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Celastrina Tutt, 1906
Celastrina Tutt, 1906; Ent. Rec. J. Var. 18 (5) : 131, TS: Papilio argiolus Linnaeus
Cyaniriodes Matsumura, 1919; Thous. Ins. Japan. Addit. 3 : 736 (preocc. de Nicéville, 1890)
= ; [NL4A, 138]
Maslowskia Kurenzov, 1974; , TS: Celastrina filipjevi Riley
= ; [NL4A, 138]
Celastrina, NSG Voucher Specimen [Wahlberg]
Celastrina, Butterflies and Moths of North America [BAMONA]
Cyaniroides[sic] ; Hemming, 1967, Bull. Br. Mus. (Ent.) Suppl. 9 : 132 (missp.)
Celastrina (Lycaenidae) ; [BRU2, 143]
Celastrina (Polyommatini) ; Eliot, 1973, Bull. Br. Mus. nat. Hist. (Ent.) 28 (6) : 449; [Opler]; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 162
Celastrina (Polyommatinae) ; [NL4A, 138]
Misidentified/misapplied:
Cyaniris; auth., not Dalman, 1816
Lycaenopsis; auth., not C & R. Felder, 1865


[sp-1v.jpg] 600x400 (~80Kb) underside India: Subansiri, 5.11.2005, Photo © Kedar Tokekar [sp-2v.jpg] 600x400 (~65Kb) underside India: Subansiri, 2.11.2005, Photo © Kedar Tokekar

The exact identification of these species is still unknown, but tentatively assumed to belong into this group.



fiCelastrina argiolus (Linnaeus, 1758)
NAF, EU, Siberia, Turkey, C.Asia, Japan. See [About maps] Paatsamasinisiipi Holly Blue Tosteblåvinge Skogsblåvinge Faulbaumfalter Najade Modraszek wieszczek
Papilio argiolus Linnaeus, 1758; Syst. Nat. (Edn 10) 1 : 483, TL: England
Papilio cleobis Sulzer, 1776; Gesch. Ins. nach linn. Syst.: pl. 18, f. 13-14
Papilio marginatus Retzius, 1783; Gen. Spec. Ins. : 30
Papilio acis Fabricius, 1787; Mantissa Insectorum 2: 73, n. 687
paraleuca (Röber, 1897)
Lycaenopsis argiolus r. britanna Verity, 1919; Ent. Rec. J. Var. 31 : 46
Lycaenopsis argiolus f. calidogenita Verity, 1919; Ent. Rec. J. Var. 31 : 46
Celastrina argiolus, NSG Voucher Specimen [Wahlberg]
Celastrina argiolus, Larvae of North-European Lepidoptera [Kimmo Silvonen]
Celastrina argiolus, Svenska fjärilar [SVF]
Celastrina argiolus, Norges Sommerfugler [NORLEP]
Celastrina argiolus, Moths and Butterflies of Europe [MBE]
Celastrina argiolus, Europaische Schmetterlinge [Mario Maier]
Celastrina argiolus, Captain's European Butterfly Guide [Simon Coombes]
Lycaenopsis argiolus ; Chapman, 1909, Proc. zool. Soc. Lond. 1909 (2) : 443; Fruhstorfer, 1917, Archiv Naturg. (A) 82 (1) : 27
Celastrina argiolus ; [BARU, #358]; [EBW]; [BOW: pl. 9, f. 13-14]; [BRU2: 143, pl. 63, f. 16-27, 31-36]
Celastrina argiola ; [MRS, 680]
Celastrina argiolus ; [NL4A, #1086]; [Otakar Kudrna]
[argiolus-1v.jpg] 500x605 (~31Kb) underside Finland, Tampere, Hervanta, 681:33, 12.5.1996, Photo © Tero Piirainen [argiolus-2mv.jpg] 533x400 (~41Kb) underside male A road at a birch grove edge near the village Mel'nichikha (Shadrikha), Novosibirsk District and Province, West Siberia, Russia. 18th May 1997, Photo © Oleg Kosterin [argiolus-3v.jpg] 1000x1186 (~160Kb) underside Russia, Udmurtia, river banks of Kama, near village Doksha east of Izhevsk, 20.8.2004, Photo © Markku Savela
[argiolus-4v.jpg] 800x600 (~115Kb) underside Russia, Kaluga area, 19.05.2005, Photo © D. Smirnov [argiolus-5m.jpg] 701x851 (~121Kb) male Russia: Kaluga area (35°45'E, 54°23'N), 19.5.2005, Photo © D. Smirnov [argiolus.jpg] 500x417 (~56Kb) Turunc, SW Turkey 19/05/92, Photo © Simon Coombes

Larva on Ribes, R. nigrum, R. rubrum, R. uva-crispa, Rhamnus frangula, R. cathartica [SPRK], Caragana frutex, Chamaecystis, Lupinus, Vaccinium [BARU], Berberis, Cuscuta engelmannii, C. lehmanniana, (C. Asia) [BRU2, 144] Sophora flavescens, Lespedeza spp., Desmodium oxyphyllum, Vicia sativa, Pueraria lobata, Wisteria floribunda, Cornus controversa, Aralia elata, Euodia meliifolia, Malus pumila, Prunus salicina, Polygonum cuspidatum, Quercus aliena, Staphylea bumalda [MRS]


EU, Caucasus, Armenia, Talysh Mts., Kazakhstan, Atai, Sayan, W.Siberia, C.Siberia Lycaenopsis argiolus argiolus ; Fruhstorfer, 1917, Archiv Naturg. (A) 82 (1) : 28
Celastrina argiolus argiolus ; [BRU2, 143]
C. a. mauretanica (Rothschild, 1925)
Morocco, Algeria, Tunisia Lycaena argiolus mauretanica Rothschild, 1925; Novit. Zool. 32 : 214 (repl. algirica Oberthür), TL: Algeria
Lycaena argiolus var. algirica Oberthür, 1915; Étud. Lépid. Comp. 10 : 401, pl. 288, f. 2343, TL: Alger (Algeria)
Celastrina argiolus mauretanica ; [BMAT: 35, pl. 11, f. 25-30]

Larva on Arbutus unedo, Rubus sp., Discolores sp., Cornus sp., Rhamnus sp., Colutea sp. [BMAT]

C. a. bieneri Forster, 1941
Yakutia, Transbaikalia, Far East, Kamchatka Celastrina bieneri Forster, 1941; Mitt. münchn. ent. Ges. 31 : 614
Celastrina argiolus bieneri ; [BARU, (note)]; [BRU2, 144]
C. a. caphis (Fruhstorfer, 1922)
Yunnan, ...? Lycaenopsis caphis Fruhstorfer, 1922; in Seitz, Gross-Schmett. Erde 9 : 871
Celastrina argiola caphis ; [MRS, 680]
Celastrina argiolus caphis ; Huang, 2003, Neue Ent. Nachr. 55: 106 (note)
C. a. sikkima (Moore, [1884])
Sikkim - Assam, Karens Hill Hedge Blue
Cyaniris sikkima Moore, [1884]; Proc. zool. Soc. Lond. 1883 (4) : 524, pl. 48, f. 11, TL: Sikkim
Lycaenopsis argiolus sikkima ; Fruhstorfer, 1917, Archiv Naturg. (A) 82 (1) : 29; [BIR, 280]

Larva on Prinsepia utilis [BIR]

?C. a. kollari (Westwood, [1852])
Chitral - Kumaon Hill Hedge Blue
Lycaena coelestina Kollar, 1848; in Hügel, Kaschmir und das Reich der Siek, 4: 423 (preocc. Lycaena coelestina Eversmann, 1843)
= Lycaenopsis argiolus ; Chapman, 1909, Proc. zool. Soc. Lond. 1909 (2) : 444
Lycaena kollari Westwood, [1852]; Gen. diurn. Lep. (2): 491 (repl. Lycaena coelestina Kollar, 1848)
Lycaenopsis argiolus coelestina ; Fruhstorfer, 1917, Archiv Naturg. (A) 82 (1) : 29
Lycaenopsis argiolus coelestina f. coelestina ; Fruhstorfer, 1917, Archiv Naturg. (A) 82 (1) : 29
Lycaenopsis argiolus kollari ; [BIR, 279(key), 280]

Larva on Prinsepia utilis [BIR]

C. a. iynteana (de Nicéville, 1884)
Nepal - Burma, Thailand, Vietnam, Tibet, S.Yunnan Jyntea Hedge Blue
Cyaniris iynteana de Nicéville, [1884]; J. Asiat. Soc. Bengal 52 Pt.II (2/4) : 69, pl. 9, f. 7 ♂, 7a ♀, TL: Sikkim
Cyaniris jynteana Moore, [1884]; Proc. zool. Soc. Lond. 1883 (4) : 524, pl. 48, f. 10, TL: Khasi Hills
= Lycaenopsis limbata ; Chapman, 1909, Proc. zool. Soc. Lond. 1909 (2) : 447
Cyaniris victoria Swinhoe, 1893; Trans. ent. Soc. Lond. 1893 (3) : 293, TL: Shillong, Assam
= Lycaenopsis argiolus ; Chapman, 1909, Proc. zool. Soc. Lond. 1909 (2) : 444
Lycaenopsis argiolus var. bothrinoides Chapman, 1909; Proc. zool. Soc. Lond. 1909 (2) : 446
Lycaenopsis argiolus var. puspargiolus Chapman, 1909; Proc. zool. Soc. Lond. 1909 (2) : 446, TL: Assam
Lycaenopsis argiolus var. albocaeruleoides Chapman, 1909; Proc. zool. Soc. Lond. 1909 (2) : 446
Cyaniris jynteana[sic] herophilus Fruhstorfer, 1910; Stettin ent. Ztg. 71 (2) : 293, TL: Vietnam
Cyaniris coelestina cition Fruhstorfer, 1910; Stettin ent. Ztg. 71 (2) : 294, TL: Vietnam
Celastrina argiolus iynteana, Butterflies in Indo-China [Yutaka Inayoshi]
Lycaenopsis argiolus herophilus ; Fruhstorfer, 1917, Archiv Naturg. (A) 82 (1) : 29
Lycaenopsis argiolus victoriae[sic, recte victoria] ; Fruhstorfer, 1917, Archiv Naturg. (A) 82 (1) : 29
Lycaenopsis jynteana ; [BIR, 280]
Celastrina argiolus iynteana ; Huang, 2001; Huang, 2003, Neue Ent. Nachr. 55: 106 (note)
C. a. hypoleuca (Kollar, [1849])
Kopet-Dagh, Tian-Shan, Ghissar-Darvaz, Pamirs-Alai Lycaena hypoleuca Kollar, [1849]; Denkschr. Akad. Wiss. Wien. 1 : 52
Celastrina argiolus hypoleuca ; [BRU2, 143]
C. a. ladonides (d'Orza, 1869)
Amur, Ussuri, Sakhalin, Kuriles, Japan Silvery Hedge Blue
Lycaena ladonides d'Orza, 1869; Lep. Jap.: 20
= Lycaenopsis argiolus ; Chapman, 1909, Proc. zool. Soc. Lond. 1909 (2) : 444
Lycaena levettii Butler, 1883; Ann. Mag. nat. Hist. (5) 11 (62) : 111
= Lycaenopsis argiolus ; Chapman, 1909, Proc. zool. Soc. Lond. 1909 (2) : 444
Celastrina sachalinensis Matsumura, 1925; J. Coll. Agric. Hokkaido imp. Univ. 15 : 105 (preocc.), TL: Sakhalin
Cyaniris heringi Kardakov, 1928; Entomol. Mitt. 17 (4): 272, TL: Primorye
Lycaenopsis manchurica Matsumura, 1939; Bull. biogeogr. Soc. Japan 9 (20): 358, TL: Manchuria
Celastrina ussuriensis Forster, 1941; Mitt. münchn. ent. Ges. 31 : 610
Celastrina ladonides, NSG Voucher Specimen [Wahlberg]

Lycaenopsis argiolus levetti ; Fruhstorfer, 1917, Archiv Naturg. (A) 82 (1) : 28
Lycaenopsis argiolus ladonides ; Fruhstorfer, 1917, Archiv Naturg. (A) 82 (1) : 28
?Lycanopsis ladonides ; [BIR, 279 (key), 280 (Murree - Kumaon)]
Celastrina heringi ; [BARU, #361]
Celastrina ladon ladonides ; [BARU, (note)]
Celastrina argiolus ladonides ; [BRU2, 144]
Celastrina ladonides ; [Oleg Kosterin]
[ladonides-1v.jpg] 802x642 (~54Kb) underside Deockcheom-ri, Gujwa-eup, Jeju-do Island, South Korea. 31st July 2002, Photo © Oleg Kosterin [ladonides-2v.jpg] 744x570 (~38Kb) underside The Shibu River (Shibukawa) shingle right bank at Hidaka village (Hidakamura), Kochi Prefecture, Shikoku, Japan, 5th August 2002, Photo © Oleg Kosterin [ladonides-3v.jpg] 638x636 (~52Kb) underside The Shibu River (Shibukawa) shingle right bank at Hidaka village (Hidakamura), Kochi Prefecture, Shikoku, Japan, 5th August 2002, Photo © Oleg Kosterin

Larva on Prinsepia utilis [BIR], Lepedeza bicolor, Vicia amoena, (Ussuri) [BRU2, 144]


C. a. crimissa (Fruhstorfer, 1917)
Formosa Lycaenopsis argiolus crimissa Fruhstorfer, 1917; Archiv Naturg. (A) 82 (1) : 28, TL: Formosa
C. a. sugurui Eliot & Kawazoé, 1983
Philippines (Luzon) Celastrina argiolus sugurui Eliot & Kawazoé, 1983; Butts. Lycaenopsis group: 215, f. 159, TL: Philippines, Luzon, 40km from Baguio, Asin hot spring

Celastrina phellodendroni Omelko, 1987
Ussuri. See [About maps] Celastrina phellodendroni Omelko, 1987; Lepid. Far East: 116, TL: [Gornotaezhnoe, Ussuri]
Celastrina phellodendroni ; [BRU2: 144, pl. 63, f. 37-42]

Larva on Phellodendron amurensis [BARU, Omelko]


Celastrina sugitanii (Matsumura, 1919)
Japan, Korea, NE.China. See [About maps] Lycaena[?] sugitanii Matsumura, 1919; Zool. Mag. Tokyo, 31: 173, TL: Honshu

Larva on (flowers of) Aesculus turbinata, Bothrocaryum controversum [BRU2, 145]

C. s. ainonica Murayama, 1952
Japan Celastrina sugitanii ainonica Murayama, 1952; Tyô to Ga 3 (1): 1, f. 2, 4-5, TL: Japan
C. s. kyushuensis Shirôzu, 1943
Japan Celastrina sugitanii kyushuensis Shirôzu, 1943; Zephyrus 9: 189
C. s. sachalinensis (Esaki, 1922)
Sakhalin Cyaniris sachalinensis Esaki, 1922; Zephyrus 9: 265
Celastrina ladon r. sachalinensis ; [BARU, (note)]
Celastrina sugitanii sachalinensis ; [BRU2, 145 (?)]
C. s. lenzeni Forster, 1941
C. s. leei Eliot & Kawazoé, 1983
South Korea Celastrina sugitanii leei Eliot & Kawazoé, 1983; Butts. Lycaenopsis group: 229, f. 446-449, TL: South Korea, Taesung, Mt Juli San
C. s. shirozui Hsu, 1987
Taiwan Celastrina sugitanii shirozui Hsu, 1987; Tyô to Ga 38 (1): 5, TL: between Pain, Taoyuan and Chihtuan, Ilan, N.Formosa

Celastrina ogasawaraensis (Pryer, 1886)
Japan. See [About maps] Lycaena ogasawaraensis Pryer, 1886; Rhop. Nihon 1: 10, pl. 11, f. 13

Celastrina ladon (Cramer, [1780])
NA, CA. See [About maps] Spring Azure
Papilio ladon Cramer, [1780]; , TL: REF#; pl. 270, f. D;N.America
Argus pseudargiolus Boisduval & Leconte, 1833; Hist. Lép. Am. Sept. (11-22): 118, pl. 36, f. 1-5
Lycaena violacea Edwards, 1866; Proc. ent. Soc. Philad. 6 (4) : 201
deutargiolus (Scudder, 1869); (summer f.)
intermedia (Strecker, 1870); (female)
Celastrina obsoletalunulata Tutt, 1908; (summer f.)
?Celastrina neglectamajor Tutt, 1908; (summer f.)
Celastrina ladon, NSG Voucher Specimen [Wahlberg]
Celastrina neglecta-major, Butterflies and Moths of North America [BAMONA]
Celastrina ladon, Georgia Lepidoptera (USA) [GALEPS]
Lycaena pseudargiolus ; Edwards, 1866, Proc. ent. Soc. Philad. 6 (4) : 204 (redescription)
?Celastrina neglecta-major ;
?Cyaniris pseudargiolus ; [BOW: pl. 19, f. 11]
Cyaniris ladon ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 45
Cyaniris ladon violacea ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 45
Celastrina ladon ; [NACL, #4363]; [BARU, #359]; [Opler]
[ladon-1vU.jpg] identification uncertain 600x440 (~43Kb) underside USA: Bridge Creek Trailhead, North Cascades Highway, (48°30'30N 120°43'00W), Chelan Co., WA, 28.7.1999, Photo © Markku Savela [ladon-2vU.jpg] identification uncertain 800x940 (~96Kb) underside USA: Miller Canyon, Sierra Vista, Cochise Co., Arizona, 8.8.2005, Photo © Markku Savela

Larva on Ceanothus, Cimicifuga racemosa [NAB], Malus, Physocarpus, Prunus, Rubus, Spiraea, Lespedeza, Lupinus, Melilotus, Ledum, Arctostaphylos, Vaccinium, Cimicifuga, Lonicera, Viburnum, Ligustrum, Humulus, Alnus, Cornus, Rhus [BARU]


Celastrina argiolus ladon ; [BNA, 397]
C. l. argentata (Fletcher, 1903)
Lycaena pseudargiolus var. argentata Fletcher, 1903; Trans. R. Soc. Canad. (2) 9 (4) : 213
Lycaena pseudargiolus var. argentata Fletcher, 1904; Can. Ent. 36 (5) : 127
Celastrina ladon argentata ; [NACL, #4363b]
C. l. nigrescens (Fletcher, 1903)
Lycaena pseudargiolus var. nigrescens Fletcher, 1903; Trans. R. Soc. Canad. (2) 9 (4) : 213
Lycaena pseudargiolus var. nigrescens Fletcher, 1904; Can. Ent. 36 (5) : 127
maculatasuffusa (Cockle, 1910); (emend.)
Celastrina ladon nigrescens, E.H. Strickland Ent. Mus, U of [ALBERTA]
Celastrina ladon nigrescens ; [NACL, #4363c]
C. l. sidara (Clench, 1944)
Colorado Lycaenopsis pseudargiolus sidara Clench, 1944; J. N.Y. ent. Soc. 52 (3): 273, TL: Manitou, Colorado
Celastrina ladon sidara ; [NACL, #4363d]
C. l. cinerea (Edwards, 1883)
Arizona Lycaena cinerea Edwards, 1883; Papilio 3 (1) : 8, TL: Arizona
Lycaena pseudargiolus var. arizonensis Edwards, 1884; Trans. Amer. Ent. Soc. 11 : 306 (summer f.), TL: Arizona
= Celastrina argiolus cinerea ; [NL4A, #1086b]
Cyaniris ladon cinerea ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 45
Cyaniris ladon arizonensis ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 45
Celastrina ladon cinerea ; [NACL, #4363e]
Celastrina argiolus cinerea ; [NL4A, #1086b]

Celastrina lucia (Kirby, 1837)
Lycaena lucia Kirby, 1837; in Richardson, Fauna Boreal Amer.: 299, pl. 3, f. 8-9
Lycaena marginata Edwards, 1883; Papilio 3 (5/6) : 86
fumida (Scudder, 1889); (nom. nud.?)
pseudora (Scudder, 1889); (nom. nud.?)
brunnea Tutt, 1908
subtusjuncta Tutt, 1908; (emend.)
inaequalis Tutt, 1908; (emend.)
Cyaniris ladon var. quesnellii Cockle, 1910; Can. Ent. 42 (6): (203-204) , TL: [Ahbau] Lake, [2950ft], 35mil NW of Quesnelle [Forks], B.C., [Canada]
= ; Guppy & Kondla, 2003, J. Lep. Soc. 57 (2): 150-152
Celastrina ladon lucia, E.H. Strickland Ent. Mus, U of [ALBERTA]
Celastrina lucia lucia, Butterflies of America [BOA]
Celastrina lucia, NSG Voucher Specimen [Wahlberg]
Celastrina lucia, Butterflies and Moths of North America [BAMONA]

quesnelli ; auth., (missp.)
quesnelii ; auth., (missp.)
Cyaniris ladon lucia ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 45
Cyaniris ladon marginata ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 45
Celastrina ladon lucia ; [NACL, #4363a]
Celastrina argiolus lucia ; [BNA, 397]
Celastrina lucia ; [Opler]

Celastrina neglecta (Edwards, 1862)
Lycaena neglecta Edwards, 1862; Proc. Acad. nat. Sci. Philad. 14: 56
Celastrina neglecta, E.H. Strickland Ent. Mus, U of [ALBERTA]
Celastrina neglecta, Butterflies of America [BOA]
Celastrina neglecta, NSG Voucher Specimen [Wahlberg]
Celastrina neglecta, Butterflies and Moths of North America [BAMONA]
Cyaniris ladon neglecta ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 45
Celastrina neglecta ; [Opler]

Celastrina echo (Edwards, 1864)
California, Oregon. See [About maps] Lycaena echo Edwards, 1864; Proc. ent. Soc. Philad. 2 (4): 506, TL: California
Cyaniris ladon ab. nunenmacheri Strand, 1915; , TL: REF#b: 159;California
= ; [NL4A, #1086a]
Lycaenopsis pseudargiolus bakeri Clench, 1944; J. N.Y. ent. Soc. 52 (3): 274, TL: Baker, Oregon
= ; [NL4A, #1086a]
Celastrina echo echo, Listing of Mexican Butterflies [ILMB]
Celastrina echo echo, Butterflies of America [BOA]
Celastrina echo, NSG Voucher Specimen [Wahlberg]
Celastrina echo, Butterflies and Moths of North America [BAMONA]
Celastrina ladon echo ; [NACL, #4363f]
Celastrina argiolus echo ; [NL4A, #1086a]
Celastrina echo ; [Opler]

Celastrina gozora (Boisduval, 1870)
Mexico, Panama, Honduras. See [About maps] Lycaena gozora Boisduval, 1870; Considérations Lépid. Guatemala: 17, TL: Guatemala (?Honduras, ?Mexico)
Celastrina gozora, Listing of Mexican Butterflies [ILMB]
Celastrina gozora, Butterflies of America [BOA]
Celastrina gozora, Butterflies and Moths of North America [BAMONA]
gazora ; Holland, 1931, (missp.)
Lycaena gozora ; Godman & Salvin, [1887], Biol. centr.-amer., Lep. Rhop. 2: 104; Godman & Salvin, [1901], Biol. centr.-amer., Lep. Rhop. 2: 722
Cyaniris ladon gozora ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 45
Celastrina ladon gozora ; [NACL, #4363g]
Celastrina argiolus gozora ; [BNA, 397]; [NL4A, #1086c]

Celastrina nigra (Forbes, 1960)
E.USA. See [About maps] Dusky Azure Spring Sooty Sooty Azure
Celastrina ebenina Clench, 1972; Ann. Carnegie Mus. 44 (4): (33-44), TL: Coalburgh, West Virginia
nig (Strecker, 1878); (ab., unavailable)
nigra (Edwards, 1887); (ab., unavailable)
Celastrina nigra, NSG Voucher Specimen [Wahlberg]
Celastrina nigra, Butterflies and Moths of North America [BAMONA]
Celastrina ebenina ; [NACL, #4364]
Celastrina nigra ; [BNA: 398, pl. 34, f. 393]; [Opler]

Larva on Aruncus dioicus [BNA]


Celastrina neglectamajor Opler & Krizek, 1984
Celastrina neglectamajor Opler & Krizek, 1984; Butts E of Great Plains: 117, pl. 18, f. 106
Celastrina neglectamajor, Butterflies of America [BOA]
Celastrina neglectamajor, NSG Voucher Specimen [Wahlberg]
C. neglectamajor, Georgia Lepidoptera (USA) [GALEPS]
Celastrina neglectamajor ; [Opler]

Celastrina humulus Scott & Wright, 1998
Celastrina humulus Scott & Wright, 1998; Papilio (N.S) 9: (1-15)
Celastrina humulus, Butterflies and Moths of North America [BAMONA]
Celastrina humulus ; [Opler]

Celastrina idella Wright & Pavulaan, 1999
Celastrina idella Wright & Pavulaan, 1999; Taxonomic Report 1 (9): (1-11)
Celastrina idella, Butterflies of America [BOA]
Celastrina idella, NSG Voucher Specimen [Wahlberg]
Celastrina idella, Butterflies and Moths of North America [BAMONA]
Celastrina idella ; [Opler]

Celastrina serotina Pavulaan & Wright, 2005
Celastrina serotina, Butterflies of America [BOA]
Celastrina serotina, NSG Voucher Specimen [Wahlberg]
Celastrina serotina, Butterflies and Moths of North America [BAMONA]

Celastrina fedoseevi Korshunov & Ivonin, 1990
Transbaikalia, Amur. See [About maps] Celastrina fedoseevi Korshunov & Ivonin, 1990; Taxonom. Ins. Helmint.: 72, TL: Amur region, near Zeya
Celastrina fedoseevi, Lepidoptera collection [SZMN]
Celastrina fedoseevi ; [BARU, #360]; [BRU2: 145, pl. 63, f. 28-30]

Celastrina gigas (Hemming, 1928)
W.Himalayas. See [About maps] Lycaenopsis gigas Hemming, 1928; Proc. Ent. Soc. Lond. 3: 29
Celastrina gigas ; [MRS, 682]
C. g. fujianensis Huang, 1994
Fujian Celastrina gigas fujianensis Huang, 1994; in Chou, Monographia Rhopalocerum Sinensium 1-2: 771, 682, f. 74, TL: Fujian

Celastrina hersilia (Leech, [1893])
Nepal - Chinal. See [About maps] Cyaniris hersilia Leech, [1893]; Butts China Japan Corea (2): 319, pl. 31, f. 16
Lycaenopsis hersilia ; Chapman, 1909, Proc. zool. Soc. Lond. 1909 (2) : 471
Celastrina hersilia ; [MRS, 681]
Lycaenopsis transpectus hersilia ; Fruhstorfer, 1917, Archiv Naturg. (A) 82 (1) : 20
C. h. evansi Toxopeus, 1927
Celastrina evansi Toxopeus, 1927; Tijdschr. Ent. 70 (3/4) : 247 n
C. h. vipia Cantile & Norman, 1960
Nepal, .. Celastrina hersilia vipia ; Morishita & Inomata, 1998, Trans. lepid. Soc. Japan 49 (1): 2

The ? species-group

Celastrina algernoni (Fruhstorfer, 1917)
-. See [About maps] Lycaenopsis oreas algernoni Fruhstorfer, 1917; Archiv Naturg. (A) 82 (1) : 30 (nom. nov)
Philippines (Luzon) Celastrina kawazoei nuydai Hayashi & Iwanaga, 1974; REF#a: 22, f. 1B-5B, 10-11, TL: Montalban, Luzon
C. a. kadazanensis Barlow, Banks & Holloway, 1971
Borneo Celastrina kawazoei Hayashi & Iwanaga, 1974; Kontyû 42 (1) : 17, f. 1-5 A, 6-8, TL: Mt. Kina-Balu, Sabah, Borneo

Celastrina huegelii (Moore, 1882)
Himalayas (Kashmir - Assam), Ceylon. See [About maps] Large Hedge Blue
Cyaniris huegelii Moore, 1882; Proc. zool. Soc. Lond. 1882 (1) : 244
= Lycaenopsis argiolus ; Chapman, 1909, Proc. zool. Soc. Lond. 1909 (2) : 444
Lycaenopsis argiolus coelestina f. huegeli ; Fruhstorfer, 1917, Archiv Naturg. (A) 82 (1) : 29
Lycaenopsis huegelii ; [BIR, 279 (key), 280]

Larva on Prinsepia utilis [BIR]

C. h. dipora (Moore, 1865)
Himalayas - Karens, Burma, N.Thailand Dusky Blue Cupid
Lycaena dipora Moore, 1865; Proc. zool. Soc. Lond. 1865 (2) : 506, pl. 31, f. 8
Everes huegelii dipora, Butterflies in Indo-China [Yutaka Inayoshi]
Everes huegelii dipora ;
Everes dipora ; [BOR, 270 (key)]

Larva on Desmodium parviflorum, Flemingia fruticulosa [BIR]

C. h. oreoides (Evans, 1925)

Celastrina lavendularis (Moore, 1877)
Dun - Assam, Burma, Bengal, S.India, Ceylon, Malaya, Philippines, Celebes. See [About maps] Plain Hedge Blue
Polyommatus lavendularis Moore, 1877; Ann. Mag. nat. Hist. (4) 20 (118) : 341, TL: Ceylon
Cyaniris lavendularis ; Moore, [1881], Lepid. Ceylon 1 (2): 75, pl. 34, f. 6, 6a, 7
Lycaenopsis lavendularis ; [BIR, 278 (key), 279]
Celastrina lavendularis ; [BOW: pl. 175, f. 16]; [MRS, 681]; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 162

Larva on Millettia, Acer, Celtis Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 163

Lycaenopsis puspa lavendularis ; Fruhstorfer, 1917, Archiv Naturg. (A) 82 (1) : 25
C. l. limbata (Moore, 1879)
N.India - Burma, Thailand, Laos, Vietnam, Yunnan Polyommatus limbatus Moore, 1879; Proc. zool. Soc. Lond. 1879 (1) : 139, TL: India
?limbata (Moore, 1877); [?]34
Cyaniris placida de Nicéville, [1884]; J. Asiat. Soc. Bengal 52 Pt.II (2/4) : 68, pl. 9, f. 8 ♂, TL: Sikkim
= Lycaenopsis limbata ; Chapman, 1909, Proc. zool. Soc. Lond. 1909 (2) : 447
Cyaniris placida hegesander Fruhstorfer, 1910; Stettin ent. Ztg. 71 (2) : 292, TL: Tonkin
Cyaniris placida pellax Fruhstorfer, 1910; Stettin ent. Ztg. 71 (2) : 292, TL: W.Java
Celastrina lavendularis limbata, Butterflies in Indo-China [Yutaka Inayoshi]
Lycaenopsis limbata ; Chapman, 1909, Proc. zool. Soc. Lond. 1909 (2) : 447
Lycaenopsis limbatus ; Fruhstorfer, 1917, Archiv Naturg. (A) 82 (1) : 31
Lycaenopsis limbatus hegesander ; Fruhstorfer, 1917, Archiv Naturg. (A) 82 (1) : 31
Lycaenopsis limbatus pellax ; Fruhstorfer, 1917, Archiv Naturg. (A) 82 (1) : 31
Celastrina lavendularis limbata ; [MRS, 681]; Huang, 2003, Neue Ent. Nachr. 55: 106 (note)
C. l. lyce (Grose-Smith, 1895)
Sulawesi Cyaniris lyce Grose-Smith, 1895; Novit. Zool. 2 (4): 506, TL: S.Celebew
= Lycaenopsis limbata ; Chapman, 1909, Proc. zool. Soc. Lond. 1909 (2) : 447
Cyaniris pellonia Fruhstorfer, 1910; Stettin ent. Ztg. 71 (2) : 301, TL: Sulawesi
= ; Cassidy, 1995, Trans. lepid. Soc. Japan 46 (2) : 56
Lycaenopsis limbatus lyce ; Fruhstorfer, 1917, Archiv Naturg. (A) 82 (1) : 32
Celastrina lavendularis lyce ; Cassidy, 1995, Trans. lepid. Soc. Japan 46 (2) : 56; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 163
C. l. himilcon (Fruhstorfer, 1909)
Taiwan Cyaniris himilcon Fruhstorfer, 1909; Ent. Zs. 23 (25) : 117, TL: Formosa, Taihanroku
Lycaenopsis limbatus himilcon ; Fruhstorfer, 1917, Archiv Naturg. (A) 82 (1) : 31
Celastrina lavendularis himilcon ; [MRS, 681]
C. l. hermesianax (Fruhstorfer, 1910)
Philippines (Mindanao) Cyaniris philippina hermesianax Fruhstorfer, 1910; Stettin ent. Ztg. 71 (2) : 300, TL: Mindanao
Lycaenopsis limbatus hermesianax ; Fruhstorfer, 1917, Archiv Naturg. (A) 82 (1) : 32
C. l. gadara (Fruhstorfer, 1910)
Obi Cyaniris gadara Fruhstorfer, 1910; Stettin ent. Ztg. 71 (2) : 301, TL: Obi
Lycaenopsis limbatus gadara ; Fruhstorfer, 1917, Archiv Naturg. (A) 82 (1) : 32
C. l. floresiana (Courvoisier, 1912)
?Cyaniris lugra f. floresiana Courvoisier, 1912; Tijdschr. Ent. 55 : 16, pl. 3, TL: Mt. Ungaran, 800-1000m
?Lycaenopsis cardia astarga f. floresiana ; Fruhstorfer, 1917, Archiv Naturg. (A) 82 (1) : 16
C. l. placidina (Fruhstorfer, 1917)
Sumatra Lycaenopsis limbatus placidina Fruhstorfer, 1917; Archiv Naturg. (A) 82 (1) : 31, TL: Sumatra
C. l. isabella Corbet, 1937
Celastrina isabella Corbet, 1937; Trans. R. ent. Soc. Lond. 86 (2): 31
Celastrina lavendularis isabella ; [BMP: 243, pl. 35, f. 38-40]
C. l. chloe Eliot & Kawazoé, 1983
Moluccas Celastrina lavendularis chloe Eliot & Kawazoé, 1983; Butts. Lycaenopsis group: 257, f. 362-363, 378-379, TL: Moluccas, central Seram
C. l. aroana Eliot & Kawazoé, 1983
New Guinea Celastrina lavendularis aroana Eliot & Kawazoé, 1983; Butts. Lycaenopsis group: 258, f. 549, TL: Papua New Guinea, upper Aroa River
Serang Celastrina limbata ssp.? ; [BAUR, 381]

Celastrina acesina (Bethune-Baker, 1906)
SE.Papua. See [About maps] Cyaniris acesina Bethune-Baker, 1906; Ann. Mag. Nat. Hist. (7) 17 (97) : 103, TL: New Guinea
Lycaenopsis acesina ; Chapman, 1909, Proc. zool. Soc. Lond. 1909 (2) : 432; Fruhstorfer, 1917, Archiv Naturg. (A) 82 (1) : 37

Celastrina morsheadi (Evans, 1915)
Tibet? Cyaniris morsheadi Evans, 1915; J. Bombay nat. Hist. Soc. 23: 543
C. m. astynome (Oberthür, 1919)

Celastrina philippina (Semper, 1889)
-. See [About maps] Philippine Hedge Blue
Cyaniris philippina Semper, 1889; Reisen Philipp. (4): 168, pl. 32, f. 14-17 (partim), TL: Philippines
Lycaenopsis phillippina[sic] ; Chapman, 1909, Proc. zool. Soc. Lond. 1909 (2) : 432
Celastrina philippina ; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 162
Borneo, Palawan, Philippines, Talaud Cyaniris philippina philippina ; Fruhstorfer, 1910, Stettin ent. Ztg. 71 (2) : 300
Lycaenopsis cardia philippina ; Fruhstorfer, 1917, Archiv Naturg. (A) 82 (1) : 14
Celastrina philippina philippina ; Hayashi & Iwanaga, 1974, Kontyû 42 (1) : 18, f. 12-13; Cassidy, 1995, Trans. lepid. Soc. Japan 46 (2) : 56; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 162
C. p. nedda (Grose-Smith, 1894)
Moluccas, Celebes, West Irian - Papua Cyaniris nedda Grose-Smith, 1894; Novit. zool. 1 (3): 572
Lycaenopsis nedda ; Chapman, 1909, Proc. zool. Soc. Lond. 1909 (2) : 429; Fruhstorfer, 1917, Archiv Naturg. (A) 82 (1) : 35
Celastrina philippina nedda ; [BOW: pl. 175, f. 19]; [BAUR, 381 (text)]
C. p. cinctuta (Grose-Smith, 1895)
Bachan, Halmahera, Ternate Cyaniris cinctuta Grose-Smith, 1895; Novit. Zool. 2 (4): 506
= Lycaenopsis nedda ; Chapman, 1909, Proc. zool. Soc. Lond. 1909 (2) : 429
Lycaenopsis nedda cinctuta ; Fruhstorfer, 1917, Archiv Naturg. (A) 82 (1) : 36
Celastrina philippina cinctuta ; [BAUR, 381]
C. p. phuste (Druce, 1895)
Timor Cyaniris phuste Druce, 1895; Proc. zool. Soc. Lond. 1895 (3) : 573, pl. 32, f. 17, TL: Dili
= Lycaenopsis nedda ; Chapman, 1909, Proc. zool. Soc. Lond. 1909 (2) : 429
Lycaenopsis nedda phuste ; Fruhstorfer, 1917, Archiv Naturg. (A) 82 (1) : 36
Celastrina philippina phuste ; [BAUR, 381 (text)]
C. p. epicharma (Fruhstorfer, 1910)
Flores Cyaniris phuste epicharma Fruhstorfer, 1910; Stettin ent. Ztg. 71 (2) : 293, TL: Flores
Lycaenopsis limbatus epicharma ; Fruhstorfer, 1917, Archiv Naturg. (A) 82 (1) : 32
Lycaenopsis philippina epicharma ; [NHM]
C. p. gradeniga (Fruhstorfer, 1910)
Sulawesi, Sula Cyaniris gradeniga Fruhstorfer, 1910; Stettin ent. Ztg. 71 (2) : 301, TL: Sulawesi
Lycaenopsis nedda proba Fruhstorfer, 1922; in Seitz, Gross-Schmett. Erde 9 : 875, TL: Sulawesi
Lycaenopsis nedda gradeniga ; Fruhstorfer, 1917, Archiv Naturg. (A) 82 (1) : 35; Fruhstorfer, 1917, Archiv Naturg. (A) 82 (1) : 35; Fruhstorfer, 1922, in Seitz, Gross-Schmett. Erde 9 : 875, pl. 152 e
Lycaenopsis philippina gradenigra[sic] ; Toxopeus, 1926, 369
Celastrina philippina gradeniga ; Cassidy, 1995, Trans. lepid. Soc. Japan 46 (2) : 56; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 162
C. p. labranda (Fruhstorfer, 1917)
Ambon, Serang, Obi Lycaenopsis nedda labranda Fruhstorfer, 1917; Archiv Naturg. (A) 82 (1) : 36, TL: Amboina, Ceram
Celastrina philippina labranda ; [BAUR, 381]
C. p. vulcanica (Rothschild, 1915)
Manam I. Lycaenopsis vulcanica Rothschild, 1915; Novit. Zool. 22 (3): 388, TL: Vulcan Is.
Lycaenopsis vulcanica ; Fruhstorfer, 1917, Archiv Naturg. (A) 82 (1) : 37
Celastrina philippina vulcanica ; [BAUR, 381 (text)]
C. p. lychorida (Fruhstorfer, 1922)
Bismarck Archipelago Lycaenopsis lychorida Fruhstorfer, 1922; in Seitz, Gross-Schmett. Erde 9 : 875
lugra Ribbe²
Celastrina philippina lychorida ; [BAUR, 381 (text)]

Celastrina perplexa Eliot & Kawazoé, 1983
Celastrina perplexa Eliot & Kawazoé, 1983; Butts. Lycaenopsis group: 242, f. 181, 342-343, 358-359, 538, TL: China, Ta-Tsien-Lou
C. p. kuroobi Yoshino, 2002

Celastrina filipjevi (Riley, 1934)
Ussuri, NE.China, Korea. See [About maps] Lycaenopsis filipjevi Riley, 1934; Entomologist 67: 85, pl. 1, f. 1-6, TL: Ussuri
Celastrina filipjevi, NSG Voucher Specimen [Wahlberg]
Maslowskia filipjevi ; [BARU, #362]
Celastrina filipjevi ; [BRU2: 145, pl. 63, f. 43-45]

Larva on Princepia chinensis [BARU, Omelko]

C. f. admirabilis (Sugitani, 1936)
Celastrina filipjevi admirabilis ;

Celastrina oreas (Leech, [1893])
Ussuri, China, Nepal, NE.India (Assam), Burma, Taiwan. See [About maps] Cyaniris oreas Leech, [1893]; Butts China Japan Corea (2): 321, pl. 31, f. 12, 15, TL: China, Ta-Chien-Lu
Maslowskia oreas, Lepidoptera collection [SZMN]
Lycaenopsis oreas ; Chapman, 1909, Proc. zool. Soc. Lond. 1909 (2) : 434; Fruhstorfer, 1917, Archiv Naturg. (A) 82 (1) : 29
Celastrina huegeli oreas ;
?Celastrina orcas[sic] ; [BOW: pl. 175, f. 25 (text)]
Maslowskia oreas ; [BARU, #363]
Celastrina oreas ; [MRS, 681]

Larva on Princepia chinensis [BARU, Omelko], Eurya strigillosa, E. acuminata, E. groffii, Prinsepia scandens [MRS]

Lycaenopsis oreas oreas ; Fruhstorfer, 1917, Archiv Naturg. (A) 82 (1) : 30
C. o. oreana (Swinhoe, 1910)
Lycaenopsis oreana Swinhoe, 1910; in Moore, Lepidoptera Indica 7 : 206, pl. 619, f. 4, TL: Khasia Hills
Lycaenopsis oreas oreana ; Fruhstorfer, 1917, Archiv Naturg. (A) 82 (1) : 30
Celastrina oreas oreana ;
C. o. arisana (Matsumura, 1910)
Taiwan Cyaniris arisanus Matsumura, 1910; Ent. Zs. 23 (50): 221, TL: Formosa
Lycaenopsis oreas arisanus ; Fruhstorfer, 1917, Archiv Naturg. (A) 82 (1) : 30
Celastrina oreas arisana ; [MRS, 681]
C. o. mirificus (Sugitani, 1936)
Korea, Ussuri Lycaenopsis mirificus Sugitani, 1936; Zephyrus 6: 167, TL: Korea
Maslowskia oreas mirificus, Lepidoptera collection [SZMN]
Celastrina oreas mirificus ;
Maslowskia oreas mirificus ; [BARU, (note)]
C. o. septentrionis Forster, 1941
C. o. hoenei Forster, 1941
C. o. baileyi Eliot & Kawazoé, 1983
SE.Tibet (Yigong) Celastrina oreas baileyi Eliot & Kawazoé, 1983; Butts. Lycaenopsis group: 236, f. 340-341, 356-357, TL: China, Tibet, Trulung
C. o. yunnana Eliot & Kawazoé, 1983
NW.Yunnan Celastrina oreas yunnana Eliot & Kawazoé, 1983; Butts. Lycaenopsis group: 237, f. 535, TL: China, Yunnan, Mekong Valley, Wei-Hsi-Ting
Celastrina oreas yunnana ; [MRS, 681]
C. o. limingani Huang, 2001
Celastrina oreas limingani Huang, 2001; Neue Entomologische Nachrichten, 51: 72, pl. 2, f. 16, TL: Gamagou, below Yela Pass, S of Bongda, E.Tibet
C. o. lixianica Sugiyama, 2004


Unknown or unplaced taxa

Lycaenopsis nedda artinia Fruhstorfer, 1917; Archiv Naturg. (A) 82 (1) : 36, TL: New Guinea


27.4.2013 (26)

References:
[ALBERTA]
University of Alberta, E.H. Strickland Entomological Museum; Browse Entomology Collection
[About maps]
Warning! The maps are automatically generated from the textual information, and the process does not always produce acceptable result; See about maps for more info.
[BAMONA]
Opler, Pavulaan, Stanford & Pogue; Butterflies and Moths of North America; Mountain Prairie Information Node
[BARU]
Korshunov & Gorbunov; Butterflies of the Asian part of Russia; English translation by Oleg Kosterin; [Dnevnye babochki aziatskoi chasti Rossii. Spravochnik]
[BAUR]
D'Abrera, 1977; Butterflies of the Australian Region (2nd edn)
[BCA]
electronic; BIOLOGIA CENTRALI-AMERICANA
[BHL]
The; Biodiversity Heritage Library; and on; Internet Archive; Note: The linking to the content of Internet Archive is somewhat delicate -- notify me when links break!
[BIR]
Wynter-Blyth, 1957; Butterflies of the Indian Region; (1982 Reprint)
[BMAT]
Tennent, 1996; The butterflies of Morocco, Algeria and Tunisia; ISBN 0 906802 05 9
[BMP]
Eliot (Ed.), Corbet & Pendlebury, 1992; The Butterflies of The Malay Peninsula; 4th edition; ISBN 983-9681-05-2
[BNA]
Scott, 1986; The Butterflies of North America
[BOA]
Interactive Listing of American Butterflies; Listado Interactivo de la Mariposas Americanas; Butterflies of America
[BOR]
D'Abrera, 1982, 1985, 1986; Butterflies of the Oriental Region, Part I-III
[BOW]
Lewis, 1974; Butterflies of the World; ISBN 0 245 52097 X
[BRU2]
Tuzov (ed.), Bogdanov, Churkin, Devyatkin, Dantchenko, Murzin, Samodurov, Zhdanko, 2000; Guide to the Butterflies of Russia and adjacent territories; Libytheidae, Danaidae, Nymphalidae, Riodinidae, Lycaenidae; Volume 2
[CiNii]
http://ci.nii.ac.jp/
[D. Smirnov]
Russia, Moscow; e-mail: D. Smirnov
[EBW]
Smart, Paul, 1976; The Illustrated Encyclopedia of the Butterfly World; ISBN 0 600 31381 6
[GALEPS]
Moths and Butterflies of Georgia and the Southeastern United States; James K. Adams; Dept. of Natural Science, Dalton State College, Dalton, GA ; http://www.daltonstate.edu/galeps/index.htm
[ILMB]
Interactive Listing of Mexican Butterflies
[Kedar Tokekar]
India
[Kimmo Silvonen]
Espoo, Finland; Kimmo's Lep Site
[LAMAS-2005]
Gerardo Lamas; Entomofauna 26 (6): 57-100; A Bibliography of the Zoological Publications of Hans Fruhstorfer (1886* - 1922+)
[LAMAS-NEO]
Gerardo Lamas; Bibliography of butterflies; An Annocated Bibliography of the Neotropical Butterflies and Skipper (Lepididoptera: Papilionoidea and Hesperioidea); Revised Electronic Edition (2007)
[MBE]
Moths and Butteflies of Europe; Paolo Mazzei, Diego Reggianti and Ilaria Pimpinelli; http://www.leps.it/
[MRS]
Chou Io (Ed.); Monographia Rhopalocerum Sinensium, 1-2
[Mario Maier]
Europäische Schmetterlinge; e-mail: Mario Maier
[NAB]
Pyle, 1981; Field Guide to North American Butterflies, National Audubon Society; ISBN 0-394-51914-0
[NACL]
Hodges, 1983; Check List of the Lepidoptera of America North of Mexico
[NHM]
Natural History Museum; The Global Lepidoptera Names Index
[NL4A]
Gerardo Lamas, 2004; Atlas of Neotropical Lepidoptera; Checklist: Part 4A; Hesperioidea - Papilionoidea
[NORLEP]
Norges Sommerfugler
[Oleg Kosterin]
Institue of Cytology & Genetics; Siberian Division of Russian Academy of Sciences; Novosibirsk; e-mail: Oleg Kosterin; http://pisum.bionet.nsc.ru/kosterin/
[Opler]
Opler & Warren, 2003 (April); Butterflies of North America. 2. Scientific Names List for Butterfly Species of North America, north of Mexico.
[SPRK]
Seppänen, E. J, 1970; Suomen suurperhostoukkien ravintokasvit, Animalia Fennica 14
[SVF]
Svenska fjärilar; Naturhistoriska riksmuseet; (Sweden)
[SZMN]
Lepidoptera; Siberian Zoological Museum
[Simon Coombes]
1 Park Street, Plymouth, PL3 4BL, UK; 01752 607854; e-mail: Simon Coombes; http://www.butterfly-guide.co.uk
[Tero Piirainen]
Tampere, Finland; e-mail: Tero Piirainen
[Wahlberg]
Niklas Wahlberg; The NSG's voucher specimen database; Nymphalidae Systematics Group
[Yutaka Inayoshi]
A Check List of Butterflies in Indo-China; Chiefly from THAILAND, LAOS & VIETNAM
[²]
This may require parentheses or not. I don't have the necessary information for this taxon.


Some related literature:
Bethune-Baker, 1906
New species of Lycaenidae from British New Guinea Ann. Mag. Nat. Hist. (7) 17 (97) : 100-104
Boisduval, 1870
Considerations sur des Lépidoptères envoyés du Guatemala à M. de l'Orza Considérations Lépid. Guatemala: 100pp
Boisduval & Leconte, 1833
Histoire générale et iconographie des lépidoptères et des chenilles de l'Amérique septentrionale Hist. Lép. Am. Sept. (1): 1-16. pl. 1-3 (1829), (2): 17-24, pl. 4-6 (1829), (3/4): 25-40, pl. 7-12 ([1830]), (5/6): 41-56, pl. 13-18 ([1830]), (7/8): 57-80, pl. 19-24 ([1830]), (9/10): 81-100, pl. 25-30 (1833), (11-22): 101-196, pl. 31-65 ([1835]), (23-26): 197-228, pl. 66-78 ([1837])
Butler, 1883
On Lepidoptera from Manchuria and the Corea Ann. Mag. nat. Hist. (5) 11 (62) : 109-117
Cassidy, 1995
On the Lycaenopsis Group of Genera (Lepidoptera, Lycaenidae) of the Sulawesi Region Trans. lepid. Soc. Japan 46 (2) : 45-62
Chapman, 1909
A review of the species of the lepidopterous genus Lycaenopsis Feld. (Cyaniris auct. nec Dalm.) on examination of the male ancilliary appendages Proc. zool. Soc. Lond. 1909 (2) : 419-476
Clench, 1944
Two new subspecies of Lycaenopsis pseudargiolus Bdv. & LeC. J. N.Y. ent. Soc. 52 (3): 273-276
Clench, 1972
Celastrina ebenina, a new species of Lycaenidae (Lepidoptera) Ann. Carnegie Mus. 44 (4): 33-44
Cockle, 1910
Notes on a few butterflies found at Kaslo and in northern British Columbia Can. Ent. 42 (6) : 203-204
Corbet, 1937
A Revision on the Malayan species of Celastrina Trans. R. ent. Soc. Lond. 86 (2): 19-33, 12 figs
Courvoisier, 1912
Javanische Lycaeniden gesammelt von Edw. Jacobson Tijdschr. Ent. 55 : 15-19, pl. 3
de Nicéville, [1884]
On new and little known Rhopalocera from the Indian Region J. Asiat. Soc. Bengal 52 Pt.II (2/4) : 65-91, pl. 1, 9-10
Westwood, [1852]
The genera of diurnal Lepidoptera, comprising their generic characters, a notice of their habitats and transformations, and a catalogue of the species of each genus; illustrated with 86 plates by W. C. Hewitson Gen. diurn. Lep. (1): 1-18, pl. A, 1-4 (1-2) (1846), (1): 19-132, pl. 4*, 5-25, 28 (3-14) (1847), (1): 133-200, pl. 1*, 26, 27, 29, 31-44 (15-23) (1848), (1): 201-242, pl. 30, 45-52, 56-58, 60-62, 64 (24-31) (1849), (1): 243-250 [?], (2): 243-326, pl. 53-55, 63, 65, 66 (32-38) (1850), (2): 327-466, pl. 59, 67-77 (39-50) (1851), (2): 467-534, 143, 144, pl. 78-80 (Index) (51-54) (1852)
Druce, 1895
A Monograph of the Bornean Lycaenidae Proc. zool. Soc. Lond. 1895 (3) : 556-627, pl. 31-34
Dyar, 1903
A List of North American Lepidoptera and Key to the Literature of this Order of Insects Bull. U.S. nat. Mus. 52 : 1-723
Eliot, 1973
The higher classification of the Lycaenidae (Lepidoptera): a tentative Arrangement Bull. Br. Mus. nat. Hist. (Ent.) 28 (6) : 373-505, 162fgs, pl. 1-6
Eliot & Kawazoé, 1983
Blue butterflies of the Lycaenopsis group. British Museum (Natural History) Butts. Lycaenopsis group: 309 pp., 560 figs
Edwards, 1862
Descriptions of certain Species of Diurnal Lepidoptera found within the limits of the United States and of British America. - No. [1]-3 Proc. Acad. nat. Sci. Philad. 13: 160-164 (1861), 14: 54-58, 221-226 (1862)
Edwards, 1864
Descriptions of certain species of Diurnal Lepidoptera, found within the limits of the United States and British America. (No. 1-3) Proc. ent. Soc. Philad. 2 (1): 14-22, pl. 1, 11 (1863), (2): 78-82 (1863), (4): 501-507 (1864)
Edwards, 1866
Descriptions of certain species of Diurnal Lepidoptera found within the limits of the United States and British America. No. 5 Proc. ent. Soc. Philad. 6 (4) : 200-208
Edwards, 1883
Notes on the collection of butterflies made by Mr. H. K. Morrison in Arizona, 1882 Papilio 3 (1) : 1-10
Edwards, 1883
On the polymorphism of Lycaena pseudargiolus Boisd. Papilio 3 (5/6) : 85-97
Edwards, 1884
Revised catalogue of the Diurnal Lepidoptera of America north of Mexico Trans. Amer. Ent. Soc. 11 : 245-337, : (Separate, Philadelphia, 1884 [1885] 95pp)
Evans, 1915
A list of butterflies caught by Capt. F. M. Bailey in S. E. Thibet during 1913 J. Bombay nat. Hist. Soc. 23: 532-546, 1pl, 1map
Fletcher, 1904
Descriptions of some new species and varieties of Canadian butterflies Can. Ent. 36 (5) : 121-129
Forster, 1941
Neue Lycaeniden-Formen aus China. 2 Mitt. münchn. ent. Ges. 31 : 593-627, pl. 19-22
Fruhstorfer, 1909
Lepidopterologisches Pêle-Mêle. [XVI]. Neues aus Formosa Ent. Zs. 23 (25) : 116-117 (18 September)
Fruhstorfer, 1910
Neue Cyaniris-Rassen und Übersicht der bekannten Arten Stettin ent. Ztg. 71 (2) : 282-305 (15 March)
Fruhstorfer, 1917
Revision der Lycaenidengattung Lycaenopsis auf Grund morphologischer Vergleiche der Klammerorgane Archiv Naturg. (A) 82 (1) : 1-42, 2 pls. (January)
Fruhstorfer, 1922
Gattungen (8-19): Parelodina - Niphanda in Seitz, Gross-Schmett. Erde 9 : 856 (14 February 1921), : 857-864 (5 January 1922), : 865-872 (21 January, 1922), : 873-880 (15 April 1922), : 881-888 (18 April 1922), : 889-896 (16 May 1922), : 897-901 (16 March 1923)
Godman & Salvin, [1887]; Godman & Salvin, [1901]
Biologia Centrali-Americana. Rhopalocera. (1887-1901) Biol. centr.-amer., Lep. Rhop. 2: 1-782, 3: pl. 1-112
Grose-Smith, 1894
An Account of a Collection of Diurnal Lepidoptera from New Guinea (Parts I-III) - made by Mr. W. Doherty at Humboldt Bay, Dutch New Guinea, and in neighbouring islands, in the Museum of the honourable Walter Rothschild at Tring, with descriptions of new species Novit. zool. 1 (2): 331-365, (2): 543-551, (3): 571-584
Grose-Smith, 1895
Descriptions of new species of butterflies, captured by Mr. Doherty in the islands of the Eastern Archipelago, and now in the Museum of the Hon. Walter Rothschild at Tring Novit. Zool. 2 (2): 75-85, (4): 505-514
Guppy & Kondla, 2003
The correct type locality of Cyaniris ladon var. quesnellii Cockle, 1910 (Lycaenidae), with designation of a lectotype J. Lep. Soc. 57 (2): 150-152
Hayashi & Iwanaga, 1974
A new species of Celastrina (Lepidoptera: Lycaenidae) from Borneo and Luzon Kontyû 42 (1) : 17-23
Hemming, 1967
Generic names of the butterflies and their type-species (Lepidoptera: Rhopalocera) Bull. Br. Mus. (Ent.) Suppl. 9 : 5-509
Hsu, 1987
A New Subspecies of Celastrina sugitanii MATSUMURA from Taiwan (Lepidoptera, Lycaenidae) Tyô to Ga 38 (1): 5-8
Huang, 2001
Report of H. Huang's 2000 Expedition to SE. Tibet for Rhopalocera Neue Entomologische Nachrichten, 51: 65-152
Huang, 2003
A list of butterflies collected from Nujiang (Lou Tse Kiang) and Dulongiang, China with descriptions of new species, and revisional notes Neue Ent. Nachr. 55: 3-114
Kardakov, 1928
Zur Kenntnis der Lepidopteren des Ussuri-Gebietes Entomol. Mitt. 17 (4): 261-273
Kirby, 1837
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Leech, [1893]
Butterflies from China, Japan, and Corea Butts China Japan Corea (1): 1-297, pl. 1-28 (1892-1894), (2): 297-681, pl. 29-43 (1893-1894)
Linnaeus, 1758
Systema Naturae per Regna Tria Naturae, Secundum Clases, Ordines, Genera, Species, cum Characteribus, Differentiis, Symonymis, Locis. Tomis I. 10th Edition Syst. Nat. (Edn 10) 1 : 1-338, : 339-824
Matsumura, 1910
Lycaeniden Japans Ent. Zs. 23 (49): 217-218, (50): 221-222
Matsumura, 1919
Thousand Insects of Japan. Additamenta 3 [Shin Nihon senchu zukai] Thous. Ins. Japan. Addit. 3 : 475-742, pl. 26-53
Matsumura, 1925
An ennumeration of the butterflies and moths from Saghalien, with descriptions of new species and subspecies J. Coll. Agric. Hokkaido imp. Univ. 15 : 83-196, pl. 8-11
Matsumura, 1939
The butterflies from Jehol (Nekka) Manchoukuo, collected by Marqis Y. Yamashina Bull. biogeogr. Soc. Japan 9 (20): 343-359, pl. XIII-XIV
Moore, 1865
List of diurnal Lepidoptera collected by Capt. A.M. Lang in the N. W. Himalayas Proc. zool. Soc. Lond. 1865 (2) : 486-509, pl. 30-31
Moore, 1877
Descriptions of Ceylon Lepidoptera Ann. Mag. nat. Hist. (4) 20 (118) : 339-348
Moore, 1879
Descriptions of new Asiatic Diurnal Lepidoptera Proc. zool. Soc. Lond. 1879 (1) : 136-144
Moore, [1881]
The Lepidoptera of Ceylon Lepid. Ceylon 1 (1): 1-40, pl. 1-18 (Dec 1880), (2): 41-80, pl. 19-36 (Jan 1881), (3): 81-136, pl. 37-54 (Jun 1881), (4): 137-190, pl. 55-71 (Dec 1881)
Moore, 1882
List of the Lepidoptera collected by the Rev. J.H. Hocking, chiefly in the Kangra Discrict, N.W Hiamalaya; with Descriptions of new Genera and Species Proc. zool. Soc. Lond. 1882 (1) : 234-263, pl. 11-12
Moore, [1884]
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Morishita & Inomata, 1998
New or little known butterflies from Nepal Trans. lepid. Soc. Japan 49 (1): 1-8
Murayama, 1952
On Celastrina sugitanii Mats. from Japan, with description of a new species and two new aberrant forms [in Japanese] Tyô to Ga 3 (1): 1-3
Oberthür, 1915
Faune des Lépidoptères de la Barbarie, Algerie Étud. Lépid. Comp. 10 : 1-459, pl. 276-277,277bis,278-315
Omelko, 1987
[Celastrina phellodendroni sp. n., a new sibling species of Celastrina argiolus L. (Lepidoptera, Lycaenidae) from the southern Primorie] Lepid. Far East: 116-122 (Russian)
Opler & Krizek, 1984
Butterflies east of the Great Plains, an illustrated natural history Butts E of Great Plains: 294pp
Retzius, 1783
Genera et Species Insectorum Gen. Spec. Ins. : 7-220
Riley, 1934
A new Palaearctic lycaenid butterfly Entomologist 67: 85-86
Rothschild, 1915
On the Lepidoptera in the Tring Museum sent by Mr. A. S. Meek from the Admiralty Islands, Dampier and Vulcan Islands Novit. Zool. 22 (2): 192-208, (3): 387-402
Rothschild, 1925
Critical List of the collection of Algerian Lepidoptera of the late Captain N. J. E. Holl. Novit. Zool. 32 : 195-229
Scott & Wright, 1998
A new Celastrina from the Front Range of Colorado Papilio (N.S) 9: 1-15
Semper, 1889
Die Schmetterlinge der Philippinischen Inseln. Reisen im Archipel der Philippinen Reisen Philipp. (1): 1-46, pl. 1-8, A (1886), (2): 47-86, pl. 9-16 (1887), (3): 87-134, pl. 17-24 (1888), (4): 135-174, pl. 25-32 (1889), (5): 175-238, pl. 33-38, B (1890), (6): 239-270, pl. 39-46 (1891), (7): 271-380, pl. 47-49 [iv] (1892)
Sulzer, 1776
Dr. Sulzers Abgekürzte Geschichte der Insecten nach dem Linaeischen System Gesch. Ins. nach linn. Syst.: -
Swinhoe, 1893
A List of the Lepidoptera of the Khasi Hills. Part I Trans. ent. Soc. Lond. 1893 (3) : 267-330
Toxopeus, 1927
Eine Revision der javanischen, zu Lycaenopsis Felder und verwandten Genera gehörigen Arten. (Mit Einführung von 6 neuen Genera, Beschreibung von 2 neuen Species un 5 neuen Subspecies). Lycaenidae Australasiae II Tijdschr. Ent. 70 (3/4) : 232-302, pl. 1, 27 text figs.
Tutt, 1906
A Study of the generic Names of the British Lycaenides and their close Allies Ent. Rec. J. Var. 18 (5) : 129-132
Vane-Wright & de Jong, 2003
The butterflies of Sulawesi: annotated checklist for a critical island faunda Zool. Verh. Leiden 343: 3-268, pl. 1-16
Verity, 1919
Seasonal polymorphism and races of some European Grypocera and Rhopalocera Ent. Rec. J. Var. 31 : 26-31,43-48,87-89,121-129,178-184,193-201
Wright & Pavulaan, 1999
Celastrina idella (Lycaenidae: Polyommatinae): a new butterfly species from the Atlantic coastal plain Taxonomic Report 1 (9): 1-11


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Keep in mind that the taxonomic information is copied from various sources, and may include many inaccuracies. Expert help is welcome.