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Belenois Hübner, [1819]
Caper Whites
Belenois Hübner, [1819]; Verz. bek. Schmett. (6): 92, TS: Papilio calypso Drury
Anaphaeis Hübner, [1819]; Verz. bek. Schmett. (6): 93, TS: Papilio creona Cramer
Glycestha Billberg, 1820; Enum. Ins. Mus. Billb. : 76, TS: Papilio coronea Cramer
Pseudohuphina Stoneham, 1940; Bull. Stoneham Mus. (40): 4, TS: Pieris raffrayi Oberthür
Pseudanaphaeis Bernardi, 1953; Rev. franc. Ent. 20 (1) : 50, TS: Pieris gidica Godart
Belenois, NSG Voucher Specimen [Wahlberg]
Belenois, Pieridae Holarcticae [Heiner Ziegler]
Annaphaeis ; Butler, 1870, (missp.)
Anapheis[sic] ; [BAUR, 153 (missp)]
Belenois (Pierinae) ; Moore, [1881], Lepid. Ceylon 1 (4): 137; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 112
Belenois ; Winhard, 2000, Butterflies of the world, 10: 27; [AfrL]

The "pseudohuphina" species-group

Belenois raffrayi (Oberthür, 1878)
Ethiopia, Sudan, W.Kenya, Zaire, Rwanda, W.Tanzania. See [About maps] Raffray's White
Pieris raffrayi Oberthür, 1878; Étud. d'Ent. 3 : 17, pl. 1, f. 3
Pseudohuphina raffrayi ;
Belenois raffrayi ; [EBW]; [AfrL]

Larva on Capparis, Rhus [BK]

S.Ethiopia, S.Sudan Belenois raffrayi raffrayi ; [BAFR, 76 (text)]; [AfrL]
B. r. extendens (Joicey & Talbot, 1927)
W.Kenya, Uganda, Rwanda, Burundi, E.Zaire, W.Tanzania? Belenois raffrayi extendens ; [BAFR, 76]; [BK, 138]; Winhard, 2000, Butterflies of the world, 10: 28, pl. 44, f. 12; [AfrL]
B. r. similis Kielland, 1978
W.Tanzania Belenois raffrayi similis Kielland, 1978; Tijdschr. Ent. 121 (4) : 163, pl. 7, f. 25-26, TL: Tanzania, Mpanda, Mt. Sitebi, 2000m
Belenois raffrayi similis ; [AfrL]

Belenois margaritacea Sharpe, 1891
Kenya, Tanzania. See [About maps] Margarita's Caper White
Belenois margaritacea Sharpe, 1891; Proc. Zool. Soc. Lond. 1891 : 191, pl. 16, f. 4, TL: Sotik
Pseudohuphina margaritacea ;
Belenois margaritacea ; [BK, 138]; [AfrL]

Larva on Capparis, Maerua, Ritchiea [BK]

SW.Kenya (Sotik) Belenois margaritacea margaritacea ; [BAFR, 76]; [BK, 138 (note)]; [AfrL]
B. m. kenyensis (Joicey & Talbot, 1927)
SE.Kenya (Teita Hills, Chyulu Hills) Belenois margaritacea kenyensis ; [BAFR, 76]; [BK, 138 (note)]; [AfrL]
B. m. plutonica (Joicey & Talbot, 1927)
N.Tanzania (Mt. Kilimanjaro, Usambara Mts., Ngorongoro Crater) Belenois margaritacea plutonica ; [BAFR, 76]; [AfrL]
B. m. somereni (Talbot, 1928)
Kenya (Meru, Matthew's Range) Belenois margaritacea somereni ; [BK, 138 (note)]; [AfrL]
B. m. intermedia Kielland, 1982
Belenois margaritacea intermedia ; [AfrL]

The ? species-group

Belenois albomaculatus (Goeze, 1779)
Papilio albomaculatus Goeze, 1779; Ent. Beyträge 3 (1) : 182
Belenois albomaculatus ; [AfrL]

Belenois grandidieri (Mabille, 1878)
Madagascar, Aldabra Is.. See [About maps] Pieris grandidieri Mabille, 1878; , TL: Madagascar
Belenois grandidieri ; [BAFR, 76]; Winhard, 2000, Butterflies of the world, 10: 27, pl. 43, f. 6; [AfrL]
Belenois (Belenois) grandidieri ; [MadL]

Belenois larima (Boisduval, 1836)
Senegal?, Congo. See [About maps] Pieris larima Boisduval, 1836; Hist. nat. Ins., Spec. gén. Lépid. 1 : 524
Belenois larima ; [BAFR, 82 (text)]; [AfrL]

Belenois zochalia (Boisduval, 1836)
Cape - Moçambique, Rhodesia - Ethiopia, Uganda. See [About maps] Forest White Forest Caper White
Pieris zochalia Boisduval, 1836; Hist. nat. Ins., Spec. gén. Lépid. 1 : 506
Pieris hyoma Boisduval, 1836; Hist. nat. Ins., Spec. gén. Lépid. 1 : 508
Belenois zochalia ; [AfrL]

Larva on Niebuhria pedunculosa, Maerua triphylla [PBSA]

South Africa, Rhodesia Belenois zochalia zochalia ; [BAFR, 76]; [AfrL]
B. z. agrippinides (Holland, 1896)
Malawi, Tanzania, Kenya, Uganda Pieris agrippinides Holland, 1896; Proc. U.S. Nat. Mus. 18 (1098) : 758
Belenois zochalia agrippinides ; [BAFR, 76]; [BK, 139]; Winhard, 2000, Butterflies of the world, 10: 27, pl. 43, f. 12; [AfrL]

Larva on Capparidaceae, Brassica, Salvadora [BK]

Belenois zochalia var. tanganjikae Lanz, 1896; Dt. ent. Z. Iris 9 (1) : 123
Belenois formosa Butler, 1898; Trans. ent. Soc. 1898 : 434
= ; [NHM]
B. z. connexiva (Joicey & Talbot, 1927)
Cameroun (highlands) Belenois zochalia connexiva ; [BAFR, 76 (text)]; [AfrL]
B. z. galla (Ungemach, 1932)
Ethiopia Belenois zochalia galla ; [BAFR, 76 (text)]; [AfrL]
?B. z. camerounica Bernardi, 1966²
Nigeria - Cameroun (hihlands) ?Belenois zochalia camerounica ; [BK, 139 (note)]

Belenois ?sp.
Belenois ?sp. ; Winhard, 2000, Butterflies of the world, 10: 27, pl. 43, f. 13 (Mpanda, Ntakatta, Kigoma, Tanzania)

Belenois crawshayi Butler, 1894
Angola, Zambia - E.Zaire, Uganda, W.Kenya, Tanzania. See [About maps] Crawshay's Caper White
Belenois crawshayi Butler, 1894; Proc. zool. Soc. Lond. 1893 : 665, TL: Lake Mweru
Pieris continentalis Heyn, 1904; Ent. Zs. 18 : 57
Belenois crawshayi ; [BAFR, 78]; [BK, 139]; [AfrL]

Larva on Capparis [BK]


Belenois diminuta Butler, 1894
Tanzania. See [About maps] Belenois diminuta Butler, 1894; Proc. zool. Soc. Lond. 1893 : 666, pl. 60, f. 7, TL: Lake Mweru
Belenois diminuta ; Winhard, 2000, Butterflies of the world, 10: 27, pl. 43, f. 14; [AfrL]

Belenois victoria Dixey, 1915
Uganda, Burundi, Rwanda, Tanzania, E.Zaire, W.Kenya. See [About maps] Victoria White
Belenois victoria Dixey, 1915; Trans. Ent. Soc. 1915 (1) : 4, pl. 2, f. 5-8, TL: Tiriki, Victoria Nyanza; Toro, W.Uganda
Belenois victoria ; [AfrL]

Larva on Capparis, Maerua, Ritchiea [BK]

W.Kenya (highlands, W of Rift Valley), Uganda Belenois victoria victoria ; [BK, 139]; [BAFR, 78]; [AfrL]
B. v. hecqi Berger, 1953
NE.Zaire Belenois victoria hecqi ; [BAFR, 78 (text)]; [AfrL]
B. v. schoutedeni Berger, 1953
W.Uganda, E.Zaire (Kivu) Belenois victoria schoutedeni ; [BAFR, 78]; Winhard, 2000, Butterflies of the world, 10: 27, pl. 43, f. 15, 18; [AfrL]

The ? species-group

Belenois subeida (C. & R. Felder, 1865)
Guinea - Ethiopia, W.Kenya, Tanzania. See [About maps] Belenois instabilis Butler, 1888; Proc. Zool. Soc. Lond. 1888 : 76, TL: Fóda
Belenois subeida ab. lutea Rebel, 1914; Ann. k.k. naturh. Hofmus. Wien. 28 : 227
Belenois subeida ab. unilutea Rebel, 1914; Ann. k.k. naturh. Hofmus. Wien. 28 : 227
Belenois subeida ab. infumata Rebel, 1914; Ann. k.k. naturh. Hofmus. Wien. 28 : 227
Belenois subeida ; [AfrL]
B. s. frobeniusi (Strand, 1909)
W.Africa Belenois subeida frobeniusi ; [BAFR, 80 (text)]; [AfrL]
Ethiopia Belenois subeida hiemalis ; [BAFR, 80 (text)]
B. s. sylvander Grose-Smith, 1890
W.Kenya, Uganda, Tanzania, N.Zaire, C.A.R. Belenois sylvander Grose-Smith, 1890; Proc. Zool. Soc. Lond. 1890 : 464
Belenois subeida sylvander ; [BAFR, 80]; [BK, 140]; Winhard, 2000, Butterflies of the world, 10: 27, pl. 42, f. 9; [AfrL]

Larva on Capparis, Maerua, Ritchiea [BK]

B. s. hailo (Ungemach, 1932)
S.Ethiopia, SE.Sudan Belenois calypso hailo ; [BAFR, 82 (text)]
Belenois subeida hailo ; [AfrL]

Belenois calypso (Drury, [1773])
Gambia - Angola, W.Kenya, Tanzania. See [About maps] Calypso Caper White
Papilio calypso Drury, [1773]; Illust. Nat. Hist. Exot. Insects 2: index, : 30, pl. 17, f. 3-4
Papilio nigronotatus Goeze, 1779; Ent. Beyträge 3 (1) : 182
= ; [NHM]
Papilio nigropictus Goeze, 1779; Ent. Beyträge 3 (1) : 183
= ; [NHM]
Belenois calypso ; [EBW]; [AfrL]
Nigeria - Ghana, Ivory Coast, Liberia, Sierra Leone Belenois calypso calypso ; [BAFR, 82]; [AfrL]
B. c. dentigera Butler, 1888
Cameroun - Zaire, NW.Angola Belenois dentigera Butler, 1888; Proc. Zool. Soc. Lond. 1888 : 78, TL: Angola, Stazione Gadda
Belenois calypso dentigera ; [BAFR, 82 (text)]; Winhard, 2000, Butterflies of the world, 10: 27, pl. 42, f. 5-6; [AfrL]
B. c. minor Talbot, 1943
Uganda, W.Kenya (W of Rift Valley) Belenois calypso minor ; [BAFR, 82 (text)]; [BK, 140]; [AfrL]

Larva on Capparis, Cadaba, Maerua, Ritchiea [BK]

B. c. marlieri Berger, 1981
Belenois calypso marlieri ; [AfrL]

Belenois welwitschii Rogenhofer, 1890
-. See [About maps] ?Pieris dentigera ratidengi Suffert, 1904; Dt. Ent. Z. Iris 17 (1) : 128, TL: Lindi, Deutsch-Ost-Africa
Belenois welwitschii ; [AfrL]
NE.Angola, S.Zaire (Shaba), Zambia, Malawi, S.Tanzania, W.Tanzania Belenois calypso welwitschii ; [BAFR, 82]
Belenois welwitschii welwitschii ; [AfrL]
B. w. shaba Berger, 1981
Belenois calypso shaba Berger, 1981
Belenois calypso shaba ; Winhard, 2000, Butterflies of the world, 10: 27, pl. 42, f. 7-10
Belenois welwitschii shaba ; [AfrL]

Belenois sudanensis (Talbot, 1929)
Cameroun - W.Kenya, Tanzania. See [About maps] Sudan Caper White
Belenois sudanensis ; [BK, 140]; [AfrL]

Larva on Clerodendrum [BK]

B. s. sudanensis (Talbot, 1929)
Belenois sudanensis sudanensis ; [AfrL]
Belenois sudanensis pseudodentigera ; Winhard, 2000, Butterflies of the world, 10: 27, pl. 42, f. 12; [AfrL]
B. s. katalensis Berger, 1981
Belenois sudanensis katalensis ; Winhard, 2000, Butterflies of the world, 10: 27, pl. 42, f. 15; [AfrL]
B. s. mayumbana Berger, 1981
Belenois sudanensis mayumbana ; [AfrL]

Belenois rubrosignata (Weymer, 1901)
Cameroun - S.Sudan, Uganda, W.Kenya, Tanzania. See [About maps] Red-Edged White
Belenois rubrosignata ; [EBW]; [AfrL]
Angola, S.Zaire (Shaba), Zambia Belenois rubrosignata rubrosignata ; [BAFR, 80]; [BK, 141]; [AfrL]
B. r. kongwana Talbot, 1943
Malawi, Tanzania, S.Uganda Belenois rubrosignata kongwana ; [BAFR, 80 (text)]; Winhard, 2000, Butterflies of the world, 10: 27, pl. 44, f. 2-3; [AfrL]
B. r. peeli Dixey, 1900
Kenya, Somalia Belenois rubrosignata peeli ; [AfrL]; [BAFR, 80 (text)]

Belenois thysa (Hopffer, 1855)
Natal, Moçambique, Rhodesia - Kenya, Ethiopia, Somalia. See [About maps] False Dotted Border
Pieris thysa Hopffer, 1855; Ber. Verh. Akad. Berlin 1855 : 639
?Pieris sabrata Doubleday, [1847]; Gen. diurn. Lep. (1): 47, TL: Congo
?Belenois sabrata Butler, 1870; in Dunning, Trans. ent. Soc. Lond. 1870 (4) : 526
Pieris larima rimala Suffert, 1904; Dt. Ent. Z. Iris 17 (1) : 80, TL: Bugundi, S. Albert Njansa
Pieris thysa malaria Suffert, 1904; Dt. Ent. Z. Iris 17 (1) : 81, TL: Loango
Pieris larima alarmi Suffert, 1904; Dt. Ent. Z. Iris 17 (1) : 81, TL: Muanza
thysa f. malaria ; [wet season f.]
Belenois thysa ; [EBW]; [BK, 141]; [AfrL]

Larva on Capparis sp. [PBSA], Maerua, Ritchiea [BK]

E.Cape - Natal, Mozambique, Rhodesia, Zambia, Malawi, Tanzania, Kenya (coast) Belenois thysa thysa ; [BAFR, 82]; Winhard, 2000, Butterflies of the world, 10: 27, pl. 43, f. 16, pl. 44, f. 1, back f. 5; [AfrL]
B. t. tricolor Talbot, 1943
Ethiopia Belenois thysa tricolor ; [BAFR, 82]; [AfrL]
B. t. meldolae Butler, 1872
Angola, S.Zaire - Uganda, W.Kenya, S.Sudan Belenois meldolae Butler, [1872]; Proc. zool. Soc. Lond. 1871 (3) : 725
Belenois thysa meldolae ; [BAFR, 82 (text)]; [AfrL]

Belenois solilucis Butler, 1874
S.Ethiopia, S.Sudan - Uganda, NW.Tanzania, W.Kenya, Cameroun, Angola. See [About maps] Belenois solilucis Butler, 1874; Trans. ent. Soc. Lond. 1874 (4) : 433, TL: Angola
Phrissura narcissus Butler, 1898; Trans. ent. Soc. 1898 : 431, TL: Angola
= ; [NHM]
Pieris glucki Suffert, 1904; Dt. Ent. Z. Iris 17 (1) : 77, TL: Mukenge
= ; [NHM]
Pieris abti Suffert, 1904; Dt. Ent. Z. Iris 17 (1) : 77, TL: Central Africa
= ; [NHM]
Pieris solilucis f. sabulosa Joicey & Talbot, 1921; Bull. Hill Mus. 1 (1) : 45, TL: Cartouche Village, near Lesse, W bank of Semliki
Belenois solilucis ; [EBW]; [BK, 141]; [AfrL]
N.Angola, Zaire - W.Tanzania, Cameroun Belenois solilucis solilucis ; [BAFR, 85 (text)]; [AfrL]
B. s. agylla (Rogenhofer, 1889)
solilucis agylla (Rogenhofer, 1889) (Mylothris); Ann. Mus. Wien. 4 : 549, pl. 23, f. 4
Pieris sulphurescens Capronnier, 1889; Bull. Ent. Soc. Belg. 33 : cxx
= ; [NHM]
Belenois solilucis agylla ; [NHM]
B. s. loveni (Aurivillius, 1921)
Uganda, NE.Zaire, S.Sudan, SW.Ethiopia Belenois solilucis loveni ; [BAFR, 85]; Winhard, 2000, Butterflies of the world, 10: 28, pl. 44, f. 10-11; [AfrL]

Belenois aldabrensis (Holland, 1896)
Aldabra, Astove, Assumption Is. Teracolus aldabrensis Holland, 1896; Proc. U.S. Nat. Mus. 18 (1064) : 269, pl. 8, f. 7-8
Teracolus aldabrensis ; Butler, 1897, Ann. Mag. nat. Hist. (6) 20 (119): 464
Belenois aldabrensis ; [AfrL]; [BAFR, 85]

Belenois anomala (Butler, 1881)
Socotra I. Synchloe anomala Butler, 1881; Proc. zool. Soc. Lond. 1881 : 178, pl. 18, f. 3, TL: Sokotra
Belenois anomalis[sic?] ; [BAFR, 86]
Belenois anomala ; [AfrL]

?Belenois ogygia (Trimen, 1883)
Natal. See [About maps] Pieris ogygia Trimen, 1883; Trans. ent. Soc. Lond. 1883 (4) : 356
Belenois ogygia ; [BAFR, 79]

The ? species-group

Belenois antsianaka (Ward, 1870)
Madagascar. See [About maps] Pieris antsianaka Ward, 1870; Ent. mon. Mag. 7 : 30, TL: Madagascar
Pieris ramona Grose-Smith, 1891; Ann. Mag. nat. Hist. (6) 7 (37) : 123, TL: NW.Madagascar
Nepheronia[sic] antsianaka var. lasti Grose-Smith & Kirby, 1892; Rhop. Exot. [1] 1: (Belenois) pl. 1, f. 4-5 (preocc.,nom. nud.?)
Belenois antsianaka var. drurii Grose-Smith & Kirby, 1892; Rhop. Exot. [1] 1: (Belenois) 1 (repl. Nepheronia antsianaka var. lasti Grose-Smith & Kirby, 1892), TL: Slary, Madagascar
Belenois antsianaka var. persimilis Grose-Smith & Kirby, 1892; Rhop. Exot. [1] 1: (Belenois) 2, pl. 1, f. 6-7, TL: Salary, Madagascar
Belenois ramona ; Grose-Smith & Kirby, 1892, Rhop. Exot. [1] 1: (Belenois) 2, pl. 1, f. 1-3
Belenois antsiaka[sic?] ; [EBW]; [BAFR, 86]
Belenois antsianaka ; Grose-Smith & Kirby, 1892, Rhop. Exot. [1] 1: (Belenois) 1; Winhard, 2000, Butterflies of the world, 10: 28, pl. 44, f. 4; [AfrL]
Belenois (Belenois) antsianaka ; [MadL]

Belenois mabella Grose-Smith, 1891
Madagascar. See [About maps] Belenois mabella Grose-Smith, 1891; Ann. Mag. nat. Hist. (6) 8 (43) : 79, TL: Madagascar
Belenois mahobo Grose-Smith, 1891; Ann. Mag. nat. Hist. (6) 8 (43) : 80, TL: Madagascar
Pinacopteryx mahobo ; Grose-Smith & Kirby, 1893, Rhop. Exot. [1] 2: (Pinacopteryx) 1, pl. 1, f. 1-3
Pinacopteryx mabella ; Grose-Smith & Kirby, 1893, Rhop. Exot. [1] 2: (Pinacopteryx) 1, pl. 1, f. 4-5
Belenois mabella ; [BAFR, 86]; Winhard, 2000, Butterflies of the world, 10: 28, pl. 44, f. 9; [AfrL]
Belenois (Pseudanaphaeis) mabella ; [MadL]

Belenois java (Linnaeus, 1768)
Indonesia - Samoa, Australia. See [About maps] Caper White
Papilio java Linnaeus, 1768; in Sparrman, Amoenit. acad., 7 (150): 504, nota
Papilio coronea Cramer, [1775]; Uitl. Kapellen 1 (1-7): 106, pl. 68, f. B, C
Papilio clytie Donovan, 1805; Epitome nat. Hist. Ins. New Holland: pl. 19, f. 2
Papilio deiopea Donovan, 1805; Epitome nat. Hist. Ins. New Holland: pl. 21, f. 2
Pieris niseia MacLeay, [1826]; in King, Narr. Surv. Austr. 2: 459 ([?] niscia)
Belenois java ab. magniplaga Fruhstorfer, 1898; Berl. Ent. Zs. 42 (3/4) : 325, TL: Sumba
Anaphaeis nigrita Joicey & Talbot, 1928
Belenois java, NSG Voucher Specimen [Wahlberg]
Belenois java, Lepidoptera Larvae of Australia [Don Herbison-Evans]
Anaphaeis java ; [BOW: pl. 156, f. 11]
Anapheis java ; [EBW]; [BOR, 144]
Belenois java ; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 113

Larva on Capparis mitchelli, C. nummularia, Apophyllum anomalum [Don Herbison-Evans], Capparis nobilis, C. lasiantha [BAUR], Apophyllum anomalum, Capparis arborea, C. lasiantha, C. mitchellii, C. nobilis, C. spinosa var. nummularia, C. canescens?, C. spp. K. L. & L. E. Dunn, 1991, Review Austr. Butts. (1-4): (120-140)

Java, Kangean - Sumbawa, Sumba, Sulawesi Anapheis java java
Anapheis java java ; [BOR, 144]
Belenois java java ; Winhard, 2000, Butterflies of the world, 10: 27, pl. 43, f. 10; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 113
Belenois java ?ssp. ; Winhard, 2000, Butterflies of the world, 10: 27, pl. 43, f. 11 (Gaibobo I., Papua New Guinea)
B. j. savuana (Fruhstorfer, 1898)
Sawu I. (Savu) Pieris (Belenois) teutonia savuana Fruhstorfer, 1898; Berl. Ent. Zs. 42 (3/4) : 326, TL: Savu I.
Anapheis java savuana ; [BOR, 144]
B. j. teutonia (Fabricius, 1775)
Australia Papilio teutonia Fabricius, 1775; Syst. Ent.: 474 [?]199
Anapheis java teutonia ; [BAUR, 153]
B. j. ina Fruhstorfer²
Timor?, Wetar, Batar, Kissar Belenois teutonia anita Fruhstorfer, 1903; Berl. ent. Zs. 48 (1/2) : 107, TL: Wetter I.
Anapheis java ina ; [BAUR, 153 (text)]
B. j. peristhene (Boisduval, 1859)
New Hebrides, New Caledonia, Loyalty Is., Norfolk I., Woodlark I. Pieris peristhene Boisduval, 1859; Bull. Soc. ent. Fr. (3) 7 : clv
Anapheis java peristhene ; [BAUR, 153 (text)]
B. j. picata Butler, 1882
New Britain, New Ireland, Duke of York I. Belenois picata Butler, 1882; Ann. Mag. Nat. Hist. (5) 10 (56): 153, TL: New Britain
Anapheis java picata ; [BAUR, 153 (text)]
B. j. micronesia (Fruhstorfer, 1902)
Fiji Pieris teutonia micronesia Fruhstorfer, 1902; Stettin ent. Ztg 63 (1) : 357, TL: Viti
Pieris peristhene vitiensis Fruhstorfer, 1902; Stettin ent. Ztg 63 (1) : 358, TL: Viti
Anapheis java micronesia ; [BAUR, 153 (text)]

Belenois helcida (Boisduval, 1833)
Madagascar. See [About maps] Pieris helcida Boisduval, 1833; Faun. ent. Madagascar, Lépid. : 17, pl. 2, f. 1-2, TL: Madagascar
Belenois helcida ; [BAFR, 84]; [AfrL]
Belenois (Belenois) helcida ; [MadL]

Belenois hedyle (Cramer, [1777])
-. See [About maps] Papilio hedyle Cramer, [1777]; Uitl. Kapellen 2 (9-16): 137, pl. 186, f. C, D
?Papilio sordide-flavicans Goeze, 1779; Ent. Beyträge 3 (1) : 182
Belenois hedyle ; [AfrL]
Sierra Leone, Liberia, Ivory Coast, Ghana, Benin Belenois hedyle hedyle ; [BAFR, 84 (text)]; [AfrL]
B. h. rhena (Doubleday, 1846)
Ghana - W.Nigeria Pieris rhena Doubleday, 1846; Ann. Mag. nat. Hist. 17 (109) : 24
Belenois hedyle rhena ; [BAFR, 84]; [AfrL]
B. h. ianthe (Doubleday, 1842)
Sierra Leone, Togo, Guinea, Guinea-Bissau Pieris ianthe Doubleday, 1842; in Gray, Zool. Misc.: 77
Belenois ianthe ; [BAFR, 84]
Belenois hedyle ianthe ; [AfrL]

Belenois theora (Doubleday, 1846)
-. See [About maps] Pieris theora Doubleday, 1846; Ann. Mag. nat. Hist. 17 (109) : 25
Pieris paroreia Hewitson, 1869; Trans. ent. Soc. Lond. 1869 (1) : 72, TL: Cape Coast
= ; [NHM]
Pieris lortzingi Suffert, 1904; Dt. Ent. Z. Iris 17 (1) : 79, TL: Sesse I., Deutch-Ost-Afrika
= ; [NHM]
Belenois theora ; [EBW]; [AfrL]
Sierra Leone - Nigeria Belenois theora theora ; [BAFR, 81]; [AfrL]
?B. t. concolor Aurivillius²
Cameroun - C.A.R., C.Zaire, S.Zaire ?Belenois theora concolor ; [BAFR, 81 (text)]
B. t. ratheo (Suffert, 1904)
C.A.R Pieris theora ratheo Suffert, 1904; Dt. Ent. Z. Iris 17 (1) : 81, TL: Johann Albrechthöhe, Camerun
Belenois theora ratheo ; Winhard, 2000, Butterflies of the world, 10: 28, pl. 44, f. 7-8; [AfrL]
B. t. laeta (Weymer, 1903)
W.Uganda, E.Zaire, S.Sudan Belenois theora laeta ; [BAFR, 81 (text)]; [AfrL]

Belenois theuszi (Dewitz, 1889)
S.Angola, NW.Zambia, S.Zaire - Gabon, Cameroun, W.Nigeria. See [About maps] ?Pieris theuszi var. semialba Aurivillius, 1895; Ent. Tidskr. 16 : 261
Belenois theuszi ; [BAFR, 82]; Winhard, 2000, Butterflies of the world, 10: 28, pl. 43, f. 17; [AfrL]

Subgenus Anaphaeis Hübner, [1819]
Anaphaeis Hübner, [1819]; Verz. bek. Schmett. (6): 93

Belenois aurota (Fabricius, 1793)
Ceylon, India, NW.India, Pakistan, Afghanistan, Baluchistan, Arabia, SAF, Tropical Africa (dry zones). See [About maps] Brown-veined White Caper White Pioneer
Papilio aurota Fabricius, 1793; Ent. Syst. 3 (1): 197, TL: Tranquebar, S.India
Papilio mesentina Cramer, [1780]; Uitl. Kapellen 3 (23-24): 140, pl. 270, f. A, B (preocc. Papilio menentina Cramer, [1777])
Pinacopteryx syrinx Wallengren, 1863; Wien ent. Monats. 4: 34
= ; [NHM]
Anapheis aurota ; [BRU, 160]
Belenois aurota ; [EBW]; [BOR, 146]; [AfrL]
Anaphaeis aurota ; [BMAT: 18, pl. 6, f. 27-29]
mesentina ; [BIR, 425]
[aurota-1u.jpg] 745x923 (~326Kb) upperside Ethiopia, Amhara, Jara River shingle bank at the bridge 9 km SSW of Hayk. 6th August 2012, Photo © Oleg Kosterin [aurota-1v.jpg] 972x935 (~394Kb) underside Ethiopia, Amhara, Jara River shingle bank at the bridge 9 km SSW of Hayk. 6th August 2012, Photo © Oleg Kosterin

Larva on Capparis oleoides, Maerua triphylla [PBSA], Capparis herbacea [BRU], Capparis spinosa, C. sepiaria, Capparis atphylla, C. heyneana, C. galeata, Maerua crassifolia, M. arenaria, Boscia senegalensis, B. albitrunca [BMAT]


India, Sikkim, Tropical Africa, Madagascar Belenois aurota aurota ; [BOR, 146]; [BAFR, 78]; [BK, 137]; Winhard, 2000, Butterflies of the world, 10: 27, pl. 42, f. 13, pl. 43, f. 1
?mesentina r. mesentina ; [BIR, 425]
Belenois (Belenois) aurota aurota ; [MadL]
B. a. augusta (Olivier, 1801)
Papilio augusta Olivier, 1801; Voy. L'Emp. Othoman. Atl. 4 (1801?): pl. 33, f. 3a-b
Belenois aurota augusta ; Winhard, 2000, Butterflies of the world, 10: 27, pl. 43, f. 4
B. a. taprobana (Moore, 1872)
Ceylon Pieris taprobana Moore, 1872; Proc. zool. Soc. Lond. 1872 (2) : 565, TL: Ceylon
Belenois mesentina fervidior Fruhstorfer, 1898; Berl. Ent. Zs. 42 (3/4) : 326, TL: Ceylon
Belenois taprobana ; Moore, [1881], Lepid. Ceylon 1 (4): 137, (3): pl. 53, f. 3, 3a
mesentina r. taprobana ; [BIR, 425]
Belenois aurota taprobana ; [BOR, 146]
Belenois aurota ?ssp. ; Winhard, 2000, Butterflies of the world, 10: 27, pl. 42, f. 14, pl. 43, f. 2-3 (Lmandraka, Madagascar)
B. a. turanica (Sheljuzhko, 1908)
S.Turkmenia, Tajikistan

Belenois creona (Cramer, [1776])
Tropical Africa (dry parts), Comoros, Madagascar, SW.Arabia. See [About maps] African Common White African Caper
Papilio creona Cramer, [1776]; Uitl. Kapellen 1 (8): 148, pl. 95, f. B, C
Colias ernestius Laporte, 1830; Bull. Soc. Linn. Bordeaux 4: 141
Belenois creona ; [EBW]; [AfrL]
Senegal - Nigeria, Sudan, W.Ethiopia Belenois creona creona ; [BAFR, 79 (text)]; [AfrL]
B. c. benadirensis (Storace, 1948)
Somalia Belenois creona benadirensis ; [BAFR, 79 (text)]; [AfrL]
B. c. boguensis (C. & R. Felder, 1865)
Agamsa?, Ethiopia Pieris boguensis C. & R. Felder, 1865; Reise Fregatte Novara, Bd 2 (Abth. 2) (2): 173
Belenois creona boguensis ; [BAFR, 79 (text)]; [AfrL]
B. c. elisa (Vollenhoven, 1869)
Comoro Is. Pieris elisa Vollenhoven, 1869; Faune Madag. 5 (Ins.) : 12
Belenois creona elisa ; [BAFR, 79 (text)]; [AfrL]
B. c. leucogyne Butler, 1885
SW.Arabia Belenois creona leucogyne ; [BAFR, 79 (text)]; [AfrL]
B. c. prorsus (Talbot, 1943)
Madagascar Anapheis creona prorsus Talbot, 1943; Ann. Mag. nat. Hist. (11) 10: 305, TL: Madagascar, Maintirano
Belenois creona prorsus ; [BAFR, 79 (text)]; [AfrL]
Belenois creona prorsa ; Winhard, 2000, Butterflies of the world, 10: 27, pl. 43, f. 5
Belenois (Belenois) creona prorsus ; [MadL]
B. c. severina (Stoll, [1781])
South Africa - E.Africa, Zaire Papilio severina Stoll, [1781]; in Cramer, Uitl. Kapellen 4 (29-31): 95, pl. 338, f. G, H
Belenois severina ; [EBW]
Belenois creona severina ; [BAFR, 79]; [BK, 137]; Winhard, 2000, Butterflies of the world, 10: 27, pl. 43, f. 7-9; [AfrL]

Larva on Capparis oleoides, C. zeyheri, Maerua triphylla [PBSA]

?B. c. infida Butler, 1888
Belenois infida Butler, 1888; Proc. Zool. Soc. Lond. 1888 : 78, TL: Wadelai

Belenois gidica (Godart, 1819)
Natal, Swaziland, Transvaal, Moçambique, Rhodesia, South West Africa, Tropical Africa (dry zones). See [About maps] African Veined White Pointed Caper
Pieris gidica Godart, 1819; Encyclopédie Méthodique. 9 (1): 131
Belenois occidentis Butler, 1898; Trans. ent. Soc. 1898 : 438, TL: Congo; Loanda
?Pieris gidica var. westwoodi Wallengren
?Pieris gidica ab. simplex Rebel, 1914; Ann. k.k. naturh. Hofmus. Wien. 28 : 226
Belenois gidica ; [EBW]; [BAFR, 86]; [BK, 138]; [AfrL]
[gidica-1x.jpg] 592x608 (~190Kb) Ethiopia, Amhara, a rapidous brook, Blue Nile right tributary just upstream the bridge of the road Addis Ababa - Bahir Dar, 1100 m a.s.l. 1st August 2012, Photo © Oleg Kosterin [gidica-1y.jpg] 666x678 (~189Kb) Ethiopia, Amhara, a rapidous brook, Blue Nile right tributary just upstream the bridge of the road Addis Ababa - Bahir Dar, 1100 m a.s.l. 1st August 2012, Photo © Oleg Kosterin [gidica-1zu.jpg] 689x646 (~219Kb) upperside Ethiopia, Amhara, a rapidous brook, Blue Nile right tributary just upstream the bridge of the road Addis Ababa - Bahir Dar, 1100 m a.s.l. 1st August 2012, Photo © Oleg Kosterin
[gidica-2aU.jpg] identification uncertain 714x799 (~282Kb) Ethiopia, Amhara, a rapidous brook, Blue Nile right tributary just upstream the bridge of the road Addis Ababa - Bahir Dar, 1100 m a.s.l. 1st August 2012, Photo © Oleg Kosterin [gidica-2bU.jpg] identification uncertain 935x859 (~316Kb) Ethiopia, Amhara, a rapidous brook, Blue Nile right tributary just upstream the bridge of the road Addis Ababa - Bahir Dar, 1100 m a.s.l. 1st August 2012, Photo © Oleg Kosterin

Larva on Capparis citrifolia [PBSA], Boscia, Maerua [BK]


Belenois gidica gidica ; [AfrL]
B. g. hypoxantha (Ungemach, 1932)
Belenois gidica hypoxantha ; [AfrL]
B. g. abyssinica (Lucas, 1852)
Ethiopia Pieris abyssinica Lucas, 1852; Revue Mag. Zool. (2) 4 (7): 328
Belenois gidica abyssinica ; [AfrL]


Unknown or unplaced taxa

Belenois johannae Butler, 1879; Ann. Mag. nat. Hist. (5) 3 (15) : 191, TL: Johanna I.


29.4.2013 (29)

References:
[About maps]
Warning! The maps are automatically generated from the textual information, and the process does not always produce acceptable result; See about maps for more info.
[AfrL]
Mark C. Williams; Checklist of Afrotropical Papilionoidea and Hesperoidea; Compiled by Mark C. Williams, 7th ed. (2008) (April 2007); Afrototropical Butterflies; e-mail: Mark C. Williams
[BAFR]
D'Abrera, 1980; Butterflies of the Afrotropical Region
[BAUR]
D'Abrera, 1977; Butterflies of the Australian Region (2nd edn)
[BHL]
The; Biodiversity Heritage Library; and on; Internet Archive; Note: The linking to the content of Internet Archive is somewhat delicate -- notify me when links break!
[BIR]
Wynter-Blyth, 1957; Butterflies of the Indian Region; (1982 Reprint)
[BK]
Larsen, 1996; The Butterflies of Kenya and their natural history
[BMAT]
Tennent, 1996; The butterflies of Morocco, Algeria and Tunisia; ISBN 0 906802 05 9
[BOR]
D'Abrera, 1982, 1985, 1986; Butterflies of the Oriental Region, Part I-III
[BOW]
Lewis, 1974; Butterflies of the World; ISBN 0 245 52097 X
[BRU]
Tuzov, Bogdanov, Devyatkin, Kaabak, Korolev, Murzin, Samodurov, Tarasov, 1997; Guide to the Butterflies of Russia and adjacent territories; Hesperiidae, Papilionidae, Pieridae, Satyridae; Volume 1
[Don Herbison-Evans]
and Stella A. Crossley; Lepidoptera Larvae of Australia; e-mail: Don Herbison-Evans; e-mail: Don Herbison-Evans
[EBW]
Smart, Paul, 1976; The Illustrated Encyclopedia of the Butterfly World; ISBN 0 600 31381 6
[HBN]
Francis Hemming, 1937; HÜBNER, A bibliographical and systematic account of the entomological works of Jacob Hübner and of the supplements thereto by Carl Geyer, Gottfried Franz von Frölich and Gottlieb August Wilhelm Herrich-Schäffer, Vol 1 & 2
[Heiner Ziegler]
Switzerland / Schweiz; www.pieris.ch
[LAMAS-2005]
Gerardo Lamas; Entomofauna 26 (6): 57-100; A Bibliography of the Zoological Publications of Hans Fruhstorfer (1886* - 1922+)
[LAMAS-NEO]
Gerardo Lamas; Bibliography of butterflies; An Annocated Bibliography of the Neotropical Butterflies and Skipper (Lepididoptera: Papilionoidea and Hesperioidea); Revised Electronic Edition (2007)
[MadL]
Lees, D.C., Kremen, C. and Raharitsimba, H. 2003; in S.M. Goodman & J. P. Benstead (eds.), The Natural History of Madagascar; Classification, Diversity and Endemism of the Butterflies (Papilionoidea and Hesperioidea): A Revised Species Checklist: : pp. 762-793; University of Chicago Press. 1709 pp.
[NHM]
Natural History Museum; The Global Lepidoptera Names Index
[Oleg Kosterin]
Institue of Cytology & Genetics; Siberian Division of Russian Academy of Sciences; Novosibirsk; e-mail: Oleg Kosterin; http://pisum.bionet.nsc.ru/kosterin/
[PBSA]
Dickson, C.G.C (ed.), Kroon, D.M.; Pennington's Butterflies of Southern Africa; AD. DONKER 1978
[Wahlberg]
Niklas Wahlberg; The NSG's voucher specimen database; Nymphalidae Systematics Group
[²]
This may require parentheses or not. I don't have the necessary information for this taxon.


Some related literature:
Aurivillius, 1895
Beiträge zur Kenntniss der Insektenfaun von Kamerun. 2. Tagfalter Ent. Tidskr. 16 : 195-220, : 255-268
Bernardi, 1953
Note sur la taxonomie des genres Belenois Hb. et Anaphaeis Hb., [Lep. Pieridae] Rev. franc. Ent. 20 (1) : 47-53
Billberg, 1820
Enumeratio Insectorum Enum. Ins. Mus. Billb. : 1-138
Boisduval, 1833
Faune Entomologique de Madagascar, Bourbon et Maurice. Lépidoptéres Faun. ent. Madagascar, Lépid. : 1-122, pl. 1-16, : (Repaged offprint from Nouv. Ann. Mus. Hist. nat. Paris 2 (2) : 149-270)
Boisduval, 1859
[communications] Bull. Soc. ent. Fr. (3) 7 : cliv-clviii
Boisduval, 1836
Histoire Naturelle des Insectes. Species Général des Lépidoptéres. Tome Premier Hist. nat. Ins., Spec. gén. Lépid. 1 : 1-690, pl. 1-24
Butler, [1872]
On a small Collection of Butterflies from Angola Proc. zool. Soc. Lond. 1871 (3) : 721-725
Butler, 1874
Descriptions of some new species and a new genus of diurnal Lepidoptera in the collection of Herbert Druce Esq. Trans. ent. Soc. Lond. 1874 (4) : 423-436, pl. 6
Butler, 1879
On a Collection of Lepidoptera from the Island of Johanna Ann. Mag. nat. Hist. (5) 3 (15) : 186-192
Butler, 1881
On the Lepidoptera collected in Socotra by I. Bayley Balfour Proc. zool. Soc. Lond. 1881 : 175-180, pl. 18
Butler, 1882
Descriptions of new species of Lepidoptera, chiefly from Duke of York Island and New Britain Ann. Mag. Nat. Hist. (5) 10 (55): 36-43, (56): 149-160, (57): 226-238
Butler, 1888
On the Lepidoptera received from Dr. Emin Pascha [from Equatorial Africa] Proc. Zool. Soc. Lond. 1888 : 56-85
Butler, 1894
On two collection of Lepidoptera sent by H. H. Johnston, Esq., C.B., from British Central Africa Proc. zool. Soc. Lond. 1893 : 643-684, pl. 60 (1894)
Butler, 1897
A revision of the Species of Butterflies belonging to the Genus Teracolus Swains. Ann. Mag. nat. Hist. (6) 20 (118): 385-399, (119): 451-473, (120): 495-507
Butler, 1898
On some new species of African Pierinae in the collection of the British Museum, with notes on seasonal forms of Belenois Trans. ent. Soc. 1898 : 431-438
Capronnier, 1889
Liste des Lépidoptères capturés au Congo par Messieurs Thys, Legat, Martini et Machado en 1887 Bull. Ent. Soc. Belg. 33 : cxviii-cxxvii [118-127]
Cramer, [1775]; Cramer, [1776]
Uitlandsche Kapellen (Papillons exotiques) Uitl. Kapellen 1 (1-7): 1-132, pl. 1-84 (1775), (8): 133-155, pl. 85-96 (1776)
Cramer, [1777]
Uitlandsche Kapellen (Papillons exotiques) Uitl. Kapellen 2 (9-16): 1-152, pl. 97-192 (1777)
Cramer, [1780]
Uitlandsche Kapellen (Papillons exotiques) Uitl. Kapellen 3 (17-21): 1-104, pl. 193-252 (1779), (22): 105-128, pl. 253-264 ([1780]), (23-24): 129-176, pl. 265-288 (1780)
Dixey, 1915
New species and subspecies of Pierinae Trans. Ent. Soc. 1915 (1) : 1-15, pl. 1-2
Donovan, 1805
An Epitome of the Natural History of the Insects of New Holland, New Zealand, New Guinea, Otaheite and other Islands in the Indian, Sothern and Pacific Oceans ... Epitome nat. Hist. Ins. New Holland: 41pls
Doubleday, 1842
Characters of undescribed Lepidoptera in Gray, Zool. Misc.: 73-78
Doubleday, 1846
Descriptions of new or imperfectly described Diurnal Lepidoptera Ann. Mag. nat. Hist. 17 (109) : 22-26
Doubleday, [1847]
The genera of diurnal Lepidoptera, comprising their generic characters, a notice of their habitats and transformations, and a catalogue of the species of each genus; illustrated with 86 plates by W. C. Hewitson Gen. diurn. Lep. (1): 1-18, pl. A, 1-4 (1-2) (1846), (1): 19-132, pl. 4*, 5-25, 28 (3-14) (1847), (1): 133-200, pl. 1*, 26, 27, 29, 31-44 (15-23) (1848), (1): 201-242, pl. 30, 45-52, 56-58, 60-62, 64 (24-31) (1849), (1): 243-250 [?], (2): 243-326, pl. 53-55, 63, 65, 66 (32-38) (1850), (2): 327-466, pl. 59, 67-77 (39-50) (1851), (2): 467-534, 143, 144, pl. 78-80 (Index) (51-54) (1852)
Drury, [1773]
Illustrations of natural history; wherein are exhibited ... Illust. Nat. Hist. Exot. Insects 2 : 1-90, pl. 1-50
K. L. & L. E. Dunn, 1991
Review of Australian Butterflies: distribution, life history and taxonomy. Pushlished by authors, Melbourne Review Austr. Butts. (1-4):
Butler in Dunning, 1870
Notes on a Collection of Insects sent by Mr. Ansell from South-West Africa Trans. ent. Soc. Lond. 1870 (4) : 521-532
Fabricius, 1775
Systema Entomologiae, sistens Insectorum Classes, Ordines, Genera, Species, Adiectis Synonymis, Locis, Descriptionibus, Observationibus. Syst. Ent.: 832pp
C. & R. Felder, 1865
Reise der österreichischen Fregatte Novara um die Erde in den Jahren 1857, 1858, 1859 unter den Behilfen des Commodore B. von Wüllerstorf-Urbair. Zoologischer Theil. Band 2. Abtheilung 2. Lepidoptera. Rhopalocera Reise Fregatte Novara, Bd 2 (Abth. 2) (1): 1-136, pl. 1-21 (Rhop.) (C. & R. Felder, [1865]), (2): 137-378, pl. 22-47 (Rhop.) (C. & R. Felder, 1865), (3): 379-536, pl. 48-74 (Rhop.) (C. & R. Felder, 1867), (4): 537-548, pl. 75-120, (Inhalts-Verz.) 1-9 (pl. 1-74), (Felder & Rogenhofer, 1874) , (5): pl. 121-140, (erklärung) 1-10 (pl. 75-107), (Inhalts-Verz.) 1-20 (pl. 108-140), (Felder & Rogenhofer, 1875)
Fruhstorfer, 1898
Neue Rhopaloceren aus dem Indo-malayischen Archipel Berl. Ent. Zs. 42 (3/4) : 311-343 (mid July)
Fruhstorfer, 1902
Beitrag zur Kenntniss der Lepidopteren der Viti-Inseln Stettin ent. Ztg 63 (1) : 350-359 (1 July)
Fruhstorfer, 1903
Neue Pieriden und Uebersicht verwandter Formen Berl. ent. Zs. 48 (1/2) : 97-111 (early May)
Godart, 1819
in Latreille & Godart, Encyclopédie Méthodique. Histoire naturelle Entomologie, ou histoire naturelle des crustacés, des arachnides et des insectes Encyclopédie Méthodique. 9 (1): 3-328 (1819), (2): 329-828 ([1824])
Goeze, 1779
Entomologische Beyträge zu des Ritter Linné zwölften Ausgabe des Natursystems Ent. Beyträge 3 (1) : 1-390
Grose-Smith, 1890
A list of the Butterflies collected by Mr. William Bonny on the Journey with Mr. Stanley from Yambuya on the Aruwimi River through the Great Forest of Central Africa; with Descriptions of nine new Species Proc. Zool. Soc. Lond. 1890 : 463-473
Grose-Smith, 1891
Descriptions of Ten new Species of Butterflies from the North-west Coast of Madagascar, captured by Mr. J. T. Last, in the Collection of Mr. H. Grose Smith Ann. Mag. nat. Hist. (6) 7 (37) : 122-128
Grose-Smith, 1891
Descriptions of Four new Species of Butterflies from South-west Madagascar, captured by Mr. J. T. Last, in the Collection of H. Grose Smith Ann. Mag. nat. Hist. (6) 8 (43) : 78-81
Grose-Smith & Kirby, 1892; Grose-Smith & Kirby, 1893
Rhopalocera Exotica, being Illustrations of New, Rare, and Unfigured Species of Butterflies Rhop. Exot. [1] 1: (Ornithoptera) 1-2, pl. 1 (1887), [1] 2: (Ornithoptera) 3-4, pl. 2 (1892), [1] 3: (Ornithoptera) [5-6] 1-2, pl. 3 (1900), [1] 3: (Ornithoptera) 7-8, pl. 4 (1901), [1] 1: (Papilio) 1-2, pl. 1 (1887), [1] 1: (Papilio) 3-5,7-14, pl. 2-6 (1888), [1] 1: (Papilio) 15-20, pl. 7-9 (1890), [1] 1: (Papilio) 21-25, pl. 10-11 (1891), [1] 2: (Papilio) 27-31, pl. 12-13 (1893), [1] 2: (Papilio) 33-35, pl. 14 (1894), [1] 2: (Papilio) 37-38, pl. 15 (1895), [1] 2: (Papilio) 39-40, pl. 16 (1897), [1] 3: (Papilio) 41-46, pl. 17-19 (1899), [1] 3: (Papilio) 47-48, pl. 20 (1900), [1] 3: (Papilio) 49-50, pl. 21 (1901), [1] 3: (Papilio) 51-55, pl. 22-23 (1902), [1] 1: (Delias) 1-3, pl. 1 (1889), [1] 2: (Delias) 5-7, 9-11, pl. 2-3 (1893), [1] 2: (Delias) 12-18, pl. 4-5 (1895), [1] 2: (Delias) 20-22, pl. 6 (1896), [1] 3: (Delias) 24-26, pl. 7 (1897), [1] 3: (Delias) 28-35, pl. 8-9 (1901), [1] 3: (Delias) 36-38, pl. 10 (1902), [1] 1: (Pieridae) 1-3, pl. 1 (1888), [1] 2: (Pinacopteryx) 1-4, pl. 1 (1893), [1] 1: (Callosune) 1-3, pl. 1 (1889), [1] 3: (Huphina) 1-7, pl. 1-2 (1898), [1] 3: (Huphina) [8],9-10, pl. 3 (1899), [1] 1: (Elodina) 1-4, pl. 1 (1890), [1] 1: (Belenois) 1-4, pl. 1 (1892), [1] 2: (Belenois) 5-8, pl. 2 (1892), [1] 3: (Mylothris) 1-3, 5-7, pl. 1-2 (1900), [1] 1: (Ixias) 1-3, pl. 1 (1888), [1] 2: (Appias) 1-3, pl. 1 (1894), [1] 1: (Pierinae) 5-7, pl. 2 (1889), [1] 2: (Dismorphia) 1-4, pl. 1 (1896), [1] 2: (Dismorphia) 5-9, 11-15, pl. 2-3 (1897), [2] 1: (Asthipa) 1-3, pl. 1 (1890), [2] 2: (Asthipa) 5-6, pl. 2 (1896), [2] 2: (Ravadeba) 7, pl. 1 (1896), [2] 2: (Mycalesis) 1-4, pl. 1 (1895), [2] 2: (Mycalesis) 5-12, pl. 2-3 (1896), [2] 3: (Mycalesis) 13-18, pl. 4-5 (1902), [2] 3: (Libythea) 1-2, pl. 1 (1902), : (Ragadia-Argyronympha-Hypocysta) : 1-6, pl. [1] (1895), [2] 2: (Pedaliodes) 1-5, pl. 1 (1893), [2] 2: (Pedaliodes) 7-9, pl. 2 (1894), [2] 2: (Pedaliodes) 10-14, pl. 3 (1895), [2] 3: (Pedaliodes) 15-20, pl. 4-5, (Oxeoschistus) 1 (1900), [2] 3: (Pronophila) 1-2, pl. 1, (Oxeoschistus) 2-3, pl. 1 (1900), [2] 2: (Elymnias) 1-4, pl. 1 (1896), [2] 3: (Elymnias) 5-6, pl. 2, (Bruasa) 6-7, pl. 1 (1897), [2] 1: (Morpho) 1-2, pl. 1 (1890), [2] 3: (Morphotenaris-Hyantis) 1-3, pl. [1] (1898), [2] 2: (Tenaris) 1-3, 5-7, pl. 1-2 (1894), [2] 2: (Tenaris) 9-11, pl. 3 (1895), [2] 2: (Tenaris) 13-15, pl. 4 (1896), [2] 3: (Tenaris) 17-18, pl. 5 (1898), [2] 1: (Heliconius) 1-3, pl. 1 (1892), [2] 1: (Acraea) 1-6, pl. 1-2 (1889), [2] 1: (Acraea) 7-10, pl. 3 (1892), [2] 2: (Acraea) 11-14, pl. 4 (1892), [2] 2: (Acraea) 15-18, pl. 5 (1894), [2] 3: (Acraea) 19-29, pl. 6-8, (Planema) 29-30, pl. [1] (1901), [2] 1: (Cethosia) 1-2, pl. 1 (1889), [2] 3: (Cirrhochroa) 1, pl. [1], (Cupha) 2-4, pl. [1] (1898), [2] 2: (Terinos) 1-2, pl. 1 (1894), [2] 1: (Thaleropis) 1-3, pl. 1 (1891), [2] 2: (Doleschallia) 1-3,5-7, pl. 1-2 (1893), [2] 2: (Kallima) 1-3, pl. 1 (1892), [2] 1: (Hypolimnas) 1-2, pl. 1 (1891), [2] 3: (Hypolimnas) 3-4, pl. 2 (1897), [2] 3: (Hypolimnas) 5-6, pl. 3 (1898), [2] 1: (Euxanthe) 1-3, pl. 1 (1890), [2] 1: (Hestina) 1-3, pl. 1 (1891), [2] 1: (Cymothoe) 1-4, pl. 1 (1890), [2] 3: (Limenitis) 1-3, pl. 1 (1899), [2] 2: (Neptis) 1-4, pl. 1 (1895), [2] 3: (Neptis) 5-11, pl. 2-3 (1899), [2] 3: (Neptis) 13-17, pl. 4-5 (1900), [2] 1: (Euthalia-Tanaecia) 1-3, pl. 1 (1889), [2] 1: (Euthalia) 5-10, pl. 2-3 (1891), [2] 2: (Euthalia) 11-14, pl. 4 (1894), [2] 3: (Euthalia) 15, pl. 5, (Dichorragia) 16, pl. 1 (1898), [2] 1: (Apatura) 1-3, pl. 1 (1892), [2] 1: (Euphaedra) 1-2, pl. 1 (1889), [2] 1: (Euryphene) 1-3, pl. 1 (1889), [2] 1: (Charaxes) 1-2,[3-4] pl. 1-2 (1887), [2] 1: (Charaxes) [5], pl. 3 (1888), [2] 1: (Charaxes) 7-9, pl. 4 (1890), [2] 1: (Charaxes) 11-13, pl. 5 (1891), [2] 1: (Mynes) 1-2, pl. 1 (1888), [2] 2: (Mynes) 3-6, pl. 2 (1894), [2] 2: (Prothoe) 1-2, pl. 1 (1896), [3] 3: (Erycinidae) 1-10, pl. 1-2 (1902), [3] 2: (Dicallaneura-etc) 1-6, pl. 1 (1897), [3] 3: (Dicallaneura-etc) 7-11, pl. 2 (1901), [3] 1: (Lycaenidae) 1-4,[5-9] 1-5, pl. 1-2 (1887), [3] 1: (Lycaenidae) 11-17, 19-22, pl. 3-5 (1888), [3] 1: (Lycaenidae) 23-25, 27-29, 31-33, pl. 6-8 (1889), [3] 1: (Lycaenidae) 35-48, pl. 9-11 (1890), [3] 1: (Lycaenidae) 49-51,53-64, pl. 12-15 (1891), [3] 2: (Lycaenidae) 65-68, pl. 16 (1892), [3] 1: (Lycaenidae) [69-72] 57-60, pl. 17 (1892), [3] 2: (Lycaenidae) [73-77] 61-65,79-84, pl. 18-19 (1892), [3] 2: (Lycaenidae) 85-107, pl. 20-23 (1893), [3] 2: (Lycaenidae) 109-120, pl. 24-25 (1894), [3] 3: (Lycaenidae) 121-125, pl. 26 (1900), [3] 3: (Lycaenidae) 127-131, 133-141, pl. 27-29 (1901), [3] 3: (Lycaenidae) 143-146, pl. 30 (1902), [3] 2: (Hypochrysops) 1-6, pl. 1 (1894), [3] 2: (Hypochrysops) 7-13, 15-22, pl. 2-3 (1895), [3] 2: (Thysonotis) 23-25, 27-31, pl. 1-2 (1895), [3] 2: (Waigeum) 34-38, pl. 1 (1896), [3] 2: (Thysonotis) 39-46, pl. 3-4 (1896), [3] 2: (Holochila) 1-4, pl. 1 (1896), [3] 2: (Holochila) 5-8, pl. 2 (1897), [3] 2: (Pseudonotis) 1-3, pl. [1], (Epimastidia) 4-5, pl. [1], (Lampides) 5-8, pl. [1] (1897), [3] 2: (Lampides) 9-10, pl. 2, (Thysonotis) 47, pl. 5 (1897), [3] 3: (Holochila) 9-11, pl. 3 (1898), [3] 3: (Arhopala) 1-7, pl. 1-2 (1898), [3] 3: (Arhopala) 9-12, pl. 3, (Ilerda) 13, pl. [1], (Horaga) 14-15, pl. [1] (1902), [3] 3: (Thysonotis) 47x,48-50, pl. 6 (1898), [3] 3: (Hypochrysops) 23-26, pl. 4 (1898), [3] 3: (Holochila) 13-15, pl. 4 (1899), [3] 3: (Hypochrysops) 27-29, pl. 5, (Waigeum) [30], pl. 2 (1899), [3] 3: (Holochila) 17-19, pl. 5 (1899), [3] 3: (Thysonotis) 51-54, pl. 7 (1899), [3] 3: (Jamides) 11-14, pl. 1 (1900), [3] 3: (Hypolycaena-etc) 1-4, pl. [1] (1900), [3] 3: (Hypochrysops) 31-33, pl. 6, (Thysonotis) 55-56, pl. 8 (1900), [3]: (Additions-Corrections) 1-2,[3],vii-x
Heyn, 1904
Pieris continentalis n. sp. Ent. Zs. 18 : 57-58
Hewitson, 1869
Descriptions of new Species of Diurnal Lepidoptera Trans. ent. Soc. Lond. 1869 (1) : 71-75
Holland, 1896
List of the Lepidoptera from Aldabra, Seychelles, and other East African Islands, Collected by Dr. W. L. Abbot Proc. U.S. Nat. Mus. 18 (1064) : 265-273
Holland, 1896
List of the Lepidoptera Collected in East Africa, 1894, by Mr. William Astor Chanler and Lieut. Ludwig von Höhnel Proc. U.S. Nat. Mus. 18 (1098) : 741-767
Hopffer, 1855
[Lepidoptera Diurna] Ber. Verh. Akad. Berlin 1855 : 639-643
Hübner, [1819]
Verzeichniss bekannter Schmettlinge, 1816-[1826] Verz. bek. Schmett. (1): [1-3], 4-16 (1816), (2): 17-32 (1819), (3): 33-48 (1819), (4): 49-64 (1819), (5): 65-80 (1819), (6): 81-96 (1819), (7): 97-112 (1819), (8): 113-128 (1819), (9): 129-144 (1819), (10): 145-160 (1819), (11): 161-176 (1819), (12): 177-192 (1820), (13): 193-208 (1820), (14): 209-224 (1821), (15): 225-240 (1821), (16): 241-256 (1821), (17): 257-272 (1823), (18): 273-288 (1823), (19): 289-304 (1823), (20): 305-320 (1825), (21): 321-336 (1825), (22): 337-352 (1825), (23-27): 353-431 ([1825])
Joicey & Talbot, 1921
New Lepidoptera Collected by Mr. T. A. Barns, in East Central Africa. New Forms of Rhopalocera Bull. Hill Mus. 1 (1) : 44-118, pl. 5-16
Kielland, 1978
A provisional checklist of the Rhopalocera of the eastern side of Lake Tanganyika Tijdschr. Ent. 121 (4) : 147-237
Lanz, 1896
Besprechung der von Dr. Bumiller 1893 aus Ostafrika mitgebrachten Schmetterlinge Dt. ent. Z. Iris 9 (1) : 113-147
Linnaeus, 1768
Sparrman (Thesis), Iter Chinense in Sparrman, Amoenit. acad., 7 (150): 497-506
Lucas, 1852
Description de nouvelles Espèces de Lépidoptères appartenant aux Collections entomologiques du Musée de Paris Revue Mag. Zool. (2) 4 (3): 128-141 (1852), 4 (4): 189-198 (1852), 4 (6): 290-300 (1852), 4 (7): 324-343 (1852), 4 (9): 422-432, pl. 10 (1852), 5 (7): 310-322 (1853)
MacLeay, [1826]
in King, Narrative of a Survey of the Intertropical and Western Coasts of Australia. 2 vols. in King, Narr. Surv. Austr. 2: -
Moore, 1872
Descriptions of new Indian Lepidoptera Proc. zool. Soc. Lond. 1872 (2) : 555-583, pl. 32-34
Moore, [1881]
The Lepidoptera of Ceylon Lepid. Ceylon 1 (1): 1-40, pl. 1-18 (Dec 1880), (2): 41-80, pl. 19-36 (Jan 1881), (3): 81-136, pl. 37-54 (Jun 1881), (4): 137-190, pl. 55-71 (Dec 1881)
Oberthür, 1878
Faune des Lépidoptères de la côte orientale d'Afrique Étud. d'Ent. 3 : 11-36, pl. 1-3
Rebel, 1914
Wissenschaftliche Ergebnisse der Expedition R. Grauernach. Zentralafrika. 1909-1911. Lepidoptera Ann. k.k. naturh. Hofmus. Wien. 28 : 219-294, pl. 17-24
Rogenhofer, 1889
Afrikanische Schmetterling des k. k. naturhist. Hofmuseums Ann. Mus. Wien. 4 : 547-554, pl. 23
Sharpe, 1891
Descriptions of New Butterflies collected by Mr. F. J. Jackson, F.Z.S:, in British East Africa, during his recent Expedition. - Part I & II Proc. Zool. Soc. Lond. 1891 : 187-194, pl. 16-17, : 633-638, pl. 48
Stoll, [1781]
Uitlandsche Kapellen (Papillons exotiques) in Cramer, Uitl. Kapellen 4 (26b-28): 29-90, pl. 305-336 (1780), (29-31): 91-164, pl. 337-372 (1781), (32-32): 165-252, 1-29, pl. 373-400 (1782)
Stoneham, 1940
Notes on the African genus Colotis, Hubnr. (Teracolus Swin.) and some new forms Bull. Stoneham Mus. (40): 1-4
Suffert, 1904
Neue afrikansche Tagfalter aus dem kön. zool. Museum, Berlin, und meiner Sammlung Dt. Ent. Z. Iris 17 (1) : 12-107, pl. 1-3
Suffert, 1904
Neue Tagfalter aus Deutsch-Ost-Afrika Dt. Ent. Z. Iris 17 (1) : 124-132
Trimen, 1883
Descriptions of twelve new species of South African Lepidoptera Rhopalocera Trans. ent. Soc. Lond. 1883 (4) : 347-363
Vane-Wright & de Jong, 2003
The butterflies of Sulawesi: annotated checklist for a critical island faunda Zool. Verh. Leiden 343: 3-268, pl. 1-16
Vollenhoven, 1869
Description des espèces nouvelles de coléoptères, orthoptères, lépidoptères et hémiptères, mentionees dans da liste précédente Faune Madag. 5 (Ins.) : 7-24, pl. 1-2
Wallengren, 1863
Lepidopterologische Mittheilungen (1-3) Wien ent. Monats. 4: 33-46,161-176 (1860), 7: 137-151 (1863)
Ward, 1870
Descriptions of new species of diurnal Lepidoptera from Madagascar Ent. mon. Mag. 7 : 30-32
Winhard, 2000
Pieridae I Butterflies of the world, 10: (1) 40pp, (2) 48pls


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Keep in mind that the taxonomic information is copied from various sources, and may include many inaccuracies. Expert help is welcome.