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Pieris Schrank, 1801
Whites
Pieris Schrank, 1801; Fauna Boica 2 (1) : 152, 161, TS: Papilio brassicae Linnaeus
Mancipium Hübner, [1806]; Tentamen determinationis digestionis ...: [1] (invalid), TS: Papilio brassicae Linnaeus
= ; [NL4A, 114]
Danaus Oken, 1815; (unavail.)
= ; [NL4A, 114]
Ganoris Dalman, 1816; K. VetenskAcad. Handl. 1816 (1): 61, TS: Papilio brassicae Linnaeus
= ; [NL4A, 114]
Andropodum Hübner, 1822; Syst.-alph. Verz.: 2-5, 7-9, TS: Papilio brassicae Linnaeus
= ; [NL4A, 114]
Tachyptera Berge, 1842; Schmetterlingsbuch: 19, 92-105, TS: Papilio brassicae Linnaeus
= ; [NL4A, 114]
Artogeia Verity, 1947; Le Farfalle diurn. d'Italia 3: 192, 193, TS: Papilio napi Linnaeus
= ; [NL4A, 114]
Talbotia Bernardi, 1958; Rev. franc. Ent. 25: 125, TS: Mancipium naganum Moore
Pieris, NSG Voucher Specimen [Wahlberg]
Pieris, Pieridae Holarcticae [Heiner Ziegler]
Pieris ; Godman & Salvin, [1889], Biol. centr.-amer., Lep. Rhop. 2: 128; Winhard, 2000, Butterflies of the world, 10: 28
Pieris (Pierini) ; [NL4A, 114]
Pieris (Pierina) ; Pelham, 2008, J. Res. Lepid. 40: 177; [AfrL]

fiPieris brassicae (Linnaeus, 1758)
NAF, EU, Asia Minor - Middle Asia, W.Siberia, S.Siberia, Mongolia, China, Japan, Amur, Ussuri. See [About maps] Kaaliperhonen Large White Large Cabbage White Kålfjäril Großer Kohlweißling Piéride du Chou Blanca de la Col Bielinek kapustnik
Papilio brassicae Linnaeus, 1758; Syst. Nat. (Edn 10) 1 : 467, TL: Sweden
Pontia chariclea Stephens, 1827; Ill. Br. Ent. (Haustellata) 1 (1): 17, pl. 3, f. 1-2
Ganoris wollastoni Butler, 1886; Ann. Mag. nat. Hist. (5) 17 (101) : 430, TL: Madeira
?Pieris brassicae var. catoleuca Röber, 1896; Ent. Nachr. 22 (6) : 81, TL: Taurus
Pieris brassicae, Butterflies and Moths of North America [BAMONA]
Pieris brassicae, Larvae of North-European Lepidoptera [Kimmo Silvonen]
Pieris brassicae, Svenska fjärilar [SVF]
Pieris brassicae, Norges Sommerfugler [NORLEP]
Pieris brassicae, Moths and Butterflies of Europe [MBE]
Pieris brassicae, Extracts from [BRU] by [Ilya Osipov]
Pieris brassicae, Europaische Schmetterlinge [Mario Maier]
Pieris brassicae ; [BRU, 169]; [EBW]; [BOW: pl. 4, f. 16]; [MRS, 257]; [NL4A, #270]; [AfrL]; [Otakar Kudrna]
[brassicae-1v.jpg] 450x353 (~31Kb) underside Ansteys Cove, Devon, UK 07/08/95, Photo © Simon Coombes [brassicae-2f.jpg] 600x763 (~41Kb) female FINLAND: Ka: Virolahti, 671:53, 7.8.1999, Photo © Markku Savela [brassicae-2fv.jpg] 900x704 (~57Kb) underside female FINLAND: Ka: Virolahti, 671:53, 7.8.1999, Photo © Markku Savela
[brassicae-3l.jpg] 400x596 (~42Kb) larva Central Europe 1990-2000, Photo © Michael Borsch [brassicae-3v.jpg] 593x400 (~35Kb) underside Central Europe 1990-2000, Photo © Michael Borsch [brassicae-4l.jpg] 700x445 (~107Kb) larva Russia, Moscow, 10.7 - 15.8.2004, Photo © D. Smirnov
[brassicae-5v.jpg] 800x951 (~81Kb) underside Russia, Tatarstan, Varkljed-Bodja (56°13'N 52°49'E), about 20km S of Agriz, 7.8.2004, Photo © Markku Savela [brassicae-6.jpg] 783x1044 (~94Kb) Germany, Rügen, Baabe, 26.7.2005 (on Eryngium maritimum), Photo © Harri Arkkio [brassicae-7uf.jpg] 906x542 (~103Kb) upperside female Russia: Moscow area (36°25'E, 56°00'N), 06.08.2007, Photo © D. Smirnov

Larva on Cruciferae, Cakile maritima, Isatis tinctoria, Brassica rapa, Brassica napus v. napobrassica, B. oleracea v. botrytis, B. oleracea v. capitata, Raphanus sativus, Crambe maritima, Armoracia rusticana, Capsella bursa-pastoris, Berteroa incana, Hesperis matronalis, Erysimum hieraciifolium, Barbarea stricta, Draba incana, Bunias orientalis, Reseda odorata, Nicotiana tabacum, Tropaeolum majus, T. Peregrinum [SPRK], Brassica spp., Lepidium spp., Tropaeolum spp. [MRS]


Pieris brassicae-vazquezi Oberthür, 1914; Etud. Lépid. Comp. 9 (2) : 89, pl. 264, f. 2207, TL: Spain
Pieris brassicae gen. vern. var. chariclea f. venata Verity, 1908; Rhopalocera Palaearctica 1: 164, pl. 35, f. 24-25, TL: Morocco
Pieris brassicae verna Zeller, 1847; Isis von Oken 1847 (3): 220, TL: Italy
Pieris brassicae verna gen. aest. aestiva Zerny, 1935; Mém. Soc. Sci. nat. Maroc. 52 (?42): 25, TL: Morocco
Pieris brassicae brassicae ; [BMAT, 7, pl. 2, f. 14-20]; Winhard, 2000, Butterflies of the world, 10: 28, pl. 45, f. 1

Larva on Sisymbrium sp., Lunaria biennis, Tropaeolum majus, Capparis spinosa, Raphanus raphanistrum, Isatis tinctoria, Brassica sp. [BMAT]


venata
[venata-1v.jpg] 600x435 (~37Kb) underside Morocco, Itzer, Moyen Atlas, Photo © Jean Delacre

P. b. ottonis Röber, [1907]
Turan, Tian-Shan, Ghissar, Darvaz, Alai, W.Pamirs Pieris brassicae ottonis Röber, [1907]; in Seitz, Grossschmett. Erde 1 : 45 (repl. brassicoides Staudinger)
brassicae brassicoides Staudinger, 1901; in Staudinger & Rebel, Cat. Lep. palaearct. Faunengeb. 1: (praeocc.)
P. b. nepalensis Gray, 1846
NW.India - N.Thailand, Laos, N.Vietnam, Yunnan Pieris brassicae var. nepalensis Gray, 1846; Descr. lep. Ins. Nepal : 9, pl. 6, f. 1, 3
?brassicae nepalensis Doubleday, [1845]; List. lepid. Ins. Brit. Mus. 1 : 32
Pieris brassicae nepalensis, Butterflies in Indo-China [Yutaka Inayoshi]
Pieris brassicae nepalensis ; [MRS, 257]; Winhard, 2000, Butterflies of the world, 10: 28, pl. 45, f. 2; Huang, 2001

Pieris cheiranthi (Hübner, [1823])
Catophaga cheiranthi Hübner, [1823]; Samml. exot. Schmett. 2 : pl. [124]
Pieris cheiranthi, Moths and Butterflies of Europe [MBE]
Pieris cheiranthi, Captain's European Butterfly Guide [Simon Coombes]
Pieris cheiranthi ; Winhard, 2000, Butterflies of the world, 10: 28, pl. 45, f. 4-5, back f. 12; [Otakar Kudrna]
P. c. benchoavensis Pinker, 1968

Pieris naganum (Moore, 1884)
Naga Hills. See [About maps] Naga White
Mancipium naganum Moore, 1884; J. asiat. Soc. Bengal 53 Pt.II (1) : 45, TL: Naga Hills, Assam
Mancipium nabanum[sic?] ; Hemming, 1967, Bull. Br. Mus. (Ent.) Suppl. 9 : 428 (missp.?)
Pieris naganum ; [BIR, 434]; [EBW]
Artogeia naganum ; [BOR, 148]
Talbotia naganum ; [MRS, 261]
Assam, N.Burma Artogeia naganum naganum
Artogeia naganum naganum ; [BOR, 148]
P. n. cisseis Leech, 1890
China (Houpe, Tsekiang, Honan, Fukien, Guangxi, Guangdong, Hainan) Pieris cisseis Leech, 1890; Entomologist 23 : 192, TL: Chang-Yang
Talbotia naganum cisseis ;
P. n. karumii Ikeda, 1937
Taiwan Pieris naganum karumii Ikeda, 1937; Zephyrus 7 (1) : 37
Artogeia naganum karumii ; [BOR, 148]
Talbotia naganum karumii ; [MRS, 261]
Pieris naganum karumii ; Winhard, 2000, Butterflies of the world, 10: 29, pl. 46, f. 16

Larva on Bretschneidera sinensis [MRS]

P. n. pamsi Vitalis de Salvaza²
N.Vietnam, Laos ?Pieris naganum pamsi ?Salvin, 1921; ?Faun. Ent. Indochine (3): 18, TL: N.Vietnam
Talbotia naganum pamsi, Butterflies in Indo-China [Yutaka Inayoshi]
Talbotia naganum pamsi ;

Pieris deota (de Nicéville, [1884])
Kashmir, Pamir, Karakorum, Himalaya. See [About maps] Mancipium deota de Nicéville, [1884]; J. Asiat. Soc. Bengal 52 Pt.II (2/4) : 82, pl. 9, f. 10 ♂, TL: Ladak
Pieris deota; [BRU, 169]
Pieris deota ; [MRS, 257]

Pieris tadjika Grum-Grshimailo, 1888
Turkestan (Darwaz). See [About maps] Pieris tadjika Grum-Grshimailo, 1888; Horae Soc. ent. Ross. 22 (3-4) : 304, TL: Darwaz, Karategin, 7000-10000
Pieris tadjika, Extracts from [BRU] by [Ilya Osipov]

The "napi" species-group

Lots of confusion about which of these are "real species" and which not!


fiPieris napi (Linnaeus, 1758)
NAF, EU, AM, AC, AN, AN. See [About maps] Lanttuperhonen Green-veined White Weined White Mustard White Rapsfjäril Heckenweißling Grünader-Weißling Piéride du Navet Blanca verdinervada Bielinek bytomkowiec
Papilio napi Linnaeus, 1758; Syst. Nat. (Edn 10) 1 : 468, TL: Sweden
?Pieris napi ab. impunctata Röber, 1907; in Seitz, Grossschmett. Erde 1 : 48
Pieris napi, NSG Voucher Specimen [Wahlberg]
Pieris napi, Larvae of North-European Lepidoptera [Kimmo Silvonen]
Pieris napi, Svenska fjärilar [SVF]
Pieris napi, Norges Sommerfugler [NORLEP]
Pieris napi, Moths and Butterflies of Europe [MBE]
Pieris napi, Extracts from [BRU] by [Ilya Osipov]
Pieris napi, Europaische Schmetterlinge [Mario Maier]
napae ; Edwards, 1881, (missp.)
Pieris napi ; [BRU, 164]; [EBW]; [BOW: pl. 5, f. 4]; [MRS, 258]; [NL4A, #271]; [Otakar Kudrna]
Pieris napi f. funebris ; Winhard, 2000, Butterflies of the world, 10: 29, pl. 45, f. 11
Misidentified/misapplied:
dubiosa; auct.
Pontia napi; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 6
Artogeia napi; [NACL, #4195]
Artogeia napi; [NAB, 358]
[napi-1u.jpg] 500x380 (~46Kb) upperside Luckett, Cornwall, UK 04/06/95, Photo © Simon Coombes [napi-1v.jpg] 500x416 (~46Kb) underside Bois de Caurel, Brittany, France 20/7/96, Photo © Simon Coombes [napi-2f.jpg] 550x527 (~33Kb) female FINLAND: Ka: Virolahti, 671:53, 18.7.1997, Photo © Markku Savela
[napi-3v.jpg] 512x768 (~57Kb) underside Germany, Baden-Wuerttemberg, Oetisheim, 8° 49' E, 48° 58' N, altitude 300m, 14.04.1998 (on Salix sp.), Photo © Karl Hofsäß [napi-4mfv.jpg] 741x565 (~58Kb) underside male female Vienna, Austria, 12 June 2000, Photo © Erich Mangl [napi-5l.jpg] 597x400 (~40Kb) larva Central Europe 1990-2000, Photo © Michael Borsch
[napi-5v.jpg] 596x400 (~31Kb) underside Central Europe 1990-2000, Photo © Michael Borsch [napi-6mu.jpg] 900x654 (~55Kb) upperside male FINLAND: Ka: Virolahti, Virojoki 671:53, 29.7.2000, Photo © Markku Savela [napi-6mv.jpg] 900x696 (~61Kb) underside male FINLAND: Ka: Virolahti, Virojoki 671:53, 29.7.2000, Photo © Markku Savela
[napi-7mfv.jpg] 700x945 (~79Kb) underside male female Belgium, Antwerpen, Scheldelaan, 23.8.2003 (2nd gen.), Photo © Toon Verbruggen [napi-8fv.jpg] 1200x1050 (~157Kb) underside female Russia, Tatarstan, Varkljed-Bodja (56°13'N 52°49'E), about 20km S of Agriz, 7.8.2004, Photo © Markku Savela [napi-9fu.jpg] 1300x1020 (~185Kb) upperside female Russia, Tatarstan, Varkljed-Bodja (56°13'N 52°49'E), about 20km S of Agriz, 7.8.2004, Photo © Markku Savela
[napi-a1m.jpg] 1024x665 (~107Kb) male Russia: Moscow area (36°25'E, 56°00'N), 11.5.2009, Photo © D. Smirnov [napi.jpg] 450x387 (~20Kb) FINLAND: N: Espoo, Tapiola, June 1996, Photo © Markku Savela

Larva on Cruciferae, Thlaspi arvense, T. alpestre, Alliaria petiolata, Brassica campestris, B. rapa, Brassica napus v. napobrassica, B. oleracea, B. oleracea v. botrytis, Raphanus raphanistrum, B. sativus, Armoracia rusticana, Rorippa islandica, Cardamine amara, Arabis alpina, Hesperis matronalis, Berteroa incana, Reseda odorata, Tropaeolum majus, Calendula officinalis [SPRK], Cardamine leucantha, C. niponica, Rorippa isbandica, R. sylvestris [MRS]


EU Papilio napaeae Esper, [1804]; Die Schmett., Suppl. Th 1 (10): 119, pl. 116, f. 5
Pieris napi var. intermedia Krulikovsky, 1890; Bull. Soc. imp. Nat. Moscou (n.s.) 4 (2) : 211, pl. 8, f. a
Pieris napi napi ; Winhard, 2000, Butterflies of the world, 10: 29, pl. 45, f. 17, 21
P. n. lusitanica de Sousa, 1929
Portugal Pieris napi lusitanica de Sousa, 1929; Lep. Portug. Porto: 2
P. n. britannica Verity, 1911
Ireland.British Isles Pieris napi britannica Verity, 1911; Rhopalocera Palaearctica 1: 332, pl. 32, f. 4, 6
P. n. lappona Rangnow, 1935
N.Sweden, N.Finland Pieris napi lappona Rangnow, 1935; Ent. Rdsch. 52: 175, TL: Lappland
bicolorata Petersen, 1947; Zool. Bidr. Uppsala 26: 361
Pieris napi lappona ; Hensle, 2001, Atalanta 32 (1/2): 89-95
P. n. adalwinda Fruhstorfer, 1909
N.Europe Pieris napi adalwinda Fruhstorfer, 1909; Int. ent. Zs. 3 (16): 88
Pieris bryoniae adalwinda ; [BRU]
P. n. keskuelai Eitschberger, 2001
Polar Urals Pieris napi keskuelai Eitschberger, 2001; Atalanta 32 (1/2): 86, pl. I, f. 1-8, TL: Polar Ural, railway station
P. n. maura Verity, 1911
Algeria - W.Tunisia Pieris napi napi maura Verity, 1911; Rhopalocera Palaearctica 1: 332, pl. 59, f. 18-22, TL: Blida (Algeria)
Pieris napi-maura gen. aest. napaeae f. blidana Holl, 1914; Bull. Soc. Hist. nat. Afr. N. 6: 37, TL: Blida-les-Glaciéres, Tala Rana (Algeria)
Pieris napi atlantica Rothschild, 1917; Novit. Zool. 24 (1): 75, TL: Blida-les-Glaciéres and Kabylie (Algeria)
Pieris napi maura f. temp. mauretaniae Rothschild, 1925; Novit. Zool. 32 : 197, TL: Blida-les-Glaciéres (Algeria)
Pieris napi maura ; [BMAT, 9, pl. 3, f. 16-28]
P. n. atlantis Oberthür, 1925
Morocco Pieris napi napaeae atlantis Oberthür, 1925; Etud. Lépid. Comp. 22 (2): 58, pl. 593, f. 5014-5016, TL: Toumliline [Azrou] (Morocco)
Pieris napi atlantis ; [BMAT: 9, pl. 3, f. 29-32]
P. n. segonzaci Le Cerf, 1923
Morocco Pieris napi segonzaci Le Cerf, 1923; Bull. Soc. ent. Fr. 1923 (15) : 197, TL: Ourika, Reraïa (Morocco)
Pieris napi-segonzaci f.indiv. deflava Le Cerf, 1923; Bull. Soc. ent. Fr. 1923 (15) : 197
Pieris napi segonzaci; [BMAT: 9, pl. 4, f. 1-8], TL: Reraïa (Morocco)
[segonzaci-1u.jpg] 600x435 (~29Kb) upperside Morocco, Djebel Siroua, Anti-Atlas, Photo © Michel Tarrier [segonzaci-2v.jpg] 600x435 (~22Kb) underside Morocco, Djebel Siroua, Anti-Atlas, Photo © Michel Tarrier

Larva on Ptilotrichum spinosum [BMAT]


P. n. muchei Eitschberger, [1984]
Tian-Shan, Ghissar, S.Ghissar, Darvaz, Alai, Dzhungarsky Alatau
P. n. japonica Shirôzu, 1953
Japan Pieris napi japonica Shirôzu, 1953; Sieboldia 1 (1952) : 18
P. n. napoleon Eitschberger, 1990
P. n. meridionalis Heyne, 1895
Pieris napi var. meridionalis Heyne, [1895]; in Rühl, Die palaearktischen Grossschmetterlinge und ihre Naturgeschichte 1 (Tagfalter): 714
Sikkim - Burma ?napi r. montana ; [BIR, 432]

= Pieris melete ssp. montana Verity, 1908?

Chumbi Valley ?napi r. melaena ; [BIR, 432]

Pieris ajaka Moore, 1865
Kashmir. See [About maps] Pieris ajaka Moore, 1865; Proc. zool. Soc. Lond. 1865 (2) : 490, pl. 31, f. 16, TL: Lower Kunawur
napi r. ajaka ; [BIR, 432]
Pieris ajaka ; Winhard, 2000, Butterflies of the world, 10: 29, pl. 46, f. 10

Pieris oleracea (Harris, 1829)
-. See [About maps] Mustard White
Pontia oleracea Harris, 1829; New England Farmer 7 (51): 402, TL: New Hampshire; Massachusetts
Pieris oleracea, E.H. Strickland Ent. Mus, U of [ALBERTA]
Pieris oleracea, Butterflies and Moths of North America [BAMONA]
Pieris oleracea ; [Opler]; Pelham, 2008, J. Res. Lepid. 40: 181
Pieris cruciferarum Boisduval, 1836; Hist. nat. Ins., Spec. gén. Lépid. 1 : 519
= ; Pelham, 2008, J. Res. Lepid. 40: 181
Pontia casta Kirby, 1837; in Richardson, Fauna Boreal Amer.: 288, pl.3, f. 1
= ; Pelham, 2008, J. Res. Lepid. 40: 181
oleracea hyemalis Edwards, 1881; Papilio 1 (6) : 90, 98, pl. 2, f. 8
= ; Pelham, 2008, J. Res. Lepid. 40: 181
aestiva Edwards, 1881; Papilio 1 (6) : 93, 98, pl. 3, f. 15-16
= ; Pelham, 2008, J. Res. Lepid. 40: 181
Pieris oleracea oleracea gen.aest. postoleracea Eitschberger, [1984]; Herbipoliana 1 (1-2): 263
= ; Pelham, 2008, J. Res. Lepid. 40: 181
Pieris cruciferarum ; [EBW]; [BOW: pl. 17, f. 36]
Pontia napi cruciferarum ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 6
Pontia napi oleracea ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 6
Artogeia napi oleracea ; [NACL, #4195d]
Pieris oleracea oleracea ; Pelham, 2008, J. Res. Lepid. 40: 181
P. o. frigida Scudder, 1861
Labrador Pieris frigida Scudder, 1861; Proc. Boston Soc. Nat. Hist. 8 : 181, TL: Caribou Island, Stratis of Belle Isle
Ganoris oleracea var. borealis Grote, 1873; Bull. Buffalo Soc. nat. Sci. 1 : 185, TL: Island of Anticosti
= ; Pelham, 2008, J. Res. Lepid. 40: 182
acadica Edwards, 1881; Papilio 1 (6) : 87, TL: Southern Newfoundland
= ; Pelham, 2008, J. Res. Lepid. 40: 182
Pieris oleracea ekisi Eitschberger, [1984]; Herbipoliana 1 (1-2): 272, TL: Newfoundland
= ; Pelham, 2008, J. Res. Lepid. 40: 182
Pieris napi var. frigida f. pseudoleracea Verity, [1908]; Rhopalocera Palaearctica 1: 146, TL: Labrador
= ; Pelham, 2008, J. Res. Lepid. 40: 182
Pieris napi var. frigida f. vivida Verity, 1911; Rhopalocera Palaearctica 1: 334, TL: Labrador
= ; Pelham, 2008, J. Res. Lepid. 40: 182
Pontia napi borealis ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 6
Pontia napi frigida ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 6
Pontia napi acadica ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 6
Artogeia napi frigida ; [NACL, #4195c]
Pieris oleracea frigida ; Pelham, 2008, J. Res. Lepid. 40: 182

Pieris marginalis Scudder, 1861
Yukon - British Columbia - California, Rocky Mountains. See [About maps] Margined White
Pieris marginalis Scudder, 1861; Proc. Boston Soc. Nat. Hist. 8 : 183, TL: "Gulf of Georgia"
Pieris marginalis, E.H. Strickland Ent. Mus, U of [ALBERTA]
Pieris marginalis marginalis, Butterflies of America [BOA]
Pieris marginalis, NSG Voucher Specimen [Wahlberg]
Pieris marginalis, Butterflies and Moths of North America [BAMONA]
Artogeia napi marginalis ; [NACL, #4195f]
Pieris marginalis ; [BOC, 138]; [Opler]; Pelham, 2008, J. Res. Lepid. 40: 178
[marginalis-1fu.jpg] 1000x791 (~61Kb) upperside female USA: Wallace Falls State Park (47°53'35N 121°40'53W), Snohomish Co., WA, 13.7.1999, Photo © Markku Savela [marginalis-1fux.jpg] 520x617 (~26Kb) upperside female USA: Wallace Falls State Park (47°53'35N 121°40'53W), Snohomish Co., WA, 13.7.1999, Photo © Markku Savela [marginalis-1fv.jpg] 400x502 (~17Kb) underside female USA: Wallace Falls State Park (47°53'35N 121°40'53W), Snohomish Co., WA, 13.7.1999, Photo © Markku Savela

Larva on Dentaria angulata, D. pulcherrima, D. breweri, Rorippa nasturtium-aquaticum [BOC]


Washington Pieris pallida Scudder, 1861; Proc. Boston Soc. Nat. Hist. 8 : 183, TL: "Gulf of Georgia"
= ; Pelham, 2008, J. Res. Lepid. 40: 179
Pieris oleracea ab. flava Edwards, 1881; Papilio 1 (6) : 94
= ; Pelham, 2008, J. Res. Lepid. 40: 179 (nom. nud.)
Pontia napi pallida ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 6
Pieris marginalis marginalis ; Pelham, 2008, J. Res. Lepid. 40: 179
P. m. venosa (Scudder, 1861)
California Pieris venosa Scudder, 1861; Proc. Boston Soc. Nat. Hist. 8 : 182, TL: San Mateo & Mendocino, California
Pieris venosa ab. flava Edwards, 1881; Papilio 1 (6) : 98
= ; Pelham, 2008, J. Res. Lepid. 40: 179 (nom. nud.)
Pieris marginalis venosa, Butterflies of America [BOA]
Pontia napi venosa ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 6
Artogeia napi venosa ; [NACL, #4195e]
Pieris marginalis venosa ; [BOC, 138]; Pelham, 2008, J. Res. Lepid. 40: 179
P. m. castoria Reakirt, 1866
California Pieris castoria Reakirt, 1866; Proc. Acad. nat. Sci. Philad. 18 (3) : 238, TL: California
Pieris nasturtii Boisduval, 1869; Ann. Soc. ent. Belg. 12 : 38 (preocc. Pieris nasturtii Edwards, 1864), TL: California
= Pieris napi castoria ; [NL4A, #271a]
= ; Pelham, 2008, J. Res. Lepid. 40: 180
Pieris resedae Boisduval, 1869; Ann. Soc. ent. Belg. 12 : 39, TL: California
= Pieris napi castoria ; [NL4A, #271a]
= ; Pelham, 2008, J. Res. Lepid. 40: 180
Pieris iberidis Boisduval, 1869; Ann. Soc. ent. Belg. 12 : 39, TL: California
= Pieris napi castoria ; [NL4A, #271a]
= ; Pelham, 2008, J. Res. Lepid. 40: 180
Pieris napi microstriata Comstock, 1924; Bull. South. Calif. Acad. Sci. 23 (4): 125, TL: California
= Pieris napi castoria ; [NL4A, #271a]
= ; Pelham, 2008, J. Res. Lepid. 40: 180
Pieris napi var. castoria ab. ♂ cottlei Gunder, 1925; Ent. News 36 (7) : 197, pl. 5, f. 9, TL: Anderson Springs, Lake Co., California
= Pieris napi castoria ; [NL4A, #271a]
= ; Pelham, 2008, J. Res. Lepid. 40: 180
Pontia napi castoria ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 6
Pieris napi castoria ; [NL4A, #271a]
Pieris marginalis castoria ; [BOC, 138]; Pelham, 2008, J. Res. Lepid. 40: 180
P. m. hulda Edwards, 1869
Alaska Pieris hulda Edwards, 1869; Trans. amer. ent. Soc. 2 : 370, TL: Kodiak
Pontia napi hulda ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 6
Artogeia napi hulda ; [NACL, #4195b]
Pieris marginalis hulda ; Pelham, 2008, J. Res. Lepid. 40: 178
P. m. pseudobryoniae (Fruhstorfer, 1909)
Alaska Pieris napi pseudobryoniae Fruhstorfer, 1909; Int. ent. Zs. 3 (16): 88
Pieris napi var. frigida f. pseudobryoniae Verity, [1908]; Rhopalocera Palaearctica 1: pl. 32, f. 36
= ; Pelham, 2008, J. Res. Lepid. 40: 178
Pieris marginalis browni Eitschberger, 1984; Herbipoliana 1 (1-2): 332, TL: Alaska
= ; Pelham, 2008, J. Res. Lepid. 40: 178
arctica Verity, 1911²
napi pseudobryoniae ; Barnes & McDunngough, 1917
Artogeia napi pseudobryoniae ; [NACL, #4195a]
Pieris marginalis pseudobryoniae ; Pelham, 2008, J. Res. Lepid. 40: 178
P. m. pseudonapi Barnes & McDunnough, 1916
Rocky Mountains (Colorado) Pieris napi pseudonapi Barnes & McDunnough, 1916; Contr. nat. Hist. Lep. N. Am. 3 (2) : 57, pl. 6, f. 1-3, TL: Silverton, Colorado
Pieris napi macdunnoughii Remington, 1954; Lepid. News 8 (3/4): 75 (repl. Pieris napi pseudonapi Barnes & McDunnough, 1916), TL: Silverton, Colorado
= ; Pelham, 2008, J. Res. Lepid. 40: 180
Pieris marginalis mcdunnoughi gen.aest. postmcdunnoughii Eitschberger, [1984]; Herbipoliana 1 (1-2): 309
= ; Pelham, 2008, J. Res. Lepid. 40: 180
Artogeia napi mcdunnoughi[sic, recte macdunoughii] ; [NACL, #4195g]
Pieris marginalis mcdunnoughi[sic, recte macdunoughii] ; [BOC, 138]
Pieris marginalis pseudonapi ; Pelham, 2008, J. Res. Lepid. 40: 180
P. m. pallidissima Barnes & McDunnough, 1916
Utah Pieris napi pallidissima Barnes & McDunnough, 1916; Contr. nat. Hist. Lep. N. Am. 3 (2) : 59, pl. 6, f. 4-5, TL: Provo, Utah
Pieris marginalis pallidissima gen.vern. antepallidissima Eitschberger, [1984]; Herbipoliana 1 (1-2): 306
= ; Pelham, 2008, J. Res. Lepid. 40: 180
Artogeia napi pallidissima ; [NACL, #4195h]
Pieris marginalis pallidissima ; Pelham, 2008, J. Res. Lepid. 40: 180
P. m. mogollon Burdick, 1942
New Mexico, Arizona, ... Pieris napi mogollon Burdick, 1942; Can. Ent. 74 (8) : 154, TL: New Mexico
Pieris narina mogollon gen.aest. warreni dos Passos, 1965; J. N.Y. Ent. Soc. 73: 136, TL: White Mts., Arizona
= Pieris napi mogollon ; [NL4A, #271b]
= ; Pelham, 2008, J. Res. Lepid. 40: 181
marginalis mogollon gen.aset. postmogollon Eitschberger, [1984]; Herbipoliana 1 (1-2): 314
= Pieris napi mogollon ; [NL4A, #271b]
= ; Pelham, 2008, J. Res. Lepid. 40: 181
Pieris marginalis mogollon, Butterflies of America [BOA]
Artogeia napi mogollon ; [NACL, #4195i]
Pieris napi mogollon ; [NL4A, #271b]
Pieris marginalis mogollon ; Pelham, 2008, J. Res. Lepid. 40: 181
P. m. meckyae Eitschberger, [1984]
Alaska Pieris marginalis meckyae Eitschberger, [1984]; Herbipoliana 1 (1-2): 322, TL: Alaska, Little Susitna River, 1100ft, Talkeetna Mts., 8 mil N. of Palmer
Pieris marginalis meckyae ; Pelham, 2008, J. Res. Lepid. 40: 178
P. m. guppyi Eitschberger, [1984]
Alaska Pieris marginalis guppyi Eitschberger, [1984]; Herbipoliana 1 (1-2): 324, TL: Alaska, Skagway
Pieris marginalis guppyi gen.aest. postguppyi Eitschberger, [1984]; Herbipoliana 1 (1-2): 324, TL: Alaska, Skagway
= ; Pelham, 2008, J. Res. Lepid. 40: 179
Pieris marginalis guppyi ; Pelham, 2008, J. Res. Lepid. 40: 179
P. m. tremblayi Eitschberger, [1984]
British Columbia Pieris marginalis tremblayi Eitschberger, [1984]; Herbipoliana 1 (1-2): 227, TL: British Columbia, Pink Mtn, 5000ft, Alaska Highway, Mile 147
Pieris marginalis tremblayi ; Pelham, 2008, J. Res. Lepid. 40: 179
P. m. reicheli Eitschberger, [1984]
British Columbia Pieris marginalis reicheli Eitschberger, [1984]; Herbipoliana 1 (1-2): 301, TL: Revelstoke, British Columbia
Pieris marginalis reicheli gen.aest. postreicheli Eitschberger, [1984]; Herbipoliana 1 (1-2): 301
= ; Pelham, 2008, J. Res. Lepid. 40: 179
Pieris marginalis reicheli, Butterflies of America [BOA]
Pieris marginalis reicheli ; Winhard, 2000, Butterflies of the world, 10: 29, pl. 46, f. 11; [BOC, 138]; Pelham, 2008, J. Res. Lepid. 40: 179
P. m. shapiroi Eitschberger, [1984]
Alaska Pieris marginalis shapiroi Eitschberger, [1984]; Herbipoliana 1 (1-2): 330, TL: Yukon Delta Hooper Bay, Kikuktok Mt., 3m
Pieris marginalis shapiroi ; Pelham, 2008, J. Res. Lepid. 40: 178
P. m. ziegleri Eitschberger, 1991
Montana Pieris marginalis ziegleri Eitschberger, 1991; Atalanta 22 (2-4): 353, TL: Montana, Gallatin Co., Hwy 191 betw. West Yellowstone and Big Sky
Pieris marginalis ziegleri ; Pelham, 2008, J. Res. Lepid. 40: 180
P. m. sequoia Emmel, Emmel & Mattoon, 1998
California Pieris marginalis sequoia Emmel, Emmel & Mattoon, 1998; in Emmel, Systematics W. North Am. Butts.: 128, TL: California, Humboldt Co., Prairie Creek Redwood State Park, 50m
Pieris marginalis sequoia ; Pelham, 2008, J. Res. Lepid. 40: 179

Pieris × passosi Warren, 1968
Pieris × passosi Warren, 1968; Ent. Rec. J. Var. 80 (3) : 62, TL: Palmer, Alaska
= Pieris marginalis shapiroi ; Pelham, 2008, J. Res. Lepid. 40: 178 (hybrid)
Pieris oleracea × hulda ;

Pieris angelika Eitschberger, [1984]
Pieris angelika Eitschberger, [1984]; Herbipoliana 1 (1-2): 340, TL: Yukon, Keno, 4600ft
Pieris angelika, Butterflies and Moths of North America [BAMONA]
Pieris angelika ; Winhard, 2000, Butterflies of the world, 10: 29, pl. 46, f. 12-13; [Opler]; Pelham, 2008, J. Res. Lepid. 40: 178

Pieris bryoniae (Hübner, [1790-1793])
CEU, SEEU. See [About maps] Bielinek bryonie
Papilio bryoniae Hübner, [1790-1793]; Schmett. Lep. linn.: 14
?Papilio bryoniae Hübner, [1805-1806]; Samml. eur. Schmett. [1] : 62, TL: near Geneva, Switzerland
?bryoniae (Ochsenheimer, 1808); Schmett. Europa 1 (2) : 151
?Pieris napi bryonides Verity, 1911; Rhopalocera Palaearctica 1: 333, pl. 52, f. 18-13 (preocc)
?Pieris napi neobryoniae Sheljuzhko, 1913; Dt. Ent. Z. Iris 27 (1) : 20 (repl. Pieris napi bryonides Verity, 1911)
Pieris flavescens Müller, 1933
Pieris bryoniae, Moths and Butterflies of Europe [MBE]
Pieris bryoniae, Extracts from [BRU] by [Ilya Osipov]
Pieris bryoniae, Europaische Schmetterlinge [Mario Maier]
Pontia napi bryoniae ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 6 (misid.?)
Pieris bryoniae ; [BRU, 164]; [EBW]; [Otakar Kudrna]
Pieris napi bryoniae ; [MRS, 258]

Larva on Brassicaceae, Thlaspi, Arabis caucasica [BRU]

P. b. bryonides Sheljuzhko, 1910
Tian-Shan, Dzhungarsky Alatau, Altai, Sayan bryoniae bryonides Sheljuzhko, 1910; Rev. russ. ent. 9: 384
bryoniae heptopotamica Krulikovsky, 1904; Rev. russ. ent. 4: 90 (nomen nudum)
[bryonides-1.jpg] 488x323 (~43Kb) The middle valley of the Kora River, 18 km NE of the Tekeli City (1600 m above sea level), the North Dzhungarskiy Alatau mountain chain (NE part of Tien Shan), the Taldy-Kurgan Province, Kazakhstan. 14th June, 19934, Photo © Oleg Kosterin

P. b. schintlmeisteri Eitschberger, 1983
Far East
P. b. kamtschadalis Röber, [1907]
Kamchatka bryoniae kamtschadalis Röber, [1907]; in Seitz, Grossschmett. Erde 1 : 49, TL: Kamchatka
P. b. sheljuzhkoi Eitschberger, [1984]
Chukot Peninsula
[sheljuzhkoi-1fu.jpg] 440x446 (~26Kb) upperside female A coastal meadow on the Cape Ploskii, the Koni Peninsula, Ola District, S Magadan Province, Russia. 8th July, 1989, Photo © Oleg Kosterin

P. b. caucasica Verity, 1908
Caucasus ?Pieris napi f. caucasica Verity, 1908; Rhopalocera Palaearctica 1: 144, pl. 32, f. 21-22
P. b. turcica Eitschberger & Hesselbarth, 1977
Armenian Highland bryoniae turcica Eitschberger & Hesselbarth, 1977; Atalanta 8 (2): 148
P. b. wolfsbergeri Eitschberger, 1984
Italy Pieris bryoniae wolfsbergeri ; Winhard, 2000, Butterflies of the world, 10: 29, pl. 45, f. 12
P. b. marani Moucha, 1956
Pieris bryoniae marani Moucha, 1956; Bull. Soc. ent. Mulhouse: 62
Pieris bryoniae marani ; Winhard, 2000, Butterflies of the world, 10: 29, pl. 45, f. 13

Pieris pseudorapae (Verity, 1908)
SEU, Turkey, Iran. See [About maps] Pieris napi var. pseudorapae Verity, 1908; Rhopalocera Palaearctica 1: 144, pl. 32, f. 23-24, pl. 33, f. 1, TL: Beirut, Lebanon
Pieris pseudorapae, Extracts from [BRU] by [Ilya Osipov]
Pieris pseudorapae ; [BRU, 165]

Larva on Alliaria pentiolata [BRU, Danchenko]

P. p. suffusa Sheljuzhko, 1931
Caucasus Minor, Armenian Highland, Talysh, Kopet-Dagh pseudorapae postsuffusa Eitschberger, 1983
P. p. balearica Wojtusiak & Niesolowski, 1946
SEU pseudorapae balearica Wojtusiak & Niesolowski, 1946; Acta Mus. Hist. nat. Acad. polon., Crakow, 6: 45
P. p. mazandarana Eitschberger, 1986

Pieris persis (Verity, 1922)
Iran, Kopet-Dagh. See [About maps] Pieris napi r. persis Verity, 1922; Ent. Rec. J. Var. 34 : 140, TL: Husseinabad, Iran
higginsi Warren, 1961
Pieris persis, Lepidoptera collection [SZMN]
Pieris persis, Extracts from [BRU] by [Ilya Osipov]
Pieris persis ; [BRU, 165]; Winhard, 2000, Butterflies of the world, 10: 29, pl. 45, f. 10

Pieris bowdeni Eitschberger, 1984
Iran, Turkey, S.Transcaucasia, Kopet-Dagh. See [About maps] Pieris bowdeni Eitschberger, 1984; Herbipoliana 1 (1-2): 218, pl. 441, f. 31-32, TL: Türkei/Agri, Asagi Kent, 2200-2400m
Pieris bowdeni, Extracts from [BRU] by [Ilya Osipov]
Pieris bowdeni ; [BRU, 165]; Winhard, 2000, Butterflies of the world, 10: 29, pl. 46, f. 1

Pieris narina Verity, 1908
Tian-Shan. See [About maps] Pieris napi var. bryoniae r. narina Verity, 1908; Rhopalocera Palaearctica 1: 145, pl. 32, f. 28-29, pl. 33, f. 2, TL: Naryn [Inner Tian-Shan]
Pieris narina, Extracts from [BRU] by [Ilya Osipov]
Pieris narina ; [BRU, 165]; Winhard, 2000, Butterflies of the world, 10: 29, pl. 46, f. 2

Pieris ochsenheimeri Staudinger, 1886
Mountains of Middle Asia Pieris ochsenheimeri Staudinger, 1886; Stett. Ent. Ztg. 47 (4-6): 199, TL: Namangan, W.Tian-Shan
Pieris ochsenheimeri, Extracts from [BRU] by [Ilya Osipov]
Pontia ochsenheimeri ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 6 (missid.?)
Pieris ochsenheimeri ; [BRU, 165]
Pieris ochsenheimeri ochsenheimeri ; Winhard, 2000, Butterflies of the world, 10: 29, pl. 46, f. 3
P. o. gerhardi Eitschberger, [1984]

Pieris euorientis (Verity, 1908)
Altai Mts. - C.Yakutia. See [About maps] euorientis (Verity, 1908); Rhopalocera Palaearctica 1: 147, pl. 32, f. 44, TL: Sajan Mts., Russia
vitimesis (Verity, 1911); Rhopalocera Palaearctica 1: 332
Pieris euorientis, Extracts from [BRU] by [Ilya Osipov]
Pieris euorientis ; [BRU, 166]

Pieris dulcinea (Butler, 1882)
NE.China, Amur, Ussuri, Far East. See [About maps] Ganoris dulcinea Butler, 1882; Ann. mag. nat. Hist. (5) 9 (49) : 18, TL: Posiette Bay, Ussuri
Pieris dulcinea, Extracts from [BRU] by [Ilya Osipov]
Pieris dulcinea ; [BRU, 166]

Larva on Arabis sagittata, Cardamine leucanthe [BRU, Kurentzov, 1970, Danchenko & Nikolaevsky]

Amur, Ussuri, Far East dulcinea sichotensis Kurentzov, 1941
dulcinea sulphurea Sheljuzhko, 1961
Pieris dulcinea dulcinea ; Winhard, 2000, Butterflies of the world, 10: 29, pl. 46, f. 7 (?)
P. d. saghalensis Nakahara, 1926
Sakhalin
P. d. tomariana Matsumura, 1928
Kuriles Pieris melete tomariana Matsumura, 1928; Insecta Matsumurana 2 (4) : 197, TL: Kuriles
dulcinea kurilica Bryk, 1942

 

Pieris melete Ménétriés, 1857
N.India, Kashmir, Manchuria, China, Korea, Amur, Ussuri, Japan. See [About maps] Pieris melete Ménétriés, 1857; Cat. lep. Petersb. 2: 113, pl. 10, f. 1-2, TL: Japan
Pieris aglaope Motschulsky, 1860; Etud. Ent. 9: 28
= ; [NHM]
Synchloë megamera Butler, 1873; Cist. Ent. 1 (7) : 173, TL: Hakodati
= ; [NHM]
?Pieris melete ab. feminalis Sheljuzhko, 1929; Mitt. Münch. Ent. Ges. 19 : 347, TL: Kagoshima (Kiu-Shiu), Japan
Pieris melete montana Verity, 1908; Rhopalocera Palaearctica 1: 141, pl. 31, f. 20-21
Pieris melete, Extracts from [BRU] by [Ilya Osipov]
Pieris melete ; [EBW]; [MRS, 259]; Winhard, 2000, Butterflies of the world, 10: 29, pl. 46, f. 9
Artogeia melete ; [BOR, 148]

Larva on Arabis sagittata, A. glauca, Cardamine leucanthe, Sisymbrium officinale [BRU, Kurentzov, 1970, Danchenko], Cardamine scutata, Arabis hirsuta, Sisymbrium luteum, Rorippa indica [MRS]

P. m. orientis Oberthür, 1880
Ussuri Pieris napi var. orientis Oberthür, 1880; Étud. d'Ent. 5 : 13
melete transiens Verity, 1908; Rhopalocera Palaearctica 1: 138, pl. 31, f. 11-12
melete amuri Sheljuzhko, 1960; Z. wien. ent. Ges. 45: 47
[orientis-1f.jpg] 642x491 (~32Kb) female A dragonfly station, Gurye County, Jeonnam Province, S Korea, 28th July 2002, Photo © Oleg Kosterin


Pieris nesis Fruhstorfer, 1909
Japan, Sakhalin. See [About maps] Pieris nesis Fruhstorfer, 1909; Int. ent. Zs. 3 (16): 88, TL: Sapporo, Japan
Pieris nesis, Extracts from [BRU] by [Ilya Osipov]
Pieris nesis ; [BRU, 167]; [BRU2: 215, pl. 83, f. 1-3]
napi nesis ;

Pieris canidia (Linnaeus, 1768)
NW.India, Himalayas, Assam (hills), Burma, Nilgiris, Palnis, Travancore (hills), Cochin, Tibet, W.China, Riu Kiu. See [About maps] Indian Cabbage White
Papilio canidia Linnaeus, 1768; in Sparrman, Amoenit. acad., 7 (150): 504n, TL: S.China
Papilio gliciria Cramer, [1777]; Uitl. Kapellen 2 (9-16): 115, pl. 171, f. E, F
= ; [NHM]
Pieris canidia, Extracts from [BRU] by [Ilya Osipov]
Pieris canidia ; [BOW: pl. 161, f. 5]; [BRU, 167]; [MRS, 258]
NW.India - N.Burma, SW.China, Hainan, Taiwan Pieris glaphyra Godart, 1819; Encyclopédie Méthodique. 9 (1): 160
Synchloe claripennis Butler, 1877; Ann. Mag. nat. Hist. (4) 19 (109) : 96, TL: Shanghai
ab. aestiva Rühl, [1893]; Palaearktischen Grosschmetterlinge: 125
Pieris canidia ab. marginalis Avinoff, 1910; Horae Soc. Ent. Ross. 39 : 247, pl. 14, f. 2, TL: Ferghana
f. nerissoides Mell, 1943; Zoologica Stuttgart, 36 (100): 104, TL: S.China
f. reducta Mell, 1943; Zoologica Stuttgart, 36 (100): 104, TL: S.China
Artogeia canidia canidia ; [BOR, 148]
Pieris canidia canidia ; Winhard, 2000, Butterflies of the world, 10: 28, pl. 45, f. 7-8

Larva on Cardamine flexuosa, C. scutata, Lepidium virginicum, Raphanus acanthiformis, Brassica alboglabra, Cleome gynandra, C. spinosa, Alstonia scholaris, Arabis hirsuta, A. stelleri, Rorippa indica [MRS]

P. c. malayica Martin, 1909
?Hongkong, Singapore Pieris melete f. malayica Martin, 1909; Ent. Zs. 23 (36) : 161
Artogeia canidia malayica ; [BOR, 148]
Pieris (Artogeia) canidia malayica ; [BMP: 86, pl. 8, f. 2]
P. c. palaearctica (Staudinger, 1886)
Middle Asia
P. c. juba Fruhstorfer, 1910
Japan Pieris canidia juba Fruhstorfer, 1910; in Seitz, Gross-Schmett. Erde 9 : 140
P. c. mars Bang-Haas, 1927
Pieris canidia mars Bang-Haas, 1927; Horae Macrolep. Palaearct. 1 : 40, pl. 5, f. 21
Pieris canidia mars ; [MRS, 258]
P. c. sordida (Butler, 1877)
Synchloe sordida Butler, 1877; Ann. Mag. nat. Hist. (4) 19 (109) : 96, TL: Shanghai
Pieris canidia sordida ; [MRS, 258]
P. c. indica Evans, 1926
Chitral, Yunnan Pieris canidia indica Evans, 1926; J. Bombay nat. Hist. Soc. 31 (3): 712, TL: Chitral, NW.Himalayas
Pieris canidia indica ; Huang, 2001
Artogeiga canidia indica ; [Torben B. Larsen]

Pieris extensa Poujade, 1888
China. See [About maps] Pieris erutae var. extensa Poujade, 1888; Bull. Soc. Ent. Fr. (6) 8 : xix
Pieris eurydice Leech, 1891; Entomologist 24 (Suppl.) : 5, TL: Wa-Shan; Chia-Kou-Ho, Huang-Mu-Chang
Pieris euridice[sic, recte eurydice] ; [EBW, 162, f. 9]
Pieris extensa ; [MRS, 259]
Pieris extensa extensa ; Winhard, 2000, Butterflies of the world, 10: 29, pl. 46, f. 8

Pieris dubernardi Oberthür, 1884
-. See [About maps] Pieris dubernardi Oberthür, 1884; Étud. d'Ent. 9 : 13, pl. 1, f. 6
Pieris dubernardi ; [MRS, 259]
N.Yunnan, W.Sichuan Pieris dubernardi dubernardi ; Winhard, 2000, Butterflies of the world, 10: 29, pl. 46, f. 15
P. d. kozlowi Alphéraky, 1897
Nanshan Pieris dubernardi var. kozlowi Alphéraky, 1897; in Romanoff, Mém. Lép. 9 : 232, pl. 12, f. 1a-c
P. d. chumbiensis (de Nicéville, 1897)
S.Tibet Parapieris chumbiensis de Nicéville, 1897; J. Asiat. Soc. Bengal 66 Pt.II (3) : 563, pl. 1, f. 6
P. d. pomiensis Yoshino, 1998
E.Tibet Pieris dubernardi pomiensis Yoshino, 1998; Neo Lepidoptera 3: 1, f. 1, 5, TL: Pomi, E. Tibet

Pieris davidis Oberthür, 1876
Pieris davidis Oberthür, 1876; Étud. d'Ent. 2 : 18, pl. 1, f. 5a-b
?Pieris davidis var. venata Leech, 1891; Entomologist 24 (Suppl.) : 57, TL: Ta-chien-lu, 8000ft
Pieris davidis ; [MRS, 260]
P. d. diluta (Verity, 1911)
Pieris davidis diluta ; [MRS, 260]; Winhard, 2000, Butterflies of the world, 10: 29, pl. 46, f. 4

fiPieris rapae (Linnaeus, 1758)
Palearctic, NA, AU. See [About maps] Naurisperhonen Small White Small Cabbage White Cabbage White Rovfjäril Rovefjäril Kleiner Kohlweißling Rübenweißling Petit Blanc Du Chou Blanquita de la Col Bielinek rzepnik
Papilio rapae Linnaeus, 1758; Syst. Nat. (Edn 10) 1 : 468, TL: Sweden
?Pieris rapae var. similis Krulikovsky, 1890; Bull. Soc. imp. Nat. Moscou (n.s.) 4 (2) : 210, pl. 8, f. b
?Pieris rapae ab. praeteria Krulikowsky, 1908; Soc. Ent. 23 (1): 3, TL: Wiatka; Kasan
Pieris rapae f. atomaria Fruhstorfer, 1909; Ent. Zs. 23 (8) : 42, TL: Dalmatia
Pieris rapae, E.H. Strickland Ent. Mus, U of [ALBERTA]
Pieris rapae rapae, Listing of Mexican Butterflies [ILMB]
Pieris rapae rapae, Butterflies of America [BOA]
Pieris rapae, Butterflies and Moths of North America [BAMONA]
Pieris rapae, Larvae of North-European Lepidoptera [Kimmo Silvonen]
Pieris rapae, Svenska fjärilar [SVF]
Pieris rapae, Norges Sommerfugler [NORLEP]
Pieris rapae, Lepidoptera Larvae of Australia [Don Herbison-Evans]
Pieris rapae, Moths and Butterflies of Europe [MBE]
Pieris rapae, Butterflies of Southeastern Arizon [Bruce Walsh]
Pieris rapae, Extracts from [BRU] by [Ilya Osipov]
Pieris rapae, Europaische Schmetterlinge [Mario Maier]
Artogeia rapae, Captain's European Butterfly Guide [Simon Coombes]
Pontia rapae ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 6
Pieris rapae ; [BRU, 168]; [EBW]; [BOW: pl. 5, f. 5]; [BAFR, 86]; [MRS, 257]; [BOC, 140]; [Opler]; [AfrL]; [NL4A, #272]; [Otakar Kudrna]; Pelham, 2008, J. Res. Lepid. 40: 182
Artogeia rapae ; [NACL, #4197]; [NAB, 360]
[rapae-1v.jpg] 500x507 (~28Kb) underside Italy: Firenze, Sesto Fiorentino, 15.5.1997, Photo © Markku Savela [rapae-2f.jpg] 600x647 (~27Kb) female USA: Wallingford, Seattle, King Co., WA, 4.7.1999, Photo © Markku Savela [rapae-3f.jpg] 700x983 (~61Kb) female USA: Wallingford, Seattle, King Co., WA, 4.7.2000, Photo © Valentina Anderson
[rapae-3fv.jpg] 800x635 (~59Kb) underside female USA: Wallingford, Seattle, King Co., WA, 4.7.2000, Photo © Valentina Anderson [rapae-4l.jpg] 550x371 (~42Kb) larva Central Europe 1990-2000, Photo © Michael Borsch [rapae-4v.jpg] 597x400 (~29Kb) underside Central Europe 1990-2000, Photo © Michael Borsch
[rapae-5mfv.jpg] 1200x819 (~112Kb) underside male female USA: Alabama, Jeff., Co., 4.3.2000, Photo © Vitaly Charny [rapae-6mu.jpg] 961x1357 (~107Kb) upperside male USA: Alabama, Jeff., Co., 12.3.2000, Photo © Vitaly Charny [rapae-7f.jpg] 1400x1050 (~130Kb) female USA: Alabama, 7.6.2003, Photo © Vitaly Charny
[rapae-8fv.jpg] 1500x975 (~112Kb) underside female USA: Alabama, 24.7.2005, Photo © Vitaly Charny [rapae-9v.jpg] 750x1106 (~84Kb) underside Japan: Tokyo, 20.09.2007, Photo © Peter Payzant [rapae.jpg] 721x619 (~57Kb) SPAIN: Teneriffa, 1-5.4.1996, Photo © Valentina Anderson

Larva on Cruciferae, Brassica spp., Brassica napus v. napobrassica, B. oleracea, Barbarea vulgaris, Cardamine pratensis, Arabis alpina, Alyssum saxatile, Hesperis matronalis, Reseda odorata, Tropaeolum majus, Nicotiana tabacum [SPRK], Brassica spp., Reseda spp. [MRS]

Larva on Cleome spp., Armoracia lapathifolia, Brassica oleracea, B. napus, B. rapa, Brassica sp. nigra/juncea, B. spp., Lepidium hyssopifolium, Raphanus sativus, Reseda sp., Tropaeolum sp. K. L. & L. E. Dunn, 1991, Review Austr. Butts. (1-4): (120-140)


Pontia metra Stephens, 1827; Ill. Br. Ent. (Haustellata) 1 (1): 19
= ; Pelham, 2008, J. Res. Lepid. 40: 183
Pieris rapae var. immaculata de Selys-Longchamps, 1857; Ann. Soc. Ent. Belge 1 : 5
= ; Pelham, 2008, J. Res. Lepid. 40: 183
Pieris yreka Reakirt, 1866; Proc. Acad. nat. Sci. Philad. 18 (3) : 238, TL: California
= ; [NL4A, #272a]; Pelham, 2008, J. Res. Lepid. 40: 183
Ganoris rapae var. novangliae Scudder, 1872; Can. Ent. 4: 79, TL: New Hampshire
= ; [NL4A, #272a]; Pelham, 2008, J. Res. Lepid. 40: 183
Pieris rapae immaculata Cockerell, 1889; Entomologist 22 : 99 (preocc. DeSelys, 1857), TL: Ireland
= ; [NL4A, #272a]; Pelham, 2008, J. Res. Lepid. 40: 183
Pieris rapae var. immaculata Skinner & Aaron, 1889; Can. Ent. 21 : 128 (preocc. DeSelys, 1857), TL: ;Pennsylvania
= ; [NL4A, #272a]; Pelham, 2008, J. Res. Lepid. 40: 183
aestivus Verity, 1913; J. Linn. Soc. Zool. Lond. 32 : 177
= ; Pelham, 2008, J. Res. Lepid. 40: 183
atomaria ; Abadjiev, 2002, Neue Ent. Nachr. 23: 17
Pontia rapae novangliae ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 6
Pontia rapae immaculata ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 6
Pieris rapae rapae ; [NL4A, #272a]; Winhard, 2000, Butterflies of the world, 10: 28, pl. 45, f. 15, 19; Pelham, 2008, J. Res. Lepid. 40: 182
P. r. deleta (Strand, 1901)
NEU rapae flavida Petersen, 1902
P. r. mauretanica Verity, 1908
N.Africa Pieris rapae mauretanica Verity, 1908; Rhopalocera Palaearctica 1: 155, pl. 33, f. 43-44, TL: Algeria and Tangier
Pieris rapae var. leucotera Stefanelli, 1869; Boll. Soc. ent. ital [1]: 147, TL: Italy
Pontia rapae var. minor Costa, 1836; Fauna Reg. Nap. Lepid.: [32], pl. 3, f. 3-4, TL: Italy
= Pieris rapae rapae ; Pelham, 2008, J. Res. Lepid. 40: 183
Ganoris rapae mauretanica f. temp. barbara Rothschild, 1925; Novit. Zool. 32 : 197, TL: Algeria
Ganoris rapae mauritanica f. gen. vern. leucoteroides Rothschild, 1926; Bull. Soc. Sci. nat. Maroc. 5: 128-129, TL: Rabat (Morocco)
Pieris rapae ab. flavoapicalis Lempke, 1934; Lambillionea 34: 81, TL: Italy [& Algeria]
Pieris rapae minor var. nana Chnéour, 1950; Bull. Soc. Hist. nat. Afr. N. 41: 41, TL: N.Tunisia
Pieris rapae messanensis Zeller, 1847; Isis, Jena 31: 221, TL: Italy
Pieris rapae mauretanica ; [BMAT, 8, pl. 3, f. 1-11]

Larva on Tropaeolum majus, Brassicaceae, Moricandia arvensis, Brassia sp. [BMAT]

P. r. transcaucasica (Stauder, 1925)
Caucasus, Armenian Highland rapae transcaucasica (Stauder, 1925); Ent. Anz.: 4
P. r. meleager Hemming, 1934
Talysh, Kopet-Dagh rapae meleager Hemming, 1934; Stylops 3 : 98 (repl. var. iranica Le Cerf)
rapae iranica Le Cerf, 1913; Ann. Hist. Nat. Délégation en Perse. 2: 25 (preocc.)
P. r. kenteana Rühl, [1893]
Altai, Sayan, Transbaikalia, Far East, Kamchatka rapae kenteana Rühl, [1893]; Palaearktischen Grosschmetterlinge: 125
rapae viluiensis Röber, [1907]; in Seitz, Grossschmett. Erde 1 : 46
P. r. crucivora Boisduval, 1836
Amur, Ussuri, Sakhalin Pieris crucivora Boisduval, 1836; Hist. nat. Ins., Spec. gén. Lépid. 1 : 522
Pieris rapae var. orientalis Oberthür, 1880; Étud. d'Ent. 5 : 13
rapae mandschurica Speyer, 1882; Stettin ent. Ztg 43 : 379
rapae wladiwostokensis Kardakov, 1928; Entomol. Mitt. 17 (4): 268, pl. 6, f. 7-8
Pieris rapae orientalis ; [MRS, 257]; Huang, 2001
Pieris rapae crucivora ; [MRS, 257]

Larva on Brassica alboglabra, B. campestris, B. oleracea, Raphanus acanthiformis, Crateva adansonii, Cleome spinosa, Rtopaeolum majus [MRS, (for crucivora)]

P. r. debilis (Alphéraky, 1889)
SEEU, Turan, Tian-Shan, Gissar, Darvaz, Alai, Pamirs rapae debilis (Alphéraky, 1889); in Romanoff, Mém. lép. 5 : 70
rapae eumorpha Fruhstorfer, 1911; Soc. ent. 26 (6) : 23
rapae pulverea Verity, 1911; Rhopalocera Palaearctica 1: 335, pl. 67, f. 25
rapae accrescens Stauder, 1924; Mitt. münch. ent. Ges. 14: 61
Pieris rapae eumorpha ; [MRS, 257]
P. r. yunnana Mell, 1943
rapae yunnana Mell, 1943; Zoologica Stuttgart, 36 (100): 104
Pieris rapae yunnaensis[sic?] ; [MRS, 257]

Pieris mannii (Mayer, 1851)
SEU, Asia Minor, Morocco, Syria. See [About maps] Southern Small White Blanca Catalana
Pontia mannii Mayer, 1851; Stettin. ent. Ztg. 1851 : 151, TL: Europe
Pieris mannii, Moths and Butterflies of Europe [MBE]
Pontia manni ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 6 (missid.?)
Pieris manni ; [EBW]; [BOW: pl. 4, f. 13]
Pieris mannii ; [Otakar Kudrna]
[mannii-1.jpg] 1138x868 (~201Kb) Italy, Ravenna, September 2002, Photo © Maurizio Elia

Larva on Cruciferae, Iberis sempervivum, Sinapis [H&R]


Pieris mannii mannii ; Winhard, 2000, Butterflies of the world, 10: 28, pl. 45, f. 16
P. m. haroldi Wyatt, 1952
Morocco (Middle Atlas) Pieris mannii haroldi Wyatt, 1952; Z. wien. ent. Ges. 37: 173, pl. 21, f. B5-B6, TL: Taghzeft Pass (Morocco)
Pieris rapae [mannii] rossii Stefanelli, 1900; Boll. Soc. ent. ital. 32 : 178, TL: Italy
Pieris mannii haroldi ; [BMAT: 8, pl. 3, f. 12-15]
P. m. todaroana Marott, 1882
Pieris mannii todaroana Marott, 1882; Giorn. Sc. Palerm. 14: 51, pl. 3
P. m. alpigena Verity, 1911
Pieris mannii alpigena Verity, 1911; Rhopalocera Palaearctica 1: 336

Pieris ergane (Geyer, [1828])
SEU, Asia Minor, Syria, Iraq, Iran, Transcaucasia. See [About maps] Mountain Small White Blanca escasa
Papilio ergane Geyer, [1828]; in Hübner, Samml. eur. Schmett. [1] : pl. 184, f. 904-907
?Papilio nelo Bergsträsser, [1779]; Nomen. Ins., 2: 47, pl. 32, f. 2
= Belenois charina ; Butler, 1872, Proc. zool. Soc. Lond. 1872 (3) : 54
= ; [NHM]
Pontia narcaea Freyer, 1828; Beitr. eur. Schmett. 1: pl. 43, f. 3
Pieris ergane gen. vern. var. stefanellii Verity, [1909]; Rhopalocera Palaearctica 1: 153, pl. 33, f. 32-36, pl. 34, f. 1, f. 23, TL: Greece
Pieris ergane, Moths and Butterflies of Europe [MBE]
Pieris ergane, Extracts from [BRU] by [Ilya Osipov]
stefanellii ; Abadjiev, 2002, Neue Ent. Nachr. 23: 18
Pieris ergane ; [BRU, 167]; [EBW]; [BOW: pl. 5, f. 1]; [Otakar Kudrna]

Larva on Aethionema saxatile [H&R]

P. e. detersa Verity, 1911
Caucasus Minor, Armenian Highland ergane detersa Verity, 1911; Rhopalocera Palaearctica 1: 153, 166, pl. 33, f. 26
P. e. elbursina Bytinski-Salz, 1937
Talysh ergane elbursina Bytinski-Salz, 1937; Ent. Rec. 49: 1
ergane alterelbursina Verity, 1937; Ent. Rec. J. Var. 49 (Suppl.) : 8

Pieris krueperi Staudinger, 1860
Balkans, Iran, Baluchistan, Kopet-Dagh, Asia Minor - Middle Asia. See [About maps] Krueper's Small White
Pieris krueperi Staudinger, 1860; Wien. ent. Monats. 4 (1) : 19, TL: "Graecia"
Pieris krueperi var. vernalis Staudinger, 1870; Horae Soc. ent. ross. 7 (1870) : 34
Pieris krueperi, Moths and Butterflies of Europe [MBE]
Pieris krueperi, Extracts from [BRU] by [Ilya Osipov]
Pieris krueperi ; [BRU, 167]; [EBW]; [BOW: pl. 5, f. 2]; [AfrL]; [Otakar Kudrna]

Larva on Alyssum montanum [H&R], Alyssum, Aurinia [BRU, Nekrutenko, Heeselbarth & al., 1995]

Pieris krueperi krueperi ; Winhard, 2000, Butterflies of the world, 10: 28, pl. 45, f. 14, 18
P. k. devta (de Nicéville, [1884])
Tian-Shan, Ghissar, S.Ghissar, Darvaz, Alai, W.Pamirs Mancipium devta de Nicéville, [1884]; J. Asiat. Soc. Bengal 52 Pt.II (2/4) : 82, pl. 9, f. 10 ♂, TL: Ladak

Pieris mahometana Grum-Grshimailo, 1888
NE.Afghanistan, W.Pamirs. See [About maps] Pieris krueperi var. mahometana Grum-Grshimailo, 1888; Horae Soc. ent. Ross. 22 (3-4) : 304, TL: Darwaz
Pieris mahometana, Extracts from [BRU] by [Ilya Osipov]

Pieris balcana Lorkovic, 1970
SEEU. See [About maps]
 

Pieris virginiensis Edwards, 1870
-. See [About maps] West Virginia White
Pieris virginiensis Edwards, 1870; Trans. amer. ent. Soc. 3 (1) : 13, TL: [Kanawha River valley, vic. Coalburgh, West Virginia]
Pieris virginiensis virginiensis, Butterflies of America [BOA]
Pieris virginiensis, Butterflies and Moths of North America [BAMONA]
Pontia napi virginiensis ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 6
Artogeia virginiensis ; [NACL, #4196]
Pieris virginiensis ; [EBW]; [Opler]; Pelham, 2008, J. Res. Lepid. 40: 182
[virginiensis-1v.jpg] 925x750 (~178Kb) underside USA: Alabama, 2.4.2005, Photo © Vitaly Charny [virginiensis-2va.jpg] 756x916 (~80Kb) underside USA: Alabama, 9.4.2005, Photo © Vitaly Charny [virginiensis-2vb.jpg] 1609x1069 (~235Kb) underside USA: Alabama, 9.4.2005, Photo © Vitaly Charny
[virginiensis-2vx.jpg] 1400x935 (~102Kb) underside USA: Alabama, 9.4.2005, Photo © Vitaly Charny

Larva on Dentaria diphylla [NAB]


West Virginia Pieris virginiensis virginiensis ; Pelham, 2008, J. Res. Lepid. 40: 182
P. v. hyatti Eitschberger, [1984]
Ontario Pieris virginiensis hyatti Eitschberger, [1984]; Herbipoliana 1 (1-2): 358, TL: Ontario, Halton Co., Forest. Campbelville
Pieris virginiensis hyatti ; Winhard, 2000, Butterflies of the world, 10: 29, pl. 46, f. 14; Pelham, 2008, J. Res. Lepid. 40: 182

The ? species-group

Pieris erutae Poujade, 1888
-. See [About maps] Pieris erutae Poujade, 1888; Bull. Soc. Ent. Fr. (6) 8 : xix
Pieris erutae Boisduval, ?MS
Yunnan, ...? Pieris erutae erutae ; Winhard, 2000, Butterflies of the world, 10: 29, pl. 46, f. 6; Huang, 2001
P. e. reissingeri Eitschberger, 1984

Pieris brassicoides Guérin-Méneville, 1849
-. See [About maps] Pieris brassicoides Guérin-Méneville, 1849; Voyage en Abyssinie 6: 365, pl. 9, f. 3-6
Pieris brassicoides ; [EBW]; [AfrL]
Ethiopia (highlands) Pieris brassicoides brassicoides ; [BAFR, 86]; Winhard, 2000, Butterflies of the world, 10: 28, pl. 45, f. 3; [AfrL]
P. b. marghanita Hemming, 1941
N.Tanzania (Mt. Meru, Ngorongoro Crater) Pieris brassicoides meridionalis Joicey & Talbot, 1922; Bull. Hill Mus. 1 (2) : 339 (nom. praeocc), TL: Great Craters, Arusha distr., Tangnyika
Pieris brassicoides marghanita ; [BAFR, 86 (text)]; [AfrL]


Unknown or unplaced taxa

Papilio sabellicae Stephens, 1827; Ill. Br. Ent. (Haustellata) 1 (1): 21, pl. 3, f. 3

Pieris napi var. nigrovenosa Selys-Longchamps, 1845; Mém. Soc. roy. Sci. Liége 2: 3 (nom. nud.?)

Pieris napi gen. II. napaeae ab. sulphurea Schöyen, 1885; Ent. Tidskr. 6 : 140

Pieris napi ab. sulphureotincta Reuter, 1893; Acta Soc. F. F. Fenn. 9 (6): 10

Pieris napi var. flava Kane, 1893; Entomologist 26: 119

Pieris napi f. (gen. I) verna Strand, 1901; Nyt Mag. Naturvid. 39: 44

Pieris napi var. bryoniae ab. immaculata Strand, 1901; Nyt Mag. Naturvid. 39: 44

Pieris napi ab. flavescens Wagner, 1903; Verh. zool.-bot. Ges. Wien 53 : 176, pl. 1, f. 1

Pieris napi ab. meta Wagner, 1903; Verh. zool.-bot. Ges. Wien 53 : 177, pl. 1, f. 3-4

Pieris napi ab. nana Röber, 1907; in Seitz, Grossschmett. Erde 1 : 48

Pieris napi f. interjecta Röber, 1907; in Seitz, Grossschmett. Erde 1 : 48

Pieris napi ab. radiata Röber, 1907; in Seitz, Grossschmett. Erde 1 : 48, pl. 21 c

Pieris napi f. meta Röber, 1907; in Seitz, Grossschmett. Erde 1 : 48, pl. 21 c

Pieris napi ab. virilis Röber, 1907; in Seitz, Grossschmett. Erde 1 : 48

Pieris napi ab. obsoleta Röber, 1907; in Seitz, Grossschmett. Erde 1 : 49

Pieris napi ab. concolor Röber, 1907; in Seitz, Grossschmett. Erde 1 : 49

Pieris napi f. suffusa Verity, 1908; Rhopalocera Palaearctica 1: 143

Pieris napi ab. carnea Verity, 1908; Rhopalocera Palaearctica 1: 149

Pieris napi. ab. nigrans Verity, 1908; Rhopalocera Palaearctica 1: 150

Pieris napi karafutonis Matsumura, 1928; Insecta Matsumurana 2 (4) : 198, TL: Saghalien


5.2.2013 (33)

References:
[ALBERTA]
University of Alberta, E.H. Strickland Entomological Museum; Browse Entomology Collection
[About maps]
Warning! The maps are automatically generated from the textual information, and the process does not always produce acceptable result; See about maps for more info.
[AfrL]
Mark C. Williams; Checklist of Afrotropical Papilionoidea and Hesperoidea; Compiled by Mark C. Williams, 7th ed. (2008) (April 2007); Afrototropical Butterflies; e-mail: Mark C. Williams
[BAFR]
D'Abrera, 1980; Butterflies of the Afrotropical Region
[BAMONA]
Opler, Pavulaan, Stanford & Pogue; Butterflies and Moths of North America; Mountain Prairie Information Node
[BCA]
electronic; BIOLOGIA CENTRALI-AMERICANA
[BHL]
The; Biodiversity Heritage Library; and on; Internet Archive; Note: The linking to the content of Internet Archive is somewhat delicate -- notify me when links break!
[BIR]
Wynter-Blyth, 1957; Butterflies of the Indian Region; (1982 Reprint)
[BMAT]
Tennent, 1996; The butterflies of Morocco, Algeria and Tunisia; ISBN 0 906802 05 9
[BMP]
Eliot (Ed.), Corbet & Pendlebury, 1992; The Butterflies of The Malay Peninsula; 4th edition; ISBN 983-9681-05-2
[BOA]
Interactive Listing of American Butterflies; Listado Interactivo de la Mariposas Americanas; Butterflies of America
[BOC]
Robert Michael Pyle, 2002; The Butterflies of Cascadia
[BOR]
D'Abrera, 1982, 1985, 1986; Butterflies of the Oriental Region, Part I-III
[BOW]
Lewis, 1974; Butterflies of the World; ISBN 0 245 52097 X
[BRU]
Tuzov, Bogdanov, Devyatkin, Kaabak, Korolev, Murzin, Samodurov, Tarasov, 1997; Guide to the Butterflies of Russia and adjacent territories; Hesperiidae, Papilionidae, Pieridae, Satyridae; Volume 1
[BRU2]
Tuzov (ed.), Bogdanov, Churkin, Devyatkin, Dantchenko, Murzin, Samodurov, Zhdanko, 2000; Guide to the Butterflies of Russia and adjacent territories; Libytheidae, Danaidae, Nymphalidae, Riodinidae, Lycaenidae; Volume 2
[BnF]
Gallica / Bibliothèque nationale de France; Annales de la Société entomologique de France; Bulletin de la Sociéte entomologique de France
[Bruce Walsh]
Bruce Walsh; e-mail: Bruce Walsh; The Moths of Southeastern Arizona; Butterflies of Southeastern Arizona
[D. Smirnov]
Russia, Moscow; e-mail: D. Smirnov
[Don Herbison-Evans]
and Stella A. Crossley; Lepidoptera Larvae of Australia; e-mail: Don Herbison-Evans; e-mail: Don Herbison-Evans
[EBW]
Smart, Paul, 1976; The Illustrated Encyclopedia of the Butterfly World; ISBN 0 600 31381 6
[ESP]
Heppner, 1981; The dates of E. J. C. Esper's Die Schmetterlinge in Abblidungen... 1776-[1830]; Archives of Natural History (1981) 10 (2): 251-254
[Erich Mangl]
http://www.mangl.at
[H&R]
Higgins, L.G, Riley, N.D, 1970; A Field Guide to the Butterflies of Britain and Europe
[HBN]
Francis Hemming, 1937; HÜBNER, A bibliographical and systematic account of the entomological works of Jacob Hübner and of the supplements thereto by Carl Geyer, Gottfried Franz von Frölich and Gottlieb August Wilhelm Herrich-Schäffer, Vol 1 & 2
[Harri Arkkio]
Finland; For the time being these pictures are only for this site. Other usage not allowed.
[Heiner Ziegler]
Switzerland / Schweiz; www.pieris.ch
[ILMB]
Interactive Listing of Mexican Butterflies
[Ilya Osipov]
The insects from former USSR for collectors Web site; e-mail: Ilya Osipov; http://www.osipov.org/insects/
[Jean Delacre]
http://homepage.mac.com/jdelacre
[Karl Hofsäß]
Karl Hofsaess; e-mail: Karl Hofsäß
[Kimmo Silvonen]
Espoo, Finland; Kimmo's Lep Site
[LAMAS-2005]
Gerardo Lamas; Entomofauna 26 (6): 57-100; A Bibliography of the Zoological Publications of Hans Fruhstorfer (1886* - 1922+)
[LAMAS-NEO]
Gerardo Lamas; Bibliography of butterflies; An Annocated Bibliography of the Neotropical Butterflies and Skipper (Lepididoptera: Papilionoidea and Hesperioidea); Revised Electronic Edition (2007)
[MBE]
Moths and Butteflies of Europe; Paolo Mazzei, Diego Reggianti and Ilaria Pimpinelli; http://www.leps.it/
[MRS]
Chou Io (Ed.); Monographia Rhopalocerum Sinensium, 1-2
[Mario Maier]
Europäische Schmetterlinge; e-mail: Mario Maier
[Maurizio Elia]
Ravenna, Italy; e-mail: Maurizio Elia
[Michael Borsch]
Geesthacht, Germany; contact through; e-mail: Francisco Welter-Schultes
[Michel Tarrier]
Michel R. Tarrier; Malaga, Spain; http://www.tarrier.org
[NAB]
Pyle, 1981; Field Guide to North American Butterflies, National Audubon Society; ISBN 0-394-51914-0
[NACL]
Hodges, 1983; Check List of the Lepidoptera of America North of Mexico
[NHM]
Natural History Museum; The Global Lepidoptera Names Index
[NL4A]
Gerardo Lamas, 2004; Atlas of Neotropical Lepidoptera; Checklist: Part 4A; Hesperioidea - Papilionoidea
[NORLEP]
Norges Sommerfugler
[Oleg Kosterin]
Institue of Cytology & Genetics; Siberian Division of Russian Academy of Sciences; Novosibirsk; e-mail: Oleg Kosterin; http://pisum.bionet.nsc.ru/kosterin/
[Opler]
Opler & Warren, 2003 (April); Butterflies of North America. 2. Scientific Names List for Butterfly Species of North America, north of Mexico.
[Peter Payzant]
Canada; http://www.pbase.com/ppayzant/profile
[SPRK]
Seppänen, E. J, 1970; Suomen suurperhostoukkien ravintokasvit, Animalia Fennica 14
[SVF]
Svenska fjärilar; Naturhistoriska riksmuseet; (Sweden)
[SZMN]
Lepidoptera; Siberian Zoological Museum
[Simon Coombes]
1 Park Street, Plymouth, PL3 4BL, UK; 01752 607854; e-mail: Simon Coombes; http://www.butterfly-guide.co.uk
[Toon Verbruggen]
Toon Verbruggen
[Vitaly Charny]
Birmingham, Alabama, USA
[Wahlberg]
Niklas Wahlberg; The NSG's voucher specimen database; Nymphalidae Systematics Group
[Yutaka Inayoshi]
A Check List of Butterflies in Indo-China; Chiefly from THAILAND, LAOS & VIETNAM
[²]
This may require parentheses or not. I don't have the necessary information for this taxon.


Some related literature:
Abadjiev, 2002
Types of Balkan butterflies in the collection of The Natural History Museum, London Neue Ent. Nachr. 23: 3-53
Alphéraky, 1889
Lépidoptères rapportés du Thibet par le Général N. M. Przewalsky de son voyage de 1884-1885 in Romanoff, Mém. lép. 5 : 59-89, pl. 4
Alphéraky, 1897
Sur quelques Lépidoptéres rapportés de l'Asie, en 1893-1895, par l'expédition de Mrs Roborowsky et Kozlov in Romanoff, Mém. Lép. 9 : 229-237, pl. 12
Avinoff, 1910
[New forms of Rhopalocera from Fergana] (in Russian) Horae Soc. Ent. Ross. 39 : 247-250, pl. 14
Bang-Haas, 1927
Horae Macrolepidopterologicae Regionis Palaearcticae Horae Macrolep. Palaearct. 1 : 1-128, pl. 1-11
Barnes & McDunnough, 1916
Notes on North American diurnal Lepidoptera Contr. nat. Hist. Lep. N. Am. 3 (2) : 49-152, pl. 4-11
Bergsträsser, [1779]
Nomenclatur und Beschreibung der Insecten in der Graftschaft Hanau-Münzenburg Nomen. Ins., 2: 79pp, pl. 15-48
Boisduval, 1869
Lépidoptères de la Californie Ann. Soc. ent. Belg. 12 : 5-28,37-94
Boisduval, 1836
Histoire Naturelle des Insectes. Species Général des Lépidoptéres. Tome Premier Hist. nat. Ins., Spec. gén. Lépid. 1 : 1-690, pl. 1-24
Butler, 1872
A synonymic List of the Species formerly included in the Genus Pieris; with all others described since Subdivision of the Group by recent Authors Proc. zool. Soc. Lond. 1872 (3) : 26-67
Butler, 1873
Descriptions of new species of Lepidoptera Cist. Ent. 1 (7) : 151-177
Butler, 1877
On Rhopalocera from Japan and Shanghai, with Descriptions of new Species Ann. Mag. nat. Hist. (4) 19 (109) : 91-97
Butler, 1882
On Lepidoptera collected in Japan and the Corea by Mr. W. Wykeham Perry Ann. mag. nat. Hist. (5) 9 (49) : 13-20
Butler, 1886
Description of a hitherto unnamed Butterfly from Madeira Ann. Mag. nat. Hist. (5) 17 (101) : 430
Le Cerf, 1913
Contribution à la faune Lépidoptérologique de la Perse Ann. Hist. Nat. Délégation en Perse. 2: -
Le Cerf, 1923
Lépidoptères nouveaux recueillis au Maroc par M. H. Ungemach Bull. Soc. ent. Fr. 1923 (15) : 197-200
Cockerell, 1889
On the variation of insects Entomologist 22 : 1-6, 26-29, 54-56, 98-100, 125-130, 147-149, 176-178, 198-202, 226-229, 243-245
Cramer, [1777]
Uitlandsche Kapellen (Papillons exotiques) Uitl. Kapellen 2 (9-16): 1-152, pl. 97-192 (1777)
Dalman, 1816
Försök till systematiks Uppställing af Sveriges Fjärilar K. VetenskAcad. Handl. 1816 (1): 48-101, (2): 199-225
de Nicéville, [1884]
On new and little known Rhopalocera from the Indian Region J. Asiat. Soc. Bengal 52 Pt.II (2/4) : 65-91, pl. 1, 9-10
de Nicéville, 1897
On new or little-known Butterflies from the Indo- and Austro-Malayan Regions J. Asiat. Soc. Bengal 66 Pt.II (3) : 543-577, pl. 1-4
dos Passos, 1965
Review of the Nearctic species of Pieris "napi" as classified by androconial scales and description of a new seasonal form J. N.Y. Ent. Soc. 73: 135-137
Doubleday, [1845]
List of the Specimens of Lepidopterous insects in The Collection of the British Museum List. lepid. Ins. Brit. Mus. 1 : 1-150
K. L. & L. E. Dunn, 1991
Review of Australian Butterflies: distribution, life history and taxonomy. Pushlished by authors, Melbourne Review Austr. Butts. (1-4):
Dyar, 1903
A List of North American Lepidoptera and Key to the Literature of this Order of Insects Bull. U.S. nat. Mus. 52 : 1-723
Edwards, 1869
Descriptions of new species of diurnal Lepidoptera found within the United States Trans. amer. ent. Soc. 2 : 369-376
Edwards, 1870
Descriptions of new species of diurnal Lepidoptera found within the United States Trans. amer. ent. Soc. 3 (1) : 10-22
Edwards, 1881
On Pieris bryoniae Ochsenheimer, and its derivative forms in Europe and America Papilio 1 (6) : 83-99
Eitschberger, [1984]
Systematische Untersuchungen am Pieris napi-bryoniae-Komplex (s.l.) Herbipoliana 1 (1-2): (1) i-xxii, 1-504, (2) 1-601
Eitschberger, 2001
Eine neue Unterart von Pieris napi (Linnaeus, 1758) vom Polar Ural Atalanta 32 (1/2): 85-88
Emmel, Emmel & Mattoon, 1998
New subspecies of Pieridae (Lepidoptera) from California, Nevada and Baja California in Emmel, Systematics W. North Am. Butts.: 127-138
Esper, [1804]
Die Schmetterlinge in Abbildungen nach der Natur mit Beschreibungen. Theil I. Die Tagschmetterlinge. Supplement Theil 1. Abschnitt 1 Die Schmett., Suppl. Th 1 (1-2): 1-8, pl. 94 (1789), (3-4): 9-60, pl. 95-101 (1789), (5-6): 61-68, pl. 102-106 (1794), ([7]): [69-88], pl. [107-109] (1798), (8-9): 89-104, pl. 110-112 (1800), (10): 105-120, pl. 113-116 ([1803-1804])
Freyer, 1828
Beiträge zur Geschichte europäischer Schmetterlinge mit Abbildungen nach der Natur Beitr. eur. Schmett. 1: 164pp, pl. 1-48
Fruhstorfer, 1909
Neues über Pieris manni [sic] Ent. Zs. 23 (8) : 41-42 (22 May)
Fruhstorfer, 1909
Neue palaearktische Pieriden Int. ent. Zs. 3 (16): 88 (17 July)
Fruhstorfer, 1910
2. Familie: Pieridae, Weisslinge in Seitz, Gross-Schmett. Erde 9 : 119-120 (1909), : 121-144 (14 January, 1910), : 145-152 (11 February, 1910), : 153-168 (2 April, 1910), : 169-176 (4 April, 1910), : 177-190 (20 May, 1910)
Fruhstorfer, 1911
Neue palaearktische Rhopaloceren Soc. ent. 26 (6) : 23-24 (17 June)
Godart, 1819
in Latreille & Godart, Encyclopédie Méthodique. Histoire naturelle Entomologie, ou histoire naturelle des crustacés, des arachnides et des insectes Encyclopédie Méthodique. 9 (1): 3-328 (1819), (2): 329-828 ([1824])
Godman & Salvin, [1889]
Biologia Centrali-Americana. Rhopalocera. (1887-1901) Biol. centr.-amer., Lep. Rhop. 2: 1-782, 3: pl. 1-112
Gray, 1846
Descriptions and Figures of some new Lepidopterous Insects chiefly from Nepal Descr. lep. Ins. Nepal : 1-16, pl. 1-14
Grote, 1873
On the Butterflies of Anticosti Bull. Buffalo Soc. nat. Sci. 1 : 185
Grum-Grshimailo, 1888
Novae species et varietates Rhopalocerum e Pamir Horae Soc. ent. Ross. 22 (3-4) : 303-307
Gunder, 1925
Several New Aberrant Lepidoptera (Rhopalocera) Ent. News 36 (7) : 193-200, pl. 5
Hemming, 1934
Some notes in the nomenclature of Palaearctic and African Rhopalocera Stylops 3 : 97-99
Hemming, 1967
Generic names of the butterflies and their type-species (Lepidoptera: Rhopalocera) Bull. Br. Mus. (Ent.) Suppl. 9 : 5-509
Hensle, 2001
Zur Frage der subspezifischen Zuordnung von "Pieris bryoniae lappona" Rangnow, 1935 Atalanta 32 (1/2): 89-95
Heyne, [1895]
Satyridae, Lycaenidae in Rühl, Die palaearktischen Grossschmetterlinge und ihre Naturgeschichte 1 (Tagfalter): 385-857
Hübner, [1790-1793]
Der Schmetterlinge Lepidoptera Linnei, Europäisches Heer, [1790-1793] Schmett. Lep. linn.: [1-3], 4-16
Hübner, [1806]
Tentamen determinationis digestionis alque denominationis singlarum stripium Lepidopterorum, peritisad inspiciendum et dijudicandum communicatum, a Jacob Hübner Tentamen determinationis digestionis ...: -
Geyer in Hübner, [1828]; Hübner, [1805-1806]
Sammlung europäischer Schmetterlinge. I. Papiliones - Falter ("Erste Band") Samml. eur. Schmett. [1] : 3-74, : pl. 1-88, f. 1-446 (1799-1800), : pl. 89-96, f. 447-490 (1800-1803), : pl. 97-114, f. 491-588 (1803-1804), : pl. 115-119, f. 589-612 (1805), : pl. 120-124, f. 613-636 (1805-1806), : pl. 125-128, f. 637-652 (1806-1808), : pl. 129-144, f. 653-727 (1808-1813), : pl. 145-147, f. 728-741 (1814-1816), : pl. 148-150, f. 742-756 (1816-1817), : pl. 151, f. 757-762 (1818-1819), : pl. 152-154, f. 757-774 (1819-1822), : pl. 155, f. 775-778 (1822-1823), : pl. 156-161, f. 779-806 (1823), : pl. 162-175, f. 807-874 (1823-1824), : pl. 176, f. 869-874 (1824-1825), : pl. 177-181, f. 875-895 (1825-1826), : pl. 182-187, f. 896-981 (1827-1828), : pl. 188-195, f. 922-968 (1828-1832), : pl. 196, f. 969-974 (1832-1833), : pl. 197-201, f. 975-997 (1834-1836), : pl. 202-203, f. 998-1009 (1836-1838), : pl. 204-207, f. 1010-1029 (1837-1838)
Hübner, [1823]
Sammlung exotischer Schmetterlinge, Vol. 2 ([1819] - [1827]) Samml. exot. Schmett. 2 : pl. [90], [180], [223] (1819), : pl. [2], [12], [16], [78], [141], [182] (1820), : pl. [11], [13], [24], [77], [83], [97], [102-103], [108-109], [116-117], [122], [147], [150], [164], [183], [195], [197], [212-214] (1821), : pl. [14], [18-19], [26], [38-39], [43], [68], [70-71], [74], [84], [86], [92], [94-95], [106], [114], [121], [161], [169], [174], [177-178], [184], [194], [200], [204-205], [207-208], [210] (1821), : pl. [33], [72], [76], [115], [131], [138-139], [153], [198], [211], [218], [220] (1822), : pl. [32], [75], [81], [93], [105], [129] (1822), : pl. [1], [6], [22-23], [25], [27], [29-31], [34-37], [40-42], [48], [50-53], [56], [58-59], [64-66], [69], [73], [87], [89], [91], [96], [98-101], [110], [124], [128], [130], [132-137], [142], [155-158], [160], [162-163], [165-167], [170-173], [175-176], [187-188], [196], [199], [201-202], [206], [215-216], [219], [222], [224] (1823), : pl. [54], [118], [143], [149], [159] [179], [181], [185], [203], [217] (1824), : pl. [3-5], [8-10], [15], [20], [45], [49], [55], [60], [62-63], [67], [79-80], [88], [104], [111-113], [120], [125], [127], [140], [144-146], [148], [151-152], [154], [168], [189-193], [221], [225] (1825), : pl. [7], [17], [21], [28], [44], [46-47], [57], [61], [82], [123], [126] (1826), : pl. [85], [119], [186], [209] (1827), : (Index Systematicus) [1-4] (1827-1832)
Hübner, 1822
Systematisch-alphabetisches Verzeichniss aller bisher bey den Fürbildungen zur Sammlung europäischer Schmetterlinge angegebenen Gattungsbenennungen; mit Vormerkung auch augsburgischer Gattungen, von Jacob Hübner Syst.-alph. Verz.: [I-III], IV-VI, [1]-2-81
Ikeda, 1937
On a new subspecies of Pieris cisseis Leech from Formosa Zephyrus 7 (1) : 37-40
Joicey & Talbot, 1922
New Forms of Butterflies from Africa Bull. Hill Mus. 1 (2) : 339-342
Kardakov, 1928
Zur Kenntnis der Lepidopteren des Ussuri-Gebietes Entomol. Mitt. 17 (4): 261-273
Kirby, 1837
Fauna Boreali-Americana or the Zoology of the northern parts of British America in Richardson, Fauna Boreal Amer.: 325pp, 8pls
Krulikovsky, 1890
[russian: ? "experimental catalog of lepidoptera of Kazakh province"] Bull. Soc. imp. Nat. Moscou (n.s.) 4 (2) : 200-251, pl. 8
Krulikowsky, 1908
Einige neue Varietäten und Aberrationen der Lepidopteren des östlichen Russlands Soc. Ent. 23 (1): 2-3, (2): 11-12, (3): 18
Leech, 1890
New species of Lepidoptera from China Entomologist 23 : 26-50, 81-83, : 109-114, : 187-192, pl. 1
Leech, 1891
New Species of Lepidoptera from China Entomologist 24 (Suppl.) : 1-6
Leech, 1891
New Species of Rhopalocera from Western China Entomologist 24 (Suppl.) : 57-61, 66-68
Linnaeus, 1758
Systema Naturae per Regna Tria Naturae, Secundum Clases, Ordines, Genera, Species, cum Characteribus, Differentiis, Symonymis, Locis. Tomis I. 10th Edition Syst. Nat. (Edn 10) 1 : 1-338, : 339-824
Linnaeus, 1768
Sparrman (Thesis), Iter Chinense in Sparrman, Amoenit. acad., 7 (150): 497-506
Martin, 1909
Eine paläarkt. Pieride unter dem Aequator Ent. Zs. 23 (36) : 161
Matsumura, 1928
New Butterflies especially from the Kuriles Insecta Matsumurana 2 (4) : 191-201
Mayer, 1851
Pont. Mannii n. sp. Stettin. ent. Ztg. 1851 : 151
Ménétriés, 1857
Enumeratio corporum animalium Musei Imperialis Academiae Scientiarum Petropilitanae. Classis Insectorum, Ordo Lepidopterorum Cat. lep. Petersb. 1: 1-66, pl. 1-6, (supplement) 67-112 (1855), 2: 67-97,99-144, pl. 7-14 (1857), 3: 145-161, pl. 15-18 (1863)
Moore, 1865
List of diurnal Lepidoptera collected by Capt. A.M. Lang in the N. W. Himalayas Proc. zool. Soc. Lond. 1865 (2) : 486-509, pl. 30-31
Moore, 1884
Descriptions of some new Asiatic diurnal Lepidoptera; chiefly from specimens contained in the Indian Museum, Calcutta J. asiat. Soc. Bengal 53 Pt.II (1) : 16-52
Motschulsky, 1860
Insectes du Japon Etud. Ent. 9: 4-39
Oberthür, 1876
Espèces nouvelles de Lépidopterès recueillis en Chine par M. l'abbé A. David / Lépidoptères nouveaux de la Chine Étud. d'Ent. 2 : 13-34, pl. 1-4
Oberthür, 1880
Faune des Lépidoptères de l'ile Askold. Premiere Partie Étud. d'Ent. 5 : 1-88, pl. 1-9
Oberthür, 1884
Lépidoptères du Thibet Étud. d'Ent. 9 : 7-22, pl. 1-2
Ochsenheimer, 1808
Die Schmetterlinge von Europa. Erster Theil, Zweyte Abtheilung. Falter, oder Tagschmetterlinge Schmett. Europa 1 (2) : 1-240
Pelham, 2008
A catalogue of the butterflies of the United States and Canada J. Res. Lepid. 40: 1-652
Poujade, 1888
[Piéride et de Noctuélide] Bull. Soc. Ent. Fr. (6) 8 : xix-xx,[19-20]
Reakirt, 1866
Descriptions of some new species of Diurnal Lepidoptera Proc. Acad. nat. Sci. Philad. 18 (3) : 238-249
Reuter, 1893
Förteckning öfver Macrolepidoptera fauna i Finland Acta Soc. F. F. Fenn. 9 (6): 85pp
Rothschild, 1917
Supplemental notes to Mr. Charles Oberthür's Faune des Lépidoptères de la Barbarie, with lists of the specimens contained in the Tring Museum Novit. Zool. 24 (1): 61-120, : 325-373, : 393-409
Rothschild, 1925
Critical List of the collection of Algerian Lepidoptera of the late Captain N. J. E. Holl. Novit. Zool. 32 : 195-229
Röber, 1896
Neue Schmetterlinge aus dem cilicischen Taurus Ent. Nachr. 22 (6) : 81-84
Schrank, 1801
Fauna Boica. Durchgedachte Geschichte der in Baiern einheimischen und zahmen Thiere Fauna Boica 2 (1) : 1-374
Schöyen, 1885
Bemærkninger om enkelte variationer af vore Rhopalocera Ent. Tidskr. 6 : 139-144
Scudder, 1861
Notice of some North American species of Pieris Proc. Boston Soc. Nat. Hist. 8 : 178-186
de Selys-Longchamps, 1857
Catalogue des insected lepidoptères de la Belgique Ann. Soc. Ent. Belge 1 : 1-111
Sheljuzhko, 1913
Lepidopterologische Notizen Dt. Ent. Z. Iris 27 (1) : 13-22
Sheljuzhko, 1929
Einige neue palaearktische Lepidopteren-Formen Mitt. Münch. Ent. Ges. 19 : 347-362, pl. 27-28
Shirôzu, 1953
New or little know butterflies from the North-Eastern Asia, with some synonymic notes. (1) & (2) Sieboldia 1 (1952) : 11-37, 11pls, : 149-159
Skinner & Aaron, 1889
A list of the butterflies of Philadelphia, PA. Can. Ent. 21 : 126-131
Stauder, 1924
Neue Palaearktenformen. II Mitt. münch. ent. Ges. 14: 59-66
Staudinger, 1860
Pieris krueperi, Ein neuer europäischer Tagfalter Wien. ent. Monats. 4 (1) : 19-20
Staudinger, 1870
Beitrag zur Lepidopterenfauna Griechenlands Horae Soc. ent. ross. 7 (1870) : 3-304, pl. 1-3
Staudinger, 1886
Centralasiatische Lepidopteren Stett. Ent. Ztg. 47 (4-6): 193-215, (7-9): 225-256
Staudinger in Staudinger & Rebel, 1901
Catalog der Lepidopteren des palaearktischen Faunengebiets. I. Famil. Papilionidae - Hepialidae Cat. Lep. palaearct. Faunengeb. 1 : 1-411
Stephens, 1827
Illustrations of British Entomology; or, a synopsis of indigenous insects: containing their generic and specific distinctions; with an account of their metamorphoses, times of appearance, localities, food, and economy, as far as practicable Ill. Br. Ent. (Haustellata) 1 (1): 1-56, pl. 1-9 (1827), (2): 57-150, pl. 10-12 (1828)
Verity, 1908; Verity, [1909]; Verity, 1911
Rhopalocera Palaearctica Iconographie et Description des Papillons diurnes de la région paléarctique. Papilionidae et Pieridae Rhopalocera Palaearctica 1: 86+368pp, 2+12+72pls
Verity, 1913
Revision of the Linnean Types of Palaearctic Rhopalocera J. Linn. Soc. Zool. Lond. 32 : 173-191
Verity, 1922
Seasonal polymorphism and races of some European Grypocera and Rhopalocera Ent. Rec. J. Var. 34 : 12-15, 68-73, 89-93, 124-142
Verity, 1937
Variation of some butterflies in anterior Asia and in Morocco Ent. Rec. J. Var. 49 (Suppl.) : 1-8,?
Wagner, 1903
Zur Kenntnis einiger Formen von Pieris Napi L. Verh. zool.-bot. Ges. Wien 53 : 174-178, pl. 1
Winhard, 2000
Pieridae I Butterflies of the world, 10: (1) 40pp, (2) 48pls
Wojtusiak & Niesolowski, 1946
Lepidoptera of the Central Caucasus, collected during the Polish Alpine Expedition in 1935, with ecological and zoogeographical remarks. I Part. Macrolepidoptera Acta Mus. Hist. nat. Acad. polon., Crakow, 6: 1-74
Yoshino, 1998
New Butterflies from China 4 Neo Lepidoptera 3: 1-7
Zeller, 1847
Bemerkungen über die auf einer Reise Nach Italien und Sicilien Beobachteten Schmetterlingsarten Isis von Oken 1847 (2): 121-159, (3): 213-233, (4): 284-308, (6): 401-457, (7): 481-522, (8): 561-594, (9): 641-673, (10): 721-771, (11): 801-859, (12): 881-914


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