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Agraulis Boisduval & Le Conte, [1835]
Agraulis Boisduval & Le Conte, [1835]; Hist. Lép. Am. Sept. (11-22): 142, TS: Papilio vanillae Linnaeus
Agraulis, NSG Voucher Specimen [Wahlberg]
 Agraulis ; Godman & Salvin, [1882], Biol. centr.-amer., Lep. Rhop. 1: 169
Agraulis (Heliconiini) ; [Wahlberg]; [NL4A, 264]
Agraulis vanillae (Linnaeus, 1758)
S.USA, Neotropical. See [About maps] Gulf Fritillary
Papilio vanillae Linnaeus, 1758; Syst. Nat. (Edn 10) 1 : 482, TL: "In Epdendro Vanilla Americes" [S.America, Surinam?]
Agraulis vanillae, NSG Voucher Specimen [Wahlberg]
Agraulis vanillae, Butterflies and Moths of North America [BAMONA]
Agraulis vanillae, Georgia Lepidoptera (USA) [GALEPS]
Agraulis vanillae vanillae, Lepidoptera of French Antilles [Pierre Zagatti]
 Dione vanillae ; Butler & Druce, 1874, Proc. Zool. Soc. Lond. 1874 (3) : 350; [BOW: pl. 12, f. 16]
Agraulis vanillae ; Godman & Salvin, [1882], Biol. centr.-amer., Lep. Rhop. 1: 171; Godman & Salvin, [1901], Biol. centr.-amer., Lep. Rhop. 2: 671; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 12; [NACL, #4413]; [BCR, 189]; [Opler]; [NL4A, #2362]
[vanillae-1.jpg] 600x851 (~83Kb) USA: LepSoc field trip to Patagonia region, Santa Cruz Co., Arizona, 3.8.2005, Photo © Markku Savela [vanillae-1l.jpg] 1300x1045 (~200Kb) larva USA: LepSoc field trip to Patagonia region, Santa Cruz Co., Arizona, 3.8.2005, Photo © Markku Savela [vanillae-2u.jpg] 1776x1162 (~273Kb) upperside USA: Alabama, Jeff., Co. 22.8.1999, Photo © Vitaly Charny
[vanillae-2v.jpg] 1315x1162 (~206Kb) underside USA: Alabama, Jeff., Co. 22.8.1999, Photo © Vitaly Charny [vanillae-3u.jpg] 1400x1050 (~162Kb) upperside USA: Alabama, 20.9.2003, Photo © Vitaly Charny [vanillae-4v.jpg] 1800x1195 (~328Kb) underside USA: Alabama, 17.9.2004, Photo © Vitaly Charny
[vanillae-5a.jpg] 700x1041 (~81Kb) USA: Alabama, 11.9.2005, Photo © Vitaly Charny [vanillae-5px.jpg] 700x1017 (~112Kb) pupa USA: Alabama, 11.9.2005, Photo © Vitaly Charny [vanillae-5py.jpg] 700x972 (~112Kb) pupa USA: Alabama, 11.9.2005, Photo © Vitaly Charny
[vanillae-6la.jpg] 1000x788 (~103Kb) larva USA: Alabama, 11.9.2005, Photo © Vitaly Charny [vanillae-6lb.jpg] 1500x1125 (~286Kb) larva USA: Alabama, 28.9.2005, Photo © Vitaly Charny [vanillae.jpg] 567x659 (~69Kb) Plate No. 25 © MERIAN

Larva on Passiflora incarnata [NAB], P. foetida, P. quadrangularis, P. ligularis, Passiflora costaricensis, P. auriculata [BCR]


A. v. vanillae
Papilio passiflorae Fabricius, 1793; Ent. Syst. 3 (1): 60 (repl. name), TL: America
= ; [NL4A, #2362a]
Agraulis vanillae vanillae ; [NL4A, #2362a]
A. v. lucina C. & R. Felder, 1862
Peru Agraulis lucina C. & R. Felder, 1862; Wien. ent. Monats. 6 (4): 110, TL: "Rio Negro" [= Peru, San Martin]
Dione vanillae catella Stichel, [1908]; (hyb.), TL: Peru
= ; [NL4A, #2362f]
Agraulis vanillae lucina ; [EBW]; [EcuL]; [NL4A, #2362f]
A. v. insularis Maynard, 1889
Antilles, Bahamas Agraulis vanillae insularis Maynard, 1889; , TL: Bahamas
Agraulis vanillae insularis, Lepidoptera of French Antilles [Pierre Zagatti]
 Agraulis vanillae insularis ; [NL4A, #2362e]; [EBW, (name only)]
A. v. galapagensis Holland, 1889
Galapagos Galapagos Fritillary
Agraulis vanillae galapagensis Holland, 1889; Proc. U.S. Natl. Mus. 12: 194, TL: [Ecuador, Galápagos Is.] Clatham Island
Dione vanillae galapagensis ; Stichel, 1938, Lep. Cat. 86: 92; Michener, 1942, Amer. Mus. Novit. No. 1215: 4; Linsley & Usinger, 1966, Proc. Cal. Acad. Sci. (4) 33: 156
Agraulis vanillae galapagensis ; Williams, 1911, Proc. Calif. Acad. Sci. 1: 298; Beebe, 1923, Zoologica 5: 55; Van Duzee, 1933, Proc. Calif. Acad. Sci. 21 (12) : 141; [EBW, (name only)]; [EcuL]; Roque-Albelo, 2004, J. Lep. Soc. 58 (1): 40, f. 9-10; [NL4A, #2362c]

Larva on Passiflora suberosa, Passiflora tridactylites Roque-Albelo, 2004, J. Lep. Soc. 58 (1): 40

A. v. maculosa (Stichel, [1908])
Brazil(Espírito Santo), Paraguay, Uruguay, Argentina vanillae maculosa (Stichel, [1908]) (Dione); , TL: Brazil, Espírito Santo; Paraguay
Papilio argentea Larrañaga, 1923; , TL: Uruguay
= ; [NL4A, #2362g]
Dione vanillae f. superargentata Giacomelli, 1925; , TL: Argentina
= ; [NL4A, #2362g]
Agraulis vanillae maculosa ; [EBW, (name only)]; Brown & Mielke, 1967, J. Lep. Soc. 21 (2): 97; [NL4A, #2362g]
A. v. forbesi Michener, 1942
Peru vanillae forbesi Michener, 1942; , TL: Lima, Peru
Agraulis vanillae forbesi ; [EBW, (name only)]; [NL4A, #2362b]
A. v. nigrior Michener, 1942
Florida vanillae nigrior Michener, 1942; , TL: Florida
Agraulis vanillae nigrior, Butterflies of America [BOA]
 Agraulis vanillae nigrior ; [NACL, #4413a]; [NL4A, #2362h]
A. v. incarnata (Riley, 1926)
Mexico, California vanillae incarnata (Riley, 1926) (Dione); , TL: Mexico, Durango
Dione vanillae ab. comstocki Gunder, 1925; , TL: California
= ; [NL4A, #2362d]
Dione vanillae insularis tr.f. fumosus Gunder, 1927; , TL: California
= ; [NL4A, #2362d]
Dione vanillae f. incarnata tr.f. margineapertus Gunder, 1928; , TL: California
= ; [NL4A, #2362d]
Dione vanillae f. incarnata tr.f. hewlettae Gunder, 1930; , TL: California
= ; [NL4A, #2362d]
Agraulis vanillae incarnata, Listing of Mexican Butterflies [ILMB]
Agraulis vanillae incarnata, Butterflies of America [BOA]
Agraulis vanillae incarnata, Butterflies of Southeastern Arizon [Bruce Walsh]
 Agraulis vanillae incarnata ; [NACL, #4413b]; [NL4A, #2362d]
Agraulis sp.
Peru. See [About maps] Agraulis sp. ; Lamas, MS, [NL4A, #2363]

17.9.2011 (1)

References:
[About maps]
Warning! The maps are automatically generated from the textual information, and the process does not always produce acceptable result; See about maps for more info.
[BAMONA]
Paul Opler, Harry Pavulaan, Ray Stanford, Michael Pogue; Butterflies and Moths of North America; Mountain Prairie Information Node
[BCA]
electronic; BIOLOGIA CENTRALI-AMERICANA
[BCR]
DeVries, Philip J.; The Butterflies of Costa Rica; Vol. I: Papilionidae, Pieridae, Nymphalidae
[BOA]
Interactive Listing of American Butterflies; Listado Interactivo de la Mariposas Americanas; Butterflies of America
[BOW]
Lewis, H. L., 1974; Butterflies of the World; ISBN 0 245 52097 X
[Bruce Walsh]
Bruce Walsh; e-mail: Bruce Walsh; The Moths of Southeastern Arizona; Butterflies of Southeastern Arizona
[EBW]
Smart, Paul, 1976; The Illustrated Encyclopedia of the Butterfly World; ISBN 0 600 31381 6
[EcuL]
Tommaso & Luigi Racheli, 2001; An annotated list of Ecuadorian Butterflies (Lepidoptera: Papilionidae, Pieridae, Nymphalidae); Fragmenta entomologica 33 (2): 213-380
[GALEPS]
Moths and Butterflies of Georgia and the Southeastern United States; James K. Adams; Dept. of Natural Science, Dalton State College, Dalton, GA ; http://www.daltonstate.edu/galeps/index.htm
[ILMB]
Interactive Listing of Mexican Butterflies
[JLS]
Journal of the Lepidopterists' Society; the original J. Lep. Soc. articles
[LAMAS-NEO]
Gerardo Lamas; Bibliography of butterflies; An Annocated Bibliography of the Neotropical Butterflies and Skipper (Lepididoptera: Papilionoidea and Hesperioidea); Revised Electronic Edition (2007)
[MERIAN]
Merian, M. S., 1705; Metamorphosis insectorum Surinamensium. Of te verandering der Surinaamsche insecten. Waar in de Surinaamsche rupsen en wormen met alle des zelfs veranderingen, na het leven afgebeeld en beschreeven worden, zynde elk geplaast op die gewassen, bloemen en vruchten, daar sy op gevonden zyn; waar in ook de generatie der kirkvorschen, wonderbaare padden, hagedissen, slangen, spinnen en mieren werden vertoond en beschreeven, alles in America na het leven en levensgroote geschildert en beschreeven; pp. 1-60, pl. 1-60; Image from Francisco Welter-Schultes
[NAB]
Pyle, R. M.; National Audubon Society: Field Guide to North American Butterflies; 1981; ISBN 0-394-51914-0
[NACL]
Hodges, Ronald W. (ed.), 1983; Check List of the Lepidoptera of America North of Mexico
[NL4A]
Gerardo Lamas, 2004; Atlas of Neotropical Lepidoptera; Checklist: Part 4A; Hesperioidea - Papilionoidea
[Opler]
Opler & Warren, 2003 (April); Butterflies of North America. 2. Scientific Names List for Butterfly Species of North America, north of Mexico.
[Pierre Zagatti]
WWW pages; Catalogue of the Lepidoptera of the French Antilles; e-mail: Pierre Zagatti
[Vitaly Charny]
Birmingham, Alabama, USA
[Wahlberg]
Niklas Wahlberg; The NSG's voucher specimen database; Nymphalidae Systematics Group

Some related literature:
Beebe, 1923
Notes on Galápagos Lepidoptera Zoologica 5: 51-59
Boisduval & Le Conte, [1835]
Histoire générale et iconographie des lépidoptères et des chenilles de l'Amérique septentrionale Hist. Lép. Am. Sept. (1): 1-16. pl. 1-3 (1829), (2): 17-24, pl. 4-6 (1829), (3/4): 25-40, pl. 7-12 ([1830]), (5/6): 41-56, pl. 13-18 ([1830]), (7/8): 57-80, pl. 19-24 ([1830]), (9/10): 81-100, pl. 25-30 (1833), (11-22): 101-196, pl. 31-65 ([1835]), (23-26): 197-228, pl. 66-78 ([1837])
Brown & Mielke, 1967
Lepidoptera of the Central Brazil Plateau. I. Preliminary list of Rhopalocera J. Lep. Soc. 21 (2): 77-106
Butler & Druce, 1874
List of the butterflies of Costa Rica, with descriptions of new species Proc. Zool. Soc. Lond. 1874 (3) : 330-370
Van Duzee, 1933
The Templeton Crocker Expedition of the California Academy of Sciences, 1932. No. 12. The Diurnal Lepidoptera of the Expedition Proc. Calif. Acad. Sci. 21 (12) : 139-146
Dyar, 1903
A List of North American Lepidoptera and Key to the Literature of this Order of Insects Bull. U.S. nat. Mus. 52 : 1-723
C. & R. Felder, 1862
Specimen faunae lepidopterologicae riparum fluminis Negro superioris in Brasilia septentrionali Wien. ent. Monats. 6 (3): 65-80, (4): 109-126, (6): 175-192, (7): 229-235
Godman & Salvin, [1882]
Biologia Centrali-Americana. Rhopalocera. Vol. 1. (1879-1886) Biol. centr.-amer., Lep. Rhop. 1: 1-487, 3: pl. 1-24,24a,25-112
Godman & Salvin, [1901]
Biologia Centrali-Americana. Rhopalocera. (1887-1901) Biol. centr.-amer., Lep. Rhop. 2: 1-782, 3: pl. 1-112
Holland, 1889
In Howard, Scientific results of explorations by the U.S. fish commission steamer Albatross. No. V. Annotated catalogue of the insects collected in 1877-'88 Proc. U.S. Natl. Mus. 12: 185-216
Linnaeus, 1758
Systema Naturae per Regna Tria Naturae, Secundum Clases, Ordines, Genera, Species, cum Characteribus, Differentiis, Symonymis, Locis. Tomis I. 10th Edition Syst. Nat. (Edn 10) 1 : 1-338, : 339-824
Linsley & Usinger, 1966
Insects of Galápagos islands Proc. Cal. Acad. Sci. (4) 33: 113-196
Michener, 1942
A review of the subspecies of Agraulis vanillae (L.) (Lepidoptera: Nymphalidae) Amer. Mus. Novit. No. 1215: 1-7
Roque-Albelo, 2004
The butterflies (Papilionoidea, Hesperoidea) of the Galápagos Islands, Ecuador: distribution, hostplants and biology J. Lep. Soc. 58 (1): 33-43
Stichel, 1938
Nymphalidae I: Subfam.: Dioninae, Anetiinae, Apaturinae Lep. Cat. 86: 1-374
Williams, 1911
Expedition of the California Academy of Sciences to the Galápagos Islands, 1905-1906. II. The butterflies and hawk-moths of the Galápagos Islands Proc. Calif. Acad. Sci. 1: 289-322

If you have corrections, comments or information to add into these pages, just send mail to Markku Savela
Keep in mind that the taxonomic information is copied from various sources, and may include many inaccuracies. Expert help is welcome.