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Hepialus Fabricius, 1775
Hepialus Fabricius, 1775; Syst. Ent.: 589, TS: Phalaena humuli Linnaeus
Hepiolus Illiger, 1801; Mag. Insektenk. (Illiger) 1: 138 (emend.)
= ; [NACL, 2]; [NHM]
Epialus Agassiz, 1847; Nom. zool. (Index univ.) : 140, 178 (emend.)
= ; [NHM]
Epiolus Agassiz, 1847; Nom. zool. (Index univ.) : 141, 178 (emend.)
Epialus Lederer, 1852; (emend.)
Hepialus, NSG Voucher Specimen [Wahlberg]
 Hegialus ; Fabricius, 1787, Mantissa Insectorum 2: 134 (missp.)
Trepialus ; Latreille, 1805, Hist. nat. Crust. Ins. 14: 174 (missp.)
Hepialus ; Godman & Salvin, 1887, Biol. centr.-amer., Lep. Heterocera 1: 231
Hepialus (Hepialidae) ; [NACL, 2]; [NHM]
fiHepialus humuli (Linnaeus, 1758)
EU. See [About maps] Humalaperho Ghost Moth Großer Hopfen-Wurzelbohrer Hopfenspinner Geistermotte
Phalaena (Noctua) humuli Linnaeus, 1758; Syst. Nat. (Edn 10) 1 : 508, TL: Europe
Hepialus thulensis Newman, 1865; Entomologist 2 : 162, TL: Shetlands
= ; [NHM]
hethlandica Staudinger, 1871; in Staudinger & Wocke, Cat. Lepid. eur. Faunegeb. (Edn. 2) : 60
= ; [NHM]
rosea Petersen, 1902; Beitr. Kunde Est-Liv- Kurlands: 167, TL: Estonia
= ; [NHM]
Hepialus humuli var. azuga Pfitzner, 1912; in Seitz, Gross-Schmett. Erde 2 : 434
= ; [NHM]
grandis Pfitzner, 1912; in Seitz, Gross-Schmett. Erde 2 : 434
= ; [NHM]
dannenbergi Stephan, 1923; Soc. ent. 38: 46, TL: Europe
= ; [NHM]
pusillus Stephan, 1923; Soc. ent. 38: 46, TL: Europe
= ; [NHM]
Hepialus humuli ab. ♂ rufomaculata Lempke, 1938; Tijdschr. Ent. 82 : 303, TL: Holland
= ; [NHM]
Hepialus humuli ab. roseoornata Bytinski-Salz, 1939; Ent. Rec. 51 : 81, TL: Tavistock, Great Britain
= ; [NHM]
Hepialus humuli thulensis f. uniformis Bytinski-Salz, 1939; Ent. Rec. 51 : 82, TL: Shetland
= ; [NHM]
Hepialus humuli thulensis f. albida Bytinski-Salz, 1939; Ent. Rec. 51 : 82, TL: Shetland
= ; [NHM]
faeroensis Dahl, 1954; Fródsskaparrit 3: 134, 153, TL: Faroes
= ; [NHM]
fumosa Cockayne, 1955; Ent. Gazette 6: 5, TL: England, Shetland
= ; [NHM]
radiata Cockayne, 1955; Ent. Gazette 6: 5, TL: England, Shetland
= ; [NHM]
Hepialus humuli f. ♀ postrufescens Lempke, 1961; Tijdschr. Ent. 104 (8) : 180, TL: Holland
= ; [NHM]
Hepialus humuli f. ♀ postnigrescens Lempke, 1961; Tijdschr. Ent. 104 (8) : 180, TL: Holland
= ; [NHM]
griseomaculata van Wisselingh, 1965; Ent. Berichten 25: 146, TL: Holland
= ; [NHM]
Hepialus humuli, NSG Voucher Specimen [Wahlberg]
Hepialus humuli, Svenska fjärilar [SVF]
Hepialus humuli, Moths and Butterflies of Europe [MBE]
Hepialus humuli, UK Moths [Ian Kimber]
 Hepialus humuli ; [NHM]
[humuli-1l.jpg] 698x472 (~104Kb) larva Germany, Baden-Württemberg, Ötisheim, Scharrkessel (8°49'E, 48°58'N, 320m), 01.11.2000, Photo © Karl Hofsäß [humuli-2f.jpg] 510x657 (~34Kb) female Germany, Baden-Württemberg, Schönenberg, Siedlungsbereich (8°49'E, 48°57'N, 230m), 02.06.2000, Photo © Karl Hofsäß [humuli-3ma.jpg] 1024x448 (~90Kb) male Russia, Moscow area, 17.06.2007, Photo © D. Smirnov
[humuli-3mb.jpg] 561x702 (~79Kb) male Russia, Moscow area, 17.06.2007, Photo © D. Smirnov [humuli-4f.jpg] 902x482 (~84Kb) female Russia, Moscow area, 12.06.2007, Photo © D. Smirnov [humuli-5lU.jpg] identification uncertain 857x337 (~55Kb) larva Russia, Moscow area, 30.5.2010 (36°25'E, 56°00'N), Photo © D. Smirnov
[humuli.jpg] 700x502 (~44Kb) FINLAND: Ka: Virolahti, 671:53, m 10.7.1976, f 25-1.7.1983, Markku Savela leg.

Larva on Urtica dioeca, Arctium tomentosum


The ? species-group

Placed in Hepialus based on [NHM]. Move somewhere?

Hepialus arizanus Matsumura, 1931
Japan. See [About maps] arizanus Matsumura, 1931; 6000 Illust. Insects Japan.-Empire: 1022, TL: Japan
Hepialus arizanus ; [NHM]
Hepialus dobrogensis Caradja, 1932
Romania. See [About maps] dobrogensis Caradja, 1932; Acad. Roumanine. Bull. Sci. 15 : 41, TL: Romania
Hepialus dobrogensis ; [NHM]
Hepialus ebba Bryk, 1950
Burma. See [About maps] ebba Bryk, 1950; Arkiv Zool., 42 (A) 19: 49
Hepialus ebba ; [NHM]
Hepialus malaisei Bryk, 1946
Burma. See [About maps] malaisei Bryk, 1946; Ent. Tidskr. 67: 213, TL: N.Burma
Hepialus malaisei ; [NHM]
Hepialus mlokossevitschi Romanoff, 1884
Armenia. See [About maps] Hepialus mlokossevitschi Romanoff, 1884; in Romanoff, Mém. Lépid. 1 : 91, pl. 4, f. 7, TL: Armenia
Hepialus mlokossevitschi ; [NHM]
Hepialus nebulosus Alphéraky, 1889
Tibet, .... See [About maps] Hepialus nebulosus Alphéraky, 1889; in Romanoff, Mém. lép. 5 : 85, TL: NE.Tibet
Hepialus luteus Grum-Grshimailo, 1891; Horae Soc. ent. Ross. 25 (3-4) : 463, TL: Hsi-ning, China
= ; [NHM]
Hepialus varians Staudinger, [1896]; Dt. Ent. Z. Iris 8 (2) : 302, pl. 5, f. 12, TL: Tibet
= ; [NHM]
Hepialus nebulosus ; [NHM]
Hepialus nubifer (Lederer, 1853)
Altai. See [About maps] Epialus nubifer Lederer, 1853; Verh. zool.-bot. Ver. Wien 3 : 362, pl. 2, f. 1, TL: Alai
Hepialus nubifer ; [NHM]
Hepialus okninskyi Ermolajev, 1937
okninskyi Ermolajev, 1937; Lambillionea 37: 219, TL: Krasnojarsk
Hepialus okninskyi ; [NHM]
Hepialus sinarabesca Bryk, 1942
Kansu. See [About maps] sinarabesca Bryk, 1942; Ent. Tidskr. 63 : 153, TL: Kansu
Hepialus sinarabesca ; [NHM]
Hepialus unimacula (Daniel, 1940)
Kiangsu. See [About maps] Gorgopis unimacula Daniel, 1940; Mitt. Münch. Ent. Ges. 30 : 1022, pl. 31, f. 7, 13, TL: Nanking, Kiangsu, China
Hepialus unimacula ; [NHM]

3.6.2012 (11)

References:
[About maps]
Warning! The maps are automatically generated from the textual information, and the process does not always produce acceptable result; See about maps for more info.
[BCA]
electronic; BIOLOGIA CENTRALI-AMERICANA
[BHL]
The; Biodiversity Heritage Library; and on; Internet Archive; Note: The linking to the content of Internet Archive is somewhat delicate -- notify me when links break!
[D. Smirnov]
Russia, Moscow; e-mail: D. Smirnov
[Ian Kimber]
http://www.ukmoths.force9.co.uk/
[Karl Hofsäß]
Karl Hofsaess; e-mail: Karl Hofsäß
[MBE]
Moths and Butteflies of Europe; Paolo Mazzei, Diego Reggianti and Ilaria Pimpinelli; http://www.leps.it/
[NACL]
Hodges, 1983; Check List of the Lepidoptera of America North of Mexico
[NHM]
Natural History Museum; The Global Lepidoptera Names Index
[SVF]
Svenska fjärilar; Naturhistoriska riksmuseet; (Sweden)
[Wahlberg]
Niklas Wahlberg; The NSG's voucher specimen database; Nymphalidae Systematics Group

Some related literature:
Agassiz, 1847
Nomenclatoris zoologici. Index Universalis Nom. zool. (Index univ.) : 1-393
Alphéraky, 1889
Lépidoptères rapportés du Thibet par le Général N. M. Przewalsky de son voyage de 1884-1885 in Romanoff, Mém. lép. 5 : 59-89, pl. 4
Bryk, 1942
Neue Schmetterlinge aus den Reichsmuseum in Stockholm. Nachtrag zur Lepidopteren-ausbeute der Sven Hedinschen Expedition (1927-1930) Ent. Tidskr. 63 : 147-154
Bytinski-Salz, 1939
New and little known forms of Hepialus mostly from Great Britain Ent. Rec. 51 : 81-85
Caradja, 1932
Beiträge zur Lepidoperen-fauna Grossrumänien für das Jahr 1931 Acad. Roumanine. Bull. Sci. 15 : 35-46
Daniel, 1940
Die Cossidae und Hepialidae der Ausbeute Höne (Lep. Het.) Mitt. Münch. Ent. Ges. 30 : 1004-1025, pl. 28-31
Fabricius, 1775
Systema Entomologiae, sistens Insectorum Classes, Ordines, Genera, Species, Adiectis Synonymis, Locis, Descriptionibus, Observationibus. Syst. Ent.: 832pp
Godman & Salvin, 1887
Biologia Centrali-Americana; or Contributions to the Knowledge of the Fauna of Mexico and Central America. Zoology. Lepidoptera. Heterocera Biol. centr.-amer., Lep. Heterocera 1: 1-490, 3: pl. 1-101
Grum-Grshimailo, 1891
Lepidoptera nova in Asia centrali novissime lecta et descripta Horae Soc. ent. Ross. 25 (3-4) : 445-465
Lederer, 1853
Lepidopterologisches aus Sibirien Verh. zool.-bot. Ver. Wien 3 : 351-386, pl. 1-7
Lempke, 1961
Catalogus der Nederlandse Macrolepitoptera (Achtse Supplement) Tijdschr. Ent. 104 (8) : 111-186, pl. 9-12
Linnaeus, 1758
Systema Naturae per Regna Tria Naturae, Secundum Clases, Ordines, Genera, Species, cum Characteribus, Differentiis, Symonymis, Locis. Tomis I. 10th Edition Syst. Nat. (Edn 10) 1 : 1-338, : 339-824
Newman, 1865
Singular Geographical Race of Hepialus Humuli Entomologist 2 : 162
Romanoff, 1884
Les Lépidoptères de la Transcaucasie. Première partie in Romanoff, Mém. Lépid. 1 : 1-92, pl. 1-5
Pfitzner in Seitz, 1912
Die Gross-Schmetterlinge des Palaearktischen Faunengebietes. 2. Die Palaearktischen Spinner & Schwärmer Gross-Schmett. Erde 2 : 1-479 [Macrolep.], pl. 1-56
Staudinger, [1896]
Beschreibungen neuer Lepidopteren aus Tibet Dt. Ent. Z. Iris 8 (2) : 300-343 (1896)
Staudinger in Staudinger & Wocke, 1871
Catalog der Lepidopteren des europaeischen Faunegebiets (Edn. 2): - Cat. Lepid. eur. Faunegeb. (Edn. 2) : 1-426, : 422-426 (Corrigenda et Addenda)

If you have corrections, comments or information to add into these pages, just send mail to Markku Savela
Keep in mind that the taxonomic information is copied from various sources, and may include many inaccuracies. Expert help is welcome.