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Plebejus Kluk, 1780
Plebejus Kluk, 1780; Hist. nat. pocz. gospod. 4: 89, TS: Papilio argus Linnaeus
Plebeius Kirby, 1871; Syn. Cat. diurn. Lep. : 653, TS: Papilio argus Linnaeus
Rusticus Hübner, [1806]; Tentamen determinationis digestionis ...: [1] (suppr. ICZN Op. 278), TS: Papilio argus Linnaeus
Lycaeides Hübner, [1819]; Verz. bek. Schmett. (5): 69, TS: Papilio argyrognomon Bergstrasser
= ; [NL4A, 140]
Lycoena Nicholl, 1901
= ; [NL4A, 140]
Plebulina Nabokov, 1945; Psyche 51 (3/4): 104, TS: Lycaena emigdionis Grinnel
= ; [NL4A, 140]
Alpherakya Zhdanko
= ; [NL4A, 140]
Plebejides Sauter, 1968
= ; [NL4A, 140]
Maurus Bálint, 1991
= ; [NL4A, 140]

Plebulina ; [NACL]; [NEN40]
Plebulina (Polyommatini) ; Eliot, 1973, Bull. Br. Mus. nat. Hist. (Ent.) 28 (6) : 450
Plebeius (Lycaenidae) ; [BRU2, 158]
Alpherakya (Lycaenidae) ; [BRU2, 176]
Plebejus (Polyommatinae) ; [NL4A, 140]
Plebeius ; Bálint, Guppy, Kondla, Johnson & Durden, 2001, Folia ent. hung. 62: (177-184)
Lycaeides (Polyommatini) ; Eliot, 1973, Bull. Br. Mus. nat. Hist. (Ent.) 28 (6) : 450
Plebejus (Polyommatini) ; Eliot, 1973, Bull. Br. Mus. nat. Hist. (Ent.) 28 (6) : 450


[sp-1.jpg] 504x502 (~51Kb) Vienna, Austria, 20 July 2000, Photo © Erich Mangl
argus? argyrognomon?

The exact identification of this species is still unknown, but tentatively assumed to belong into this group.


The ardis species-group

Plebejus eversmanni (Lang, 1884)
Kopet-Dagh, Ghissar, Darvaz, Pamirs-Alai, W.Tian-Shan, NE.Afghanistan. See [About maps] Lycaena eversmanni Lang, 1884; Rhopalocera Europa Descripta et Delineata: 370, TL: Samarkand
Lycaena eversmanni Staudinger, 1886; Stett. Ent. Ztg. 47 (4-6): 205 (preocc.), TL: Samarkand
Plebejus ardis Bálint & Johnson, 1997; Neue Ent. Nachr. 40: 14 (unnec. repl. Lycaena eversmanni Staudinger, 1886)
Lycaena eversmanni var. grumi Staudinger, 1901; in Staudinger & Rebel, Cat. Lepid. palaearct. Faunengeb., 1: 79
eversmanni ; [NEN46, 29]; [NEN46, 30]
Plebejus eversmanni ; [BOW: pl. 206, f. 23]
Plebeius eversmanni ; [BRU2: 160, pl. 70, f. 41-46]

Larva on Chesneya quinata, C. hissarica [BRU2, 160]

P. e. titania (Hemming, 1931)
Plebeius eversmanni titania Hemming, 1931; Ann. Mag. Nat. Hist. (10) 8: 535, TL: Samarkand, Uzbekistan
titania Hemming, 1931; [NEN46, 61]
P. e. badachshanus Forster, 1972
Pamirs Plebejus badachshana Forster, 1972; Ent. Z., Frankf. a.M. 82: 226
Plebejus badachshana ; [NEN40]
Plebeius eversmanni badachshanus ; [BRU2, 160]

Plebejus baroghila (Tytler, 1926)
W.Himalayas. See [About maps] Lycaena baroghila Tytler, 1926; J. Bombay nat. Hist. Soc. 31 (3): 583, pl. 1, f. 1, TL: Baroghil, Gilgit
baroghila ; [NEN46, 17]
Plebejus baroghila ; [NEN40]

Plebejus firuskuhi (Forster, 1940)
NW.Afghanistan. See [About maps] Polyommatus firuskuhi Forster, 1940; Dt. ent. Z. Iris 54: 73
Plebejus firuskuhi ; [NEN40]

Plebejus kwaja (Evans, 1932)
Baluchistan. See [About maps] Polyommatus morgiana kwaja Evans, 1932; Indian Butterflies (edn. 2): 227, TL: Baluchistan, W. Khojak, 6500, Pakistan
kwaja ; [NEN46, 40]
Plebejus kwaja ; [NEN40]

The argus species-group
Plebejus; (s. str)

Plebejus aegidion (Gerhard, 1853)
C.Asia (mountains). See [About maps] Lycaena aegidion Gerhard, 1853; Mon. Lyc. : pl. 23, f. 4
Plebejus aegidion ; [NEN40]

fiPlebejus argus (Linnaeus, 1758)
Europe, Temperate Asia, Japan. See [About maps] Kangassinisiipi Silver-studded Blue Allmän blĺvinge Taggblĺvinge Nińa hocecillas Modraszek argus
Papilio argus Linnaeus, 1758; Syst. Nat. (Edn 10) 1 : 483, TL: S.Sweden
sylvius (Poda, 1761)
Lycaeides argus orientaloides Verity, 1931; Dt. Ent. Z. Iris, 45: 48, TL: Hyrcana, N. Persia
Lycaeides argus altaegidon Verity, 1931; Dt. Ent. Z. Iris, 45: 59 (unavail., repl. Lycaena alpina Courvoisier, 1911), TL: Valais, Route du Simplon
Lycaena alpina Courvoisier, 1911; 92 [?]
Lycaena aegon plouharnelensis Oberthür, 1910; Čtud. Lépid. Comp. 4 : 186, pl. 38, f. 252-253, TL: Plouharnel, France
Plebeius argus, Larvae of North-European Lepidoptera [Kimmo Silvonen]
Plebeius argus, Svenska fjärilar [SVF]
Plebeius argus, Norges Sommerfugler [NORLEP]
Plebejus argus, Moths and Butterflies of Europe [MBE]
Plebejus argus, Europaische Schmetterlinge [Mario Maier]
Plebejus argus, Captain's European Butterfly Guide [Simon Coombes]
Plebejus argus ; [BARU, #404]; [EBW]; [BOW: pl. 10, f. 21]; [NEN40]; [MRS, 689]
Plebeius argus ; [BRU2: 158, pl. 73, f. 16-38]
orientaloides ; [NEN46, 50]
altaegidon ; [NEN46, 10]
plouharnelensis ; [NEN46, 52]
[argus-1um.jpg] 500x497 (~50Kb) upperside male Bolt Head, Devon, UK 17/07/96, Photo © Simon Coombes [argus-1vmf.jpg] 500x404 (~62Kb) underside male female Bolt Head, Devon, UK 17/07/96, Photo © Simon Coombes [argus-2vmf.jpg] 600x580 (~48Kb) underside male female FINLAND: Ab: Suomusjärvi, 669:31, 13.7.1997, Photo © Markku Savela
[argus-3vm.jpg] 600x680 (~67Kb) underside male FINLAND: Ab: Suomusjärvi, 669:31, 18.7.1997, Photo © Markku Savela [argus-4vf.jpg] 560x694 (~50Kb) underside female FINLAND: Ka: Virolahti Oravakorpi, 671:53, 2.8.1997, Photo © Markku Savela [argus-5mu.jpg] 900x695 (~67Kb) upperside male Finland, Sipoo 670:39, 5.7.1979, Photo © Pertti Pakkanen
[argus-6fu.jpg] 1007x881 (~156Kb) upperside female Finland: Puumala, Luonteri, Ikosensaaret, 20.7.2006, Photo © Harri Arkkio [argus-6fv.jpg] 1044x783 (~141Kb) underside female Finland: Puumala, Luonteri, Ikosensaaret, 20.7.2006, Photo © Harri Arkkio [argus-6mu.jpg] 868x832 (~126Kb) upperside male Finland: Puumala, Luonteri, Ikosensaaret, 20.7.2006, Photo © Harri Arkkio
[argus-6mv.jpg] 845x881 (~128Kb) underside male Finland: Puumala, Luonteri, Ikosensaaret, 20.7.2006, Photo © Harri Arkkio [argus-7mfv.jpg] 1067x762 (~137Kb) underside male female Russia, Moscow area (36°25'E, 56°00'N), 31.7.2009, Photo © D. Smirnov [argus-8fu.jpg] 736x514 (~95Kb) upperside female Russia, Moscow area, 3.7.2010 (36°25'E, 56°00'N), Photo © D. Smirnov
[argus-8fv.jpg] 650x630 (~106Kb) underside female Russia, Moscow area, 3.7.2010 (36°25'E, 56°00'N), Photo © D. Smirnov

Larva on Vaccinium uliginosum [SPRK], Fabaceae, Trifolium, Chamaecytisus, Genista, Lotus, Ononis, Lathyrus, Vicia [BARU], Cirsium japonicum, C. sieboldii, Artemisia montana, Hedysarum vicioides, Astragalus membranaceus, Pologonum cuspidatum, Loranthus spp. [MRS]


Scandinavia Plebeius argus argus
Plebeius argus argus ; [BRU2, 158]
P. a. aegon (Denis & Schiffermüller, 1775)
Karelia Papilio aegon Denis & Schiffermüller, 1775; Ankündung eines systematischen Werkes von den Schmetterlingen der Wienergegend: 185, n. 15
Plebeius argus aegon ; [BRU2, 158]
P. a. cleomenes (Fruhstorfer, 1910)
Carpathians Lycaena cleomenes Fruhstorfer, 1910; Int. ent. Zs. 4 (12): 62
Plebeius argus cleomenes ; [BRU2, 158]
P. a. wolgensis (Forster, 1936)
S.Europe, lowlands of? (Kazakhstan, Tian-Shan, Tarbagatai, Saur, S.Altai) Lycaena wolgensis Forster, 1936; Mitt. münch. ent. Ges. 26 (2) : 133, TL: Volga valley
Plebeius argus wolgensis ; [BRU2, 158]
P. a. bellus (Herrich-Schäffer, [1844])
Asia Minor, Kurdistan, Levant, Caucasus, Armenia, Talysh Lycaena bella Herrich-Schäffer, [1844]; Syst. Bearb. Schmett. Europ. 1 (7): f. 227-228
Lycaena tscherkessica Forster, 1936; Mitt. münch. ent. Ges. 26 (2) : 131, TL: Caucasia
Lycaena georgica Forster, 1936; Mitt. münch. ent. Ges. 26 (2) : 132, TL: Georgia
orientalis (Verity, 1931)
Plebejus bella ; [NEN40]
Plebeius argus bellus ; [BRU2, 158]
P. a. obensis (Forster, 1936)
Ural, W.Siberia Lycaena obensis Forster, 1936; Mitt. münch. ent. Ges. 26 (2) : 135
Lycaena korshunovi Shtandel, 1960; Ent. Obozr. 39: 695
Plebejus argus obensis ; [BARU, (note)]
Plebeius argus obensis ; [BRU2, 158]
[obensis-1mfv.jpg] 533x400 (~49Kb) underside male female A lawn, Akademgorodok [Academy Town], Novosibirsk, West Siberia, Russia. 22nd June 1989, Photo © Oleg Kosterin [obensis-1mfvx.jpg] 478x694 (~36Kb) underside male female A lawn, Akademgorodok [Academy Town], Novosibirsk, West Siberia, Russia. 22nd June 1989, Photo © Oleg Kosterin

P. a. clarasiaticus (Verity, 1931)
E.Altai, Sayan, Transbbaikalia, W.Amur Lycaena clarasiatica Verity, 1931; Dt. Ent. Z. Iris, 45: 62, TL: E.Sayan
Lycaena katunica Shtandel, 1960; Ent. Obozr. 39: 695
Plebejus argus clarasiatica ; [BARU, (note)]
Plebeius argus clarasiaticus ; [BRU2, 158]
P. a. pamirus (Forster, 1936)
Pamirs-Alai, ?highlands (Tian-Shan) Lycaena pamira Forster, 1936; Mitt. münch. ent. Ges. 26 (2) : 138, TL: Pamir
Plebeius argus pamirus ; [BRU2, 158]
P. a. coreanus Tutt, 1909
E.Amur, Ussuri, Korea Plebejus coreana Tutt, 1909; Nat. Hist. Br. Lepid. 10 : 201, TL: Gensan, Corea
kurentzovi Koçak, 1980
orientalis (Kurenzov, 1970); (preocc. P. argus orientalis Tutt, 1909)
coreana ; [NEN46, 24]
Plebejus argus coreana ; [BARU, (note)]
Plebeius argus coreanus ; [BRU2, 158]
P. a. micrargus (Butler, 1878)
Japan, Sakhalin Lycaena micrargus Butler, 1878; Cist. Ent. 2 (19) : 283 (Lampides micrargus?)
Lycaena aegon japonica Oberthür, 1910; Čtud. Lépid. Comp. 4 : pl. 42, f. 313-314, TL: Japan
Lycaeides argus micrargus ab. chiyodaensis Murayama, 1963; Tyô to Ga 14 (2): 49, f. 46, TL: Yamagata, Japan
micrargus ; [NEN46, 44]
japonica ; [NEN46, 38]
Plebejus micrargus ; [NEN40]
Plebeius argus micrargus ; [BRU2, 158]
P. a. asur Agenjo, 1966
Plebejus argus asur Agenjo, 1966; Rev. Esp. Ent. 41: 174, TL: Villasur, Spain
asur ; [NEN46, 16]
P. a. bejarensis (Chapman, 1902)
Lycaena argus bejarensis Chapman, 1902; Proc. ent. Soc. Lond. 1902 : xxxv, TL: Castilla-Leon, Béjar, Spain
bejarensis ; [NEN46, 18]
Plebeius argus branuelasensis Tutt, 1909; Nat. Hist. Br. Lepid. 10 : 198, TL: Branuelas, Spain
branuelasensis ; [NEN46, 20]
P. a. casaiacus Chapman, 1907
Plebejus argus var. casaiacus Chapman, 1907; Trans. ent. Soc. Lond. 1907 (1) : 151, pl. 5, f. 6-9, TL: Casayo, N.W. Spain
casaiacus ; [NEN46, 22]
P. a. claraobscura (Verity, 1931)
Lycaeides argus claraobscura Verity, 1931; Dt. Ent. Z. Iris, 45: 54, TL: Larche, Basses-Alpes, 1700-2000m, France
claraobscura ; [NEN46, 23]
Plebeius argus hypochionagraecea Tutt, 1909; Nat. Hist. Br. Lepid. 10 : 196, TL: "Parnass"[?]
hypochionagraecea ; [NEN46, 35]
P. a. seoki Shirozu & Sibitani, 1943
Plebejus argus seoki Shirozu & Sibitani, 1943; Trans. Kansai. ent. Soc. 13: 25, TL: Saishuto, Korea
seoki ; [NEN46, 58]
P. a. vigensis Tutt, 1909
Plebejus argus vigensis Tutt, 1909; Nat. Hist. Br. Lepid. 10 : 198, TL: Vigo, NW.Spain
vigensis ; [NEN46, 62]
?P. a. sultana (Forster, 1936)
Asia Minor Lycaena sultana Forster, 1936; Mitt. münch. ent. Ges. 26 (2) : 128

Plebejus corsicus (Bellier, 1862)
Corsica, Sardinia. See [About maps] Lycaena aegon corsica Bellier, 1862; Ann. Soc. ent. Fr. (4) 2 : 615, pl. 14, f. 5, TL: Corse, Montagnes
Plebejus corsicus ; [NEN40]
corsica ; [NEN46, 24]

Plebejus hypochiona (Rambur, 1858)
Iberia. See [About maps] Lycaena hypochiona Rambur, 1858; Cat. syst. Lépid. Andalousie (1): 35
?Plebejus argus hypochiona ;
?Plebejus hipochyonus Rambur, 1862; [NEN40, (spell.?)]

The idas species-group
Lycaeides
Plebulina

Plebejus agnatus (Rühl, 1895)
Alai, W.Tien-Shan, Inner Tien-Shan (Alaisky, E.Fergansky, Sarykolsky Mts.), Kokshaal-Too, Turugart Mts., Kunlun Mts.. See [About maps] Lycaena agnata Rühl, 1895; in Heyne, Palaearkt. Grossschmett. 1 : 299, TL: Issyk-Kul-See
Lycaena christophi var. agnata Staudinger, 1889; Stettin Ent. Ztg 50 (1-3) : 19
?Lycaena christophi var. agnata Staudinger, 1901; in Staudinger & Rebel, Cat. Lepid. palaearct. Faunengeb., 1: 80
Plebejus agnata ; [NEN40]
Plebeius agnatus ; [BRU2: 167, pl. 74, f. 16-18]

Plebejus argyrognomon (Bergstrasser, [1779])
CEU, SEU, Caucasus, S.Siberia (mountains), Mongolia, Amur. See [About maps] Reverdin's Blue Northern Blue Kronärtblĺvinge Modraszek srebroplamek
Papilio argyrognomon Bergstrasser, [1779]; Nomen. Ins., 2: 76, pl. 46, f. 1-2, TL: Germany, Hanau-Munzberg
ismenias (Meigen, 1829)
aegus (Chapman, 1917)
Lycaeides argyrognomon, Lepidoptera collection [SZMN]
Plebeius argyrognomon, Larvae of North-European Lepidoptera [Kimmo Silvonen]
Plebeius argyrognomon, Svenska fjärilar [SVF]
Plebeius argyrognomon, Norges Sommerfugler [NORLEP]
Plebejus argyrognomon, Moths and Butterflies of Europe [MBE]
Plebejus argyrognomon, Europaische Schmetterlinge [Mario Maier]
Plebejus argyrognomon, Captain's European Butterfly Guide [Simon Coombes]
Lycaeides argyrognomon ; [BOW: pl. 9, f. 35]; [MRS, 688]
Plebejus argyrognomon ; [NEN40]; [BARU, #405]; [Otakar Kudrna]
Plebeius argyrognomon ; [BRU2: 164, pl. 72, f. 43-45]

Larva on Coronilla [H&R], Indigofera kirilowii, Fabaceae [MRS], Securigera varia, Medicago sativa, Melilotus officinalis, Trifolium repens, Astragalus glycyphyllos, Lotus corniculatus, Onobrychis viciifolia, Vicia sativa [BRU2, 164]

P. a. euergetes (Stauder, 1914)
P. a. danapriensis (Stempfer-Schmidt, 1932)
W.Europe, S.Europe Plebeius argyrognomon danapriensis ; [BRU2, 164]
P. a. caerulescens (Grum-Grshimailo, 1893)
S.Ural, SW.Siberia, Altai, Prisayanye Lycaena argus caerulescens Grum-Grshimailo, 1893; Horae Soc. ent. Ross. 27 (1-2): 127, TL: Ural; Nishnaya Salda
= ?Plebejus idas (Linnaeus, 1761); [BRU2, 161 (?)]
caerulescens ; [NEN46, 21]
Plebejus argyrognomon caerulescens ; [BARU, (note)]
[caerulescens-1m.jpg] 533x400 (~65Kb) male An subalpine meadow, at 2000 m above sea level, on the eastern spurs of the Katunskiy Mt. Range, the valley of the Argem (Direntai) stream, Central Altai Mts., West Siberia, Russia. 13th July, 1988, Photo © Oleg Kosterin

P. a. buchara (Forster, 1936)
Lycaena buchara Forster, 1936; Mitt. münch. ent. Ges. 26 (2) : 75, pl. 10-11, TL: Buchara, Kurgan-Tjube, Uzbekistan
buchara ; [NEN46, 21]
P. a. letitia (Hemming, 1934)
Lycaeides ismenias letitia Hemming, 1934; Stylops 3 : 98 (repl. ligurica Oberthür)
Lycaena argus ligurica Oberthür, 1910; Čtud. Lépid. Comp. 4 : 201, pl. 41, f. 293-294, TL: Lago di Como, Italy
P. a. praeterinsularis (Verity, 1921)
Plebeius insularis praeterinsularis Verity, 1921; Ent. Rec. J. Var. 33 : 175, pl. 3, f. 5-6
praeterinsularis ; [NEN46, 53]

Include into Plebejus pseudaegon?


Plebejus pseudaegon (Butler, [1882])
Ussuri, NE.China, N.Korea, Japan. See [About maps] Lycaena pseudaegon Butler, [1882]; in Butler & Fenton, Proc. Zool. Soc. Lond. 1881 (4) : 851, TL: Iburi, Hokkaido
pseudaegon ; [NEN46, 53]
Plebejus argus pseudoaegon[sic, recte pseudaegon] ; [BARU, (note)]
Plebeius pseudoaegon[sic, recte pseudaegon] ; [BRU2: 164, pl. 72, f. 46-48]
P. p. ussuricus (Forster, 1936)
Ussuri Lycaena ussurica Forster, 1936; Mitt. münch. ent. Ges. 26 (2) : 79, TL: Ussuri
Plebejus argyrognomon ussurica ; [BARU, (note)]
Plebeius pseudoaegon ussuricus ; [BRU2, 164]

Larva on Vicia amoena [BRU2, 164]

P. p. insularis (Leech, 1893)
Japan Lycaena argus var. insularis Leech, 1893; Butts China Japan Corea (2): 302, pl. 31, f. 5, 8, TL: Hakodate
insularis ; [NEN46, 37]
Plebeius pseudoaegon insularis ; [BRU2, 164]
P. p. sinicus (Forster, 1936)
Sichuan Lycaena sinica Forster, 1936; Mitt. münch. ent. Ges. 26 (2) : 77, pl. 10-11, f. 15-16, TL: Szetschwan, China
?argyrognomon sinica ;
sinica ; [NEN46, 59]
Plebeius pseudoaegon sinicus ; [BRU2, 164]

Plebejus maracandicus (Erschoff, 1874)
-. See [About maps] Lycaena argus var. maracandica Erschoff, 1874; in Fedschenko, Travels in Turkestan. 2 (5) : 10, TL: [Vicinity of Samarkand, Uzbekistan]
Lycaena dschagatai Grum-Grshimailo, 1887; in Romanoff, Mém. Lép. 3 : 400, TL: Shir-Abad, Buchara
= ; [BRU2, 165]
Polyommatus orbitulus dschagatai ; Bridges, 1994, Cat. Family and Genus group Riodinidae & Lycaenidae:
Plebejus argyrognomon dschagatai ;
Plebejus argus maracandica ;
Plebeius maracandicus ; [BRU2: 165, pl. 73, f. 1-14]
SW.Russia, Turan Plebeius maracandicus maracandicus
Plebeius maracandicus maracandicus ; [BRU2, 165]
P. m. planorum (Alphéraky, 1881)
Ili River valley Lycaena argus var. planorum Alphéraky, 1881; Horae Soc. ent. Ross. 16 (3-4): 379
Lycaena argus var. calmuca Grum-Grshimailo, 1891; Horae Soc. ent. Ross. 25 (3-4) : 450, TL: Tien Shan, Krygyzstan
?calmucca; (spell.?)
?calmuca ; [NEN46, 21]
Plebejus planorum ; [NEN40]
Plebeius maracandicus planorum ; [BRU2, 165]

Larva on Astragalus sogotensis [BRU2, 165]

P. m. caspicus (Forster, 1936)
S.Urals Lycaena caspica Forster, 1936; Mitt. münch. ent. Ges. 26 (2) : 74
Plebeius maracandicus caspicus ; [BRU2, 165]
P. m. chalcha Korshunov, 1982
Tuva, S.Transbaikalia, Mongolia Plebejus argyrognomon chalcha ; [BARU, (note)]
Plebeius maracandicus chalcha ; [BRU2, 165]
P. m. transbaicalensis (Kurenzov, 1970)
N.Transbaikalia Plebejus argyrognomon transbaicalensis ; [BARU, (note)]
Plebeius maracandicus transbaicalensis ; [BRU2, 165]
P. m. jacuticus (Kurenzov, 1970)
Yakutia, Magadan Plebejus argyrognomon jacutica ; [BARU, (note)]
Plebejus maracandicus jacuticus ; [BRU2, 165]

Plebejus lepidus Zhdanko, 2000
E.Kazakhstan. See [About maps] Plebeius lepidus Zhdanko, 2000; [BRU2: 165, pl. 74, f. 31-33], TL: E.Kazakhstan, 15km E of Aksuat

Plebejus uiguricus Zhdanko, 2000
S.Altai, Saur, Tarbagatai Mts., Dzhungarsky Alatau?. See [About maps] Plebeius uiguricus Zhdanko, 2000; Helios, 1: 101, TL: Bukombai, 20km SE Karatogai, Kurchum Mts., S. Altai
Plebeius uiguricus ; [BRU2: 166, pl. 83, f. 34-36]

Plebejus mongolicus Rühl, [1893]
Transbaikalia, Amur, Mongolia. See [About maps] mongolicus Rühl, [1893]; Palaearktischen Grosschmetterlinge: 235, TL: N.Mongolia, Darkhan [neotype]
?Lycaena argus var. mongolica Grum-Grshimailo; (nom. nud.?)
Plebeius mongolicus ; [BRU2: 166, pl. 72, f. 40-42, pl. 83, f. 26-27 [neotype]]

Plebejus baldur (Hemming, 1934)
Kurdistan, Levant Lycaena argyrognomon baldur Hemming, 1934; Stylops 3 : 193 (repl. Lycaena argyrognomon balcanica Züllich, 1929)
Lycaena argyrognomon f. balcanica Züllich, 1929; Z. öst. ent. Ver. 14 (5): 52 (preocc. Lycaena semiargus balcanica Tutt, 1909), TL: Monastery Rilski, Bulgaria
Plebejus baldur ; [NEN40]
balcanica ; [NEN46, 17]; Abadjiev, 2002, Neue Ent. Nachr. 23: 22

Plebejus bellieri (Oberthür, 1910)
Corsica. See [About maps] Lycaena argus bellieri Oberthür, 1910; Čtud. Lépid. Comp. 4 : 191, pl. 40, f. 280-281, TL: Bastelica
Plebejus bellieri ; [NEN40]
bellieri ; [NEN46, 18]
Plebejus idas bellieri ;
Plebejus bellieri ; [Otakar Kudrna]

Plebejus christophi (Staudinger, 1874)
N.Iran, Caucasus, Kurdistan, Turkestan, Chitral, N.Afghanistan. See [About maps] Lycaena christophi Staudinger, 1874; Stettin ent. Ztg 35 (1-3) : 87, TL: Schachrud [NE.Iran]
argiva (Staudinger, 1886); (Middle Asia)
Lycaena bracteata Butler, 1880; Proc. zool. Soc. Lond. 1880 (3) : 407, pl. 39, f. 4, TL: Candahar, Afghanistan
christophi ; [NEN46, 23]
Lycaena christophi ; [BOW: pl. 206, f. 11]
bracteata ; [NEN46, 20]
Plebejus christophi ; [NEN40]
Plebeius christophi ; [BRU2: 167, pl. 74, f. 13-15, 25-27]

Larva on Alhagi spp. [BRU2, 167]

Turan, S.Ghissar Plebeius christophi christophi
Plebeius christophi christophi ; [BRU2, 167]
P. c. transcaucasicus (Rebel, 1901)
S.Armenia, Turkey Lycaena argyrognomon var. transcaucasica Rebel, 1901; Ann. Hofmus. Wien. 16 : 166
Plebeius christophi transcaucasicus ; [BRU2, 167]
?P. c. nanshanica (Forster, 1936)
Kansu Lycaena nanshanica Forster, 1936; Mitt. münch. ent. Ges. 26 (2) : 84

Plebejus roxane (Grum-Grshimailo, 1887)
Tajikistan, N.Afghanistan. See [About maps] Lycaena roxane Grum-Grshimailo, 1887; in Romanoff, Mém. Lép. 3 : 400, TL: Shir-Abad, Surchan-Darya
Lycaena roxane var. bactriana Grum-Grshimailo, 1890; in Romanoff, Mém. Lép. 4 : 374, TL: Surchan-Darja, Buchara
= ; [BRU2, 168]
submontana (Shchetkin, 1960)
= ; [BRU2, 168]

submontanus ; [BRU2, 168]
Lycaeides christophi submontana ;
bactriana ; [NEN46, 16]
bactrianus ; [BRU2, 168]
Plebejus christophhi roxane ;
roxane ; [NEN46, 55]
Plebeius roxane ; [BRU2: 168, pl. 74, f. 22-24]

Larva on Hippophae rhamnoideus, Alhagi spp. [BRU2, 168]


Plebejus emigdionis (Grinnell, 1905)
California. See [About maps] San Emigdio Blue
Lycaena emigdionis Grinnell, 1905; Ent. News 16 (4) : 115
Lycaena melimona Wright, 1906; Butts. West Coast (2nd edn.): (1905?)
Plebejus emigdionis, Butterflies of America [BOA]
Plebejus emigdionis, Butterflies and Moths of North America [BAMONA]
Plebulina emigdionis ; [NACL, #4377]
Plebejus emigdionis ; [NEN40]; [Opler]

Larva on Atriplex canescens, Hosachia purschiana [NAB]


fiPlebejus idas (Linnaeus, 1761)
Europe, Siberia, S.Siberia (mountains), Yakutia, Alaska. See [About maps] Ketosinisiipi Idas Blue Föränderlig blĺvinge Backblĺvinge Nińa Esmaltada Modraszek idas
Papilio idas Linnaeus, 1761; Fauna Suecica (Edn 2): 284, TL: Sweden
?Papilio idas Linnaeus, 1758; Syst. Nat. (Edn 10) 1 : 488
Papilio argus Denis & Schiffermüller, 1775
Papilio leodorus Esper, 1782; Die Schmett. Th. I, Bd. 2 (5): 137, pl. 80, f. 1-2
?acdestis (Grum-Grhsimailo, 1890)
= ?; [BRU2, 161 (?)]
Lycaena argus armoricana Oberthür, 1910; Čtud. Lépid. Comp. 4 : 189, pl. 40, f. 275-276, TL: Monterfil, France
Lycaeides idas, E.H. Strickland Ent. Mus, U of [ALBERTA]
Plebejus idas, Butterflies and Moths of North America [BAMONA]
Plebeius idas, Larvae of North-European Lepidoptera [Kimmo Silvonen]
Plebeius idas, Svenska fjärilar [SVF]
Plebeius idas, Norges Sommerfugler [NORLEP]
Plebejus idas, Moths and Butterflies of Europe [MBE]
Plebejus idas, Europaische Schmetterlinge [Mario Maier]
Plebejus idas, Captain's European Butterfly Guide [Simon Coombes]
armoricana ; [NEN46, 14]
Plebejus idas ; [BARU, #406]; [NEN40]
Plebeius (idas) idas ; [BRU2: 161, pl. 73, f. 46-57]
Lycaeides idas ; [BOW: pl. 9, f. 35 (text only)]
Plebejus idas ; [Opler]; [Otakar Kudrna]
Misidentified/misapplied:
Lycaeides argyrognomon; [NACL, #4374]
[idas.jpg] 321x296 (~11Kb) FINLAND: Li: Utsjoki, Karigasniemi, 770:45, 17.7.1995, Photo © Valentina Anderson

Larva on Leguminosae, Lotus corniculatus, Trifolium pratense, Calluna vulgaris, Vaccinium uliginosum, Empetrum nigrum [SPRK], Hippocrepis emeroides, Cercis siliquastrum, Melilotus albus, Chrysaspis campestris, Genista tinctoria, Astragalus alpinus, Cicer flexuosum [BRU2, 162]


P. i. acreon (Fabricius, 1787)
CEU, SEU, W.Kazakhstan, N.Kasakhstan Plebeius idas acreon ; [BRU2, 161]
P. i. altarmenus (Forster, 1936)
Armenia Lycaena idas altarmena Forster, 1936; Mitt. münch. ent. Ges. 26 (2) : 103, TL: Armenia
Plebeius idas altarmenus ; [BRU2, 161]
P. i. lotis Lintner, 1879˛
lotis Lintner, 1879; 30 Rept. N.Y. Mus. Nat. Hist.: 169
Rusticus lotis ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 44
Lycaeides argyrognomon lotis ; [NACL, #4374c]
P. i. alaskensis (Chermock, 1945)
idas alaskensis (Chermock, 1945) (Plebeius); Can. Ent. 76 (11): 213
Lycaeides argyrognomon alaskensis ; [NACL, #4374d]
Misidentified/misapplied:
kodiak; auth.
P. i. scudderi (Edwards, 1861)
British Columbia Lycaena scudderi Edwards, 1861; Proc. Acad. nat. Sci. Philad. 13: 164, TL: Saskatchewan R. Manitoba
Plebeius subarcticus Chermock, 1945; Can. Ent. 76 (11): 213
Rusticus scudderi ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 44
Lycaeides argyrognomon scudderi ; [NACL,#4374e]
Lycaeides idas scudderi ; [BOC, 242]
P. i. aster (Edwards, 1882)
Newfoundland Lycaena aster Edwards, 1882; Can. Ent. 14 (10) : 194, TL: Newfoundland
Rusticus aster ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 44
Lycaeides argyrognomon aster ; [NACL, #4374f]
Lycaeides aster ; [BOW: pl. 19, f. 38]
P. i. empetri Freeman, 1938˛
Lycaeides argyrognomon empetri ; [NACL, #4374g]
N.Idaho, NE.Washington, SE.British Columbia Plebejus atrapraetextus Field, 1939; J. Kansas Ent. Soc. 12: 135, TL: Priest River, Idaho
Plebeius ferniensis Chermock, 1945; Can. Ent. 76 (11): 215, TL: Fernie, British Columbia
= Lycaeides idas atrapratextus Field, 1939; [BOC, 242]
Plebeius sweadneri Chermock, 1945; Can. Ent. 76 (11): 214
Lycaeides idas ferniensis ;
Lycaeides argyrognomon ferniensis ; [NACL, #4374h]
Lycaeides argyrognomon atrapraetextus ; [NACL, #4374i]
Lycaeides idas atrapraetextus ; [BOC, 142]
P. i. sublivens (Nabokov, 1949)
Lycaeides sublivens Nabokov, 1949; Bull. Mus. Comp. Zool. 101: 513
Lycaeides argyrognomon sublivens ; [NACL, #4374j]
sublivent ; F. M. Brown, Eff & Roger, 1955, (missp.)
P. i. longinus (Nabokov, 1949)
Lycaeides longinus Nabokov, 1949
Plebejus idas longinus, Butterflies of America [BOA]
Lycaeides argyrognomon longinus ; [NACL, #4374k]
P. i. nabokovi (Masters, 1972)
Lycaeides nabokovi Masters, 1972
Lycaeides argyrognomon nabokovi ; [NACL, #4374l]
P. i. argulus (Frey, 1882)
Lycaena argus var. argulus Frey, 1882; Mitt. schweiz. ent. Ges. 6 (7) : 350, TL: Switzerland
argulus ; [NEN46, 14]
P. i. bavarica (Forster, 1936)
Lycaena idas bavarica Forster, 1936; Mitt. münch. ent. Ges. 26 (2) : 90, pl. 12-13, f. 9-10, TL: Bavaria, Germany
bavarica ; [NEN46, 18]
P. i. sareptensis Chapman, 1917
Plebeius sareptensis Chapman, 1917; Étud. Lép. comp. 14 : 52, TL: Sarepta
= ?Plebeius maracandicus caspicus (Forster, 1936); [BRU2, 165 (?)]
sareptensis ; [NEN46, 57]
Plebejus idas sareptensis ; [NEN46, 57]
P. i. tshimganus (Forster, 1936)
W.Tian-Shan Lycaena idas tshimgana Forster, 1936; Mitt. münch. ent. Ges. 26 (2) : 104
Plebeius idas tshimganus ; [BRU2, 161]

Plebejus ganssuensis (Grum-Grshimailo, 1891)
Lycaena argus var. ganssuensis Grum-Grshimailo, 1891; Horae Soc. ent. Ross. 25 (3-4) : 450, TL: Amda, Kuku-Noor, China
Plebejus obscurolunulata Huang, 1999; Lambillionea XCIX (3): 327, TL: S of Tsinghai Lake, Qinghai
= ; Huang & Xue, 2004, Neue Ent. Nachr. 57: 171, pl. 14, f. 6
Plebejus idas ganssuensis ;
ganssuensis ; [NEN46, 33]
Plebejus ganssuensis ; Huang & Xue, 2004, Neue Ent. Nachr. 57: 171, pl. 14, f. 5-6

Plebejus nushibi Zhdanko, 2000
SE.Kazakhstan. See [About maps] Plebeius nushibi Zhdanko, 2000; [BRU2: 163, pl. 73, f. 43-45], TL: SE. Kazakhstan, Zailiisky Alatau Mts., Kaskelen Gorgo, 1500m

Larva on Onobrychis sp.? [BRU2, 163]


Plebejus calliopis (Boisduval, 1832)
European highlands (Alps), boreal Europe (N.Europe?), Transcaucasia. See [About maps] Argus calliopis Boisduval, 1832; Icon. hist. Lépid. Europe 1 (7-8): 58, pl. 15, f. 4-5, TL: Grenoble, France
Plebejus idas calliopis ;
calliopis ; [NEN46, 21]
Plebeius (idas) calliopis ; [BRU2: 162, pl. 73, f. 58-60, pl. 74, f. 1-12]

Larva on Hedysarum armenum, (Transcaucasia) Cicer flexuosum, (Ghissar) [BRU2, 162]

P. c. armoricanellus (Beuret, 1934)
N.Europe? Plebeius calliopis armoricanellus ; [BRU2, 162]
P. c. polaris (Nordström, 1928)
Kamchatka kamtchatica Kurenzov, 1970
Plebejus tancrei kamtchatica ; [BARU, (note)]
Plebeius calliopis polaris ; [BRU2, 162]

[kamtchatica-1mfu.jpg] 400x300 (~61Kb) upperside male female A detritous montane tundra on the SW side of a ring mountain range circling the Ksudach Caldera, Elizovo District, South Kamchatka, Russia. 21 August 1991 (playing male and female), Photo © Oleg Kosterin

P. c. aborigeni Zhdanko, 1999
Far East Plebeius calliops aborigeni ; [BRU2, 162]
P. c. sailjugemiaeginus Zhdanko & Samodurov, 1999
Altai Plebeius calliopis sailjugemiaeginus ; [BRU2, 162]
P. c. aegina (Grum-Grshimailo, 1891)
N.Tian-Shan, Inner Tian-Shan Lycaena aegina Grum-Grshimailo, 1891; Horae Soc. ent. Ross. 25 (3-4) : 451, TL: Tien Chan, Boro
Lycaena argus f. naruena Courvoisier, 1913; Ent. Mitt. 2 (10) : 289
= ; [BRU2, 162]
aegina ; [NEN46, 8]
Plebejus naruena ; [NEN40]
Plebeius idas naruenus ; [BRU2, 161]
Plebeius idas aegina ; [BRU2, 162]
P. c. gissari Zhdanko, 1999
Ghissar Plebeius calliopis gissari ; [BRU2, 162]

Plebejus anna (Edwards, 1861)
-. See [About maps] Northern Blue Anna Blue
Lycaena anna Edwards, 1861; Proc. Acad. nat. Sci. Philad. 13: 163, TL: Donner Pass, Nevada Co., California
cajona (Reakirt, 1866); Proc. Ent. Soc. Philad. 6 (1) : 147
Lycaena argyrotoxus Behr, 1867; Proc. Calif. Acad. Sci. 3 (4) : 281
Lycaena philemon Boisduval, 1869; Ann. Soc. ent. Belg. 12 : 47
Plebejus anna anna, Butterflies of America [BOA]
Plebejus anna, Butterflies and Moths of North America [BAMONA]
Rusticus anna ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 44
Lycaeides argyrognomon anna ; [NACL, #4374a]
Lycaeides anna ; [BOC, 242]
Plebejus anna ; [NEN40]; [Opler]

Larva on Lathyrus torreyi, Vicia exigua, Lupinus latifolius [BOC, 243]

California, Oregon Lycaeides anna anna
Lycaeides anna anna ; [BOC, 242]
P. a. ricei Cross, 1937
Oregon, Washington, SW.British Columbia Plebejus ricei Cross, 1937; Pan-Pacific Ent. 13: 88, TL: Big Cultus lake, Deschutes Co., Oregon
Plebeius fretchini Chermock, 1945; Can. Ent. 76 (11): 214, TL: Paradise Valley, Mt. Rainier, Washington
Lycaeides argyrognomon ricei ; [NACL, #4374b]
Lycaeides anna ricei ; [BOC, 243]
[ricei-1.jpg] 600x691 (~43Kb) USA: FR9705, Table Mtn, Kittitas Co., WA, 30.7.1999, Photo © Markku Savela
Lycaeides idas ricei det. Norbert Kondla
[ricei-1v.jpg] 500x462 (~25Kb) underside USA: FR9705, Table Mtn, Kittitas Co., WA, 30.7.1999, Photo © Markku Savela
Lycaeides idas ricei det. Norbert Kondla
[ricei-2u.jpg] 600x600 (~53Kb) upperside USA: FR9708, Table Mtn, Kittitas Co., WA, 19.7.1999, Photo © Markku Savela
Lycaeides idas ricei det. Norbert Kondla
[ricei-2v.jpg] 500x585 (~42Kb) underside USA: FR9708, Table Mtn, Kittitas Co., WA, 19.7.1999, Photo © Markku Savela
Lycaeides idas ricei det. Norbert Kondla
[ricei-3fu.jpg] 600x576 (~40Kb) upperside female USA: Near Lion Rock, Table Mtn, Kittitas Co., WA, 30.7.1999, Photo © Markku Savela
Lycaeides idas ricei det. Norbert Kondla
[ricei-4vm.jpg] 700x955 (~87Kb) underside male USA: FR 9712 nr. Liberty, Lion Gulch, (47°15'47N 120°40'15W ±0.5km), Kittitas Co., WA, 14.8.2001, Photo © Markku Savela

P. a. vancouverense (Guppy & Shepard, 2001)
Vancouver I. Lycaeides anna vancouverense Guppy & Shepard, 2001; , TL: Cream Lake, Strathcona Park, Vancouver I.
Lycaeides anna vancouverense ; [BOC, 243]

Plebejus iburiensis (Butler, [1882])
Japan. See [About maps] Lycaena iburiensis Butler, [1882]; in Butler & Fenton, Proc. Zool. Soc. Lond. 1881 (4) : 852, TL: Hokkaido
iburiensis Butler, [1882]; [NEN46, 35]
Plebejus iburiensis ; [NEN40]

Plebejus melissa (Edwards, 1873)
WNA - C.Mexico. See [About maps] Orange-bordered Blue
Lycaena melissa Edwards, 1873; Trans. amer. ent. Soc. 4 : 346, TL: La Plata Peak, lake Co., Colorado
Lycaeides melissa, E.H. Strickland Ent. Mus, U of [ALBERTA]
Plebejus melissa melissa, Butterflies of America [BOA]
Plebejus melissa, Butterflies and Moths of North America [BAMONA]
Rusticus melissa ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 44
Lycaeides melissa ; [NACL, #4375]; [BOW: pl. 19, f. 35-36]; [BOC, 244]
Plebejus melissa ; [NEN40]; [Opler]; [NL4A, #1179]
[melissa-1mvU.jpg] identification uncertain 600x457 (~46Kb) underside male USA: Robinson Canyon Rd, (47°01'00N 120°43'40W), Kittitas Co., WA, 26.7.1999, Photo © Markku Savela [melissa-2f.jpg] 500x588 (~35Kb) female USA: Dry Falls / Sun Lakes State Park, Grant Co., WA, 9.7.2000, Photo © Markku Savela

Larva on Lupinus, Medicago sativa, Oxytropis, Glycyrrhiza [NAB]

Larva on Astragalus inflexus, A. lentiginosus, Swainsona salsula, Medicago sativa, Glycyrrhiza lepidota [BOC, 244]


W.Minnesota, Colorado Lycaeides melissa pseudosamuelis Nabokov, 1949; , TL: Colorado
= ; [NL4A, #1179a]
Lycaeides melissa melissa ;
Lycaeides melissa pseudosamuelis ; [NACL, #4375a]
Plebejus melissa melissa ; [NL4A, #1179a]

Larva on Astragalus crassicarpus, A. adsurgens, A. missouriensis, A. flexuosus, A. lotiflorus, Oxytropis lambertii, Medicago sativa, Coronilla varia Dana, 2005, J. Lep. Soc. 59 (3): 175

P. m. fridayi Chermock, 1945
S.Oregon Plebeius fridayi Chermock, 1945; Can. Ent. 76 (11): 215, TL: Mammoth, California
Lycaeides melissa fridayi ; [BOC, 244]
P. m. paradoxa Chermock, 1945˛
melissa paradoxa Chermock, 1945˛
inyoensis Gunder, 1927˛; (ab., unavail.)
Plebejus melissa paradoxa, Listing of Mexican Butterflies [ILMB]
Plebejus melissa paradoxa, Butterflies of America [BOA]
Lycaeides melissa paradoxa ; [NACL, #4375b]
P. m. annetta (Edwards, 1882)
Lycaena annetta Edwards, 1882; Papilio 2 (3) : 48
Rusticus annetta ?Mead, 1882
Rusticus annetta ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 44
Lycaeides melissa annetta ; [NACL, #4375c]
P. m. samuelis (Nabokov, 1944)
Lycaeides melissa samuelis Nabokov, 1944
Plebejus melissa samuelis, Butterflies of America [BOA]
Lycaeides melissa samuelis ; [NACL, #4375d]
Misidentified/misapplied:
scudderii; auth.
P. m. mexicana (Clench, 1965)
Mexico Lycaeides melissa mexicana Clench, 1965; Ent. News 76: (157-162) , TL: Upper Rio Piedras Verdes, W. Chihuahua, Mexico, 7100-7300 ft
Plebejus melissa mexicana ; [NL4A, #1179b]

Plebejus nevadensis (Oberthür, 1910)
Iberia. See [About maps] Lycaena argus nevadensis Oberthür, 1910; Čtud. Lépid. Comp. 4 : 191, pl. 39, f. 259-260, TL: Alfacar, Sierra Nevada, Spain
Lycaena argus nevadensis Oberthür, 1896; Étud. d'Ent. 20 : pl. 4, f. 54 (nom. nud.)
nevadensis ; [NEN46, 46]
Plebejus nevadensis ; [NEN40]

Plebejus samudra (Moore, [1875])
-. See [About maps] Polyommatus samudra Moore, [1875]; Proc. zool. Soc. Lond. 1874 (4) : 574, pl. 67, f. 2, TL: Skardu, India
tomyris (Grum-Grshimailo, 1890)
micropunctulatus Shchetkin, 1975
samudra ; [NEN46, 56]
Lycaeides christophi samudra ; [BOW: pl. 206, f. 11 (text)]
Plebejus samudra ; [NEN40]
Plebeius samudra ; [BRU2: 168, pl. 74, f. 19-21]

Larva on Hippophae rhamnoideus [BRU2, 168]

NW.India Plebeius samudra samudra
Plebeius samudra samudra ; [BRU2, 168]
P. s. rognedus (Grum-Grshimailo, 1890)
Kashgar, Darvaz, Pamirs Lycaena christophi rogneda Grum-Grshimailo, 1890; in Romanoff, Mém. Lép. 4 : 376, TL: Kashgar, Chingoob-Darya, China
rogneda ; [NEN46, 55]
Plebeius samudra rognedus ; [BRU2, 168]
?P. s. lesliei (Tytler, 1926)
NW.Hindu Kush Lycaena lesliei Tytler, 1926; J. Bombay nat. Hist. Soc. 31 (3): 584, pl. 1, f. 2, TL: Chitral
= ?Plebejus samudra (Moore, [1875]); [BRU2, 168 (?)]
Plebejus christophi lesliei ;
lesliei ; [NEN46, 41]
leslei ; [BRU2, 168 (missp.)]

Plebejus subsolanus (Eversmann, 1851)
Transbaikalia, Amur, Ussuri, N.Mongolia, Korea, Japan. See [About maps] Lycaena subsolanus Eversmann, 1851; Bull. Soc. imp. Nat. Moscou 24 (2) : 620, TL: Irkutsk
?imanishii (Takeuchi, 1936); , TL: Sakhalin
Plebejus subsolanus ; [NEN40]; [BARU, #407]; [BHR]
Plebeius subsolanus ; [BRU2: 160, pl. 72, f. 29-34]

Larva on Vicia, Astragalus, Hedysarum, (and others) [BARU]

Transbaikalia, Amur, Ussuri montanus Yagi, 1915
Lycaena putealis Matsumura, 1927; Insecta Matsumurana 1 (4) : 167, pl. 5, f. 8, TL: Corea, Mt. Hakuto
Plebejus subsolanus subsolanus ; [BARU, (note)]
Plebeius subsolanus subsolanus ; [BRU2, 160]
P. s. ida (Grum-Grshimailo, 1891)
SW.Transbaikalia Lycaena ida Grum-Grshimailo, 1891; Horae Soc. ent. Ross. 25 (3-4) : 451, TL: Amdo, Kuku-Noor, China
= ?Plebejus subsolanus sifanica ;
ida ; [NEN46, 36]
Plebejus ida ; [NEN40]
Plebejus cleobis ida ;
Lycaeides cleobis qiliashanus (Murayama); [MRS, 688]
P. s. sifanica (Grum-Grshimailo, 1891)
S.Mongolia (mountains), E.Himalayas (Tibet) Lycaena argus var. sifanica Grum-Grshimailo, 1891; Horae Soc. ent. Ross. 25 (3-4) : 450, TL: Kansu
Plebejus subsolanus sifanica ; [NEN46, 58]
sifanica ; [NEN46, 59]
P. s. tagakusiensis (Murayama, 1964)
Lycaeides subsolanus tagakusiensis Murayama, 1964; Z. wien. ent. Ges. 49: 36, pl. 4, f. 6-9, TL: Tagakusi, Japan
tagakusiensis ; [NEN46, 62]
togakusiensis[sic?] ;
P. s. myokoensis (Murayama, 1969)
Lycaeides subsolanus myokoensis Murayama, 1969; Tyô to Ga 20 (3-4): 69, f. 7-10, 12-14

Plebejus cleobis (Bremer, 1861)
S.Siberia, Altai, Transbaikalia - Far East, Amur, Ussuri, Japan. See [About maps] Lycaena cleobis Bremer, 1861; Bull. Acad. Imp. Sci. St. Petersb. 3 : 472, TL: Nord. Seite Baikal
Lycaena aegonides Bremer, 1864; Mém. Acad. Sci. St. Pétersb. (7) 8 (1) : 28, n. 128, pl. 3, f. 8 (unnec. repl. Lycaena cleobis Bremer, 1861) [is ref. correct?]
Lycaeides cleobis ; [MRS, 688]; [BOW: pl. 206, f. 12, 13]
Plebeius cleobis ; [BRU2: 161, pl. 72, f. 19-21, 26-28, 35-39]
P. c. tancrei (Graeser, 1888)
Amur, Ussuri ?ternejanus Kurenzov, 1970
= ?; [BRU2, 161 (?)]
Plebejus tancrei (Graeser, 1888); [NEN40]
?Plebejus subsolanus ternejana ; [BARU, (note)]
Plebejus tancrei ; [BARU, #408]
Plebeius cleobis tancrei ; [BRU2, 161]

Larva on Empetrum nigrum, Ledum, Vaccinium, Astragalus [BARU]

P. c. kenteanus (Staudinger, 1892)
W.Sayan, Tuva, N.Mongolia Lycaena subsolanus kenteana Staudinger, 1892; Dt. Ent. Z. Iris 5 (2) : 316
Plebeius cleobis kenteanus ; [BRU2, 161]
P. c. ongodai Tutt, 1909
Altai Plebejus argus ongodai Tutt, 1909; Nat. Hist. Br. Lepid. 10 : 199, TL: Ongodai, Altai Mts
ongodai ; [NEN46, 49]
Plebejus subsolanus ongodai ; [BARU, (note)]
Plebeius cleobis ongodai ; [BRU2, 161]
[ongodai-1mv.jpg] 300x400 (~27Kb) underside male A meadow, with Medicago dominating, on the Tes-Khem River right bank in front of the Shivilig-Khem valley, the Ubsu-Nur intermontane hollow, Tes-Khem District, Tuva Republic, Siberia, Russia. 9th July 1990, Photo © Oleg Kosterin [ongodai-2fu.jpg] 513x518 (~52Kb) upperside female A dry Kobresia tundra patch among dwarf birch tundras on a ledge between the Chikty Rivulet and a next upstream right tributary of the Dzhazator River upper flow, 2400 m above sea level, the southern principal slope of the Yuzhno-Chuiskii mountain range, Kosh-Agach District, Altai Republic, West Siberia, Russia. 15th July 1998 (on Veronica porphyriana), Photo © Oleg Kosterin [ongodai-2mv.jpg] 483x600 (~58Kb) underside male A dry Kobresia tundra patch among dwarf birch tundras on a ledge between the Chikty Rivulet and a next upstream right tributary of the Dzhazator River upper flow, 2400 m above sea level, the southern principal slope of the Yuzhno-Chuiskii mountain range, Kosh-Agach District, Altai Republic, West Siberia, Russia. 15th July 1998, Photo © Oleg Kosterin

?P. c. verchojanicus Kurenzov, 1970
NE.Sibera, Ochot coast Plebejus argus caerulea Gorbunov, 1995; [BARU], TL: C.Yakutia
Plebejus tancrei verchojanicus ; [BARU, (note)]
Plebeius cleobis verchojanicus ; [BRU2, 161 (?)]
[verchojanicus-1mu.jpg] 400x300 (~37Kb) upperside male A flowery slope of the coastal terrace at the Burgauli River mouth on the southern coast of the Koni Peninsula, Ola District, S Magadan Province, Russia. 17th July, 1989 (on a flower of Geranium erianthum DC.), Photo © Oleg Kosterin [verchojanicus-1mv.jpg] 400x533 (~44Kb) underside male A flowery slope of the coastal terrace at the Burgauli River mouth on the southern coast of the Koni Peninsula, Ola District, S Magadan Province, Russia. 17th July, 1989 (on its probable foodlpant Oxytropis czukotica Jurtz.), Photo © Oleg Kosterin

The lucifera species-group

Plebejus lucifera (Staudinger, 1867)
S.Siberia (mountains), NE.Kazakhstan, Mongolia, W.China. See [About maps] Lycaena lucifera Staudinger, 1867; Stettin Ent. Ztg 28 : 100, TL: ;Ust'-Kamenogorsk, W. Altai
Polyommatus lucifera ;
Plebejus lucifera ; [BOW: pl. 206, f. 23 (text)]; [NEN40]
Plebejus lucifer[sic, recte lucifera] ; [BARU, #403]
Patricius lucifer[sic, recte lucifera] ; [BRU2: 158, pl. 72, f. 13-18]
Altai, Sayan Patricius lucifer lucifer[sic, recte lucifera] ; [BRU2, 158]
P. l. selengensis Forster, 1940
Transbaikalia Lycaena biton Bremer, 1861; Bull. Acad. Imp. Sci. St. Petersb. 3 : 472 (?preocc. Papilio biton Sulzer, 1776)
Plebejus lucifera selengensis Forster, 1940
Vacciniina lornex Higgins, 1981
otton Korshunov, 1996
Plebejus lucifer selengensis ; [BARU, (note)]
Patricius lucifer selengensis ; [BRU2, 158]

Plebejus thermis (Grum-Grshimailo, 1891)
C.Asia (mountains). See [About maps] Plebejus thermis ; [NEN40]

The pylaon species-group
Plebejides
Maurus

Plebejus allardi (Oberthür, 1874)
SE.Morocco, AlgeriaTunisia, W.Libya. See [About maps] Lycaena allardii Oberthür, 1874; Petites Nouv. Ent. 1: 412-413, TL: Sebdou (Algeria)
Lycaena allardi ab. stigmatica Stetter-Stättermayer, 1937; Ent. Rdsch. 54 (42): 531, TL: Sebdou, Algeria
Lycaena allardi ab. lunulata Stetter-Stättermayer, 1937; Ent. Rdsch. 54 (42): 531, TL: Sebdou, Algeria
Lycaena allardi ab. punctifera Stetter-Stättermayer, 1937; Ent. Rdsch. 54 (42): 531, TL: Sebdou, Algeria
Lycaena allardi ab. depuncta Stetter-Stättermayer, 1937; Ent. Rdsch. 54 (42): 531, TL: Sebdou, Algeria
Lycaena allardi ab. communa Stetter-Stättermayer, 1937; Ent. Rdsch. 54 (42): 531, TL: Sebdou, Algeria
Lycaena allardi ab. tristis Stetter-Stättermayer, 1937; Ent. Rdsch. 54 (42): 531, TL: Sebdou, Algeria
Lycaena allardi ab. brunnea Stetter-Stättermayer, 1937; Ent. Rdsch. 54 (42): 531, TL: Sebdou, Algeria
Lycaena allardi ab. fusca Stetter-Stättermayer, 1937; Ent. Rdsch. 54 (42): 531, TL: Sebdou, Algeria
Lycaena allardi ab. oberthüri Stetter-Stättermayer, 1937; Ent. Rdsch. 54 (42): 531, TL: Sebdou, Algeria
Lycaena allardi ab. suffusa-argentea Stetter-Stättermayer, 1937; Ent. Rdsch. 54 (42): 532, TL: Sebdou, Algeria
Lycaena allardi ab. luxurians Stetter-Stättermayer, 1937; Ent. Rdsch. 54 (42): 532, TL: Sebdou, Algeria
Lycaena allardi ab. rufomarginata Stetter-Stättermayer, 1937; Ent. Rdsch. 54 (42): 532, TL: Sebdou, Algeria
Lycaena allardi ab. elliptica Stetter-Stättermayer, 1937; Ent. Rdsch. 54 (42): 532, TL: Sebdou, Algeria
Lycaena allardi ab. pluripuncta Stetter-Stättermayer, 1937; Ent. Rdsch. 54 (42): 532, TL: Sebdou, Algeria
Lycaena allardi ab. polyphemus Stetter-Stättermayer, 1937; Ent. Rdsch. 54 (42): 532, TL: Sebdou, Algeria
Lycaena allardi ab. parvipuncta Stetter-Stättermayer, 1937; Ent. Rdsch. 54 (42): 532, TL: Sebdou, Algeria
Plebejus allardii antiatlasicus Tarrier, 1993; (nom. nud.?), TL: Tizi-n-Taraktine
Plebejus allardi ; [BMAT: 40, pl. 12, f. 26-38]; [NEN40]
Plebejus martini allardi ; [H&R, 275]
allardi Oberthür, 1874; [NEN46, 10]

Larva on Astragalus caprinus [BMAT]


antiatlasicus
[antiatlasicus-1uf.jpg] 600x435 (~34Kb) upperside female Morocco, Tizi-n-Tarakatine, Anti-Atlas, Photo © Michel Tarrier [antiatlasicus-2v.jpg] 600x435 (~44Kb) underside Morocco, Tizi-n-Tarakatine, Anti-Atlas, Photo © Jean Delacre


Plebejus beani Bálint & Johnson, 1997
C.Asia (mountains), Himalayas. See [About maps] Plebejus beani Bálint & Johnson, 1997; Neue Ent. Nachr. 40, (repl. for Lycaena indica Evans, 1925)
Lycaena pylaon indica Evans, 1925; J. Bombay nat. Hist. Soc. 30 (2): 346, pl. 27, f. 234 (homonym of Lycaena indica, Murray, 1874), TL: Baluchistan, Hanna and W.Tibet
indica ; [NEN46, 36]

Plebejus hespericus (Rambur, 1840)
Iberia. See [About maps] Lycaena hesperica Rambur, 1840; Faune ent. Andal. 2 (5): 270, 2 (4): pl. 10, f. 1-4
Plebejus hespericus, Moths and Butterflies of Europe [MBE]
Plebejus pylaon hespericus ; [H&R, 275]
Plebejus hespericus ; [NEN40]; [Otakar Kudrna]

Larva on Astragalus arragonensis [H&R]


Plebejus martini (Allard, 1867)
Morocco, Algeria. See [About maps] Martin's Blue
Lycaena martini Allard, 1867; Ann. Soc. ent. Fr. (4) 7 : 314, 319, pl. 6, f. 2a-b, TL: Lambessa (Algeria)
Plebejus martini, Moths and Butterflies of Europe [MBE]
martini ; [NEN46, 43]
Plebejus martini ; [NEN40]; [H&R, 274]
Algeria (Batna, Lambessa, Khenchala, Kabylie) Plebejus martini martini ; [BMAT: 39, pl. 12, f. 1-8]

Larva on Astragalus spp., Astragalus massiliensis, Acanthyllis numidia [BMAT]

P. m. ungemachi (Rothschild, 1926)
Morocco (High Atlas, Middle Atlas) Lycaena allardi ungemachi Rothschild, 1926; Bull. Soc. Sci. nat. Maroc. 5: 333, TL: Aselda (High Atlas), Aghbalu Larbi, Taghzeft Pass (Middle Atlas) (Morocco)
ungemachi ; [NEN46, 62]
Plebejus martini ungemachi ; [BMAT: 39, pl. 12, f. 9-17]
[ungemachi-1um.jpg] 600x435 (~39Kb) upperside male Morocco, Tizi-n-Talrhemt, Haut Atlas , Photo © Jean Delacre [ungemachi-2uf.jpg] 600x435 (~57Kb) upperside female Morocco, Tirhboula, Moyen Atlas (ovipositing on Astragalus nemorosus), Photo © Jean Delacre [ungemachi-2vf.jpg] 600x435 (~57Kb) underside female Morocco, Tirhboula, Moyen Atlas (ovipositing on Astragalus nemorosus), Photo © Jean Delacre
[ungemachi-3v.jpg] 599x435 (~38Kb) underside Morocco, Djebel Ayachi, Haut Atlas, Photo © Jean Delacre

Larva on Astragalus incanus ssp. incurvus, Astragalus spp.? [BMAT]


P. m. regularis Tennent, 1995
Morocco (Dj. Lakra, ?western Rif Mts.) Plebejus martini regularis Tennent, 1995; Entomologist's Gaz. 46 (2): 121, TL: Dj. Lakraa (Morocco)
regularis ; [NEN46, 54]
Plebejus martini regularis ; [BMAT: 40, pl. 12, f. 18-25]

Larva on Astragalus armatus [BMAT]

P. m. mgouna Tarrier, 2002
Morocco Plebejus martini mgouna Tarrier, 2002; , TL: Morocco
[mgouna-1v.jpg] 600x435 (~43Kb) underside Morocco, Djebel Azourki, Haut Atlas, Photo © Jean Delacre


Plebejus nichollae (Elwes, 1901)
Levant Lycaena nicholli Elwes, 1901; in Nicholl, Trans. ent. Soc. Lond. 1901 (1) : 96, TL: Baalbek, Lebanon
?Lycaena zephyrus nicholli ;
Plebejus pylaon nicholli ;
nicholli ; [NEN46, 47]
Plebejus nichollae ; [NEN40, (spell.?)]

Plebejus patriarcha Bálint, 1992
Darvaz Plebejus patriarcha Bálint, 1992; Atalanta 23: 134, pl. VII, f. 1-4, TL: Karnak, Darwas
patriarcha ; [NEN46, 50]
Plebejus patriarcha ; [NEN40]
Plebejides patriarchus ; [BRU2: 170, pl. 71, f. 31-33]

Larva on Astragalus sp. [BRU2, 170]


Plebejus philbyi Graves, 1925
Levant (deserts) Plebeius sephirus philbyi Graves, 1925; Trans. ent. Soc. Lond. 1925: 55, pl. 5, f. 1-2, TL: Transjorda, Petra
philbyi ; [NEN46, 52]
Plebejus indica philbyi ;
Plebejus philbyi ; [NEN40]
Polyommatus (Albulina) pylaon philbyi ; [AfrL]

Plebejus pylaon (Fischer de Waldheim, 1832)
SEU, Asia Minor, S.Siberia (mountains), N.Mongolia. See [About maps] Zephyr Blue Nińa de Astragalo
Lycaena pylaon Fischer de Waldheim, 1832; Nouv. Mém. Soc. Imp. Nat. Moscou 2 : 357, pl. 19, f. 5-6, TL: Povolzhye ( ?Sarepta)
cyane (Freyer, [1845])
Plebejus pylaon, Moths and Butterflies of Europe [MBE]
Plebejus pylaon, Captain's European Butterfly Guide [Simon Coombes]
Plebejus pylaon ; [BARU, #402]; [H&R, 275]; [BOW: pl. 10, f. 20, 22]; [NEN40]; [Otakar Kudrna]
Plebejides pylaon ; [BRU2: 169, pl. 71, f. 43-45]
Polyommatus (Albulina) pylaon ; [AfrL]

Larva on Astragalus [H&R], Astragalus rupifragus, (NW.Kazakhstan) [BRU2, 169]

S.Europe, SW.Siberia, Turan, W.Altai Plebejides pylaon pylaon
Plebejides pylaon pylaon ; [BRU2, 169]
P. p. katunensis Balint & Lukhtanov, 1990
Altai, Kuznetskiy Alatau Plebejus pylaon katunensis ; [BARU, (note)]
Plebejides pylaon katunensis ; [BRU2, 169]
P. p. cleopatra Hemming, 1934
Plebejus cleopatra Hemming, 1934; Entomologist 67: 30, TL: Tel Khuweilfe, n. Beersheba
cleopatra ; [NEN46, 24]
P. p. delattini Junge, 1971
Plebejus pylaon delattini Junge, 1971; NachrBl. bayerisch. Ent. 20: 35, f. 1-4, TL: Südtirol, Vinschgau, Italy
delattini ; [NEN46, 25]
P. p. solimana (Forster, 1938)
Lycaena pylaon solimana Forster, 1938; Ent. Rundsch. 55: 336, pl. 1-2, f. 4-6, TL: Elburs, Tacht i Suleiman, Iran
solimana ; [NEN46, 59]
P. p. tadjikus Tschikolovetz, ?˛
pylaon tadjikus Tschikolovetz, ?˛

Plebejus sephirus (Frivaldszky, 1835)
E.Europe, Caucasus, Asia Minor. See [About maps] Lycaena sephirus Frivaldszky, 1835; A Magyar Tudós Társág Évkönyvei, 2: 269, pl. 7, f. 1-2, TL: Sliven, Balkan Mts, Bulgaria
Lycaena zephyrus akbesiana Oberthür, 1904; Étud. Lépid. Comp. 1: 16, pl. 2, f. 21-22, TL: Syrie, Akbés
Lycaena sephyrus uhryki Rebel, 1911; Ent. Z. 25: 191, TL: Deliblát, Flamunda, Yugoslavia
Plebejus modica Verity, 1935; Bull. Soc. ent. Fr. 40 : 245, TL: Asia Minor
= ; [BRU2, 169]
microsephyrus Verity, 1935˛
Plebeius sephirus kovacsi Szabó, 1954; Folia ent. hung. (N.S) 7: 39, TL: Csomád Fót, Hungary
Plebeius sephirus proximus Szabó, 1954; Folia ent. hung. (N.S) 7: 39
Plebeius foticus Szabó, 1956; Folia ent. hung. (N.S) 9: 276
Plebejus brethertoni Brown, 1976; Entomol. Ber., Amst. 36 (3): 46
Plebejides magnificus Bálint, 1987

sephirus ; [NEN46, 58]; Abadjiev, 2002, Neue Ent. Nachr. 23: 22
akbesiana ; [NEN46, 9]
uhryki ; [NEN46, 62]
kovacsi ; [NEN46, 40]
zephyrus ; (missp.)
sephyrus ; (missp.)
Plebejus pylaon sephirus ; [H&R, 275]
Plebeius pylaon sephirus ;
Plebejus sephirus ; [NEN40]; [Otakar Kudrna]
Plebejides sephirus ; [BRU2: 169, pl. 71, f. 37-42]

Larva on Astragalus spp., (ants Lasius alienus, Formica pratensis) [BRU2, 170]

P. s. kubanensis Shchurov, 1999˛
NW.Caucasus sephirus kubanensis Shchurov, 1999˛
Plebejides sephirus kubanensis ; [BRU2, 169]

Plebejus trappi Verity, 1927
Alps. See [About maps] Plebeius trappi Verity, 1927; Ann. Soc. ent. Fr. 96 : 16, TL: Simplon
lycidas Trapp, 1863
Plebejus trappi ; [NEN40]; [Otakar Kudrna]
Plebejus pylaon trappi ; [H&R, 275]

Larva on Astragalus excapus [H&R]


Plebejus vogelii (Oberthür, 1920)
Morocco (high Atlas). See [About maps] Vogel's Blue
Lycaena vogelii Oberthür, 1920; Étud. Lépid. Comp. 17 : 55, pl. G, TL: Tizi-n-Taghzeft (Morocco)
vogelii ; [NEN46, 63]
Plebejus vogelii ; [NEN40]; [H&R, 274]
Morocco (near Taghzeft Pass) Maurus vogelii vogelii
Maurus vogelii vogelii ; [BMAT: 38, pl. 11, f. 69-75]
[vogelii_vogelii-1u.jpg] 600x435 (~36Kb) upperside Morocco, Aďt-Kermouss, Moyen Atlas, Photo © Michel Tarrier

Larva on Erodium cheilanthifolium [BMAT]


P. v. insperatus (Tennent, 1996)
Morocco (Tizi-n-Test) Maurus vogelii insperatus Tennent, 1996; The Butterflies of Morocco, Algeria and Tunisia: 38, pl. 11, f. 76-81, TL: Tizi-n-Test, High Atlas Mts., Morocco
insperatus ; [NEN46, 36]

Larva on Erodium cheilanthifolium [BMAT]


Plebejus zephyrinus (Christoph, 1884)
C.Asia (mountains). See [About maps] Lycaena zephyrinus Christoph, 1884; in Romanoff, Mém. Lépid. 1 : 102, pl. 6, f. 3a-b, TL: Askhabad and Nuchur, Iran
zephyrinus ; [NEN46, 64]
Plebejus zephyrinus ; [NEN40]
Plebejides zephyrinus ; [BRU2: 170, pl. 71, f. 19-30]

Larva on Astragalus sp. [BRU2, 170]

Kopet-Dagh Plebejides zephyrinus zephyrinus
Lycaena turcmenica Forster, 1936; Mitt. münch. ent. Ges. 26 (2) : 55
Plebejides zephyrinus zephyrinus ; [BRU2, 170]
P. z. usbekus Forster, 1939
W.Tian-Shan, N.Tian-Shan Plebejus usbekus Forster, 1939; in Muksgaard, Lepidopteren aus Iran. (1): 6 (repl. zephyrinus Staudinger, 1886)
Lycaena zephyrinuus zephyrinus Staudinger, 1886; Stett. Ent. Ztg. 47 (4-6): 204 (preocc. Lycaena zephyrinus Christoph, 1884), TL: Samarkand, Uzbekistan
zephyrinus ; [NEN46, 65]
Plebeius pylaon usbeka ;
Plebejus usbekus ; [NEN40]
Plebejides zephyrinus usbecus[sic, recte usbekus] ; [BRU2, 170]
P. z. forsteri (Bálint, 1900)
Dzhungarsky Alatau Mts. Plebejides zephyrinus forsteri Bálint, 1900
Plebejides zephyrinus forsteri ; [BRU2, 170]
P. z. tarbagataiensis (Bálint, 1992)
Saur, Tarbagatai, S.Altai Plebeius zephyrinus tarbagataiensis Bálint, 1992; in Bálint, Kertéz & Lukhtanov, Ent. oboz. 71 (4): 875, f. 69-70, TL: Altai, Kenderlik
tarbagataiensis ; [NEN46, 60]
Plebejides zephyrinus tarbagataiensis ; [BRU2, 170]
P. z. ordubadi (Forster, 1938)
Caucasus Major, Armenia (highlands), Talysh Lycaena ordubadi Forster, 1938; Ent. Rundsch. 55: 238, TL: Armenia
Lycaena solimana Forster, 1938; Ent. Rundsch. 55: 336, TL: Elburs Mts.
= ; [BRU2, 170]
Lycaena iranica Forster, 1938; Ent. Rundsch. 55: 216, TL: N.Iran
= ; [BRU2, 170]
Plebejus (Plebejides) pylaon albertii Nekrutenko, 1975; J. Lep. Soc. 29 (3): (151-155)
Plebejus (Plebejides) pylaon abchasicus Nekrutenko, 1975; J. Lep. Soc. 29 (3): (151-155)
iranicola Koçak, 1980
Plebejus sephirus semiturcmenicus Bálint, 1991; A Janus Pannonius Múzeum Évkönyve (Pécs) 35: 51, f. 51-52, TL: Baku, Azerbaijan
= ; [BRU2, 170]
Plebejus pylaon semiturcmenicus ;
semiturcmenicus ; [NEN46, 58]
Plebejides zephyrinus ordubadi ; [BRU2, 170]

Plebejus klausrosei (Bálint, 1992)
S.Ghissar. See [About maps] Plebejides klausrosei Bálint, 1992; in Bálint, Kertéz & Lukhtanov, Ent. oboz. 71 (4): 875, f. 55-58, TL: Tadshikistan, Wachschski Chrebet
Plebejides klausrosei ; [BRU2: 170, pl. 71, f. 34-36]

The sarta species-group
Alpherakya

Plebejus devanica (Moore, [1875])
Pamirs, Hindu Kush, Karakorum. See [About maps] Polyommatus devanica Moore, [1875]; Proc. zool. Soc. Lond. 1874 (4) : 573, pl. 66, f. 4, TL: Ladak, Dras Valley
Polyommatus sarta gooraisica Tytler, 1926; J. Bombay nat. Hist. Soc. 31 (3): 586, TL: Goorais Valley, India
devanica ; [NEN46, 26]
Polyommatus devanica ; [BOW: pl. 206, f. 25]
gooraisica ; [NEN46, 33]
?Alpherakya sartoides gooraisica ; [BRU2, 177 (?)]
Plebejus devanicus ; [NEN40]
Alpherakya devanica ; [BRU2: 177, pl. 72, f. 1-6]
P. d. tshikolovetsi (Bálint, 1999)
W.Pamirs, S.Pamirs Polyommatus devanica tshikolovetsi Bálint, 1999; Neue Entomologische Nachrichten, Band 46: 34 (repl. Polyommatus devanica evansi Tshikolovets, 1977)
Polyommatus devanica evansi Tshikolovets, 1997; The Butterflies of Pamir: 141 (repl. Lycaena sarta gracilis Evans, 1912, preocc. Lycaena gracilis Miskin, 1890)
Lycaena sarta gracilis Evans, 1912; J. Bombay nat. Hist. Soc. 21 (2): 559, TL: Chitral, 12000
gracilis ; [NEN46, 34]
Alpherakya devanica evansii ; [BRU2, 177]
P. d. bellona (Grum-Grshimailo, 1888)
NW.Pamirs Lycaena bellona Grum-Grshimailo, 1888; Horae Soc. ent. Ross. 22 (3-4) : 306, TL: Koudara
Polyommatus devanica kohibabaensis Sakai, 1978; Atalanta 9 (1): 116, TL: Paghman Mts, 3500-3800m, Afghanistan
bellona ; [NEN46, 19]
kohibabaensis ; [NEN46, 40]
Plebejus bellona ; [NEN40]
Polyommatus bellona ;
Alpherakya devanica bellona ; [BRU2, 177]
P. d. rupala (Evans, 1925)
Polyommatus sarta rupala Evans, 1925; J. Bombay nat. Hist. Soc. 30 (2): 350, TL: Rupal, N.Astor, 11000
Polyommatus sarta rupala Tytler, 1926; J. Bombay nat. Hist. Soc. 31 (3): 586, pl. 1, f. 14-15, TL: Rupal, N.Astor, 11000
rupala ; [NEN46, 55]; [NEN46, 56]
?Alpherakya sartoides rupala ; [BRU2, 177 (?)]

Plebejus pilgram Bálint & Johnson, 1997
C.Asia (mountains). See [About maps] Plebejus pilgram Bálint & Johnson, 1997; Neue Ent. Nachr. 40: 16 (replacement for Lycaena serica Grum-Grshimailo, 1902)
Lycaena serica Grum-Grshimailo, 1902; (homonym Lycaena serica C. Felder, 1862)

Plebejus sarta (Alphéraky, 1881)
Afghanistan, Pakistan, Ghissar, Alai, Tian-Shan, Tarbagatai, Saur, S.Altai. See [About maps] Lycaena sarta Alphéraky, 1881; Horae Soc. ent. Ross. 16 (3-4): 387, pl. 14, f. 8, TL: Kuldja
Lycaena phryxis Lang, 1884; Rhopalocera Europa Descripta et Delineata: 372, TL: Samarkand
Lycaena phryxis Staudinger, 1886; Stett. Ent. Ztg. 47 (4-6): 212
?Lycaena sarta ab. ♀ pupillata Courvoisier, 1913; Ent. Mitt. 2 (10) : 292
Plebejus sarta ; [NEN40]
Polyommatus sarta ;
Alpherakya sarta ; [BRU2: 176, pl. 72, f. 7-9]

Larva on Pseudosedum longidentatum, Rosularia kokanica, Orostachys thyrsiflora [BRU2, 176]

P. s. laziensis (Huang, 1998)
C.Tibet Polyommatus sarta laziensis Huang, 1998; Neue Entomologische Nachrichten, 41: 266, pl. f. 3a-b, TL: Lazi, C.Tibet, 4500m

Plebejus sartoides (Swinhoe, 1910)
Darvaz, Alai, Pamirs, Afghanistan, Pakistan, N.India. See [About maps] Polyommatus devanica sartoides Swinhoe, 1910; in Moore, Lepidoptera Indica 8 : 21, pl. 645, f. 1, 1a-b, TL: Jhela Drosh, Chitral
Polyommtus sarta sartoides ;
Alpherakya sartoides ; [BRU2: 177, pl. 72, f. 10-12]

The eurypilus species-group
Kretania

Plebejus psylorita (Freyer, 1845)
Crete Cretan Argus
Lycaena psylorita Freyer, 1845; Neuere Beitr. Schmett. 5 : 146, pl. 469, f. 3-4, TL: Mt. Ida, Crete
Polyommatus psylorita ; [NEN40]
Kretania psylorita ; [H&R, 280]; [BOW: pl. 9, f. 32]; Shchurov & Lukhtanov, 2001, Atalanta 32 (1/2): 218
Plebejus psyloritus ; [Otakar Kudrna]

Larva on Astragalus [BHR]


Plebejus carmon (Gerhard, [1851])
Balkans, Asia Minor, Iraq, Iran. See [About maps] Eastern Brown Argus
Lycaena carmon Gerhard, [1851]; Versuch Mon. europ. Schmett.: 15, pl. 25, f. a-b, TL: Turkey
Lycaena eurypilus Freyer, 1852; Neuere Beitr. Schmett. 6 : 46, pl. 573, f. 4, TL: Amasia, Turkey
= ; [Carbonell]
eurypilus Freyer, 1852; [NEN46, 29]
Kretania eurypilus ; [H&R, 280]; [BRU2: 182, pl. 68, f. 57-62]
Plebejus euripilus[sic] ; [BOW: pl. 206, f. 23 (text)]
Polyommatus eurypilus ; [NEN40]
Plebejus eurypilus ; [Otakar Kudrna]
Plebeius (Kretania) carmon ; [Carbonell]

Larva on (Transcaucasia) Astracantha arnacanthoides, (Greece) A. rumelica, (Turkey) A. gummifera, (S.Transcaucasia) A. aurea, A. caucasica, (ants Camponotus kiesenwetteri) [BRU2, 182]

Asia Minor, Transcaucasia, W.Iran Kretania eurypilus eurypilus ; Shchurov & Lukhtanov, 2001, Atalanta 32 (1/2): 218, pl. XV, f. 7-8
P. c. euaemon (Hemming, 1931)
Polyommatus eurypilus euaemon Hemming, 1931; Ann. Mag. Nat. Hist. (10) 8: 536, TL: Hermon, 5600ft, Lebanon
euaemon ; [NEN46, 29]
Polyommatus euaemon ; [NEN40]
Kretania eurypilus euaemon ; Shchurov & Lukhtanov, 2001, Atalanta 32 (1/2): 219
P. c. pelopides (van der Poorten, 1984)
S.Greece Kretania eurypilus pelopides van der Poorten, 1984; Nota Lepid. 7 (2): 164, TL: S.Peloponnesos
Kretania eurypilus pelopides ; Shchurov & Lukhtanov, 2001, Atalanta 32 (1/2): 219

Plebejus iranicus (Forster, 1938)
Kurdistan, Elburs, C.Asia? (mountains). See [About maps] Lycaena eurypilus iranica Forster, 1938; Ent. Rundsch. 55: 215, pl. 1-2, f. a5, a6, TL: Elburs, Tacht i Suleiman
iranica ; [NEN46, 37]
Polyommatus iranicus ; [NEN40]
Kretania iranica ; Shchurov & Lukhtanov, 2001, Atalanta 32 (1/2): 219

Plebejus csomai (Bálint, 1992)
W.Himalayas. See [About maps] Polyommatus csomai Bálint, 1992; Linn. Belg. 13: 434, f. 6-7, TL: Chitral, Pakistan
csomai ; [NEN46, 24]
Polyommatus csomai ; [NEN40]
Kretania csomai ; Shchurov & Lukhtanov, 2001, Atalanta 32 (1/2): 220

Plebejus zamotajlovi (Shchurov & Lukhtanov, 2001)
NW.Caucasus. See [About maps] Kretania zamotajlovi Shchurov & Lukhtanov, 2001; Atalanta 32 (1/2): 220, f. 2-3, pl. XV, f. 1-6, pl. XVI, TL: Russia, Krasnodar, Black Sea Coast, Abrau peninsula, 10km SE Anapa, Navagir Mt. Range

Larva on Astragalus arnacanthoides Shchurov & Lukhtanov, 2001, Atalanta 32 (1/2): 224


The ? species-group

Plebejus shuroabadicus Shchetkin, 1963
Darvaz Plebejus shuroabadica Shchetkin, 1963; Trudy Inst. Zool. Parasit., Stalinabad 24: 57, TL: [Pass Shuroabad, Khozratishoh Mts., Tajikistan]
Plebeius shuroabadicus ; [BRU2: 159, pl. 83, f. 37-39]

Plebejus dzhizaki Zhdanko, 2000
Plebeius dzhizaki Zhdanko, 2000; [BRU2: 159, pl. 73, f. 40-42], TL: Uzbekistan, Dzhizak Region, near Buston

Plebejus bergi Kusnezov, 1908
Kazakhstan. See [About maps] Plebejus bergi Kusnezov, 1908; Izvest. Turkest. Otd. Russ. Geogr. Obsc. 4 (8): 107, pl. 4a, f. 1-3, TL: [N. coast of Aral Sea, Kazakhstan]
Plebeius bergi ; [BRU2: 168, pl. 74, f. 28-30]

Plebejus noah (Herz, 1900)
Turan. See [About maps] Lycaena noah Herz, 1900; Ann. Mus. Zool. Acad. Sci. St.-Petersb. 5 (4) : 442, TL: Noah-Pairambar... Bucharen, Noah
Vacciniina alcedo noah ;
Plebeius noah ; [BRU2: 169, pl. 83, f. 28-30]

Larva on Alhagi spp. [BRU2, 169]


Plebejus qinghaiensis (Murayama)
Lycaeides qinghaiensis Murayama
Lycaeides qinghaiensis ; [MRS, 688]

Plebejus hishikawai (Yoshino, 2003)
S.Tibet. See [About maps] Lycaeides hishikawai Yoshino, 2003; Futao (43): 7, f. 10-12, 14-16, 36, TL: 50km SE from Lhasa, 4200m, S Tibet

Plebejus callaghani (Carbonell & Naderi, 2007)
N.Iran. See [About maps] Plebeius callaghani Carbonell & Naderi, 2007; Bull. Soc. ent. Fr. 112 (1): 130, TL: Iran, NE. Qazvin, "Alamut pass" (N36°24'/E50°13'), 2200-2500m, prov. Zanjan


Unknown or unplaced taxa

regina Rühl, 1893; (Middle Asia)

Lycaena argus f. ligurica Courvoisier, 1910; Ent. Zs. 24 (16): 81
Lycaena argus f. ligurica ; Courvoisier, 1911, Dt. ent. Z. Iris 25 (9) : 103, pl. 2, f. 1

Lycaena argus f. nivea Courvoisier, 1910; Ent. Zs. 24 (16): 81
Lycaena argus f. nivea ; Courvoisier, 1911, Dt. ent. Z. Iris 25 (9) : 103, pl. 2, f. 2

Lycaena aegon f. nigrescens Courvoisier, 1910; Ent. Zs. 24 (18): 94
Lycaena aegon f. nigrescens ; Courvoisier, 1911, Dt. ent. Z. Iris 25 (9) : 103, pl. 2, f. 3

Lycaena aegon f. carinthiaca Courvoisier, 1913; Ent. Mitt. 2 (10) : 289

Lycaena aegon f. killiasii Courvoisier, 1913; Ent. Mitt. 2 (10) : 290

Lycaena aegon ab. ♀ inornata Courvoisier, 1913; Ent. Mitt. 2 (10) : 290

Lycaena lycidas ab. rubrimaculata Courvoisier, 1913; Ent. Mitt. 2 (10) : 291

Plebeius leucofasciatus Röber, 1886; Corr.-Bl. Ent. Ver. Iris 1 (3) : 59, pl. 4, f. 32, TL: Hongkong
Plebejus leucofasciatus ; [NHM]


11.2.2013 (62)

References:
[ALBERTA]
University of Alberta, E.H. Strickland Entomological Museum; Browse Entomology Collection
[About maps]
Warning! The maps are automatically generated from the textual information, and the process does not always produce acceptable result; See about maps for more info.
[AfrL]
Mark C. Williams; Checklist of Afrotropical Papilionoidea and Hesperoidea; Compiled by Mark C. Williams, 7th ed. (2008) (April 2007); Afrototropical Butterflies; e-mail: Mark C. Williams
[BAMONA]
Opler, Pavulaan, Stanford & Pogue; Butterflies and Moths of North America; Mountain Prairie Information Node
[BARU]
Korshunov & Gorbunov; Butterflies of the Asian part of Russia; English translation by Oleg Kosterin; [Dnevnye babochki aziatskoi chasti Rossii. Spravochnik]
[BHL]
The; Biodiversity Heritage Library; and on; Internet Archive; Note: The linking to the content of Internet Archive is somewhat delicate -- notify me when links break!
[BHR]
D'Abrera; Butterflies of the Holarctic Region, Part I-III
[BMAT]
Tennent, 1996; The butterflies of Morocco, Algeria and Tunisia; ISBN 0 906802 05 9
[BOA]
Interactive Listing of American Butterflies; Listado Interactivo de la Mariposas Americanas; Butterflies of America
[BOC]
Robert Michael Pyle, 2002; The Butterflies of Cascadia
[BOW]
Lewis, 1974; Butterflies of the World; ISBN 0 245 52097 X
[BRU2]
Tuzov (ed.), Bogdanov, Churkin, Devyatkin, Dantchenko, Murzin, Samodurov, Zhdanko, 2000; Guide to the Butterflies of Russia and adjacent territories; Libytheidae, Danaidae, Nymphalidae, Riodinidae, Lycaenidae; Volume 2
[BnF]
Gallica / Bibliothčque nationale de France; Annales de la Société entomologique de France; Bulletin de la Sociéte entomologique de France
[CiNii]
http://ci.nii.ac.jp/
[D. Smirnov]
Russia, Moscow; e-mail: D. Smirnov
[EBW]
Smart, Paul, 1976; The Illustrated Encyclopedia of the Butterfly World; ISBN 0 600 31381 6
[ESP]
Heppner, 1981; The dates of E. J. C. Esper's Die Schmetterlinge in Abblidungen... 1776-[1830]; Archives of Natural History (1981) 10 (2): 251-254
[Erich Mangl]
http://www.mangl.at
[H&R]
Higgins, L.G, Riley, N.D, 1970; A Field Guide to the Butterflies of Britain and Europe
[HBN]
Francis Hemming, 1937; HÜBNER, A bibliographical and systematic account of the entomological works of Jacob Hübner and of the supplements thereto by Carl Geyer, Gottfried Franz von Frölich and Gottlieb August Wilhelm Herrich-Schäffer, Vol 1 & 2
[Harri Arkkio]
Finland; For the time being these pictures are only for this site. Other usage not allowed.
[ILMB]
Interactive Listing of Mexican Butterflies
[Jean Delacre]
http://homepage.mac.com/jdelacre
[Kimmo Silvonen]
Espoo, Finland; Kimmo's Lep Site
[LAMAS-2005]
Gerardo Lamas; Entomofauna 26 (6): 57-100; A Bibliography of the Zoological Publications of Hans Fruhstorfer (1886* - 1922+)
[MBE]
Moths and Butteflies of Europe; Paolo Mazzei, Diego Reggianti and Ilaria Pimpinelli; http://www.leps.it/
[MRS]
Chou Io (Ed.); Monographia Rhopalocerum Sinensium, 1-2
[Mario Maier]
Europäische Schmetterlinge; e-mail: Mario Maier
[Michel Tarrier]
Michel R. Tarrier; Malaga, Spain; http://www.tarrier.org
[NAB]
Pyle, 1981; Field Guide to North American Butterflies, National Audubon Society; ISBN 0-394-51914-0
[NACL]
Hodges, 1983; Check List of the Lepidoptera of America North of Mexico
[NEN40]
Zsolt Bálint & Kurt Johnson, 1997; Neue Entomologische Nachrichten 40; Reformation of the Polyommatus Section with a Taxonomic and Biogeographic Overview (Lepidoptera, Lycaenidae, Polyommatini); ISSN 0722-3773; e-mail: Kurt Johnson; and his web site; http://www.edutrek.net/
[NEN46]
Zsolt Bálint, 1999; Neue Entomologische Nachrichten 46; Annotated list of type specimens of Polyommatus sensu Eliot of the Natural History Museum, London
[NHM]
Natural History Museum; The Global Lepidoptera Names Index
[NL4A]
Gerardo Lamas, 2004; Atlas of Neotropical Lepidoptera; Checklist: Part 4A; Hesperioidea - Papilionoidea
[NORLEP]
Norges Sommerfugler
[Norbert Kondla]
Canada, BC; e-mail: Norbert Kondla
[Oleg Kosterin]
Institue of Cytology & Genetics; Siberian Division of Russian Academy of Sciences; Novosibirsk; e-mail: Oleg Kosterin; http://pisum.bionet.nsc.ru/kosterin/
[Opler]
Opler & Warren, 2003 (April); Butterflies of North America. 2. Scientific Names List for Butterfly Species of North America, north of Mexico.
[Pertti Pakkanen]
e-mail: Pertti Pakkanen; Butterfly and moth observations from Finland
[SPRK]
Seppänen, E. J, 1970; Suomen suurperhostoukkien ravintokasvit, Animalia Fennica 14
[SVF]
Svenska fjärilar; Naturhistoriska riksmuseet; (Sweden)
[SZMN]
Lepidoptera; Siberian Zoological Museum
[Simon Coombes]
1 Park Street, Plymouth, PL3 4BL, UK; 01752 607854; e-mail: Simon Coombes; http://www.butterfly-guide.co.uk
[˛]
This may require parentheses or not. I don't have the necessary information for this taxon.


Some related literature:
Abadjiev, 2002
Types of Balkan butterflies in the collection of The Natural History Museum, London Neue Ent. Nachr. 23: 3-53
Agenjo, 1966
Morfologia y distribución geográfica en Espańa de la "nińa hocecillas"Plebejus (Plebejus) argus (L., 1758) (Lep. Lycaenidae) Rev. Esp. Ent. 41: 163-179
Allard, 1867
Notes sur les insectes de l'Algerie Ann. Soc. ent. Fr. (4) 7 : 311-322, pl. 6
Alphéraky, 1881
Lépidoptčres du district de Kouldjŕ et des montagnes environnantes. (1-3) Horae Soc. ent. Ross. 16 (3-4): 334-435, pl. 14-15 (1881), 17 (1-2): 15-103, pl. 1-3 (1882), 17 (3-4): 156-227, pl. 8-9 (1883)
Bálint, 1991
Egy xeromontán boglárkalepke: a Plebejus pylaon (Fischer von Waldheim, 1832) és rokonsági köre (Lepidoptera: Lycaenidae), 1 A Janus Pannonius Múzeum Évkönyve (Pécs) 35: 33-69
Bálint, 1992
A new species of Plebejides Sauter, 1968 from Central Asia (Lepidoptera, Lycaenidae) Atalanta 23: 133-138, [3] figs, pl. 8
Bálint, 1992
New taxa of the Polyommatus section (sensu Eliot, 1973) from Central Asia. Studies on Central Palaearctic Lycaenids II Linn. Belg. 13: 423-445, f. 1-12
Bálint, 1992
A Survey of the Subgenus Plebejides Sauter, 1968 (Lepidoptera, Lycaenidae) in Bálint, Kertéz & Lukhtanov, Ent. oboz. 71 (4): 863-886, 74figs
Bálint, 1999
Annotated list of type specimens of Polyommatus sensu Eliot of the Natural History Museum, London Neue Entomologische Nachrichten, Band 46: 1-87
Bálint, Guppy, Kondla, Johnson & Durden, 2001
Plebeius Kluk, 1780 or Plebejus Kluk, 1802? (Lepidoptera: Lycaenidae) Folia ent. hung. 62: 177-184
Bálint & Johnson, 1997
Reformation of the Polyommatus Section with a Taxonomic and Biogeographic Overview (Lepidoptera, Lycaenidae, Polyommatini) Neue Ent. Nachr. 40: 1-68, 4pls, 23 figs
Behr, 1867
Enumeration of the Californian species of Lycaena Proc. Calif. Acad. Sci. 3 (4) : 279-283
Bellier, 1862
Variétčs nouvelles de Lépidoptčres observées en Corse et decrites Ann. Soc. ent. Fr. (4) 2 : 615-616
Bergstrasser, [1779]
Nomenclatur und Beschreibung der Insecten in der Graftschaft Hanau-Münzenburg Nomen. Ins., 2: 79pp, pl. 15-48
Boisduval, 1832
Icones historique des Lépidoptčres nouveaux ou peu connus. Collection, avec Figures coloritées, des Papillons d'Europe nouvellement découverts; ouvrage formant le complément de tous les auteurs iconographes Icon. hist. Lépid. Europe 1 (1-2): 1-24, pl. 1-4 (1832), (3-4): 25-40, pl. 5-8 (1832), (5-6): 41-56, pl. 9-12 (1832), (7-8): 57-72, pl. 13-16 (1832), (9-10): 73-104, pl. 17-20 (1833), (11-12): 105-120, pl. 21-24 (1833), (13-14): 121-144, pl. 25-28 (1833), (15-16): 145-160, pl. 29-32 (1833), (17-18): 161-184, pl. 33-36 (1833), (19-20): 185-208, pl. 37-40 (1834), (21-22): 209-224, pl. 41-44 (1834), (23-24): 225-248, pl. 45-47 (1834), (25-28): 249-252 (1834)
Boisduval, 1869
Lépidoptčres de la Californie Ann. Soc. ent. Belg. 12 : 5-28,37-94
Bremer, 1861
Neue Lepidopteren aus Ost-Sibirien und dem Amur-Lande gesammelt von Radde und Maack, bescrieben von Otto Bremer Bull. Acad. Imp. Sci. St. Petersb. 3 : 461-496, Mélanges biol. St.-Pétersb. 3 (5-6): 538-589
Bremer, 1864
Lepidopteren Ost-Sibiriens, insbesondere der Amur-Landes, gesammelt von den Herren G. Radde, R. Maack und P. Wulffius Mém. Acad. Sci. St. Pétersb. (7) 8 (1) : 1-104, 8pls
Bridges, 1994
Catalogue of Family-group and Genus-group names of the Riodinidae & Lycaenidae of the World Cat. Family and Genus group Riodinidae & Lycaenidae: 390pp
Butler, 1878
On some Butterflies recently sent home from Japan by Mr. Montagne Fenton Cist. Ent. 2 (19) : 281-286
Butler, 1880
On a collection of Lepidoptera from Candahar Proc. zool. Soc. Lond. 1880 (3) : 403-415, pl. 39
Butler in Butler & Fenton, [1882]
On butterflies from Japan, with which are incorporated notes and descriptions of new species by Montague Fenton Proc. Zool. Soc. Lond. 1881 (4) : 846-856
Carbonell & Naderi, 2007
Plebeius callaghani n. sp. dans le nord de l'Iran (Lep., Lycaenidae) Bull. Soc. ent. Fr. 112 (1): 130
Chapman, 1902
[Remarks on two butterflies taken at Bejar, West Central Spain] Proc. ent. Soc. Lond. 1902 : xxxv-xxxvii,[35-37]
Chapman, 1907
in Chapman & Champion, Entomology in N.W. Spain (Galicia and Leon) Trans. ent. Soc. Lond. 1907 (1) : 147-171, pl. 5-11
Chapman, 1917
A new European Lycaena Plebeius Argus (Argyrognomon) Aegus sp. nov. / Étud. Lép. comp. 14 : 41-57,59-76,pl. 1-20
Un nouveau Lycaena Plabeius Argus (Argyrognomon) et Aegus_, nova species
Christoph, 1884
Lepidopteren aus dem Achal-Tekke-Gebiete. Erster Theil in Romanoff, Mém. Lépid. 1 : 93-138, pl. 6-8
Clench, 1965
A collection of butterflies from western Chihuahua, Mexico. (Made by C. H. T. Townsend in 1899) Ent. News 76 : 157-162
Courvoisier, 1910
Entdeckungsreisen und kritische Spaziergänge ins Gebiet der Lycaeniden Ent. Zs. 24 (12): 59-60, (13): 65-67, (14): 70-71, (15): 77-78, (16): 81-82, (17): 88-89, (18): 92-94, (19): 99-101, (20): 106-108, (21): 112, (23): 125-127, (24): 131-132, (26): 141-142, (27): 147-149, (28): 156, (31): 167-170, (32): 175-177, (33): 181-182, (34): 185-188, (35): 190-192, (36): 196-200, (37): 201-204, (38): 205-209, (39): 210-214, (40): 227-228, (41): 230-231, (42): 233-236, (43): 237-238, (44): 241-242, (45): 245-247, (46): 249-250, (47): 253-255, (48): 257-259, (49): 261-263, (50): 265-266, (51/52): 269-270
Courvoisier, 1911
Einige neue oder wenig bekannte Lycaeniden-Formen Dt. ent. Z. Iris 25 (9) : 103-109, pl. 2
Courvoisier, 1913
Einige neue oder wenig bekannte Lycaeniden-Formen Ent. Mitt. 2 (10) : 289-297
Dana, 2005
New larval host plant for Lycaeides melissa melissa in Wisconsin and Minnesota and potential threat to Lycaeides melissa samuelis (Lycaenidae) J. Lep. Soc. 59 (3): 175-177
Dyar, 1903
A List of North American Lepidoptera and Key to the Literature of this Order of Insects Bull. U.S. nat. Mus. 52 : 1-723
Eliot, 1973
The higher classification of the Lycaenidae (Lepidoptera): a tentative Arrangement Bull. Br. Mus. nat. Hist. (Ent.) 28 (6) : 373-505, 162fgs, pl. 1-6
Edwards, 1861
Descriptions of certain Species of Diurnal Lepidoptera found within the limits of the United States and of British America. - No. [1]-3 Proc. Acad. nat. Sci. Philad. 13: 160-164 (1861), 14: 54-58, 221-226 (1862)
Edwards, 1873
Descriptions of diurnal Lepidoptera found within the United States Trans. amer. ent. Soc. 4 : 343-348
Edwards, 1882
Description of new species of butterflies found in the United States Papilio 2 (3) : 45-49
Edwards, 1882
Description of a new species of Lycaena from Newfoundland Can. Ent. 14 (10) : 194-195
Esper, 1782
Die Schmetterlinge in Abbildungen nach der Natur mit Beschreibungen. Theil I. Die Tagschmetterlinge. Fortsetzung. Band 2 Die Schmett. Th. I, Bd. 2 (1): 1-36, pl. 51-56 (1780), (2): 37-68, pl. 57-62 (1781), (3): 69-100, pl. 63-68 (1781), (4): 101-124, pl. 69-74 (1781), (5): 125-140, pl. 75-80 (1782), (7): 141-156, pl. 81-82 (1783), (8): 157-172, pl. 83-88 (1783), (9): 173-184, pl. 89-92 (1784), (28): 185-190, pl. 93 (1786)
Erschoff, 1874
Travels in Turkestan. Volume 2. Zoogeographical Investigations. Lepidoptera [in Russian] in Fedschenko, Travels in Turkestan. 2 (5) : 1-128, pl. 1-6
Evans, 1912
A list of Indian butterflies J. Bombay nat. Hist. Soc. 21 (2): 553-584, (3): 969-1008
Evans, 1925
The identification of Indian Butterflies (5-8) J. Bombay nat. Hist. Soc. 30 (1): 72-96, (2): 322-351, pl. 26-27, (3): 610-639, pl. 28, (4): 726-776, pl. 29
Evans, 1932
The Identification of Indian Butterflies (edn. 2) Indian Butterflies (edn. 2): x + 454pp, 9 figs, 32 pls
Eversmann, 1851
Description de quelques nouvelles espéce de Lépidoptéres de la Russie Bull. Soc. imp. Nat. Moscou 24 (2) : 610-644
Fischer de Waldheim, 1832
Lepidopterorum rariorum Rossiae observationes quinque Nouv. Mém. Soc. Imp. Nat. Moscou 2 : 355-360, 2pls
Forster, 1936
Beiträge zur Systematik der Tribus Lycaenini unter besonderer Berücksichtigung der argyrognomon under der argus Gruppe Mitt. münch. ent. Ges. 26 (2) : 41-150
Forster, 1938
Die Lycaena pylaon-Gruppe Ent. Rundsch. 55: 213-219, 236-239, 334-337, 361-364, 417-420, 485-490, 2pls
Forster, 1939
in Munksgaard, Danish Scientific Investigations in Iran, Part 1 in Muksgaard, Lepidopteren aus Iran. (1): 7pp
Forster, 1940
Beiträge zur Kenntnis zentralasiatischer Lycaeniden Dt. ent. Z. Iris 54: 70-84, pl. 2-3
Frey, 1882
Zweiter Nachtrag zur Lepidopteren-Fauna der Schweiz Mitt. schweiz. ent. Ges. 6 (7) : 349-375
Freyer, 1845
Neuere Beiträge zur Schmetterlingskunde mit Abbildungen nach der Natur. (65-80) Neuere Beitr. Schmett. 5 : 1-40 (1842), : 41-92 (1844), : 93-164 (1845)
Freyer, 1852
Neuere Beiträge zur Schmetterlingskunde mit Abbildungen nach der Natur. (81-100) Neuere Beitr. Schmett. 6 : 1-20 (1846), : 21-82 (1848), : 83-104 (1849), : 105-146 (1850), : 147-198 (1851), : (1852)
Frivaldszky, 1835
A Balkány vidékén tett természettudományi utazásról A Magyar Tudós Társág Évkönyvei, 2: 235-276, pls 1-7
Fruhstorfer, 1910
Neue Lycaeniden des palaearktischen Gebietes Int. ent. Zs. 4 (10): 55 (4 June), (12): 62-63 (18 June)
Graves, 1925
The Rhopalocera and Grypocera of Palestine and Transjordania Trans. ent. Soc. Lond. 1925: 17-126, pl. 4-5
Grinnell, 1905
Lycaena emigdionis n. sp. Ent. News 16 (4) : 115-116
Grum-Grshimailo, 1887
Bericht über meine Reise in das östliche Buchara (Correspondez; mit 1 Karte). Nebst Anghang: Dianosen einger neuen Species in Romanoff, Mém. Lép. 3 : 357-402
Grum-Grshimailo, 1888
Novae species et varietates Rhopalocerum e Pamir Horae Soc. ent. Ross. 22 (3-4) : 303-307
Grum-Grshimailo, 1890
Le Pamir et sa faune lépidoptérologique in Romanoff, Mém. Lép. 4 : 1-576, pl. A,1-21
Grum-Grshimailo, 1891
Lepidoptera nova in Asia centrali novissime lecta et descripta Horae Soc. ent. Ross. 25 (3-4) : 445-465
Grum-Grshimailo, 1893
Lepidoptera Palearctica Nova Horae Soc. ent. Ross. 27 (1-2): 127-129, : 379-386
Hemming, 1931
Description of Three Palaearctic Butterflies Ann. Mag. Nat. Hist. (10) 8: 534-536
Hemming, 1934
Notes on two collections of the butterflies made in Palestine, with a note on teh occurrence in Transjordan of an unrecorded species Entomologist 67: 1-4, 29-32, 135-137, 196-199
Hemming, 1934
Some notes in the nomenclature of Palaearctic and African Rhopalocera Stylops 3 : 97-99
Hemming, 1934
Revisional notes on certain species of Rhopalocera (Lepidoptera) Stylops 3 : 193-200
Herrich-Schäffer, [1844]
Systematische Bearbeitung der Schmetterlinge von Europa, zugleich als Text, Revision und Supplement zu J. Hübner's Sammlung europäischer Schmetterlinge, 1843 (-1855), Die Tagfalter Syst. Bearb. Schmett. Europ. 1 (9): Title ("1843"), (1): [1-3], 4-16, (ii) pl. 1-5, f. 1-29 (1843), (2): 17-40, (ii) pl. 29-34, f. 131-156 (1843), (3): 41-56, (ii) pl. 6-12, f. 30-60, pl. 35-36, f. 157-164 (1844), (4): 57-80, (ii) pl. 13-22, f. 61-107 (1844), (5): 81-90, (ii) pl. 23-28, f. 108-130, pl. 37-41, f. 165-189 (1844), (6): 91-98, (ii) pl. 42-44, f. 190-203 (1844), (7): 99-114, (ii) pl. 45-52, f. 204-248 (1844), (9): 115-122 (1844), (10): 123-146, (ii) pl. 53-57, f. 249-274 (1845), (11): 147-162 (1845), (12): (iii) pl. 1-3, f. 1-17 (1845), (13): 163-164, (ii) pl. 58-65, f. 275-315 (1846), (14): (ii) pl. 66-70, f. 316-338, (iii) pl. 4-5, f. 18-30 (1846), (26): (ii) pl. 71-77, f. 339-376 (1847), (32): (ii) pl. 78-82, f. 377-400 (1848), (36): (ii) pl. 83-89, f. 401-426, (iii) pl. 6, f. 31-36 (1848), (37): (ii) pl. 90-91, f. 427-437 (1849), (43): (ii) pl. 92-94, f. 438-450, 452-453 (1850), (45): (ii) pl. 95-102, f. 453-489 (1850), (47): (ii) pl. 103, f. 490-495 (1851), (49): (ii) pl. 104-109, f. 496-528 (1851), (51): (ii) pl. 110-118, f. 529-570 (1851), (54): (ii) pl. 119-123, f. 571-596 (1852), (56): (ii) pl. 124-127, f. 597-618, pl. 130-132, f. 628-639, pl. 134, f. 646-651 (1852), (59): (ii) pl. 128-129, f. 619-627, pl. 133, f. 640-645 (1853), (65): (iii) pl. 7, f. 37-42, (index) [1], 2-20 (1854), (66): (index) 21-24 (1855)
Herz, 1900
Meine Lepidopteren-Ausbeute im nördlichen Buchara un im Seravschan-Gebiete im Jahre 1892 Ann. Mus. Zool. Acad. Sci. St.-Petersb. 5 (4) : 428-457
Huang, 1998
Two New Rhopalocera from Central Tibet Neue Entomologische Nachrichten, 41: 265-270
Huang, 1999
Plebejus obscurolunulata sp. n. from Tsinghai province of China Lambillionea XCIX (3): 327-328
Huang & Xue, 2004
Notes on some Chinese butterflies Neue Ent. Nachr. 57: 171-177, pl. 14
Hübner, [1806]
Tentamen determinationis digestionis alque denominationis singlarum stripium Lepidopterorum, peritisad inspiciendum et dijudicandum communicatum, a Jacob Hübner Tentamen determinationis digestionis ...: -
Hübner, [1819]
Verzeichniss bekannter Schmettlinge, 1816-[1826] Verz. bek. Schmett. (1): [1-3], 4-16 (1816), (2): 17-32 (1819), (3): 33-48 (1819), (4): 49-64 (1819), (5): 65-80 (1819), (6): 81-96 (1819), (7): 97-112 (1819), (8): 113-128 (1819), (9): 129-144 (1819), (10): 145-160 (1819), (11): 161-176 (1819), (12): 177-192 (1820), (13): 193-208 (1820), (14): 209-224 (1821), (15): 225-240 (1821), (16): 241-256 (1821), (17): 257-272 (1823), (18): 273-288 (1823), (19): 289-304 (1823), (20): 305-320 (1825), (21): 321-336 (1825), (22): 337-352 (1825), (23-27): 353-431 ([1825])
Junge, 1971
Eine neue Subspezies von Plebejus pyalon F. W. in Südtirol (Lep. Lycaenidae) NachrBl. bayerisch. Ent. 20: 33-35, 4figs
Kirby, 1871
A synonymic catalogue of the diurnal Lepidoptera Syn. Cat. diurn. Lep.: vii + 690pp : 1-690
Kluk, 1780
Historyja naturalna zwierzat domowych i dzikich, osobliwie kraiowych, historyi naturalney poczatki, i gospodarstwo : potrzebnych I pozytecznych donowych chowanie, rozmnozenie, chorob leczenie, dzikich lowienie, oswaienie : za·zycie; szkodliwych zas wygubienie. 4 vols. Hist. nat. pocz. gospod. 4: -
Kusnezov, 1908
[News Turkest. Russ. Geogr. Soc.] Izvest. Turkest. Otd. Russ. Geogr. Obsc. 4 (8): 107
Leech, 1893
Butterflies from China, Japan, and Corea Butts China Japan Corea (1): 1-297, pl. 1-28 (1892-1894), (2): 297-681, pl. 29-43 (1893-1894)
Linnaeus, 1758
Systema Naturae per Regna Tria Naturae, Secundum Clases, Ordines, Genera, Species, cum Characteribus, Differentiis, Symonymis, Locis. Tomis I. 10th Edition Syst. Nat. (Edn 10) 1 : 1-338, : 339-824
Matsumura, 1927
A list of the butterflies of Corea, with description of new species, subspecies and aberrations Insecta Matsumurana 1 (4) : 159-170
Moore, [1875]
Descriptions of new Asiatic Lepidoptera Proc. zool. Soc. Lond. 1874 (4) : 565-579, pl. 66-67
Murayama, 1963
Remarks on some butterflies from Japan and Korea, with descriptions of 2 races, 1 form, 4 aberrant forms Tyô to Ga 14 (2): 43-50
Murayama, 1964
Neue Tagfalterforme aus Japan und Korea Z. wien. ent. Ges. 49: 35-37, pl. 4-5
Murayama, 1969
Vier Tagfalter aus Nordkorea and Japan, mit der Beschreibung einer neuen Rasse [in Japanese] Tyô to Ga 20 (3-4): 67-70
Nabokov, 1949
The Nearctic members of the genus Lycaeides Hübner (Lycaenidae, Lepidoptera) Bull. Mus. Comp. Zool. 101: 479-541, 9pls
Nekrutenko, 1975
Two new subspecies of Plebejus (Plebejides) pylaon from the southern and northern sides of the West Caucasus (Lycaenidae) J. Lep. Soc. 29 (3): 151-155
Elwes in Nicholl, 1901
Butterflies of the Lebanon, with a Preface and Notes by Henry John Elwes Trans. ent. Soc. Lond. 1901 (1) : 75-97
Oberthür, 1874
Lépidoptčres nouveaux d'Algérie. 1. Diurnes Petites Nouv. Ent. 1: 412-413
Oberthür, 1910
Notes pour servir ŕ établir la Faune Française et Algérienne des Lépidoptčres Čtud. Lépid. Comp. 4 : 15-691, pl. 35-58
van der Poorten, 1984
On the occurrence of Kretania eurypilus Freyer on the peloponnesos Nota Lepid. 7 (2): 164-170
Rambur, 1840
Faune entomologique de l'Andalusie. 1837-1840 (Vol. 2) Faune ent. Andal. 2 (1): pl. 14-15, 18 (1837), 2 (2): 1-16, pl. 1-4 (1838), 2 (3): 17-96, pl. 6-7, 9, 11 (1838), 2 (4): 97-176, pl. 5, 8, 10, 12, 17 (1839), 2 (5): 177-336 (1840)
Rambur, 1858
Catalogue systematique des lépidoptčres de l'Andalousie Cat. syst. Lépid. Andalousie (1): 1-92 (1858), (2): 93-442 ([1866])
Reakirt, 1866
Coloradian butteflies Proc. Ent. Soc. Philad. 6 (1) : 122-151
Rebel, 1901
Ueber einige neue von Herrn Max Korb in Westasien gesammelte Lepidopterenformen Ann. Hofmus. Wien. 16 : 165-169
Rebel, 1911
Eine neue Lycaenidenform aus Südungarn Ent. Z. 25: 191-192
Röber, 1886
Neue Tagschmetterlinge der Indo-Australischen Fauna Corr.-Bl. Ent. Ver. Iris 1 (3) : 45-72, pl. 2-5
Sakai, 1978
Butterflies from the Hindukush, Karakorum, Kashmir and Ladak, with Descriptions of two new Species and six Subspecies Atalanta 9 (1): 104-131, 69figs
Shchurov & Lukhtanov, 2001
Notes on the taxononomy of the genus Kretania (Beuret, 1959) with the description of a new species from Black Sea Coast of Russia Atalanta 32 (1/2): 217-225
Shchetkin, 1963
[To the fauna of higher hlepidopterans of low-montane southern Tajikistan] Trudy Inst. Zool. Parasit., Stalinabad 24: 21-73
Shirozu & Sibitani, 1943
On the Japanese sepcies of the genera Plebejus Klug and Lycaeides Hübner (Lep.) Trans. Kansai. ent. Soc. 13: 25-35, figs
Staudinger, 1867
Einige neue Lepidopteren (sämmtlich aus der Sammlung des verstorbenen O. Gruner) Stettin Ent. Ztg 28 : 100-110
Staudinger, 1874
Einige neue Lepidopteren de europäischen Faunengebiets Stettin ent. Ztg 35 (1-3) : 87-98
Staudinger, 1886
Centralasiatische Lepidopteren Stett. Ent. Ztg. 47 (4-6): 193-215, (7-9): 225-256
Staudinger, 1889
Centralasiatische Lepidopteren Stettin Ent. Ztg 50 (1-3) : 16-60
Staudinger, 1892
Lepidopteren des Kentei-Gebirges Dt. Ent. Z. Iris 5 (2) : 300-393
Szabó, 1954
[The forms of Plebeius sephirus Friv. in the Central Danubian Basin] Folia ent. hung. (N.S) 7: 29-39 (in Hungarian)
Tennent, 1995
A new subspecies of Plebejus martini Allard, 1867 (Lepidoptera: Lycaenidae) from the Rif mountains of Morocco Entomologist's Gaz. 46 (2): 121-124
Tutt, 1909
A natural history of the British Lepidoptera. A text-book for students and collectors Nat. Hist. Br. Lepid. 10 : 1-410, pl. 1-53
Tytler, 1926
Notes on some new and interesting Butterflies from India and Burma. (1) & (2) J. Bombay nat. Hist. Soc. 31 : 248-260, (3): 578-590, pl. 1-5 [?]
Verity, 1921
Seasonal polymorphism and races of some European Grypocera and Rhopalocera Ent. Rec. J. Var. 33 : 170-176,190-193,210-214
Verity, 1927
La variation géographique dans l'Europe occidentale des Plebeius idas L. (argus Schiff. = argyrognomon Berg.) et insularis Leech. le nom du P. lycidas est de meigen et no de trapp. Ann. Soc. ent. Fr. 96 : 1-16
Verity, 1931
On the geographical variations and the evolution of Lycaeides argus L. Dt. Ent. Z. Iris, 45: 30-69
Verity, 1935
Races de Lysandra thersites [Lep. Cant.] Chapm. et de Plebeius sephyrus Friv. de l'Asie Mineure Bull. Soc. ent. Fr. 40 : 243-245
Wright, 1906
The Butterflies of the west coast of the United States Butts. West Coast (2nd edn.) : 1-257
Yoshino, 2003
New butterflies from China 8 Futao (43): 6-19
Züllich, 1929
Einige neue Lycaenidenformen aus meiner Sammlung Z. öst. ent. Ver. 14 (5): 51-53


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Keep in mind that the taxonomic information is copied from various sources, and may include many inaccuracies. Expert help is welcome.