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Anguilliformes
Muraenidae
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Gymnomuraena
Lycodontis
Gymnothorax
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gymnothorax
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gymnothorax
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Gymnothorax
Bloch, 1795
Notorabula
Whitley, 1934; (synonym)
Polyuranodon
Kaup, 1856; (synonym)
Rabula
Jordan & Davis, 1891; (synonym)
Pseudomuraena
Johnson, 1862; (synonym)
Priodonophis
Kaup, 1860; (synonym)
Serranguilla
Whitley & Phillipps, 1939; (synonym)
Thaerodontis
McClelland, 1844; (synonym)
Verdithorax
Whitley, 1931; (synonym)
Chasmenchelys
Fowler, 1944; (synonym)
Ahynnodontophis
Fowler, 1912; (synonym)
Gymnothorax afer
Muraena tristis
Kaup, 1856; (synonym)
Muraena tristis
Kaup, 1856; (unavailable, no features)
Thyrsoidea maculipinnis
Kaup, 1856; (synonym)
Thyrsoidea maculipinnis
Kaup, 1856; (unavailable, no features)
Thyrsoidea marginata
Kaup, 1860; (synonym)
Gymnothorax albimarginatus
Hatooka, 1984
Muraena albimarginata
Temminck & Schlegel, 1846; (valid as)
Gymnothorax albomaculatus
Gymnothorax albo maculatus
Bliss, 1883; (misspelling)
Gymnothorax albomaculatus
Bliss, 1883; (misspelling)
Gymnothorax angusticauda
Randall & Golani, 1995
Muraena angusticauda
Weber & de Beaufort, 1916; (valid as)
Priodonophis angusticauda
Weber & de Beaufort, 1916; (valid as)
Gymnothorax angusticeps
Priodonophis angusticeps
Hildebrand & Barton, 1949; (valid as)
Gymnothorax annasona
Francis, 1993
Gymnothorax flavimarginatus annasona
Whitley, 1937; (valid as)
Gymnothorax annulatus
Smith & Bæhlke, 1997
Gymnothorax australicola
Lavenberg, 1992
Gymnothorax bacalladoi
Bæhlke [E. B] & Brito, 1987
?Gymnothorax bathyphilus
Randall & McCosker, 1975
Gymnothorax berndti
Snyder, 1904
Gymnothorax boschii
Muraena boschii
Bleeker, 1853; (valid as)
Gymnothorax boschii
Paxton et al., 1989; (valid as)
?Gymnothorax brasiliensis
Bloch & Schneider, 1801
Gymnothorax breedeni
McCosker & Randall, 1977
Gymnothorax buroensis
Hatooka, 1984
Muraena buroensis
Bleeker, 1857; (valid as)
Muraena buro¯nsis
Bleeker, 1857; (misspelling)
Muraena atra
Ehrenberg, 1871; (unavailable, in synonymy)
Muraena corallina
Klunzinger, 1871; (synonym)
Gymnothorax marquesensis
Seale, 1906; (synonym)
Lycodontis corallinus
Dor, 1984; (synonym)
Gymnothorax castaneus
Allen & Robertson, 1994
Sidera castanea
Jordan & Gilbert, 1883; (valid as)
Gymnothorax chilospilus
Bleeker, 1865
Gymnothorax samalensis
Seale, 1910; (synonym)
Gymnothorax chlamydatus
Gymnothorax chlamydatus
Snyder, 1908; (valid, uncertain)
?Gymnothorax colubrinus
Suckow, 1799
Gymnothorax conspersus
Poey, 1867
Gymnothorax cribroris
Whitley, 1932
?Gymnothorax dakini
Whitley & Colefax, 1938
?Gymnothorax detactus
Bryan & Herre, 1903
Gymnothorax dorsalis
Seale, 1917
Gymnothorax dovii
Allen & Robertson, 1994
Muraena dovii
Gìnther, 1870; (valid as)
Gymnothorax arae
Borodin, 1928; (synonym)
?Gymnothorax echidna
Suckow, 1799
Echidna variegata
Forster, 1788; (unavailable, in synonymy)
?
Gymnothorax echidna
Forster, 1801
Echidna variegata
Forster, 1844; (unavailable, in synonymy)
Gymnothorax elegans
Bliss, 1883
Muraena flavimarginata
Kaup, 1856; (1 homonym)
Muraena flavimarginata
Kaup, 1856; (unavailable, in synonymy)
Muraena flavimarginata
Kaup, 1856; (unavailable, no features)
Gymnothorax goldsboroughi
Jordan & Evermann, 1903; (synonym)
Gymnothorax enigmaticus
McCosker & Randall, 1982
Gymnothorax equatorialis
Allen & Robertson, 1994
Priodonophis equatorialis
Hildebrand, 1946; (valid as)
Gymnothorax eurostus
Hatooka, 1984
Thyrsoidea eurosta
Abbott, 1861; (valid as)
Lycodontis parvibranchialis
Fowler, 1900; (synonym)
Muraena laysana
Steindachner, 1900; (synonym)
Gymnothorax ercodes
Jenkins, 1903; (synonym)
Gymnothorax chalazius
Waite, 1904; (synonym)
Gymnothorax gilberti
Tanaka, 1908; (synonym)
Gymnothorax dentex
de Buen, 1961; (synonym)
Gymnothorax favagineus
Bloch & Schneider, 1801
Thaerodontis reticulata
McClelland, 1844; (synonym)
Muraena python
Kaup, 1856; (synonym)
Muraena python
Kaup, 1856; (unavailable, no features)
Gymnothorax fimbriatus
Hatooka, 1984
Muraena fimbriata
Bennett, 1832; (valid as)
Muraena bullata
Richardson, 1848; (synonym)
Muraena isingleenoides
Bleeker, 1853; (synonym)
Muraena isingleeno¯des
Bleeker, 1853; (misspelling)
Muraena melanostigma
Kuhl, 1856; (unavailable, in synonymy)
Gymnothorax flavimarginata
Muraena flavimarginata
Rìppell, 1830; (valid as)
Gymnothorax flavimarginatus
Hatooka, 1984; (valid as)
Lycodontis flavimarginatus
Dor, 1984; (valid as)
Gymnothorax flavimarginata
Paxton et al., 1989; (valid as)
Gymnothorax flavimarginatus
Muraena mauritiana
Kaup, 1856; (synonym)
Muraena mauritiana
Kaup, 1856; (unavailable, no features)
Muraena batuensis
Bleeker, 1856-57; (synonym)
Gymnothorax viridipinnis
Bliss, 1883; (synonym)
Gymnothorax thalassopterus
Jenkins, 1903; (synonym)
Muraena viridipinna
Seale, 1917; (synonym)
Muraena viridipinna
Seale, 1917; (synonym, objective)
Lycodontis lemayi
Smith, 1949; (synonym)
Gymnothorax formosus
Bleeker, 1865
Gymnothorax rhodocephalus
Bleeker, 1865; (synonym)
Gymnothorax funebris
Ranzani, 1840
Gymnothorax funebris
Ranzani, 1840; (unavailable, no features)
Muraena lineo-pinnis
Richardson, 1848; (misspelling)
Muraena lineopinnis
Richardson, 1848; (synonym)
Taeniophis westphali
Kaup, 1860; (synonym)
Thyrsoidea aterrima
Kaup, 1860; (synonym)
Muraena infernalis
Poey, 1861; (synonym)
Gymnothorax fuscomaculata
Rabula fuscomaculata
Schultz, 1953; (valid as)
Gymnothorax fuscomaculatus
Castle & McCosker, 1986; (valid as)
Gymnothorax fuscomaculata
Paxton et al., 1989; (valid as)
?Gymnothorax garsiae
Whitley & Colefax, 1938
Gymnothorax gracilicauda
Jenkins, 1903
Heteromyrus atolli
Pietschmann, 1935; (synonym)
Gymnothorax hepaticus
Muraena cinerascens
Rìppell, 1830; (synonym)
Muraena hepatica
Rìppell, 1830; (valid as)
Muraena hemprichii
Klunzinger, 1871; (synonym)
Leptocephalus muraenae hepaticae
D'Ancona, 1928; (valid as)
Lycodontis hepaticus
Dor, 1984; (valid as)
Gymnothorax hepaticus
Chen et al., 1994; (valid as)
Gymnothorax herrei
Beebe & Tee-Van, 1933
Gymnothorax brunneus
Herre, 1923; (1 homonym)
Gymnothorax brunneus
Herre, 1923; (synonym)
Gymnothorax herrei
Beebe & Tee-Van, 1933; (replacement)
Gymnothorax hilonis
Gymnothorax hilonis
Jordan & Evermann, 1903; (valid, uncertain)
Gymnothorax hubbsi
Bæhlke & Bæhlke, 1977
?Gymnothorax insignis
Seale, 1917
Gymnothorax intesi
Fourmanoir & Rivaton, 1979
Lycodontis intesi
Fourmanoir & Rivaton, 1979; (valid as)
Gymnothorax isingteena
Muraena isingteena
Richardson, 1845; (valid as)
Gymnothorax javanicus
Muraena javanica
Bleeker, 1859; (valid as)
Gymnothorax javanicus
Hatooka, 1984; (valid as)
Lycodontis javanicus
Dor, 1984; (valid as)
Gymnothorax johnsoni
Castle & McCosker, 1986
Lycodontis johnsoni
Smith, 1962; (valid as)
Gymnothorax kidako
Hatooka, 1984
Muraena kidako
Temminck & Schlegel, 1846; (valid as)
Gymnothorax kolpos
Bæhlke & Bæhlke, 1980
Gymnothorax leucostigma
Jordan & Richardson, 1909
Gymnothorax longinquus
Paxton et al., 1989
Lycodontis longinquus
Whitley, 1948; (valid as)
?Gymnothorax macrosiphon
Bleeker, 1873
?Gymnothorax maculaepinnis
Bleeker, 1866
Gymnothorax maderensis
Bæhlke et al., 1989
Pseudomuraena maderensis
Johnson, 1862; (valid as)
Ahynnodontophis rosaliae
Howell Rivero, 1932; (synonym)
?Gymnothorax makassariensis
Bleeker, 1863
Gymnothorax mareei
Poll, 1953
Gymnothorax margaritophorus
Bleeker, 1865
Gymnothorax talofa
Jordan & Starks, 1906; (synonym)
Gymnothorax marshallensis
Myers, 1991
Rabula marshallensis
Schultz, 1953; (valid as)
Gymnothorax mccoskeri
Smith & Bæhlke, 1997
Gymnothorax megaspilus
Bæhlke & Randall, 1995
Gymnothorax melanospilos
Muraena melanospilos
Bleeker, 1855; (valid as)
Gymnothorax melanospilos
Hatooka, 1984; (valid as)
Muraena melanospilus
Hatooka, 1984; (misspelling)
Gymnothorax melatremus
Schultz, 1953
Gymnothorax meleagris
Muraena meleagris
Shaw, 1795; (valid as)
Thyrsoidea chlorostigma
Kaup, 1856; (synonym)
Gymnothorax leucostictus
Jenkins, 1903; (synonym)
Gymnothorax meleagris
Hatooka, 1984; (valid as)
Lycodontis meleagris
Dor, 1984; (valid as)
Gymnothorax microspila
Muraena microspila
Gìnther, 1870; (valid as)
Gymnothorax mieroszewskii
Muraena mieroszewskii
Steindachner, 1896; (valid as)
Gymnothorax miliaris
Thrysoidea miliaris
Kaup, 1856; (valid as)
Leptocephalus fuliginosus
Stræmman, 1896; (synonym)
Muraena grandimaculis
Regan, 1903; (synonym)
Muraena miliaris
Robins & Ray, 1986; (valid as)
Gymnothorax miliaris
Edwards & Glass, 1987; (valid as)
?Gymnothorax mindanoensis
Borodin, 1930
Gymnothorax minor
Smith & Bæhlke, 1997
Muraena minor
Temminck & Schlegel, 1846; (valid as)
Gymnothorax moluccensis
Randall & Golani, 1995
Priodonophis moluccensis
Bleeker, 1865; (valid as)
Gymnothorax monochrous
Hatooka, 1984
Muraena monochrous
Bleeker, 1856; (valid as)
Gymnothorax monostigma
Muraena monostigma
Regan, 1909; (valid as)
Gymnothorax mordax
Eschmeyer & Herald, 1983
Muraena mordax
Ayres, 1859; (valid as)
Rabula davisi
Fowler, 1912; (synonym)
Gymnothorax moringa
Robins & Ray, 1986
Muraena moringa
Cuvier, 1829; (valid as)
Gymnothorax rostratus
Agassiz, 1831; (synonym)
Muraena punctata
Gronow, 1854; (synonym)
Murenophis caramuru
Castelnau, 1855; (synonym)
Murenophis curvilineata
Castelnau, 1855; (synonym)
Muraena acutirostris
Abbott, 1861; (synonym)
Thyrsoidea concolor
Abbott, 1861; (synonym)
Gymnothorax flavoscriptus
Poey, 1876; (synonym)
Gymnothorax picturatus
Poey, 1880; (synonym)
Lycodontis albimentis
Evermann & Marsh, 1900; (synonym)
Gymnothorax mucifer
Snyder, 1904
Gymnothorax nasuta
de Buen, 1961
Gymnothorax neglectus
Tanaka, 1911
?Gymnothorax nigricans
Suckow, 1799
Gymnothorax nigromarginatus
Robins & Ray, 1986
Neomuraena nigromarginata
Girard, 1858; (valid as)
Gymnothorax niphostigmus
Chen, Shao & Chen, 1996
?Gymnothorax nivosus
Tanaka, 1918
Gymnothorax nubilus
Paulin, Stewart, Roberts & McMillan, 1989
Muraena nubila
Richardson, 1848; (valid as)
Gymnothorax nudivomer
Muraena nudivomer
Gìnther, 1867; (valid as)
Gymnothorax xanthostomus
Snyder, 1904; (synonym)
Gymnothorax nudivomer
Hatooka, 1984; (valid as)
Lycodontis nudivomer
Dor, 1984; (valid as)
Gymnothorax nuttingi
Snyder, 1904
Gymnothorax obesus
Gomon et al., 1994
Uropterygius obesus
Whitley, 1932; (valid as)
Gymnothorax griffini
Whitley & Phillipps, 1939; (synonym)
?Gymnothorax obscurirostris
Rendahl, 1921
Gymnothorax ocellatus
Agassiz, 1831
Muraena tricolor
Richardson (ex Banks & Solander), 1848; (unavailable, in synonymy)
Murenophis variegata
Castelnau, 1855; (synonym)
Priodonophis ocellatus
Kaup, 1860; (valid as)
Gymnothorax ocellatus
var.
januarius
Jordan & Davis, 1891; (replacement)
Gymnothorax ocellatus
var.
januarius
Jordan & Davis, 1891; (synonym)
Lycodontis jordani
Evermann & Marsh, 1899; (synonym)
Muraena ocellata
Menni, Ringuelet & Aramburu, 1984; (valid as)
Gymnothorax odiosus
Gymnothorax odiosus
Snyder, 1908; (valid, uncertain)
Gymnothorax panamensis
Allen & Robertson, 1994
Muraena panamensis
Steindachner, 1876; (valid as)
Chasmenchelys panamensis
Fowler, 1944; (synonym)
Lycodontis umbra
Fowler, 1944; (synonym)
Gymnothorax parini
Collette, Smith & Bæhlke, 1991
Gymnothorax permistus
Lycodontis permistus
Smith, 1962; (valid as)
Gymnothorax permistus
Castle & McCosker, 1986; (valid as)
Gymnothorax pescadoris
Jordan & Evermann, 1902
Gymnothorax phasmatodes
Castle & McCosker, 1986
Lycodontis phasmatodes
Smith, 1962; (valid as)
Gymnothorax philippinus
Jordan & Seale, 1907
Gymnothorax pictus
Muraena picta
Ahl, 1789; (valid as)
Muraena lita
Richardson, 1848; (synonym)
Muraena pfeifferi
Bleeker, 1853; (synonym)
Muraena polyophthalmus
Bleeker, 1858; (synonym)
Gymnothorax thomsoni
Borodin, 1928; (synonym)
Gymnothorax pictus
Hatooka, 1984; (valid as)
Siderea picta
Dor, 1984; (valid as)
?G. p. kowanensis
Aoyagi, 1943
Gymnothorax pikei
Bliss (ex Steindachner), 1883
Gymnothorax pikei
Seale, 1917; (1 HOMONYM)
Gymnothorax pindae
Smith, 1962
Gymnothorax polygonius
Poey, 1875
Gymnothorax polyuranodon
Paxton et al., 1989
Muraena polyuranodon
Bleeker, 1853; (valid as)
Uropterygius fijiensis
Fowler & Bean, 1923; (synonym)
Gymnothorax porphyreus
Francis & Randall, 1993
Muraenophis rufo-marmorato
Guichenot, 1848; (synonym)
Murenophis marmoratus
Guichenot, 1848; (synonym)
Murenophis porphyreus
Guichenot, 1848; (valid as)
Muraena chilensis
Gìnther, 1872; (synonym)
Gymnothorax wieneri
Sauvage, 1883; (synonym)
Gymnothorax prasinus
Paxton et al., 1989
Muraena prasina
Richardson, 1848; (valid as)
Gymnothorax jacksoniensis
Bleeker, 1863; (synonym)
Muraena callorhyncha
Gìnther, 1870; (synonym)
Verdithorax prasinus
Whitley, 1931; (valid as)
Notorabula callorhyncha
Whitley, 1934; (synonym)
Gymnothorax leecote
Scott, 1965; (synonym)
Gymnothorax prionodon
Ogilby, 1895
Serranguilla prionodon
Whitley & Phillipps, 1939; (valid as)
Gymnothorax prolatus
Sasaki & Amaoka, 1991
Gymnothorax pseudothyrsoideus
Hatooka, 1984
Muraena pseudothryso¯dea
Bleeker, 1852; (misspelling)
Muraena pseudothyrsoidea
Bleeker, 1852; (valid as)
?Gymnothorax punctarenae
Fowler, 1917
Gymnothorax punctatofasciatus
Bleeker, 1863
Gymnothorax punctato-fasciatus
Bleeker, 1863; (misspelling)
Gymnothorax punctatus
Gymnothorax punctatus
Bloch & Schneider, 1801; (valid as)
Lycodontis punctatus
Dor, 1984; (valid as)
Gymnothorax punctatus
Randall & Golani, 1995; (valid as)
Gymnothorax randalli
Smith & Bæhlke, 1997
Gymnothorax reevesii
Hatooka, 1984
Muraena reevesii
Richardson, 1845; (valid as)
Gymnothorax reticularis
Bloch, 1795
Muraena bicolor
Bleeker (ex Reinwardt), 1864-65; (unavailable, in synonymy)
Gymnothorax richardsonii
Castle & McCosker, 1986
Muraena ceramensis
Bleeker, 1852; (synonym)
Muraena richardsonii
Bleeker, 1852; (valid as)
Muraena scoliodon
Bleeker, 1853; (synonym)
Muraena scoliodon
Bleeker, 1853; (unavailable, no features)
Muraena troschelii
Bleeker, 1853; (synonym)
Muraena florisiana
Bleeker, 1854; (synonym)
Muraena diplodon
Peters, 1855; (synonym)
Muraena vermicularis
Peters, 1855; (synonym)
Muraena chrysops
Kaup, 1856; (synonym)
Muraena chrysops
Kaup, 1856; (unavailable, no features)
Thyrsoidea multifasciata
Kaup, 1856; (synonym)
Muraena modesta
Kaup, 1860; (synonym)
Gymnothorax richardsoni
Hatooka, 1984; (misspelling)
Gymnothorax robinsi
Bæhlke, 1997
Gymnothorax rueppelliae
Dalophis rueppelliae
McClelland, 1844; (valid as)
Dalophis rìppelliae
McClelland, 1844; (misspelling)
Muraena umbrofasciata
Rìppell, 1852; (unavailable, in synonymy)
Muraena nigrolineata
Kaup, 1856; (synonym)
Muraena nigrolineata
Kaup, 1856; (unavailable, no features)
Sidera chlevastes
Jordan & Gilbert, 1883; (synonym)
Gymnothorax leucacme
Jenkins, 1903; (synonym)
Gymnothorax waialuae
Snyder, 1904; (synonym)
Gymnothorax ruppelliae
Hatooka, 1984; (valid as)
Lycodontis rueppelli
Dor, 1984; (valid as)
Gymnothorax ruppelliae
Muraena interrupta
Kaup, 1856; (synonym)
Muraena petelli
Bleeker, 1856; (synonym)
Gymnothorax sagmacephalus
Bæhlke, 1997
Gymnothorax saxicola
Robins & Ray, 1986
Gymnothorax ocellatus
var.
saxicola
Jordan & Davis, 1891; (valid as)
Leptocephalus gillii
Facciol , 1893; (valid as)
Leptocephalus gillii
Eigenmann & Kennedy, 1902; (synonym)
Leptocephalus latus
Eigenmann & Kennedy, 1902; (synonym)
Leptocephalus strommani
Eigenmann & Kennedy, 1902; (synonym)
Leptocephalus stræmmani
Eigenmann & Kennedy, 1902; (synonym)
Gymnothorax serratidens
Priodonophis serratidens
Hildebrand & Barton, 1949; (valid as)
?Gymnothorax signifer
Bliss, 1883
?Gymnothorax sokotrensis
Kotthaus, 1968
Gymnothorax steindachneri
Jordan & Evermann, 1903
?Gymnothorax stigmanotus
Fowler, 1912
?Gymnothorax tahitensis
Seale, 1906
Gymnothorax tile
Lycodontis literata
McClelland, 1844; (synonym)
Muraena gracilis
Richardson, 1848; (synonym)
Muraena vermiculata
Richardson, 1848; (synonym)
Thyrsoidea microdon
Kaup, 1856; (synonym)
Gymnothorax borneensis
Bleeker, 1863; (synonym)
Gymnothorax borne«nsis
Bleeker, 1863; (misspelling)
Lycodontis literata
Jordan & Evermann, 1896; (synonym)
Gymnothorax unicolor
Edwards & Glass, 1987
Muraena unicolor
Olafsen & Povelsen, 1775; (valid as)
Muraenophis unicolor
Delaroche, 1809; (valid as)
Muraenophis cristini
Risso, 1810; (synonym)
Muraena monaca
Gìnther (ex Cocco), 1870; (unavailable, in synonymy)
Lycodontis unicolor
Smith & Bæhlke, 1990; (valid? as)
Gymnothorax verrilli
McCosker & Rosenblatt, 1995
Sidera verrilli
Jordan & Gilbert, 1883; (valid as)
?Gymnothorax wakanourae
Tanaka, 1908
Gymnothorax woodwardi
McCulloch, 1912
Gymnothorax wooliensis
Paxton et al., 1989
Lycodontis wooliensis
Whitley, 1968; (valid as)
Gymnothorax ypsilon
Hatooka & Randall, 1992
Gymnothorax zonipectis
Seale, 1906
Gymnothorax indong
Seale, 1910; (synonym)
24.4.2004 (130)
Additional information sources:
The taxonomic information and the basic database has been provided by;
e-mail:
Igor Sheremetyev
; This is a conversion from a database dump, and there are probably many odd things as result of automatic conversion.