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Cepora Billberg, 1820
Gulls
Cepora Billberg, 1820; Enum. Ins. Mus. Billb. : 76, TS: Papilio coronis Cramer
Huphina Moore, [1881]; Lepid. Ceylon 1 (3): 136, TS: Papilio coronis Cramer
Huphina (Pierinae) ; Moore, [1881], Lepid. Ceylon 1 (3): 136
Cepora ; Winhard, 2000, Butterflies of the world, 10: 22
Cepora (Pierinae) ; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 110

Cepora nerissa (Fabricius, 1775)
Ceylon, India, Burma. See [About maps] Common Gull
Papilio nerissa Fabricius, 1775; Syst. Ent.: 471, no. 123, TL: China
Pieris hira Moore, 1865; Proc. zool. Soc. Lond. 1865 (2) : 490, pl. 31, f. 17, TL: Punjab; Oude
= ; [NHM]
Pieris copia Wallace, 1867; Trans. ent. Soc. Lond. (3) 4 (3) : 340, TL: Bengal
= ; [NHM]
Cepora nerissa nerissa, Butterflies in Indo-China [Yutaka Inayoshi]
Huphima nerissa ; [BIR, 421]
Cepora nerissa ; [EBW]; [BOW: pl. 158, f. 19]; [BOR, 148]; [MRS, 255]
[nerissa-1v.jpg] 600x400 (~82Kb) underside India: Arunachal Pradesh, Taley Valley WLS, 12.11.2005, Photo © Kedar Tokekar

Larva on Capparis [BIR], Capparis cantoniensis [MRS]


N.Vietnam, S.China Papilio amasene Cramer, [1775]; Uitl. Kapellen 1 (1-7): pl. 44, f. A, TL: China
Huphina praetermissa Watkins, 1927; Ann. Mag. nat. Hist. (9) 19 : 337, TL: Hong Kong
Huphina f. tairai Umeno, 1935; Zephyrus 6: 102
Cepora nerissa nerissa ; [BOR, 148]
C. n. phryne (Fabricius, 1775)
N.India - W.Burma Papilio phryne Fabricius, 1775; Syst. Ent.: 473
?Papilio phryne Fabricius, 1775; Syst. Ent.: 196 [?]
Huphina phryne ; Moore, [1881], Lepid. Ceylon 1 (3): 136, pl. 53, f. 1, 1a-b
Cepora nerissa phryne ; [BOR, 148]
C. n. dapha (Moore, [1879])
C.Burma, E.Burma - Thailand, N.Peninsular Malaya, Langkawi Appias dapha Moore, [1879]; Proc. zool. Soc. Lond. 1878 (4) : 838, TL: Moulmein, Myanmar
Cepora nerissa dapha, Butterflies in Indo-China [Yutaka Inayoshi]
Cepora nerissa dapha ; [BOR, 148]; [BMP: 87, pl. 8, f. 5-6]; Winhard, 2000, Butterflies of the world, 10: 22, pl. 32/33, f. 9
C. n. evagete (Cramer, [1779])
Ceylon, S.India Papilio evagete Cramer, [1779]; Uitl. Kapellen 3 (17-21): pl. 221, f. F, G
Papilio zeuxippe Stoll, [1781]; in Cramer, Uitl. Kapellen 4 (29-31): 141, pl. 362, f. E, F
Papilio cassida Fabricius, 1798; Ent. Syst. (Suppl.): 427
Cepora nerissa evagete ; [BOR, 148]
C. n. cibyra Fruhstorfer
Taiwan Cepora nerissa cibyra ; [MRS, 255]; [BOR, 149]
Andamans Cepora nerissa lichenosa ; [BOR, 149]
C. n. sumatrana (Hagen, 1894)
Sumatra Pieris nerissa var. sumatrana Hagen, 1894; Dt. ent. Z. Iris 7 (1) : 32, TL: Sumatra
Cepora nerissa sumatrana ; [BOR, 149]
C. n. corva (Wallace, 1867)
Java Pieris corva Wallace, 1867; Trans. ent. Soc. Lond. (3) 4 (3) : 339, TL: Java, Bali
Cepora nerissa corva ; [BOR, 149]
C. n. vaso (Doherty, 1891)
Sumbawa Huphina vaso Doherty, 1891; J. Asiat. Soc. Bengal 60 Pt.II (2) : 188, TL: Sambawa
Cepora nerissa vaso ; [BOR, 149]
C. n. physkon Fruhstorfer˛
Lombok Cepora nerissa physkon ; [BOR, 149]
Cepora nerissa yunnanensis ; [MRS, 255]
C. n. coronis (Cramer, [1775])
Papilio coronis Cramer, [1775]; Uitl. Kapellen 1 (1-7): 69, pl. 44, f. B, C, TL: China
C. n. dissimilis (Rothschild, 1892)
Bali Huphina dissimilis Rothschild, 1892; Dt. ent. Z. Iris 5 (2) : 440, pl. 5, f. 5-6, TL: S.Celebens

Cepora fora (Fruhstorfer, 1897)
-. See [About maps] ?Appias zelmira fora Fruhstorfer, 1897; Berl. Ent. Zs. 42 (1/2) : 117, TL: Bua-Kraeng, Pik von Bonthain
?Huphina fora f. vallifusca Martin, 1919; Dt. ent. Z. Iris 33 (1/2) (3/4): 69
?Huphina fora f. margilatior Martin, 1919; Dt. ent. Z. Iris 33 (1/2) (3/4): 70
Sulawesi Cepora fora fora ; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 112
C. f. papayatana Watanabe, 1987
N.Sulawesi Cepora fora papayatana ; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 112
C. f. milos Watanabe, 1987
Buton Cepora fora milos ; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 112

Cepora aspasia (Stoll, [1790])
Papilio aspasia Stoll, [1790]; Aanhangsel Werk, Uitl. Kapellen (2-5): 148, pl. 33, f. 3, 3c (preocc. Papilio aspasia Fabricius, 1787)
Cepora aspasia ; [EBW]; [BOW: pl. 158, f. 3]; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 112
Cepora aspasia aspasia ; [BAUR, 155]; Winhard, 2000, Butterflies of the world, 10: 22, pl. 32/33, f. 13
C. a. talautensis Niepelt, 1926
Talaud I. aspasia talautensis Niepelt, 1926; Int. Ent. Zs. 20: 186
Cepora aspasia talautensis ; Iwasaki & Yata, 2005, Trans. lepid. Soc. Japan 56 (4) : 305
?C. a. emma (Vollenhoven, 1865)
Morotai, Halmahera, Ternate, Bachan Pieris emma Vollenhoven, 1865; Faune ent. 2 (Monogr. Pierid.) : 24, pl. 4, f. 2
Cepora aspasia emma ; [BAUR, 155]
?C. a. hester (Vollenhoven, 1865)
Waigeu Pieris hester Vollenhoven, 1865; Faune ent. 2 (Monogr. Pierid.) : 24, pl. 4, f. 1
Cepora aspasia hester ; [BAUR, 155 (text)]
C. a. jael (Wallace, 1867)
Buru, Ambon, Serang Pieris jael Wallace, 1867; Trans. ent. Soc. Lond. (3) 4 (3) : 335, TL: Buru
Cepora aspasia jael ; Iwasaki & Yata, 2005, Trans. lepid. Soc. Japan 56 (4) : 305
?C. a. aspasina (Fruhstorfer, 1904)
Obi Huphina aspasia aspasina Fruhstorfer, 1904; Ent. Meddel. 2 (5/6) : 304, TL: Obi
Cepora aspasia aspasina ; [BAUR, 155]

Cepora judith (Fabricius, 1787)
-. See [About maps] Papilio iudith Fabricius, 1787; Mantissa Insectorum 2: 22, no. 230
?Huphina judith vaga van Eecke, 1918; Zool. Meded. 4 (7) : 74, TL: Pulu Babi
Cepora judith ; [EBW]; [BOW: pl. 158, f. 6]; [MRS, 256]
Cepora iudith ; [BOR, 150]; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 112
Cepora idith ; (missp.?)

How to split the taxa beween judith/iudith and aspasia, if separated?

Larva on Capparis [BMP]

Java Cepora iudith iudith ; [BOR, 151]
C. j. lea (Doubleday, 1846)
S.Burma Pieris lea Doubleday, 1846; Ann. Mag. nat. Hist. 17 (109) : 23, TL: Moulmein, Myanmar
Huphina lea continentalis Röber, 1927; Int. ent. Zs. 20 (44): 401, TL: N.Myanmar
Cepora iudith lea, Butterflies in Indo-China [Yutaka Inayoshi]
Cepora lea ; [EBW]; [BOW: pl. 158, f. 4]
Cepora iudith lea ; [BOR, 151]
Cepora judith lea ; Winhard, 2000, Butterflies of the world, 10: 22, pl. 32/33, f. 12
C. j. malaya (Fruhstorfer, 1899)
Peninsular Malay, Langkawi, Singaphore Orange Gull
Huphina lea malaya Fruhstorfer, 1899; Berl. ent. Zs. 44 (1/2) : 101
Cepora iudith malaya ; [BOR, 151]; [BMP: 88, pl. 8, f. 7-8]
C. j. siamensis (Butler, 1899)
Thailand, Pulau Aur Huphina siamensis Butler, 1899; Ann. Mag. nat. Hist. (7) 3 (15) : 208
Cepora aora Pendlebury, 1933; J. Fed. Malay St. Mus. 17 (2): 381
Cepora iudith siamensis ; [BOR, 151]; [BMP: 88, pl. 8, f. 9]
[siamensis-1.jpg] 887x673 (~123Kb) Thailand, Chon Buri Province, Si Racha environs, the Chan-ta-Then River valley at lower reaches, 12th January 2006, Photo © Oleg Kosterin [siamensis-2v.jpg] 808x693 (~183Kb) underside Thailand, Chon Buri Province, Si Racha environs, the Chan-ta-Then River valley at lower reaches, 12th January 2006, Photo © Oleg Kosterin [siamensis-3v.jpg] 825x669 (~139Kb) underside Thailand, Chon Buri Province, Si Racha environs, the Chan-ta-Then River valley at lower reaches, 12th January 2006, Photo © Oleg Kosterin
[siamensis-4.jpg] 1097x918 (~130Kb) Thailand, Chon Buri Province, Si Racha environs, the Chan-ta-Then River valley at lower reaches, 12th January 2006, Photo © Oleg Kosterin

C. j. talboti Corbet, 1937
Pulau Tioman Cepora talboti Corbet, 1937; Proc. R. ent. Soc. Lond. (B) 6 (3): 48
Cepora iudith talboti ; [BOR, 151]
C. j. amalia (Vollenhoven, 1865)
Sumatra Pieris amalia Vollenhoven, 1865; Faune ent. 2 (Monogr. Pierid.) : 23, pl. 3, f. 6
Cepora iudith amalia ; [BOR, 151]
C. j. montana (Fruhstorfer, 1899)
N.Borneo Hyphina lea ♀ f. montana Fruhstorfer, 1899; Berl. ent. Zs. 44 (1/2) : 102
Cepora iudith montana ; [BOR, 151]
SE.Borneo Cepora iudith meridionalis ; [BOR, 151]
Sarawak, Labuan Cepora iudith hespera ; [BOR, 151]
C. j. natuna (Fruhstorfer, 1899)
Natuna Is. Huphina lea natuna Fruhstorfer, 1899; Berl. ent. Zs. 44 (1/2) : 101, TL: Natuna Is.
Cepora iudith natuna ; [BOR, 151]
C. j. selma (Weymer, 1885)
Nias Pieris selma Weymer, 1885; Stettin ent. Ztg. 46 (4-6) : 269, pl. 2, f. 5, TL: Nias
Cepora iudith selma ; [BOR, 151]
Enggano Cepora iudith ethel ; [BOR, 151]
C. j. naomi (Wallace, 1867)
Lombok Pieris naomi Wallace, 1867; Trans. ent. Soc. Lond. (3) 4 (3) : 336, TL: Lombock, Flores
Cepora iudith naomi ; [BOR, 151]
C. j. aga (Fruhstorfer, 1903)
Sumbawa Huphina lea aga Fruhstorfer, 1903; Dt. ent. Z. Iris 15 (2) : 280, TL: Tambora, Sumbava
Cepora iudith aga ; [BOR, 151]
C. j. oberthueri (Röber, 1891)
Flores Pieris oberthüri Röber, 1891; Tijdschr. Ent. 34: 277, TL: Flores
Cepora iudith oberthueri ; [BOR, 151]
C. j. eirene (Doherty, 1891)
Sumba Huphina eirene Doherty, 1891; J. Asiat. Soc. Bengal 60 Pt.II (2) : 188, TL: Sumba
Cepora iudith eirene ; [BOR, 151]
C. j. olga (Eschscholtz, 1821)
Philippines Pieris olga olga Eschscholtz, 1821; in Kotzebue, Entdeck. Reise Süd-See 3 (App. 5): pl. 9, f. 21a-b
Cepora iudith olga ; [BOR, 151]
Cepora judith olga ; [MRS, 256]
C. j. anaitis Fruhstorfer˛
Philippines (NW.Luzon) Cepora iudith anaitis ; [BOR, 152]
C. j. rhemia Fruhstorfer˛
Philippines (Mindoro, Negros, ?Bohol) Cepora iudith rhemia ; [BOR, 152]
C. j. poetelia Fruhstorfer˛
Philippines (Cebu) Cepora iudith poetelia ; [BOR, 152]
C. j. orantia Fruhstorfer˛
Philippines (Mindanao) Cepora iudith orantia ; [BOR, 152]
C. j. olgina Staudinger˛
Philippines (Palawan) Cepora iudith olgina ; [BOR, 152]
C. j. zisca (Fruhstorfer, 1899)
Philippines (Basilan) Huphina aspasia zisca Fruhstorfer, 1899; Berl. ent. Zs. 44 (1/2) : 104, TL: Bazilan
Cepora iudith zisca ; [BOR, 152]
C. j. irma Fruhstorfer˛
Philippines (Jolo) Cepora iudith irma ; [BOR, 152]
C. j. phokaia Fruhstorfer˛
Philippines (Balabac) Cepora iudith phokaia ; [BOR, 152]

Cepora abnormis (Wallace, 1867)
New Guinea. See [About maps] Tachyris abnormis Wallace, 1867; Trans. ent. Soc. Lond. (3) 4 (3) : 368, pl. 8, f. 5 ♀, TL: NW. New Guinea
Delias abnormis ; Grose-Smith & Kirby, 1889, Rhop. Exot. [1] 1: (Pierinae) 6, pl. 2, f. 6-7; Grose-Smith, 1894, Novit. zool. 1 (2): 334
Cepora abnormis ; [EBW]; [BOW: pl. 158, f. 2]

Larva on Capparis zippeliana [BAUR]

West Irian - New Guinea Cepora abnormis abnormis ; [BAUR, 155]; Winhard, 2000, Butterflies of the world, 10: 22, pl. 34/35, f. 2
C. a. euryxantha (Honrath, 1892)
New Guinea - Papua Delias abnormis var. euryxantha Honrath, 1892; Berl. ent. Z. 36 (2) : 435, TL: New Guinea
Delias euryxantha ; Grose-Smith & Kirby, 1896, Rhop. Exot. [1] 2: (Delias) pl. 6, f. 7-8
Tachyris euryxantha ; Grose-Smith & Kirby, 1896, Rhop. Exot. [1] 2: (Delias) 22
Cepora abnormis euryxantha ; [BAUR, 156]

Cepora eperia (Boisduval, 1836)
-. See [About maps] Pieris eperia Boisduval, 1836; Hist. nat. Ins., Spec. gén. Lépid. 1 : 470
Cepora eperia ; [EBW]; [BOW: pl. 158, f. 5]; [BOR, 149]; Winhard, 2000, Butterflies of the world, 10: 22, pl. 32/33, f. 10; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 112
Sulawesi Ceporia eperia eperia
?Huphina eperia f. pallaminor Martin, 1919; Dt. ent. Z. Iris 33 (1/2) (3/4): 72
Ceporia eperia eperia ; [BOR, 149]
C. e. foa Fruhstorfer˛
Sulawesi (mountains) Ceporia eperia foa ; [BOR, 149]
C. e. flava Iwasaki & Yata, 2005
Buton I. Cepora eperia flava Iwasaki & Yata, 2005; Trans. lepid. Soc. Japan 56 (4) : 307, TL: Baubau, Buton I.

Cepora licea (Fabricius, 1787)
Nias Papilio licea Fabricius, 1787; Mantissa Insectorum 2: 20, TL: Nias
Papilio licaea[sic?] ; (spell.?)
Cepora licea ; Winhard, 2000, Butterflies of the world, 10: 22, pl. 32/33, f. 11

Cepora perimale (Donovan, 1805)
Java - Australia. See [About maps] Australian Gull
Papilio perimale Donovan, 1805; Epitome nat. Hist. Ins. New Holland: pl. 20
Pieris lanassa Boisduval, 1836; Hist. nat. Ins., Spec. gén. Lépid. 1 : 477
Pieris rachel Boisduval, 1836; Hist. nat. Ins., Spec. gén. Lépid. 1 : 469, TL: Java?
Pieris nabis Lucas, 1852; Revue Mag. Zool. (2) 4 (7): 326, TL: Australia
Pieris perithea C. & R. Felder, 1865; Reise Fregatte Novara, Bd 2 (Abth. 2) (2): 169, TL: Fiji[?]
Pieris periclea C. & R. Felder, 1865; Reise Fregatte Novara, Bd 2 (Abth. 2) (2): 169, TL: Australia
Pieris narses Wallace, 1867; Trans. ent. Soc. Lond. (3) 4 (3) : 333, pl. 6, f. 3, TL: Moreton Bay, Australia
Belenois pallida Grose-Smith, 1894; Novit. zool. 1 (2): 336, TL: Biak
Huphina pallescens Grose-Smith, 1899; in Grose-Smith & Kirby, Rhop. Exot. [1] 3: (Huphina) [8] 9 (repl. Belenois pallida Grose-Smith, 1894), TL: Biak
Pieris hartei Ribbe, 1901
Huphina pomentes hierocles Fruhstorfer, 1914; Ent. Rundschau 31 (6) : 33, TL: Yule I.
= Cepora perimale hierocles ; [NHM]
Huphina perimale yuleana Fruhstorfer, 1914; Ent. Rundschau 31 (6) : 33, TL: Yule I.
= ; [NHM]
Cepora perimale, Lepidoptera Larvae of Australia [Don Herbison-Evans]
Huphina periclea ; Grose-Smith & Kirby, 1898, Rhop. Exot. [1] 3: (Huphina) 5, pl. 2, f. 4-5
Huphina pallida ; Grose-Smith & Kirby, 1899, Rhop. Exot. [1] 3: (Huphina) pl. 3, f. 7-8
Huphina perimale rachel ; Rothschild, 1915, Novit. Zool. 22 (1): 111
Huphina perimale ; Grose-Smith & Kirby, 1898, Rhop. Exot. [1] 3: (Huphina) 6, pl. 2, f. 6; Grose-Smith & Kirby, 1899, Rhop. Exot. [1] 3: (Huphina) 9, pl. 3, f. 1-3
Cepora perimale ; [EBW]; [BOW: pl. 158, f. 7]; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 111

Larva on Capparis mitchellii, C. spp. K. L. & L. E. Dunn, 1991, Review Austr. Butts. (1-4): (120-140)

Norfolk I. Cepora perimale perimale ; [BAUR, 155 (text)]
C. p. pitys (Godart, 1819)
Timor Pieris pitys Godart, 1819; Encyclopédie Méthodique. 9 (1): 134, TL: Timor
Cepora perimale pitys ; [BAUR, 154 (text)]
C. p. scyllara (MacLeay, 1826)
N.Australia - Hunter River, New South Wales Pieris scyllara MacLeay, [1826]; in King, Narr. Surv. Austr. 2: 459, TL: [New Holland?]
Pieris lanassa Boisduval, 1836; Hist. nat. Ins., Spec. gén. Lépid. 1 : 477
Pieris scyllara ; Boisduval, 1836, Hist. nat. Ins., Spec. gén. Lépid. 1 : 482
Tachyris scyllaria[sic] ; Wallace, 1867, Trans. ent. Soc. Lond. (3) 4 (3) : 367
Cepora perimale scyllara ; [BAUR, 154]
C. p. acrisa (Boisduval, 1859)
New Caledonia, Loyalty Is. Pieris acrisa Boisduval, 1859; Bull. Soc. ent. Fr. (3) 7 : clvi, TL: Woodlark I.
Huphina perimale hiscia Fruhstorfer, 1914; Ent. Rundschau 31 (6) : 33, TL: Woodlark I.
= ; [NHM]
Cepora perimale acrisa ; [BAUR, 155 (text)]
C. p. perictione (C. & R. Felder, 1865)
Aru Pieris perictione C. & R. Felder, 1865; Reise Fregatte Novara, Bd 2 (Abth. 2) (2): 168, TL: Aru Islands
Cepora perimale perictione ; [BAUR, 154 (text)]
C. p. wallaceana (C. & R. Felder, 1865)
Waigeu Pieris wallaceana C. & R. Felder, 1865; Reise Fregatte Novara, Bd 2 (Abth. 2) (2): 168
Cepora perimale wallaceana ; [BAUR, 154 (text)]
C. p. mentes (Wallace, 1867)
Lombok I. Pieris mentes Wallace, 1867; Trans. ent. Soc. Lond. (3) 4 (3) : 332, TL: Lombok, Flores
Huphina mentes ; Grose-Smith & Kirby, 1898, Rhop. Exot. [1] 3: (Huphina) 1, pl. 1, f. 1-3
C. p. latilimbata (Butler, 1876)
Darnley I, Papua Belenois latilimbata Butler, 1876; Ann. Mag. Nat. Hist. (4) 18 (105) : 247, TL: Port Moresby, New Guinea
Huphina latilimbata ; Grose-Smith & Kirby, 1899, Rhop. Exot. [1] 3: (Huphina) 9, pl. 3, f. 4-6
Cepora perimale latilimbata ; [BAUR, 154 (text)]; Winhard, 2000, Butterflies of the world, 10: 22, pl. 34/35, f. 1
C. p. terranea (Butler, 1877)
Loyalty Is. Belenois terranea Butler, 1877; Ann. Mag. nat. Hist. (4) 20 (118) : 356, TL: Lifu, Loyalty Is.
C. p. consanguis (Butler, 1883)
Tanimber Group Belenois consanguis Butler, 1883; Proc. zool. Soc. Lond. 1883 : 369, TL: Larat I.
Huphina consanguis ; Grose-Smith & Kirby, 1898, Rhop. Exot. [1] 3: (Huphina) 1, pl. 1, f. 4-5
Cepora perimale consanguis ; [BAUR, 154 (text)]
C. p. kuehni (Röber, 1885)
Salayar ?Pieris kühni Röber, 1885; Corr.-Bl. Ent. Ver. Iris 1 (2) : 20, pl. 1, f. 2-3, TL: Moluccas
Cepora perimale kuehni ; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 111
C. p. pygmaea (Röber, 1891)
Leti I. perimale pygmaea (Röber, 1891) (Pieris); Tijdschr. Ent. 34: 279, TL: Letti I.
Cepora perimale pygmaea ; [BAUR, 154 (text)]
C. p. discolor (Mathew, 1887)
Ugi, Ulana Is. Pieris discolor Mathew, 1887; Trans. Ent. Soc. Lond. 1887 (1) : 47, TL: Ugi, Solomon Is.
Cepora perimale discolor ; [BAUR, 155 (text)]
C. p. dohertyana (Grose-Smith, 1894)
West Irian - S.New Guinea Belenois dohertyana Grose-Smith, 1894; Novit. zool. 1 (2): 337, TL: New Guinea
Huphina dohertyana ; Grose-Smith & Kirby, 1898, Rhop. Exot. [1] 3: (Huphina) 6, pl. 2, f. 7-9
Cepora perimale dohertyana ; [BAUR, 154 (text)]
C. p. leucophora (Grose-Smith, 1897)
Trobriand Is. Huphina leucophora Grose-Smith, 1897; Ann. Mag. Nat. hist. (6) 19 : 405, TL: Kirwini I., Trobriands
Huphina leucophora ; Grose-Smith & Kirby, 1898, Rhop. Exot. [1] 3: (Huphina) 2, pl. 1, f. 9-11
Cepora perimale leucophorus ; [BAUR, 154 (text)]
C. p. wetteri (Grose-Smith, 1898)
Wetar I. Huphina wetteri Grose-Smith, 1898; in Grose-Smith & Kirby, Rhop. Exot. [1] 3: (Huphina) 2, pl. 1, f. 6-8, TL: Wetter; Dili
?Cepora perimale wetterensis ; [BAUR, 154 (text)]
C. p. agnata (Grose-Smith, 1889)
Guadalcanal, Roviana Pieris agnata Grose-Smith, 1889; Ent. mon. Mag. 25 : 301, TL: Rubiana; Guadalcanar
Huphina agnata ; Grose-Smith & Kirby, 1898, Rhop. Exot. [1] 3: (Huphina) 4, pl. 2, f. 1-3
Cepora perimale agnata ; [BAUR, 155 (text)]
C. p. maculata (Grose-Smith, 1896)
New Britain Tachyris maculata Grose-Smith, 1896; Ann. Mag. Nat. Hist. (6) 17 (101) : 374
Huphina maculata ; Grose-Smith & Kirby, 1899, Rhop. Exot. [1] 3: (Huphina) 10, pl. 3, f. 9-10
Cepora perimale maculata ; [BAUR, 155 (text)]
C. p. synchroma (Röber, 1891)
Flores, Alor Pieris synchroma Röber, 1891; Tijdschr. Ent. 34: 278, TL: Flores, Alor
Cepora perimale synchroma ; [NHM]
C. p. macdonaldi (Ribbe, 1898)
Bougainville Tachyris macdonaldi Ribbe, 1898; Dt. ent. Z. Iris 11 (1): 91
Cepora perimale macdonaldi ; [BAUR, 155 (text)]
C. p. quadricolor (Salvin & Godman, 1877)
New Britain, New Ireland, New Hanover, Duke of York Group Pieris quadricolor Salvin & Godman, 1877; Proc. zool. Soc. Lond. 1877 : 147, pl. 23, f. 3-4, TL: Bismarck Is.
Cepora perimale quadricolor ; [BAUR, 155]
Cepora quadricolor ; [EBW]
C. p. mithra (Fruhstorfer, 1903)
Fergusson I. Huphina pitys mithra Fruhstorfer, 1903; Soc. Ent. 18 (3) : 17, TL: Fergusson I.
Huphina pitys mithra Fruhstorfer, 1903; Berl. ent. Zs. 48 (1/2) : 102, TL: Fergusson I.
Cepora perimale mithra ; [BAUR, 154 (text)]
C. p. pityna (Fruhstorfer, 1903)
Damar I. Huphina pitys pityna Fruhstorfer, 1903; Berl. ent. Zs. 48 (1/2) : 105, TL: Dammer I.
Cepora perimale pityna ; [BAUR, 154 (text)]
C. p. babberica (Fruhstorfer, 1903)
Babar I. Huphina pitys babberica Fruhstorfer, 1903; Berl. ent. Zs. 48 (1/2) : 106, TL: Babber
Cepora perimale babberica ; [BAUR, 154 (text)]
C. p. bolana (Fruhstorfer, 1903)
Kai I. Huphina pitys bolana Fruhstorfer, 1903; Berl. ent. Zs. 48 (1/2) : 104, TL: Key-Tual
Cepora perimale bolana ; [BAUR, 154 (text)]
C. p. chrysopis Fruhstorfer, 1910˛
Neomfor I, Geelvink Bay perimale chrysopis Fruhstorfer, 1910˛; in Seitz, Gross-Schmett. Erde 9 : 143, TL: Mefor I.
Cepora perimale chrysopis ; [BAUR, 154 (text)]
C. p. toekangbesiensis (Jurriaanse & Lindemans, 1920)
Tukangbesi Huphina perimale toekangbesiensis Jurriaanse & Lindemans, 1920; Tijdschr. Ent. 62 (Suppl.) : 7, TL: Toekan Bessi Arch.
?Huphina perimale toekangbesiensis Jurriaanse, 1920; Tijdschr. Ent. 62 : xlvii
Cepora perimale toekangbesiensis ; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 111
Cepora perimale tuekangbesiensis[sic] ; Iwasaki & Yata, 2005, Trans. lepid. Soc. Japan 56 (4) : 309
C. p. radiata Howarth, 1962
Bellona I., Somonons, Buton I. perimale radiata Howarth, 1962; The Natural History of the Rennel and Bellona Is. (4) : 64, TL: Bellona I.
Cepora perimale radiata ; [BAUR, 155 (text)]

Cepora laeta (Hewitson, 1862)
Timor. See [About maps] Pieris laeta Hewitson, 1862; Ill. exot. Butts [1] (Pieris VII): [41], pl. [22], f. 45-46, TL: Timor
Pieris laetissima Hewitson, 1866; Ill. exot. Butts [3] : iv (repl. Pieris laeta Hewitson, 1862)
Cepora laeta ; [EBW]
Cepora lacta[sic] ; [BOW: pl. 158, f. 8 (missp.)]
Cepora laeta ; [BAUR, 155]; Winhard, 2000, Butterflies of the world, 10: 22, pl. 32/33, f. 15

The ? species-group

Cepora nadina (Lucas, 1852)
Ceylon, W.Ghats, Sikkim - Assam, Burma. See [About maps] Lesser Gull
Pieris nadina Lucas, 1852; Revue Mag. Zool. (2) 4 (7): 333, TL: Khasi Hills
Huphina liquida Swinhoe, 1890; Ann. Mag. nat. Hist. (6) 5 (29) : 361, TL: Mahableshwur
= ; [NHM]
Cepora nadina nadina, Butterflies in Indo-China [Yutaka Inayoshi]
Huphina nadina ; [BIR, 421]
Cepora nadina ; [BOW: pl. 158, f. 11]; [MRS, 256]

Larva on Capparis [BIR]

Sikkim - Indo-China, Thailand, S.Yunnan Pieris amba Wallace, 1867; Trans. ent. Soc. Lond. (3) 4 (3) : 340, TL: N.India
Appias amboides Moore, 1884; J. asiat. Soc. Bengal 53 Pt.II (1) : 31, 46, TL: Silhet, Assam
Pieris nama Moore, 1857; Proc. zool. Soc. Lond. 1857 (333) : 102, pl. 44, f. 1-2
Pieris nama Moore, [1858]; in Horsfield & Moore, Cat. lep. Ins. Mus. East India Coy 1 : 76, TL: N.India
Cepora nadina nadina ; [BOR, 150]
Pieris nama ; Hewitson, 1861, Ill. exot. Butts [1] (Pieris III-VI): [39], pl. [21], f. 37
C. n. remba (Moore, [1858])
S.India Pieris remba Moore, [1858]; in Horsfield & Moore, Cat. lep. Ins. Mus. East India Coy 1 : 75
Huphina remba ; Moore, [1881], Lepid. Ceylon 1 (4): 137, (3): pl. 53, f. 2, 2a
Cepora nadina remba ; [BOR, 150]
Ceylon Cepora nadina cingala ; [BOR, 150]
Andamans Cepora nadina andamana ; [BOR, 150]
C. n. andersoni (Distant, 1885)
Peninsular Malaya Appias andersoni Distant, 1885; Entomologist 18 : 146
Cepora nadina andersoni ; [BOR, 150]; [BMP: 87, pl. 8, f. 3-4]
Sumatra Cepora nadina fawcetti ; [BOR, 150]
C. n. eunama (Fruhstorfer, 1903)
Taiwan Huphina nama eunama Fruhstorfer, 1903; Berl. ent. Zs. 48 (1/2) : 110, TL: S.Formosa, Takau
Cepora nadina eunama ab. discoloris Murayama, 1961; Tyô to Ga 11 (4): 55, f. 9, TL: Poli, Formosa
Cepora nadina eunama ; [MRS, 256]; [BOR, 150]

Larva on Capparis acutifolia, C. sikkimensis [MRS]

C. n. hainanensis (Fruhstorfer, 1913)
Huphina nama hainanensis Fruhstorfer, 1913; Ent. Rundschau 30 (16): 92, TL: Hainan
Cepora nadina hainanensis ; [MRS, 256]

Cepora temena (Hewitson, 1861)
-. See [About maps] Pieris temena Hewitson, 1861; Ill. exot. Butts [1] (Pieris III-VI): [33], pl. [18], f. 19, TL: Lombock
Cepora temena ; [BOW: pl. 158, f. 12]; [BOR, 152]
Lombok Cepora temena temena ; [BOR, 152]
C. t. tamar (Wallace, 1867)
Java, Bali Pieris tamar Wallace, 1867; Trans. ent. Soc. Lond. (3) 4 (3) : 337, TL: Bali
Cepora temena tamar ; [BOR, 152]
C. t. lenitas Fruhstorfer, 1910˛
Sumbawa Cepora temena lenitas ; [BOR, 152]; Winhard, 2000, Butterflies of the world, 10: 22, pl. 32/33, f. 14
C. t. hyele Fruhstorfer˛
Lomblen Cepora temena hyele ; [BOR, 152]

Cepora timnatha (Hewitson, 1862)
-. See [About maps] Pieris timnatha Hewitson, 1862; Ill. exot. Butts [1] (Pieris VII): [41], pl. [22], f. 47-48, TL: Tondano
Cepora timnatha ; [BOR, 153]; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 111
N.Sulawesi Cepora timnatha timnatha ; [BOR, 153]; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 111
C. t. soror (Fruhstorfer, 1899)
Sula Is. Pieris eperia soror Fruhstorfer, 1899; Soc. Ent. 14 (2) : 10, TL: Sula Is.
Pieris eperia soror Fruhstorfer, 1899; Berl. ent. Zs. 44 (1/2) : 51, TL: Sula Is.
Cepora timnatha soror ; [BOR, 153]; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 111
C. t. aurulenta (Fruhstorfer, 1899)
Bangkai Pieris eperia aurulenta Fruhstorfer, 1899; Soc. Ent. 14 (2) : 10, TL: Bangkai
Cepora timnatha aurulenta ; [BOR, 153]; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 111
C. t. filiola (Fruhstorfer, 1899)
Sula Besi Pieris eperia filiola Fruhstorfer, 1899; Berl. ent. Zs. 44 (1/2) : 52
Cepora timnatha filiola ; [BOR, 153]; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 111
C. t. filia (Fruhstorfer, 1902)
S.Sulawesi Huphina timnatha filia Fruhstorfer, 1902; Dt. ent. Z. Iris 15 (1) : 169, TL: Samanga, S.Celebes
Cepora timnatha filia ; [BOR, 153]; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 111
C. t. butona Iwasaki & Yata, 2005
Buton I. Cepora timnatha butona Iwasaki & Yata, 2005; Trans. lepid. Soc. Japan 56 (4) : 306, TL: Baubau, Buton I.

Cepora julia (Doherty, 1891)
-. See [About maps] Huphina julia Doherty, 1891; J. Asiat. Soc. Bengal 60 Pt.II (2) : 187, pl. 2, f. 12, TL: Sumba
Pieris pagenstecheri Röber, 1900; Ent. Nachr. 26 (13) : 199
= ; [NHM]
Cepora julia ; [BOR, 153]
Sumba Cepora julia julia ; [BOR, 153]
C. j. calliparga Fruhstorfer˛
Sumbawa Cepora julia calliparga ; [BOR, 153]

Cepora celebensis (Rothschild, 1892)
Sulawesi. See [About maps] Huphina celebensis Rothschild, 1892; Dt. ent. Z. Iris 5 (2) : 439, pl. 4, f. 1-2, TL: S.Celebes
Cepora celebensis ; [BOW: pl. 158, f. 5 (text)]; [BOR, 149]; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 111
Sulawesi, Salayar, Banggai Cepora celebensis celebensis ; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 111
C. c. kazuyoe Watanabe, 1987
Buton Cepora celebensis kazuyoe ; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 111

Cepora eurygonia (Hopffer, 1874)
Toga I. Pieris eurygonia Hopffer, 1874; Stettin ent. Ztg 35 (1-3) : 23, TL: Togian I.
Cepora eurygonia ; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 111

Cepora boisduvaliana (C. & R. Felder, 1862)
-. See [About maps] Pieris boisduvaliana C. & R. Felder, 1862; Wien. ent. Monats. 6 (9) : 287, TL: Luzon
Pieris nephele C. & R. Felder, 1861; Wien. ent. Monats. 5 (10) : 299, TL: Luzon
Cepora boisduvaliana ; [BOR, 150]
Philippines Cepora boisduvaliana boisduvaliana ; [BOR, 150]
C. b. semperi Staudinger˛
Mindanao Cepora boisduvaliana semperi ; [BOR, 150]
C. b. cirta Fruhstorfer˛
Bohol Cepora boisduvaliana cirta ; [BOR, 150]
Camiguin de Mindanao Cepora boisduvaliana balabonga ; [BOR, 150]
Romblon Is. Cepora boisduvaliana sibuyanensis ; [BOR, 150]
Cebu Cepora boisduvaliana cebuensis ; [BOR, 150]

Cepora pactolicus (Butler, 1865)
Borneo. See [About maps] Pieris pactolicus Butler, 1865; Proc. Zool. Soc. Lond. 1865 (2) : 455, pl. 26, f. 1, TL: "Bogota" [? error, = Borneo]


Unknown or unplaced taxa

Huphina pitys aelia Fruhstorfer, 1903; Berl. ent. Zs. 48 (1/2) : 102, TL: Kangean

Hyphina pitys lucia Fruhstorfer, 1903; Berl. ent. Zs. 48 (1/2) : 103, TL: Sumbawa, Tambora

Huphina nama var. koannania Matsumura, 1909; Ent. Zs. 23 (18) : 88, TL: Kanshirei; Horisha

Huphina timnatha lamponga Martin, 1919; Dt. ent. Z. Iris 33 (1/2) (3/4): 78

Pieris amarella Wallace, 1867; Trans. ent. Soc. Lond. (3) 4 (3) : 373, pl. 9, f. 2, TL: New Caledonia


4.5.2013 (17)

References:
[About maps]
Warning! The maps are automatically generated from the textual information, and the process does not always produce acceptable result; See about maps for more info.
[BAUR]
D'Abrera, 1977; Butterflies of the Australian Region (2nd edn)
[BHL]
The; Biodiversity Heritage Library; and on; Internet Archive; Note: The linking to the content of Internet Archive is somewhat delicate -- notify me when links break!
[BIR]
Wynter-Blyth, 1957; Butterflies of the Indian Region; (1982 Reprint)
[BMP]
Eliot (Ed.), Corbet & Pendlebury, 1992; The Butterflies of The Malay Peninsula; 4th edition; ISBN 983-9681-05-2
[BOR]
D'Abrera, 1982, 1985, 1986; Butterflies of the Oriental Region, Part I-III
[BOW]
Lewis, 1974; Butterflies of the World; ISBN 0 245 52097 X
[CiNii]
http://ci.nii.ac.jp/
[Don Herbison-Evans]
and Stella A. Crossley; Lepidoptera Larvae of Australia; e-mail: Don Herbison-Evans; e-mail: Don Herbison-Evans
[EBW]
Smart, Paul, 1976; The Illustrated Encyclopedia of the Butterfly World; ISBN 0 600 31381 6
[Kedar Tokekar]
India
[LAMAS-2005]
Gerardo Lamas; Entomofauna 26 (6): 57-100; A Bibliography of the Zoological Publications of Hans Fruhstorfer (1886* - 1922+)
[LAMAS-NEO]
Gerardo Lamas; Bibliography of butterflies; An Annocated Bibliography of the Neotropical Butterflies and Skipper (Lepididoptera: Papilionoidea and Hesperioidea); Revised Electronic Edition (2007)
[MRS]
Chou Io (Ed.); Monographia Rhopalocerum Sinensium, 1-2
[NBC]
Netherlands Centre for Biodiversity Naturalis; NBC naturalis
[NHM]
Natural History Museum; The Global Lepidoptera Names Index
[Oleg Kosterin]
Institue of Cytology & Genetics; Siberian Division of Russian Academy of Sciences; Novosibirsk; e-mail: Oleg Kosterin; http://pisum.bionet.nsc.ru/kosterin/
[Yutaka Inayoshi]
A Check List of Butterflies in Indo-China; Chiefly from THAILAND, LAOS & VIETNAM
[˛]
This may require parentheses or not. I don't have the necessary information for this taxon.


Some related literature:
Billberg, 1820
Enumeratio Insectorum Enum. Ins. Mus. Billb. : 1-138
Boisduval, 1859
[communications] Bull. Soc. ent. Fr. (3) 7 : cliv-clviii
Boisduval, 1836
Histoire Naturelle des Insectes. Species Général des Lépidoptéres. Tome Premier Hist. nat. Ins., Spec. gén. Lépid. 1 : 1-690, pl. 1-24
Butler, 1865
Description of six new species of exotic butterflies in the collection of the British Museum Proc. Zool. Soc. Lond. 1865 (2) : 455-459, pl. 26
Butler, 1876
On a Collection of Lepidoptera from Port Moresby, New Guinea Ann. Mag. Nat. Hist. (4) 18 (105) : 240-249
Butler, 1877
On a Collection of Lepidoptera obtained by the Rev. S. J. Whitmee from Lifu (Loyalty Group), with Descriptions of new Species Ann. Mag. nat. Hist. (4) 20 (118) : 348-359
Butler, 1883
List of Lepidoptera collected by Mr. H. O. Forbes in the Islands of Timor Laut Proc. zool. Soc. Lond. 1883 : 365-371, pl. 38
Butler, 1899
A Revision of the Pierine Genus Huphina, with Notes on the Seasonal Phases and Descriptions of new Species Ann. Mag. nat. Hist. (7) 3 (15) : 201-214
Corbet, 1937
Observations on Species of Papilionidae, Pieridae and Danaidae from the Malay Peninsula Proc. R. ent. Soc. Lond. (B) 6 (3): 45-52
Cramer, [1775]
Uitlandsche Kapellen (Papillons exotiques) Uitl. Kapellen 1 (1-7): 1-132, pl. 1-84 (1775), (8): 133-155, pl. 85-96 (1776)
Cramer, [1779]
Uitlandsche Kapellen (Papillons exotiques) Uitl. Kapellen 3 (17-21): 1-104, pl. 193-252 (1779), (22): 105-128, pl. 253-264 ([1780]), (23-24): 129-176, pl. 265-288 (1780)
Distant, 1885
Description of a new species of Pierinae from the Malay Peninsula Entomologist 18 : 146-147
Doherty, 1891
The Butterflies of Sumba and Sumbawa, with some Account of the Island of Sumba J. Asiat. Soc. Bengal 60 Pt.II (2) : 141-197, pl. 2
Donovan, 1805
An Epitome of the Natural History of the Insects of New Holland, New Zealand, New Guinea, Otaheite and other Islands in the Indian, Sothern and Pacific Oceans ... Epitome nat. Hist. Ins. New Holland: 41pls
Doubleday, 1846
Descriptions of new or imperfectly described Diurnal Lepidoptera Ann. Mag. nat. Hist. 17 (109) : 22-26
K. L. & L. E. Dunn, 1991
Review of Australian Butterflies: distribution, life history and taxonomy. Pushlished by authors, Melbourne Review Austr. Butts. (1-4):
van Eecke, 1918
Studies on Indo-Australian Lepidoptera--III. Some Rhopalocera and Netrocera[sic] from Simalur, Pulu Lasia, Pulu Babi and Sumatra Zool. Meded. 4 (7) : 70-101, 2pls
Eschscholtz, 1821
Beschreibung neuer ausländischer Schmetterling in Kotzebue, Entdeck. Reise Süd-See 3 (App. 5): 201-219, 11pls
Fabricius, 1775
Systema Entomologiae, sistens Insectorum Classes, Ordines, Genera, Species, Adiectis Synonymis, Locis, Descriptionibus, Observationibus. Syst. Ent.: 832pp
C. & R. Felder, 1861
Lepidoptera nova a Dr. Carolo Semper in Insulis Philippinis collecta diagnosibus. (1) Wien. ent. Monats. 5 (10) : 297-306
C. & R. Felder, 1862
Lepidoptera nova a Dr. Carolo Semper in Insulis Philippinis collecta diagnosibus. (2) Wien. ent. Monats. 6 (9) : 282-294
C. & R. Felder, 1865
Reise der österreichischen Fregatte Novara um die Erde in den Jahren 1857, 1858, 1859 unter den Behilfen des Commodore B. von Wüllerstorf-Urbair. Zoologischer Theil. Band 2. Abtheilung 2. Lepidoptera. Rhopalocera Reise Fregatte Novara, Bd 2 (Abth. 2) (1): 1-136, pl. 1-21 (Rhop.) (C. & R. Felder, [1865]), (2): 137-378, pl. 22-47 (Rhop.) (C. & R. Felder, 1865), (3): 379-536, pl. 48-74 (Rhop.) (C. & R. Felder, 1867), (4): 537-548, pl. 75-120, (Inhalts-Verz.) 1-9 (pl. 1-74), (Felder & Rogenhofer, 1874) , (5): pl. 121-140, (erklärung) 1-10 (pl. 75-107), (Inhalts-Verz.) 1-20 (pl. 108-140), (Felder & Rogenhofer, 1875)
Fruhstorfer, 1897
Neue Rhopaloceren aus dem malayischen Archipel Berl. Ent. Zs. 42 (1/2) : 113-118 (mid November)
Fruhstorfer, 1899
Eine seltene Pieride vom Kina-Balu Soc. Ent. 14 (2) : 10 (15 April)
Fruhstorfer, 1899
Neue asiatische Lepidopteren Berl. ent. Zs. 44 (1/2) : 49-64 (mid July)
Fruhstorfer, 1899
Pieriden-Studien Berl. ent. Zs. 44 (1/2) : 100-104 (mid July)
Fruhstorfer, 1902
Neue Lepidopteren aus dem Indo-Malayischen Gebiet Dt. ent. Z. Iris 15 (1) : 169-178 (early September)
Fruhstorfer, 1903
Verzeichniss der in Tonkin, Annam und Siam gesammelten Pieriden und Besprechung verwandter Formen Dt. ent. Z. Iris 15 (2) : 269-305 (1 May)
Fruhstorfer, 1903
Neue Pieriden aus dem Indo-Australischen Gebiet Soc. Ent. 18 (3) : 17-18 (1 May)
Fruhstorfer, 1903
Neue Pieriden und Uebersicht verwandter Formen Berl. ent. Zs. 48 (1/2) : 97-111 (early May)
Fruhstorfer, 1904
Neue Rhopaloceren aus dem Malayischen Archipel Ent. Meddel. 2 (5/6) : 291-332, 5 figs. (June)
Fruhstorfer, 1910
2. Familie: Pieridae, Weisslinge in Seitz, Gross-Schmett. Erde 9 : 119-120 (1909), : 121-144 (14 January, 1910), : 145-152 (11 February, 1910), : 153-168 (2 April, 1910), : 169-176 (4 April, 1910), : 177-190 (20 May, 1910)
Fruhstorfer, 1913
Neue indo-australische Rhopaloceren Ent. Rundschau 30 (16): 91-92 (23 August), (21): 124-125 (8 November)
Fruhstorfer, 1914
Neue Pieriden Ent. Rundschau 31 (6) : 32-33 (28 March)
Godart, 1819
in Latreille & Godart, Encyclopédie Méthodique. Histoire naturelle Entomologie, ou histoire naturelle des crustacés, des arachnides et des insectes Encyclopédie Méthodique. 9 (1): 3-328 (1819), (2): 329-828 ([1824])
Salvin & Godman, 1877
On a Collection of Lepidoptera made by he Rev. G. Brown of Duke-of-York Island and its Neighbourhood Proc. zool. Soc. Lond. 1877 : 139-151, pl. 22-23
Grose-Smith, 1889
Descriptions of new species of butterflies captured by Mr. C.M. Woodford in the Solomon Islands Ent. mon. Mag. 25 : 299-303
Grose-Smith, 1894
An Account of a Collection of Diurnal Lepidoptera from New Guinea (Parts I-III) - made by Mr. W. Doherty at Humboldt Bay, Dutch New Guinea, and in neighbouring islands, in the Museum of the honourable Walter Rothschild at Tring, with descriptions of new species Novit. zool. 1 (2): 331-365, (2): 543-551, (3): 571-584
Grose-Smith, 1896
Descriptions of Two new Species of Butterflies from New Britain Ann. Mag. Nat. Hist. (6) 17 (101) : 374-375
Grose-Smith, 1897
Descriptions of further new Species of Butterflies from the Pacific Islands Ann. Mag. Nat. hist. (6) 19 : 403-407
Grose-Smith & Kirby, 1889; Grose-Smith & Kirby, 1896; Grose-Smith & Kirby, 1898; Grose-Smith & Kirby, 1899; Grose-Smith in Grose-Smith & Kirby, 1898; Grose-Smith in Grose-Smith & Kirby, 1899
Rhopalocera Exotica, being Illustrations of New, Rare, and Unfigured Species of Butterflies Rhop. Exot. [1] 1: (Ornithoptera) 1-2, pl. 1 (1887), [1] 2: (Ornithoptera) 3-4, pl. 2 (1892), [1] 3: (Ornithoptera) [5-6] 1-2, pl. 3 (1900), [1] 3: (Ornithoptera) 7-8, pl. 4 (1901), [1] 1: (Papilio) 1-2, pl. 1 (1887), [1] 1: (Papilio) 3-5,7-14, pl. 2-6 (1888), [1] 1: (Papilio) 15-20, pl. 7-9 (1890), [1] 1: (Papilio) 21-25, pl. 10-11 (1891), [1] 2: (Papilio) 27-31, pl. 12-13 (1893), [1] 2: (Papilio) 33-35, pl. 14 (1894), [1] 2: (Papilio) 37-38, pl. 15 (1895), [1] 2: (Papilio) 39-40, pl. 16 (1897), [1] 3: (Papilio) 41-46, pl. 17-19 (1899), [1] 3: (Papilio) 47-48, pl. 20 (1900), [1] 3: (Papilio) 49-50, pl. 21 (1901), [1] 3: (Papilio) 51-55, pl. 22-23 (1902), [1] 1: (Delias) 1-3, pl. 1 (1889), [1] 2: (Delias) 5-7, 9-11, pl. 2-3 (1893), [1] 2: (Delias) 12-18, pl. 4-5 (1895), [1] 2: (Delias) 20-22, pl. 6 (1896), [1] 3: (Delias) 24-26, pl. 7 (1897), [1] 3: (Delias) 28-35, pl. 8-9 (1901), [1] 3: (Delias) 36-38, pl. 10 (1902), [1] 1: (Pieridae) 1-3, pl. 1 (1888), [1] 2: (Pinacopteryx) 1-4, pl. 1 (1893), [1] 1: (Callosune) 1-3, pl. 1 (1889), [1] 3: (Huphina) 1-7, pl. 1-2 (1898), [1] 3: (Huphina) [8],9-10, pl. 3 (1899), [1] 1: (Elodina) 1-4, pl. 1 (1890), [1] 1: (Belenois) 1-4, pl. 1 (1892), [1] 2: (Belenois) 5-8, pl. 2 (1892), [1] 3: (Mylothris) 1-3, 5-7, pl. 1-2 (1900), [1] 1: (Ixias) 1-3, pl. 1 (1888), [1] 2: (Appias) 1-3, pl. 1 (1894), [1] 1: (Pierinae) 5-7, pl. 2 (1889), [1] 2: (Dismorphia) 1-4, pl. 1 (1896), [1] 2: (Dismorphia) 5-9, 11-15, pl. 2-3 (1897), [2] 1: (Asthipa) 1-3, pl. 1 (1890), [2] 2: (Asthipa) 5-6, pl. 2 (1896), [2] 2: (Ravadeba) 7, pl. 1 (1896), [2] 2: (Mycalesis) 1-4, pl. 1 (1895), [2] 2: (Mycalesis) 5-12, pl. 2-3 (1896), [2] 3: (Mycalesis) 13-18, pl. 4-5 (1902), [2] 3: (Libythea) 1-2, pl. 1 (1902), : (Ragadia-Argyronympha-Hypocysta) : 1-6, pl. [1] (1895), [2] 2: (Pedaliodes) 1-5, pl. 1 (1893), [2] 2: (Pedaliodes) 7-9, pl. 2 (1894), [2] 2: (Pedaliodes) 10-14, pl. 3 (1895), [2] 3: (Pedaliodes) 15-20, pl. 4-5, (Oxeoschistus) 1 (1900), [2] 3: (Pronophila) 1-2, pl. 1, (Oxeoschistus) 2-3, pl. 1 (1900), [2] 2: (Elymnias) 1-4, pl. 1 (1896), [2] 3: (Elymnias) 5-6, pl. 2, (Bruasa) 6-7, pl. 1 (1897), [2] 1: (Morpho) 1-2, pl. 1 (1890), [2] 3: (Morphotenaris-Hyantis) 1-3, pl. [1] (1898), [2] 2: (Tenaris) 1-3, 5-7, pl. 1-2 (1894), [2] 2: (Tenaris) 9-11, pl. 3 (1895), [2] 2: (Tenaris) 13-15, pl. 4 (1896), [2] 3: (Tenaris) 17-18, pl. 5 (1898), [2] 1: (Heliconius) 1-3, pl. 1 (1892), [2] 1: (Acraea) 1-6, pl. 1-2 (1889), [2] 1: (Acraea) 7-10, pl. 3 (1892), [2] 2: (Acraea) 11-14, pl. 4 (1892), [2] 2: (Acraea) 15-18, pl. 5 (1894), [2] 3: (Acraea) 19-29, pl. 6-8, (Planema) 29-30, pl. [1] (1901), [2] 1: (Cethosia) 1-2, pl. 1 (1889), [2] 3: (Cirrhochroa) 1, pl. [1], (Cupha) 2-4, pl. [1] (1898), [2] 2: (Terinos) 1-2, pl. 1 (1894), [2] 1: (Thaleropis) 1-3, pl. 1 (1891), [2] 2: (Doleschallia) 1-3,5-7, pl. 1-2 (1893), [2] 2: (Kallima) 1-3, pl. 1 (1892), [2] 1: (Hypolimnas) 1-2, pl. 1 (1891), [2] 3: (Hypolimnas) 3-4, pl. 2 (1897), [2] 3: (Hypolimnas) 5-6, pl. 3 (1898), [2] 1: (Euxanthe) 1-3, pl. 1 (1890), [2] 1: (Hestina) 1-3, pl. 1 (1891), [2] 1: (Cymothoe) 1-4, pl. 1 (1890), [2] 3: (Limenitis) 1-3, pl. 1 (1899), [2] 2: (Neptis) 1-4, pl. 1 (1895), [2] 3: (Neptis) 5-11, pl. 2-3 (1899), [2] 3: (Neptis) 13-17, pl. 4-5 (1900), [2] 1: (Euthalia-Tanaecia) 1-3, pl. 1 (1889), [2] 1: (Euthalia) 5-10, pl. 2-3 (1891), [2] 2: (Euthalia) 11-14, pl. 4 (1894), [2] 3: (Euthalia) 15, pl. 5, (Dichorragia) 16, pl. 1 (1898), [2] 1: (Apatura) 1-3, pl. 1 (1892), [2] 1: (Euphaedra) 1-2, pl. 1 (1889), [2] 1: (Euryphene) 1-3, pl. 1 (1889), [2] 1: (Charaxes) 1-2,[3-4] pl. 1-2 (1887), [2] 1: (Charaxes) [5], pl. 3 (1888), [2] 1: (Charaxes) 7-9, pl. 4 (1890), [2] 1: (Charaxes) 11-13, pl. 5 (1891), [2] 1: (Mynes) 1-2, pl. 1 (1888), [2] 2: (Mynes) 3-6, pl. 2 (1894), [2] 2: (Prothoe) 1-2, pl. 1 (1896), [3] 3: (Erycinidae) 1-10, pl. 1-2 (1902), [3] 2: (Dicallaneura-etc) 1-6, pl. 1 (1897), [3] 3: (Dicallaneura-etc) 7-11, pl. 2 (1901), [3] 1: (Lycaenidae) 1-4,[5-9] 1-5, pl. 1-2 (1887), [3] 1: (Lycaenidae) 11-17, 19-22, pl. 3-5 (1888), [3] 1: (Lycaenidae) 23-25, 27-29, 31-33, pl. 6-8 (1889), [3] 1: (Lycaenidae) 35-48, pl. 9-11 (1890), [3] 1: (Lycaenidae) 49-51,53-64, pl. 12-15 (1891), [3] 2: (Lycaenidae) 65-68, pl. 16 (1892), [3] 1: (Lycaenidae) [69-72] 57-60, pl. 17 (1892), [3] 2: (Lycaenidae) [73-77] 61-65,79-84, pl. 18-19 (1892), [3] 2: (Lycaenidae) 85-107, pl. 20-23 (1893), [3] 2: (Lycaenidae) 109-120, pl. 24-25 (1894), [3] 3: (Lycaenidae) 121-125, pl. 26 (1900), [3] 3: (Lycaenidae) 127-131, 133-141, pl. 27-29 (1901), [3] 3: (Lycaenidae) 143-146, pl. 30 (1902), [3] 2: (Hypochrysops) 1-6, pl. 1 (1894), [3] 2: (Hypochrysops) 7-13, 15-22, pl. 2-3 (1895), [3] 2: (Thysonotis) 23-25, 27-31, pl. 1-2 (1895), [3] 2: (Waigeum) 34-38, pl. 1 (1896), [3] 2: (Thysonotis) 39-46, pl. 3-4 (1896), [3] 2: (Holochila) 1-4, pl. 1 (1896), [3] 2: (Holochila) 5-8, pl. 2 (1897), [3] 2: (Pseudonotis) 1-3, pl. [1], (Epimastidia) 4-5, pl. [1], (Lampides) 5-8, pl. [1] (1897), [3] 2: (Lampides) 9-10, pl. 2, (Thysonotis) 47, pl. 5 (1897), [3] 3: (Holochila) 9-11, pl. 3 (1898), [3] 3: (Arhopala) 1-7, pl. 1-2 (1898), [3] 3: (Arhopala) 9-12, pl. 3, (Ilerda) 13, pl. [1], (Horaga) 14-15, pl. [1] (1902), [3] 3: (Thysonotis) 47x,48-50, pl. 6 (1898), [3] 3: (Hypochrysops) 23-26, pl. 4 (1898), [3] 3: (Holochila) 13-15, pl. 4 (1899), [3] 3: (Hypochrysops) 27-29, pl. 5, (Waigeum) [30], pl. 2 (1899), [3] 3: (Holochila) 17-19, pl. 5 (1899), [3] 3: (Thysonotis) 51-54, pl. 7 (1899), [3] 3: (Jamides) 11-14, pl. 1 (1900), [3] 3: (Hypolycaena-etc) 1-4, pl. [1] (1900), [3] 3: (Hypochrysops) 31-33, pl. 6, (Thysonotis) 55-56, pl. 8 (1900), [3]: (Additions-Corrections) 1-2,[3],vii-x
Hagen, 1894
Verzeichnis der von mir auf Sumatra gefangenen Dt. ent. Z. Iris 7 (1) : 1-41
Hewitson, 1861; Hewitson, 1862
Illustrations of new species of exotic Butterflies selected chiefly from the collections of W. Wilson Saunders and William C. Hewitson. Ill. exot. Butts [1] : (Systematic Index) I, (Ornithoptera and Papilio): [1-2], pl. [1] (1855), (Papilio II): [3-4], pl. [2] (1856), (Papilio III): [5-6], pl. [3] (1858), (Papilio IV): [7-8], pl. [4] (1859), (Papilio V): [9-10], pl. [5] (1861), (Papilio VI): [11-12], pl. [6] (1864), (Papilio VII-VIII): [13-16], pl. [7-8] (1865), (Papilio IX): [17-18], pl. [9] (1868), (Papilio X-XII): [19-22], pl. [10-12] (1869), (Papilio XIII): [23-24], pl. [13] (1873), (Papilio XIV): [25-26], pl. [14] (1875), (Papilio XV): [27-28], pl. [15] (1877), (Pieris I): [29-30], pl. [16] (1852), (Pieris II): [31-32], pl. [17] (1853), (Pieris III-VI): [33-40], pl. [18-21] (1861), (Pieris VII): [41-42], pl. [22] (1862), (Pieris VIII): [43-44], pl. [23] (1866), (Euterpe): [45-46], pl. [24] (1860), (Euterpe II): [47-48], pl. [25] (1872), (Euterpe and Leptalis): [49-50], pl. [26] (1853), (Leptalis II): [51-52], pl. [27] (1857), (Leptalis III): [53-54], pl. [28] (1858), (Leptalis IV): [55-56], pl. [29] (1860), (Leptalis V-VII): [57-62], pl. [30-32] (1870), (Callidryas and Eronia): [63-64], pl. [33] (1867), (Euploea): [65-66], pl. [34] (1858), (Euploea II): [67-68], pl. [35] (1866), (Heliconia I): [69-70], pl. [36] (1853), (Heliconia II-III): [71-74], pl. [37-38] (1854), (Heliconia IV): [75-76], pl. [39] (1858), (Heliconia V): [77-78], pl. [40] (1867), (Heliconia VI): [79-80], pl. [41] (1871), (Heliconia and Tithorea VII): [81-82], pl. [42] (1873), (Heliconia VIII): [83-84], pl. [43] (1875), (Athesis): [85-86], pl. [44] (1872), (Mechanitis I): [87-88], pl. [45] (1856), (Mechanitis II-III): [89-92], pl. [46-47] (1860), (Ithomia I): [93-94], pl. [48] (1851), (Ithomia II): [95-96], pl. [49] (1852), (Ithomia III): [97-98], pl. [50] (1853), (Ithomia IV-V): [99-102], pl. [51-52] (1854), (Ithomia VI-XIII): [103-118], pl. [53-60] (1855)
Hewitson, 1866
Illustrations of new species of exotic Butterflies selected chiefly from the collections of W. Wilson Saunders and William C. Hewitson Ill. exot. Butts [3] : (Systematic Index) III, (Callithea): [1-2], pl. [1] (1857), (Callithea Agrias II): [3-4], pl. [2] (1870), (Epicalia & Myscelia I): [5-6], pl. [3] (1851), (Epicalia II): [7-8], pl. [4] (1852), (Timetes): [9-10], pl. [5] (1852), (Neptis): [11-12], pl. [6] (1868), (Limenitis I-II): [13-16], pl. [7-8] (1859), (Heterochroa): [17-18], pl. [9] (1867), (Diadema): [19-20], pl. [10] (1863), (Diadema II): [21-22], pl. [11] (1865), (Diadema III): [23-24], pl. [12] (1868), (Amechania): [25-26], pl. [13] (1861), (Romaleosoma): [27-28], pl. [14] (1864), (Romaleosoma II-III): [29-32], pl. [15-16] (1865), (Romaleosoma IV): [33-34], pl. [17] (1866), (Euryphene): [35-36], pl. [18] (1864), (Euryphene II): [37-38], pl. [19] (1865), (Euryphene III Aterica): [39-40], pl. [20] (1865), (Euryphene IV): [41-42], pl. [21] (1865), (Aterica & Euryphene V-VI): [43-46], pl. [22-23] (1866), (Euryphene VII): [47-48], pl. [24] (1871), (Euryphene VIII & Aterica): [49-50], pl. [25] (1871), (Euryphene IX): [51-52], pl. [26] (1874), (Aterica & Harma): [53-54], pl. [27] (1866), (Harma): [55-56], pl. [28] (1864), (Harma II): [57-58], pl. [29] (1866), (Harma III-IV): [59-60], pl. [30-31] (1869), (Harma V-VI & Euryphene X): [61-62], pl. [32-33] (1874), (Adolias): [63-64], pl. [34] (1861), (Adolias II): [65-66], pl. [35] (1862), (Adolias III): [67-68], pl. [36] (1863), (Adolias IV): [69-70], pl. [37] (1875), (Agrias): [71-72], pl. [38] (1860), (Agrias & Nymphalis): [73-74], pl. [39] (1855), (Agrias & Siderone): [75-76], pl. [40] (1854), (Prepona): [77-78], pl. [41] (1859), (Prepona II & Agrias): [79-80], pl. [42] (1876), (Pandora & Prepona): [81-82], pl. [43] (1854), (Apatura): [83-84], pl. [44] (1869), (Nymphalis): [85-86], pl. [45] (1854), (Nymphalis II): [87-88], pl. [46] (1859), (Charaxes III): [89-90], pl. [47] (1863), (Charaxes IV): [91-92], pl. [48] (1874), (Charaxes V): [93-94], pl. [49] (1876), (Paphia II): [95-96], pl. [50] (1869), (Paphia & Siderone): [97-98], pl. [51] (1856), (Siderone): [99-100], pl. [52] (1860), (Zeuxidia & Aemona): [101-102], pl. [53] (1868), : iv
Honrath, 1892
Neue Rhopalocera (10) Berl. ent. Z. 36 (2) : 429-440, pl. 15
Hopffer, 1874
Beitrag zur Lepidoptera-Fauna von Celebes Stettin ent. Ztg 35 (1-3) : 17-47
Howarth, 1962
Rhopalocera of the Rennel and Bellona Islands The Natural History of the Rennel and Bellona Is. (4) : 63-83
Iwasaki & Yata, 2005
A provisional checklist of the genus Cepora (Lepidoptera, Pieridae) from the Sulawesi Region, with descriptions of two new subspecies from Buton Island Trans. lepid. Soc. Japan 56 (4) : 303-310
Jurriaanse, 1920
[Verslag] Tijdschr. Ent. 62 : xlvii-li
Jurriaanse & Lindemans, 1920
Bijdrage tot de kennis der Lepidoptera van Z.O.-Celebes en omliggende eilanden Tijdschr. Ent. 62 (Suppl.) : 1-39
Lucas, 1852
Description de nouvelles Espčces de Lépidoptčres appartenant aux Collections entomologiques du Musée de Paris Revue Mag. Zool. (2) 4 (3): 128-141 (1852), 4 (4): 189-198 (1852), 4 (6): 290-300 (1852), 4 (7): 324-343 (1852), 4 (9): 422-432, pl. 10 (1852), 5 (7): 310-322 (1853)
MacLeay, [1826]
in King, Narrative of a Survey of the Intertropical and Western Coasts of Australia. 2 vols. in King, Narr. Surv. Austr. 2: -
Martin, 1919
Die Tagfalter der Insel Celebes. 3. Fortsetzung Dt. ent. Z. Iris 33 (1/2) (1/2): 48-64, (3/4): 65-98
Mathew, 1887
Descriptions of some new species of Rhopalocera from the Solomon Islands Trans. Ent. Soc. Lond. 1887 (1) : 37-49, pl. 4
Matsumura, 1909
Die Pieriden Japans Ent. Zs. 23 (18) : 87-88
Moore, 1857
Description of some new Species of Lepidopterous Insects from Northern India Proc. zool. Soc. Lond. 1857 (333) : 102-104, pl. 44-45
Moore, [1858]
A Catalogue of the Lepidopterous Insects in the Museum of the Hon. East-India Company in Horsfield & Moore, Cat. lep. Ins. Mus. East India Coy 1: : 1-278, pl. 1-12,1a,2a,3a,4a,5a,6a
Moore, 1865
List of diurnal Lepidoptera collected by Capt. A.M. Lang in the N. W. Himalayas Proc. zool. Soc. Lond. 1865 (2) : 486-509, pl. 30-31
Moore, [1879]
A list of the Lepidopterous Insects collected by Mr. Ossian Limborg in Upper Tenasserim, with Descriptions of new Species Proc. zool. Soc. Lond. 1878 (4) : 821-859, pl. 51-53 (1879)
Moore, [1881]
The Lepidoptera of Ceylon Lepid. Ceylon 1 (1): 1-40, pl. 1-18 (Dec 1880), (2): 41-80, pl. 19-36 (Jan 1881), (3): 81-136, pl. 37-54 (Jun 1881), (4): 137-190, pl. 55-71 (Dec 1881)
Moore, 1884
Descriptions of some new Asiatic diurnal Lepidoptera; chiefly from specimens contained in the Indian Museum, Calcutta J. asiat. Soc. Bengal 53 Pt.II (1) : 16-52
Murayama, 1961
An unrecorded and some aberrant butterflies from Formosa [in Japanese] Tyô to Ga 11 (4): 54-61
Pendlebury, 1933
Notes and new Records of Butterflies from the Malay Peninsula J. Fed. Malay St. Mus. 17 (2): 377-401
Ribbe, 1898
Beiträge zur Lepidopteren-Fauna des Bismarck- und Salomon-Archipels in der Süd-See Dt. ent. Z. Iris 11 (1): 35-133, 12 (1): 219-260
Rothschild, 1892
Notes on a Collection of Lepidoptera made by William Doherty in Southern Celebes during August and September 1891. Part I, Rhopalocera Dt. ent. Z. Iris 5 (2) : 429-442
Rothschild, 1915
On Lepidoptera from the islands of Ceram (Seran), Buru, Bali, and Misol Novit. Zool. 22 (1): 105-144, (2): 209-227
Röber, 1885
Zur ostindischen Schmetterlings-Fauna Corr.-Bl. Ent. Ver. Iris 1 (2) : 19-23
Röber, 1891
Beitrag zur Kenntnis der Indo-Australischen Lepidopteren-Fauna Tijdschr. Ent. 34: 261-334 (1891), 35: 85-86, pl. 3-6 (1892)
Röber, 1900
Neue Schmetterlinge Ent. Nachr. 26 (13) : 199-204
Röber, 1927
Neue exotische Falter Int. ent. Zs. 20 (44): 400-403, 1pl (22 February), (45): 412-414 (1 March), (46): 419-421 (8 March), (47): 428-430 (15 March)
Stoll, [1781]
Uitlandsche Kapellen (Papillons exotiques) in Cramer, Uitl. Kapellen 4 (26b-28): 29-90, pl. 305-336 (1780), (29-31): 91-164, pl. 337-372 (1781), (32-32): 165-252, 1-29, pl. 373-400 (1782)
Stoll, [1790]
Aanhangsel van het Werk, de Uitlandsche Kapellen Aanhangsel Werk, Uitl. Kapellen (1): 1-42, pl. 1-8 (1787), (2-5): 43-184, pl. 9-42 (1790)
Swinhoe, 1890
New Species of Indian Butterflies Ann. Mag. nat. Hist. (6) 5 (29) : 353-365
Vane-Wright & de Jong, 2003
The butterflies of Sulawesi: annotated checklist for a critical island faunda Zool. Verh. Leiden 343: 3-268, pl. 1-16
Vollenhoven, 1865
Essai d'une Faune Entomologique de l'Archipel Indo-Néerlandais, Seconde monographie: Famille des Piérides Faune ent. 2 (Monogr. Pierid.) : 1-70, pl. 1-7
Wallace, 1867
On the Pieridae of the Indian and Australian Regions Trans. ent. Soc. Lond. (3) 4 (3) : 301-416, pl. 6-9
Watkins, 1927
Butterflies from N.W. Yunnan Ann. Mag. nat. Hist. (9) 19 : 313-344, 512
Weymer, 1885
Exotische Lepidopteren. III. Beitrag zur Lepidoperen-Fauna von Nias Stettin ent. Ztg. 46 (4-6) : 257-285, pl. 1-2
Winhard, 2000
Pieridae I Butterflies of the world, 10: (1) 40pp, (2) 48pls


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Keep in mind that the taxonomic information is copied from various sources, and may include many inaccuracies. Expert help is welcome.