24a,25-112
Godman & Salvin, [1901]
Biologia Centrali-Americana. Rhopalocera. (1887-1901) Biol. centr.-amer., Lep. Rhop. 2: 1-782, 3: pl. 1-112
Grose-Smith, 1898
Descriptions of new Species of Butterflies from South Africa Ann. Mag. nat. Hist. (7) 2 (7) : 70-73
Grose-Smith & Kirby, 1892
Rhopalocera Exotica, being Illustrations of New, Rare, and Unfigured Species of Butterflies Rhop. Exot. [1] 1: (Ornithoptera) 1-2, pl. 1 (1887), [1] 2: (Ornithoptera) 3-4, pl. 2 (1892), [1] 3: (Ornithoptera) [5-6] 1-2, pl. 3 (1900), [1] 3: (Ornithoptera) 7-8, pl. 4 (1901), [1] 1: (Papilio) 1-2, pl. 1 (1887), [1] 1: (Papilio) 3-5,7-14, pl. 2-6 (1888), [1] 1: (Papilio) 15-20, pl. 7-9 (1890), [1] 1: (Papilio) 21-25, pl. 10-11 (1891), [1] 2: (Papilio) 27-31, pl. 12-13 (1893), [1] 2: (Papilio) 33-35, pl. 14 (1894), [1] 2: (Papilio) 37-38, pl. 15 (1895), [1] 2: (Papilio) 39-40, pl. 16 (1897), [1] 3: (Papilio) 41-46, pl. 17-19 (1899), [1] 3: (Papilio) 47-48, pl. 20 (1900), [1] 3: (Papilio) 49-50, pl. 21 (1901), [1] 3: (Papilio) 51-55, pl. 22-23 (1902), [1] 1: (Delias) 1-3, pl. 1 (1889), [1] 2: (Delias) 5-7, 9-11, pl. 2-3 (1893), [1] 2: (Delias) 12-18, pl. 4-5 (1895), [1] 2: (Delias) 20-22, pl. 6 (1896), [1] 3: (Delias) 24-26, pl. 7 (1897), [1] 3: (Delias) 28-35, pl. 8-9 (1901), [1] 3: (Delias) 36-38, pl. 10 (1902), [1] 1: (Pieridae) 1-3, pl. 1 (1888), [1] 2: (Pinacopteryx) 1-4, pl. 1 (1893), [1] 1: (Callosune) 1-3, pl. 1 (1889), [1] 3: (Huphina) 1-7, pl. 1-2 (1898), [1] 3: (Huphina) [8],9-10, pl. 3 (1899), [1] 1: (Elodina) 1-4, pl. 1 (1890), [1] 1: (Belenois) 1-4, pl. 1 (1892), [1] 2: (Belenois) 5-8, pl. 2 (1892), [1] 3: (Mylothris) 1-3, 5-7, pl. 1-2 (1900), [1] 1: (Ixias) 1-3, pl. 1 (1888), [1] 2: (Appias) 1-3, pl. 1 (1894), [1] 1: (Pierinae) 5-7, pl. 2 (1889), [1] 2: (Dismorphia) 1-4, pl. 1 (1896), [1] 2: (Dismorphia) 5-9, 11-15, pl. 2-3 (1897), [2] 1: (Asthipa) 1-3, pl. 1 (1890), [2] 2: (Asthipa) 5-6, pl. 2 (1896), [2] 2: (Ravadeba) 7, pl. 1 (1896), [2] 2: (Mycalesis) 1-4, pl. 1 (1895), [2] 2: (Mycalesis) 5-12, pl. 2-3 (1896), [2] 3: (Mycalesis) 13-18, pl. 4-5 (1902), [2] 3: (Libythea) 1-2, pl. 1 (1902), : (Ragadia-Argyronympha-Hypocysta) : 1-6, pl. [1] (1895), [2] 2: (Pedaliodes) 1-5, pl. 1 (1893), [2] 2: (Pedaliodes) 7-9, pl. 2 (1894), [2] 2: (Pedaliodes) 10-14, pl. 3 (1895), [2] 3: (Pedaliodes) 15-20, pl. 4-5, (Oxeoschistus) 1 (1900), [2] 3: (Pronophila) 1-2, pl. 1, (Oxeoschistus) 2-3, pl. 1 (1900), [2] 2: (Elymnias) 1-4, pl. 1 (1896), [2] 3: (Elymnias) 5-6, pl. 2, (Bruasa) 6-7, pl. 1 (1897), [2] 1: (Morpho) 1-2, pl. 1 (1890), [2] 3: (Morphotenaris-Hyantis) 1-3, pl. [1] (1898), [2] 2: (Tenaris) 1-3, 5-7, pl. 1-2 (1894), [2] 2: (Tenaris) 9-11, pl. 3 (1895), [2] 2: (Tenaris) 13-15, pl. 4 (1896), [2] 3: (Tenaris) 17-18, pl. 5 (1898), [2] 1: (Heliconius) 1-3, pl. 1 (1892), [2] 1: (Acraea) 1-6, pl. 1-2 (1889), [2] 1: (Acraea) 7-10, pl. 3 (1892), [2] 2: (Acraea) 11-14, pl. 4 (1892), [2] 2: (Acraea) 15-18, pl. 5 (1894), [2] 3: (Acraea) 19-29, pl. 6-8, (Planema) 29-30, pl. [1] (1901), [2] 1: (Cethosia) 1-2, pl. 1 (1889), [2] 3: (Cirrhochroa) 1, pl. [1], (Cupha) 2-4, pl. [1] (1898), [2] 2: (Terinos) 1-2, pl. 1 (1894), [2] 1: (Thaleropis) 1-3, pl. 1 (1891), [2] 2: (Doleschallia) 1-3,5-7, pl. 1-2 (1893), [2] 2: (Kallima) 1-3, pl. 1 (1892), [2] 1: (Hypolimnas) 1-2, pl. 1 (1891), [2] 3: (Hypolimnas) 3-4, pl. 2 (1897), [2] 3: (Hypolimnas) 5-6, pl. 3 (1898), [2] 1: (Euxanthe) 1-3, pl. 1 (1890), [2] 1: (Hestina) 1-3, pl. 1 (1891), [2] 1: (Cymothoe) 1-4, pl. 1 (1890), [2] 3: (Limenitis) 1-3, pl. 1 (1899), [2] 2: (Neptis) 1-4, pl. 1 (1895), [2] 3: (Neptis) 5-11, pl. 2-3 (1899), [2] 3: (Neptis) 13-17, pl. 4-5 (1900), [2] 1: (Euthalia-Tanaecia) 1-3, pl. 1 (1889), [2] 1: (Euthalia) 5-10, pl. 2-3 (1891), [2] 2: (Euthalia) 11-14, pl. 4 (1894), [2] 3: (Euthalia) 15, pl. 5, (Dichorragia) 16, pl. 1 (1898), [2] 1: (Apatura) 1-3, pl. 1 (1892), [2] 1: (Euphaedra) 1-2, pl. 1 (1889), [2] 1: (Euryphene) 1-3, pl. 1 (1889), [2] 1: (Charaxes) 1-2,[3-4] pl. 1-2 (1887), [2] 1: (Charaxes) [5], pl. 3 (1888), [2] 1: (Charaxes) 7-9, pl. 4 (1890), [2] 1: (Charaxes) 11-13, pl. 5 (1891), [2] 1: (Mynes) 1-2, pl. 1 (1888), [2] 2: (Mynes) 3-6, pl. 2 (1894), [2] 2: (Prothoe) 1-2, pl. 1 (1896), [3] 3: (Erycinidae) 1-10, pl. 1-2 (1902), [3] 2: (Dicallaneura-etc) 1-6, pl. 1 (1897), [3] 3: (Dicallaneura-etc) 7-11, pl. 2 (1901), [3] 1: (Lycaenidae) 1-4,[5-9] 1-5, pl. 1-2 (1887), [3] 1: (Lycaenidae) 11-17, 19-22, pl. 3-5 (1888), [3] 1: (Lycaenidae) 23-25, 27-29, 31-33, pl. 6-8 (1889), [3] 1: (Lycaenidae) 35-48, pl. 9-11 (1890), [3] 1: (Lycaenidae) 49-51,53-64, pl. 12-15 (1891), [3] 2: (Lycaenidae) 65-68, pl. 16 (1892), [3] 1: (Lycaenidae) [69-72] 57-60, pl. 17 (1892), [3] 2: (Lycaenidae) [73-77] 61-65,79-84, pl. 18-19 (1892), [3] 2: (Lycaenidae) 85-107, pl. 20-23 (1893), [3] 2: (Lycaenidae) 109-120, pl. 24-25 (1894), [3] 3: (Lycaenidae) 121-125, pl. 26 (1900), [3] 3: (Lycaenidae) 127-131, 133-141, pl. 27-29 (1901), [3] 3: (Lycaenidae) 143-146, pl. 30 (1902), [3] 2: (Hypochrysops) 1-6, pl. 1 (1894), [3] 2: (Hypochrysops) 7-13, 15-22, pl. 2-3 (1895), [3] 2: (Thysonotis) 23-25, 27-31, pl. 1-2 (1895), [3] 2: (Waigeum) 34-38, pl. 1 (1896), [3] 2: (Thysonotis) 39-46, pl. 3-4 (1896), [3] 2: (Holochila) 1-4, pl. 1 (1896), [3] 2: (Holochila) 5-8, pl. 2 (1897), [3] 2: (Pseudonotis) 1-3, pl. [1], (Epimastidia) 4-5, pl. [1], (Lampides) 5-8, pl. [1] (1897), [3] 2: (Lampides) 9-10, pl. 2, (Thysonotis) 47, pl. 5 (1897), [3] 3: (Holochila) 9-11, pl. 3 (1898), [3] 3: (Arhopala) 1-7, pl. 1-2 (1898), [3] 3: (Arhopala) 9-12, pl. 3, (Ilerda) 13, pl. [1], (Horaga) 14-15, pl. [1] (1902), [3] 3: (Thysonotis) 47x,48-50, pl. 6 (1898), [3] 3: (Hypochrysops) 23-26, pl. 4 (1898), [3] 3: (Holochila) 13-15, pl. 4 (1899), [3] 3: (Hypochrysops) 27-29, pl. 5, (Waigeum) [30], pl. 2 (1899), [3] 3: (Holochila) 17-19, pl. 5 (1899), [3] 3: (Thysonotis) 51-54, pl. 7 (1899), [3] 3: (Jamides) 11-14, pl. 1 (1900), [3] 3: (Hypolycaena-etc) 1-4, pl. [1] (1900), [3] 3: (Hypochrysops) 31-33, pl. 6, (Thysonotis) 55-56, pl. 8 (1900), [3]: (Additions-Corrections) 1-2,[3],vii-x
Guérin-Méneville, [1844]
Iconographie du Règne Animal de G. Cuvier. Insectes. Icon. Règne anim. Cuvier 3 (Insectes) : pl. 3-12 (1829), : pl. 13-14,21-24 (1830), : pl. 25-30,42-46,52-55,60,62,76,24-25,25bis,28bis (1831), : pl. 77-92,99,84bis (1832), : pl. 31,36-39,47-49,59,101,39bis (1833), : pl. 32-35,50,61,64-66,49bis (1834), : pl. 15-20,40,51,56-58,67-75,93-98 (1835), : pl. 100,102-104 (1836), : pl. 1-2,41,63 (1837), : 1-576 (1844)
Hall, 1930
New forms of Tropical Rhopalocera Entomologist 63 : 278-279
Hall, 1936
The butterflies of St. Kitts Entomologist 69 : 274-278
Hewitson, 1853; Hewitson, 1854; Hewitson, 1858; Hewitson, 1867; Hewitson, 1871; Hewitson, 1872; Hewitson, 1873; Hewitson, 1875
Illustrations of new species of exotic Butterflies selected chiefly from the collections of W. Wilson Saunders and William C. Hewitson. Ill. exot. Butts [1] : (Systematic Index) I, (Ornithoptera and Papilio): [1-2], pl. [1] (1855), (Papilio II): [3-4], pl. [2] (1856), (Papilio III): [5-6], pl. [3] (1858), (Papilio IV): [7-8], pl. [4] (1859), (Papilio V): [9-10], pl. [5] (1861), (Papilio VI): [11-12], pl. [6] (1864), (Papilio VII-VIII): [13-16], pl. [7-8] (1865), (Papilio IX): [17-18], pl. [9] (1868), (Papilio X-XII): [19-22], pl. [10-12] (1869), (Papilio XIII): [23-24], pl. [13] (1873), (Papilio XIV): [25-26], pl. [14] (1875), (Papilio XV): [27-28], pl. [15] (1877), (Pieris I): [29-30], pl. [16] (1852), (Pieris II): [31-32], pl. [17] (1853), (Pieris III-VI): [33-40], pl. [18-21] (1861), (Pieris VII): [41-42], pl. [22] (1862), (Pieris VIII): [43-44], pl. [23] (1866), (Euterpe): [45-46], pl. [24] (1860), (Euterpe II): [47-48], pl. [25] (1872), (Euterpe and Leptalis): [49-50], pl. [26] (1853), (Leptalis II): [51-52], pl. [27] (1857), (Leptalis III): [53-54], pl. [28] (1858), (Leptalis IV): [55-56], pl. [29] (1860), (Leptalis V-VII): [57-62], pl. [30-32] (1870), (Callidryas and Eronia): [63-64], pl. [33] (1867), (Euploea): [65-66], pl. [34] (1858), (Euploea II): [67-68], pl. [35] (1866), (Heliconia I): [69-70], pl. [36] (1853), (Heliconia II-III): [71-74], pl. [37-38] (1854), (Heliconia IV): [75-76], pl. [39] (1858), (Heliconia V): [77-78], pl. [40] (1867), (Heliconia VI): [79-80], pl. [41] (1871), (Heliconia and Tithorea VII): [81-82], pl. [42] (1873), (Heliconia VIII): [83-84], pl. [43] (1875), (Athesis): [85-86], pl. [44] (1872), (Mechanitis I): [87-88], pl. [45] (1856), (Mechanitis II-III): [89-92], pl. [46-47] (1860), (Ithomia I): [93-94], pl. [48] (1851), (Ithomia II): [95-96], pl. [49] (1852), (Ithomia III): [97-98], pl. [50] (1853), (Ithomia IV-V): [99-102], pl. [51-52] (1854), (Ithomia VI-XIII): [103-118], pl. [53-60] (1855)
Hewitson, 1867
Descriptions of some new species of diurnal Lepidoptera Trans. ent. Soc. Lond. (3) 5 : 561-566
Hewitson, 1869
Description of six new species of diurnal Lepidoptera from Nicaragua Trans. ent. Soc. Lond. 1869 (1) : 33-35
Hewitson, 1869
Equatorial Lepidoptera collected by Mr Buckley Equat. Lepid. Buckley (1): 1-16, (2): 17-32, (3): 33-48, (4): 49-80 (1870), (5): 81-96 (1877)
Hewitson, 1872
Descriptions of seven new exotic Rhopalocera Ent. mon. Mag. 9 : 83-86
Hewitson, 1874
Description of a new species of Heliconia from Central America Ent. mon. Mag. 10 (118) : 224
Hewitson, 1875
Descriptions of six new species of butterflies from South America, West Africa, and Borneo Ent. mon. Mag. 11 : 182-184
Hopffer, 1869
Berichte über Felder's Lepidopteren der Reise der Fregatte Novara (Fortsetzung) Stettin Ent. Ztg 30 : 427-453
Hopffer, 1874
Neue Lepidopteren von Peru und Bolivia Stettin ent. Ztg 35 (10-12) : 329-371
Hopffer, 1879
Exotische Schmetterlinge Stettin Ent. Ztg 40 (1-2): 47-95, (10-12): 413-454
Hübner, [1806]
Sammlung exotischer Schmetterlinge, Vol. 1 (1806 - [1819]) Samml. exot. Schmett. 1 : [1-4] (1807), : [5-12] (1807-1809), : [13-36] (1823-1824), : pl. [1-2], [5-8], [12], [15], [29], [47] (1806), : pl. [9], [11], [17], [21], [25-27], [32], [35], [43-44], [56], [60], [79], [102], [104], [116], [141], [148], [150-151], [155], [159], [200] (1807), : pl. [19], [39], [57], [73], [82], [114], [157], [167], [170], [173], [187] (1807), : pl. [3], [31], [37], [46], [54], [58], [62], [76], [84], [92], [95], [105], [112-113], [117], [137-138], [146], [154], [158], [180], [209] (1808), : pl. [16], [28], [65], [89], [142], [160], [162], [166], [182], [188], [193] (1809), : pl. [4], [22], [42], [53], [61], [88], [90], [97], [100], [107], [110], [118], [120], [140], [143], [149], [161], [171], [178], [184], [196], [202] (1809), : pl. [10], [18], [30], [41], [51], [80], [144], [172], [189] (1810), : pl. [36], [70], [87], [91], [101], [124], [129], [177], [183], [207], [210] (1811), : pl. [20], [38], [45], [55], [85], [98], [103], [108], [136], [139], [152-153], [156], [163], [179], [186], [190], [205-206] (1812), : pl. [13-14], [40], [48], [50], [67], [74], [86], [119], [121], [130], [135], [175-176], [197-199], [204], [212] (1813), : pl. [72], [75], [77-78], [83], [93], [109], [111], [115], [128], [131], [165], [174], [194] (1813), : pl. [24], [49], [52], [63-64], [66], [68-69], [71], [81], [123], [145], [181], [192] (1814), : pl. [126], [132-134], [168] (1815), : pl. [23], [96], [122], [125], [169], [195] (1816), : pl. [33], [99], [127], [187], [185], [213] (1817), : pl. [59], [106], [147], [164], [208], [211] (1818), : pl. [34], [94], [191], [201] (1819), : (Index Systematicus) [1-4] (1819 - April 1823)
Hübner, [1823]; Hübner, [1825]
Sammlung exotischer Schmetterlinge, Vol. 2 ([1819] - [1827]) Samml. exot. Schmett. 2 : pl. [90], [180], [223] (1819), : pl. [2], [12], [16], [78], [141], [182] (1820), : pl. [11], [13], [24], [77], [83], [97], [102-103], [108-109], [116-117], [122], [147], [150], [164], [183], [195], [197], [212-214] (1821), : pl. [14], [18-19], [26], [38-39], [43], [68], [70-71], [74], [84], [86], [92], [94-95], [106], [114], [121], [161], [169], [174], [177-178], [184], [194], [200], [204-205], [207-208], [210] (1821), : pl. [33], [72], [76], [115], [131], [138-139], [153], [198], [211], [218], [220] (1822), : pl. [32], [75], [81], [93], [105], [129] (1822), : pl. [1], [6], [22-23], [25], [27], [29-31], [34-37], [40-42], [48], [50-53], [56], [58-59], [64-66], [69], [73], [87], [89], [91], [96], [98-101], [110], [124], [128], [130], [132-137], [142], [155-158], [160], [162-163], [165-167], [170-173], [175-176], [187-188], [196], [199], [201-202], [206], [215-216], [219], [222], [224] (1823), : pl. [54], [118], [143], [149], [159] [179], [181], [185], [203], [217] (1824), : pl. [3-5], [8-10], [15], [20], [45], [49], [55], [60], [62-63], [67], [79-80], [88], [104], [111-113], [120], [125], [127], [140], [144-146], [148], [151-152], [154], [168], [189-193], [221], [225] (1825), : pl. [7], [17], [21], [28], [44], [46-47], [57], [61], [82], [123], [126] (1826), : pl. [85], [119], [186], [209] (1827), : (Index Systematicus) [1-4] (1827-1832)
Hübner, 1816
Verzeichniss bekannter Schmettlinge, 1816-[1826] Verz. bek. Schmett. (1): [1-3], 4-16 (1816), (2): 17-32 (1819), (3): 33-48 (1819), (4): 49-64 (1819), (5): 65-80 (1819), (6): 81-96 (1819), (7): 97-112 (1819), (8): 113-128 (1819), (9): 129-144 (1819), (10): 145-160 (1819), (11): 161-176 (1819), (12): 177-192 (1820), (13): 193-208 (1820), (14): 209-224 (1821), (15): 225-240 (1821), (16): 241-256 (1821), (17): 257-272 (1823), (18): 273-288 (1823), (19): 289-304 (1823), (20): 305-320 (1825), (21): 321-336 (1825), (22): 337-352 (1825), (23-27): 353-431 ([1825])
Hübner, [1826]
Anzeiger der im Verzeichniss bekannter Schmettlinge angenommenen Benennungen ihrer Horden, Rotten, Stämme, Familien, Vereine und Gattungen Verz. bek. Schmett. (Anz.) (1-9): 1-72
Hübner, [1831]
Zuträge zur Sammlung exotischer Schmettlinge, Vol. 3 [1824-] 1825 [-1831] Zuträge Samml. exot. Schmett. 3 : [1-3], 4-6, [7], 8-40, [41-48] (1827 - 1831), : pl. [70-100], f. 401-582 (1824), : pl. [101-103], f. 583-600 (1825)
Joicey & Kaye, 1917
New Races and Aberrations of Heliconius Ann. Mag. nat. Hist. (8) 20 (115) : 87-94, pl. 5-6
Joicey & Kaye, 1917
On a collection of Heliconine forms from French Guiana Trans. ent. Soc. Lond. 1916 : 412-431, pl. 107-108
Joicey & Kaye, 1919
Notes on a large Heliconine collection made in French Guiana in 1917, compared with similar collection made in 1915 Trans. ent. Soc. Lond. 1918 : 347-353
Joicey & Talbot, 1925
Notes on some Lepidoptera, with descriptions of new forms Ann. Mag. Nat. Hist. (9) 16 (96) : 633-653
Poulton in Joseph, 1910
The Collections of William John Burchell, D.C.L., in the Hope Department, Oxford University Museum. On the Lepidoptera Rhopalocera collected by W. J. Burchell in Brazil, 1825-1830 Ann. Mat. nat. Hist. (8) 5 (28) : 322-346
Kaye, 1906
New species of Guiana and Jamaican butterflies Entomologist 39 : 49-53, pl. 2
Kaye, 1916
New Forms of Heliconius Ent. Rec. J. Var. 28 : 194-195
Kaye, 1919
New South-American Butterflies Ann. Mag. nat. Hist. (9) 3 (14) : 215-218
Kaye, 1920
New specis and sub-species of S. American Lepidoptera Ent. Rec. J. Var. 32 (10) : 187-189
Kluk, 1780
Historyja naturalna zwierzat domowych i dzikich, osobliwie kraiowych, historyi naturalney poczatki, i gospodarstwo : potrzebnych I pozytecznych donowych chowanie, rozmnozenie, chorob leczenie, dzikich lowienie, oswaienie : za·zycie; szkodliwych zas wygubienie. 4 vols. Hist. nat. pocz. gospod. 4: -
Krüger, 1927
Eine neue Heliconiusform Int. ent. Z. 21: 86
Krüger, 1929
Neubeschreibungen Int. ent. Z. 22 : 376-377
Lathy, 1906
On three new forms of butterfly of the genus Heliconius Proc. zool. Soc. Lond. 1906 : 452-453, pl. 34
Latreille, [1817]
Insectes de l'Amérique équinoxiale, recueillis pendant de voyage de MM. de Humboldt et Bonpland. Seconde Partie. In Humbolt & Bonpland, aux régions équinoxiales du Nouveau Continent. 2. Recueil d'observations de zoologie et d'anatomie comparée; faites dans l'Océan Atlantique, dans l'intérieur du Nouveau Continent et dans la Mer du Sund, pendant les années 1799, 1800, 1801, 1802 et 1803 in Humboldt & Bonpland, Voy. rég. équinoxiales..., 2 (9): 65-96, pl. 31-34 (1813), (10): 97-138, pl. 35-43 (1817)
Linnaeus, 1758
Systema Naturae per Regna Tria Naturae, Secundum Clases, Ordines, Genera, Species, cum Characteribus, Differentiis, Symonymis, Locis. Tomis I. 10th Edition Syst. Nat. (Edn 10) 1 : 1-338, : 339-824
Linnaeus, 1764
Museum S'ae R'ae M'tis Ludovicae Ulricae Reginae Svecorum, Gothorum, Vandalorumque Mus. Lud. Ulr.: vi + 720 + [2]
Linnaeus, 1767
Systema Naturae per Regna tria Naturae, secundum Classes, Ordines, .... Editio Duocecima Reformata. Tom. 1. Part II. Syst. Nat. (Edn 12) 1 (2): -,533-1327
Ménétriés, 1855; Ménétriés, 1857
Enumeratio corporum animalium Musei Imperialis Academiae Scientiarum Petropilitanae. Classis Insectorum, Ordo Lepidopterorum Cat. lep. Petersb. 1: 1-66, pl. 1-6, (supplement) 67-112 (1855), 2: 67-97,99-144, pl. 7-14 (1857), 3: 145-161, pl. 15-18 (1863)
Müller, 1774
Des Ritters Carl von Linné königlich schwedischen Leib-arztes u. u. vollständiges Natursystem nach der zwölften lateinischen Ausgabe und nach Anleitung des hollän-dischen Houttuynischen Werks mit einer ausfürlichen Erklärung ausgefertigt Natursystem 5 (1): 1-758, pl. 1-22
Möschler, 1872
Neue exotische Schmetterlinge Stettin Ent. Ztg 33 (7-9) : 336-362
Möschler, 1877
Beiträge zur Schmetterlings-Fauna von Surinam Verh. zool.-bot. Ges. Wien 26 : 293-352, pl. 3-4
Neukirchen, 1991
Heliconius egeria mariasibyllae subspec. nov. von Pará, Brasilien Atalanta 22 (2/4): 87-92
Neukirchen, 1995
Vier neue Unterarten von Heliconius burneyi (Hübner, 1816) Atalanta 26 (1/2): 201-208
Neustetter, 1925
Drei neue Heliconius-Formen Int. ent. Z. 19 : 60-61
Neustetter, 1926
Beschreibung und Besprechung neuer und wenig gekannter Heliconius-Formen Int. ent. Z. 20 : 34-40
Neustetter, 1926
Bescprechung einer Heliconius-Ausbeute aus Französisch-Guayana und Beschreibung der neuen Formen aus derselben Int. ent. Z. 20 : 278-282, 289-293
Neustetter, 1927
Neue Heliconius-Formen Int. ent. Z. 21 : 227-230
Neustetter, 1928
Zwei neue Heliconius Int. ent. Zs. 21 : 442-444
Neustetter, 1928
Neue Heliconius-Formen Int. ent. Zs. 22 : 75-80
Neustetter, 1928
Neue und wenig bekannte Heliconius Int. ent. Zs. 22 : 245-248
Neustetter, 1928
Neue Heliconius Int. ent. Zs. 22 : 258-259
Niepelt, 1907
Zwei neue Lokalformen der Gattung Heliconius Latr. Soc. ent. 22 (6) : 42
Niepelt, 1908
Zwei neue Formen der Gattung Heliconius Berl. ent. Zs. 52 : 213-214
Niepelt, 1908
Neue Formen der Gattung Heliconius (Lep.) Dt. ent. Zs. 1908 : 505-506
Niepelt, 1909
Neue Heliconius-Formen Berl. ent. Zs. 54 : 106-107
Niepelt, 1921
Eine neue Heliconius Form v. Guatemala Int. Ent. Zs. 15 : 19
Niepelt, 1925
Eine neue Heliconius-Form von Bolivia Int. ent. Zs. 18 : 252
Niepelt, 1926
Eine neue Heliconius-Form von Bolivia Int. ent. Zs. 19 : 337
Nöldner, 1901
Zwei neue Heliconius Berl. ent. Z. 46 : 5-8
Oberthür, 1902
Variation des Heliconia Thelxiope et vesta Étud. d'Ent. 21 : 1-26, pl. 1-12
Oberthür, 1925
Lépidoptères de l'Amérique du Sud Etud. Lépid. Comp. 22 : 81-89
Reakirt, 1866
Descriptions of some new species of Diurnal Lepidoptera Proc. Acad. nat. Sci. Philad. 18 (3) : 238-249
Reakirt, 1868
Descriptions of some new species of diurnal Lepidoptera. Series III Proc. Acad. nat. Sci. Philad. 20 (1/2) : 87-91
Riffarth, 1899
Neue Heliconius-Formen Berl. ent. Zs. 43 (3/4) : 405-408
Riffarth, 1900
Die Gattung Heliconius Latr. Neubearbeitet und Beschreibung neuer Formen Berl. ent. Zs. 45 (3/4) : 183-214
Riffarth, 1901
Die Gattung Heliconius Latr. Neubearbeitet und Beschreibung neuer Formen Berl. ent. Zs. 46 (1/2) : 25-183
Riffarth, 1903
Nochmals Ch. Oberthür's Etudes d'Entomologie, vol. 21 Berl. ent. Zs. 47 (3/4) : 157-166
Riffarth, 1906
Eine neue Heliconius-form Insekten-Börse 23 (14): 56
Riffarth, 1907
Eine neue Lokalform von Heliconius batesi Riff. (Lep.) Dt. ent. Zs. 1907 (3) : 333-334
Riffarth, 1907
Neue und wenig bekannte Formen der Gattung Heliconius, nebst einer neuen Eueides-Form (Lep.) Dt. ent. Zs. 1907 (5) : 501-514, pl. 5
Riffarth, 1908
Über Neustetters "Neu Heliconius-Formen Dt. ent. Zs. 1908 (1) : 110-114
Riley, 1919
Some new Rhopalocera from Brazil collected by E. H. W. Wickham, Esq. Entomologist 52 : 181-186, 200-202
Salvin, 1871
Description of new species of butterflies from tropical America Ann. Mag. nat. Hist. (4) 4 (21) 4 (21): 163-181 (1869), 7 (42): 412-416 (1871)
Schaus, 1913
New Species of Rhopalocera from Costa Rica Proc. zool. Soc. Lond. 1913 (3) : 339-367, pl. 50-54
Staudinger, 1876
Neue Lepidopteren des südamerikanischen Faunengebietes Verh. zool.-bot. Ges. Wien, 25 (1) : 89-124
Staudinger, 1882
On three new and interesting species of Rhopalocera Proc. zool. Soc. Lond. 1882 (3) : 396-398, pl. 24
Staudinger, 1885
Exotische Tagfalter in sysmatischer Reihenfolge mit Berücksichtigung neuer Arten in Staudinger & Schatz, Exot. Schmett. 1 (1): 3-6, pl. 1-5 (1884), (2): 7-10, pl. 6-10 (1884), (3): 11-14, pl. 11-15 (1884), (4): 15-22, pl. 16-20 (1884), (5): 23-30, pl. 21-25 (1884), (6): 31-38, pl. 26-30 (1884), (7): 39-46, pl. 31-35 (1885), (8): 47-58, pl. 30-40 (1885), (9): 59-82, pl. 41-50 (1885), (10): 83-102, pl. 51-60 (1885), (11): 103-122, pl. 61-65 (1886), (12): 123-138, pl. 66-70 (1886), (13): 139-158, pl. 71-75 (1886), (14): 159-174, pl. 76-80 (1886), (15): 175-194, pl. 81-85 (1887), (16): 195-214, pl. 86-90 (1887), (17): 215-234, pl. 91-95 (1887), (18): 235-270, pl. 96-98 (1888), (19): 1-2,271-333, pl. 99-100 (1888)
Staudinger, 1894
Hochlandine Lepidoptera Dt. Ent. Z. Iris 7 (1) : 43-100, pl. 1-2
Staudinger, [1897]
Neue Heliconius-Arten und formen Dt. Ent. Z. Iris 9 (2) : 284-317 (1897)
Staudinger, 1900
Eine neue Heliconius-Form Dt. Ent. Z. Iris 12 (2): 404-406
Stichel, 1899
Heliconius nanna n. sp. Vorläufige Ent. Z. 12 (19): 143
Stichel, 1899
Neue Heliconius aus Süd-Brasilien Ent. Nachr. 25 (2) : 28-30
Stichel, 1902
Nachrichten aus dem Berliner Entomologischen Verein Insekten-Börse 19 (45) : 355, Berl. ent. Z. 48 (1/2): (s.b.) 14-16
Stichel, 1906
Lepidoptera Rhopalocera. Fam. Nymphalidae. Subfam. Heliconiinae Genera Insectorum 37 : 1-74, pl. 1-3
Stichel, 1909
Beschreibung einiger neuer Heliconiidae Soc. ent. 23 (23) : 178-179, f. 1-2
Stichel, 1912
Zwei neue Formen von Heliconius hermathena Hew. Int. ent. Z. 6 (1) : 1-2
Stichel, 1919
Anmerkungen und Zuträge zur Gattung Heliconius L. Neue Beitr. syst. Ins. 1 (15) : 119-120
Stichel, 1923
Kolumbische Heliconius (Lep., Rhop.) Dt. ent. Z. 1923 (3) : 260-270
Stoll, [1781]
Uitlandsche Kapellen (Papillons exotiques) in Cramer, Uitl. Kapellen 4 (26b-28): 29-90, pl. 305-336 (1780), (29-31): 91-164, pl. 337-372 (1781), (32-32): 165-252, 1-29, pl. 373-400 (1782)
Niepelt in Strand, 1916
Abbildungen und Beschreibungen neuer und wenig bekannter Lepidoptera aus der Sammlung W. Niepelt Lepidoptera Niepeltiana (2) : 1-26, pl. 13-17
Takahashi, 1977
List of the subfamily Heliconiinae (Lepidoptera: Nymphalidae) collected in Colombia and the adjacent area, South America, during the expedition "La expedicion japonesa a las zonas Amazonicas, 1973", with description of a new subspecies Tyô to Ga 28 (1): 9-24
Talbot, 1928
List of Rhopalocera collected by Mr. C. L. Collenette in Matto Grosso, Brazil Bull. Hill Mus. Witley 2 : 192-220
Talbot, 1932
New species and forms of Lepidoptera from South America Bull. Hill Mus. 4 (3) : 189-197
Turner, 1966
A little-recognised species of Heliconius butterfly (Nymphalidae) J. Res. Lepid. 5 : 97-112
Turner, 1966
A rare mimetic Heliconius (Lepidoptera: Nymphalidae) Proc. R. ent. Soc. Lond. (B) 35 : 128-132
Weeks, 1901
Descriptions of three new butterflies Proc. New England Zool. Club 2 : 71-74
Weymer, 1875
Exotische Lepidopteren. Stettin ent. Ztg 36 (7/9) : 368-385, pl. 1-2
Weymer, 1883
Exotische Lepidoperen. II. Stettin ent. Ztg 45 (1-3) : 7-28, pl. 1-2
Weymer, 1891
Exotische Lepidopteren. V Stettin ent. Ztg 51 (7-12) : 280-294
Weymer, 1894
Revision der ersten Gruppe der Gattung Heliconius Dt. ent. Z. Iris 6 (2) : 281-345, pl. 4-5
Weymer, 1897
Heliconius Tyndarus Dt. ent. Z. Iris 9 (2) : 317-319
Weymer, 1906
Zwei neue Heliconius-Formen Dt. ent. Z. Iris 19 (2) : 68-71
Weymer, 1912
Einige neue Heliconiden-Formen Ent. Rdsch. 29 (11) : 73-74
Weymer in Weymer & Maassen, 1890
Lepidoptera gesammelt auf einer Reise durch Colombia, Ecuador, Peru, Brasilien, Argentinien und Bolivien in den Jahren 1868-1877 von Alphons Stübel Reisen Süd-America : 1-182, pl. 1-9


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Keep in mind that the taxonomic information is copied from various sources, and may include many inaccuracies. Expert help is welcome.