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Colias Fabricius, 1807
Clouded Yellows
Colias Fabricius, 1807; Magazin f. Insektenk. (Illiger) 6: 284, TS: Papilio hyale Linnaeus
Eurymus Horsfield, [1829]; Descr. Cat. lep. Ins. Mus. East India Coy (2): 134, TS: Papilio hyale Linnaeus
= ; [NL4A, 102]
Ganura Zetterstedt, [1839]; (unavail.)
= ; [NL4A, 102]
Scalidoneura Butler, 1871; Proc. zool. Soc. Lond. 1871 : 250, TS: Scalidoneura hermina Butler
= ; [NL4A, 102]; [AfrL]
Eriocolias Watson, 1895; Entomologist 28 : 167, TS: Papilio edusa Fabricius
= ; [NL4A, 102]; [AfrL]
Coliastes Hemming, 1931; Entomologist 64: 273, TS: Papilio hyale Linnaeus
= ; [NL4A, 102]; [AfrL]
Protocolias Petersen, 1963
= ; [NL4A, 102]
Mesocolias Petersen, 1963
= ; [NL4A, 102]
Neocolias Berger, 1986
= ; [NL4A, 102]
Palaeocolias Berger, 1986
= ; [NL4A, 102]
Eucolias Berger, 1986
= ; [NL4A, 102]
Similicolias Berger, 1986
= ; [NL4A, 102]
Paracolias Berger, 1986
= ; [NL4A, 102]
Colias, NSG Voucher Specimen [Wahlberg]
Colias, Pieridae Holarcticae [Heiner Ziegler]
Colias ; Godman & Salvin, [1889], Biol. centr.-amer., Lep. Rhop. 2: 151; Winhard, 2000, Butterflies of the world, 10: 9; [LCG]
Colias (Coliadinae) ; [AfrL]; [NL4A, 102]

fiColias hyale (Linnaeus, 1758)
S.Russia, Altai, Mongolia, NW.China. See [About maps] Vaaleakeltaperhonen Pale Clouded Yellow Ljusgul höfjäril Vaxfjäril Goldene Acht Le Soufre Szlaczkoń siarecznik
Papilio hyale Linnaeus, 1758; Syst. Nat. (Edn 10) 1 : 469, TL: S.England
?Colias pallens Butler, 1866; J. Linn. Soc. Zool. Lond. 9 (34) : 52, TL: Hakodate, Japan
Colias hyale ab. suzukii Matsumura, 1929; Insecta Matsumurana 3 (2/3) : 90, TL: Kyoto
Colias hyale, NSG Voucher Specimen [Wahlberg]
Colias hyale, Larvae of North-European Lepidoptera [Kimmo Silvonen]
Colias hyale, Svenska fjärilar [SVF]
Colias hyale, Moths and Butterflies of Europe [MBE]
Colias hyale, Extracts from [BRU] by [Ilya Osipov]
Colias hyale, Europaische Schmetterlinge [Mario Maier]
Colias hyale, Captain's European Butterfly Guide [Simon Coombes]
Colias hyale ; [BRU, 172]; [EBW]; [BOW: pl. 4, f. 4-5]; [MRS, 220]; [Otakar Kudrna]
[hyale-1mv.jpg] 1024x768 (~234Kb) underside male Eiskeller, Berlin, Germany, 10-94, Photo © S. Shuichi Haupt [hyale-2fv.jpg] 462x400 (~33Kb) underside female The Irtysh River left bank floodland within the Omsk City, West Siberia, Russia. 27th August 1996 (on Cirsium setosum (Willd.) M.B.), Photo © Oleg Kosterin [hyale-3vU.jpg] identification uncertain 800x534 (~92Kb) underside Ukraine, Ivano Frankovsk, near Kalush, Stankova, 24.10.2001, Photo © Erich Mangl
[hyale-4vU.jpg] identification uncertain 1000x1162 (~159Kb) underside Russia, Udmurtia, Izhevsk (parklike areas along shore of "Izhevsky Prud"), 5.8.2004, Photo © Markku Savela [hyale-5fuv.jpg] 536x768 (~141Kb) upperside underside female Russia, Kaluga area, 09.09.2005, Photo © D. Smirnov [hyale-5muv.jpg] 586x768 (~141Kb) upperside underside male Russia, Kaluga area, 09.09.2005, Photo © D. Smirnov
[hyale-fv.jpg] 500x373 (~31Kb) underside female Axat, Pyrenees orientales, France 24/07/96, Photo © Simon Coombes [hyale.jpg] 600x448 (~67Kb) France, Pyrenees Orientales, Axat, Summer 1996, Photo © Simon Coombes

Larva on Vicia cracca [SPRK], Fabaceae, Vicia, Coronilla, Medicago, Lotus, Cytisus, Trifolium [BRU]


Europe, Ukraine, Kirghizia, Kazkhstan, S.Russia aegra Verity, 1947; Le Farfalle diurn. d'Italia 3: 262
= ; [LCG, 40]
alba Derenne, 1933; Lambillionea 33: 184
= ; [LCG, 40]
ab. albescens Metschl, 1922; Mitt. munchn. ent. Ges. 12: 6
= ; [LCG, 40]
albinotica Goodson, 1960; Ent. Gaz. 11: 18
= ; [LCG, 40]
alboradiata Herzig, 1935; Int. ent. Z. 29: 365
= ; [LCG, 40]
antehyale Verity, 1947; Le Farfalle diurn. d'Italia 3: 264
= ; [LCG, 40]
anteubercalida Verity, 1947; Le Farfalle diurn. d'Italia 3: 266
= ; [LCG, 40]
apicata Tutt, 1896; Brit. Butts: 253
= ; [LCG, 40]
appendiculata Verity, 1947; Le Farfalle diurn. d'Italia 3: 260
= ; [LCG, 40]
Colyas[sic] hyale ab. ♀ argentea Fritsch, 1913; Ent. Rundschau 30 : 46
= ; [LCG, 40]
argyphea Lowe, 1909; Ent. Rec.: 36
= ; [LCG, 40]
atava Reutti, 1853; Beitr. Rhein. Naturg. 3: ?
= ; [LCG, 40]
bipupillata Cabeau, 1912; Rev. mens. Soc. ent. Namur 12: 54
= ; [LCG, 40]
brabantica Strand, 1912; Ent. Zeit. 25: 49-50
= ; [LCG, 40]
brevis de Crombrugghe, 1911; Rev. mens. Soc. ent. Namur 11: 104
= ; [LCG, 40]
canarina Stauder, 1922; Mitt. münch. ent. Ges. 12 (1-6) : 25
= ; [LCG, 40]
cleopatra Latier, 1935; P.-v. Soc. linn. Bordeaux 86: 149
= ; [LCG, 40]
crassipuncta Le Charles, 1927; Bull. Soc. ent. Fr. 1927 (8) : 138
= ; [LCG, 41]
crocoexpansa Caruel, 1955; Revue Française de Lépid. 1-2: 27
= ; [LCG, 41]
deannulata Rocci, 1920; Atti Soc. Lig. 30 (4): 30
= ; [LCG, 41]
Colias hyale ab. demarginata Nitsche, 1913; in Rebel, Verh. zool.-bot. Ges. Wien 63: 21
= ; [LCG, 41]
deuara Herzig, 1935; Int. ent. Z. 29: 365
= ; [LCG, 41]
duplex Ksienschopolski, 1911; Trudy obsch. Izsl. Volyni Zitomir 8: 28, pl. 1, f. 2
= ; [LCG, 41]
elongata Vorbrodt & Müller-Rutz, 1917; Mitt. schweiz. ent. Ges. 12 (9-10): 434
= ; [LCG, 41]
emarginata Röber, [1907]; in Seitz, Grossschmett. Erde 1 : 65
= ; [LCG, 41]
eratoides Strand, 1918; Soc. ent. 33 (7): 27, TL: Turkestan
= ; [LCG, 41]
flava Husz, 1880; Jb. Ungar Karpath. Ver. 8: 272
= ; [LCG, 41]
flavida Gruber, 1932; Ent. Zeit. 46: 195, f. 14
= ; [LCG, 41]
ab. flavoapicalis Metschl, 1922; Mitt. munchn. ent. Ges. 12: 7
= ; [LCG, 41]
flavofasciata Lambillion, 1907; Cat. Lep. Belg. (2nd ed): 409
= ; [LCG, 41]
flavopupillata Soja, 1929; Ver. zool.-bot. Ges. Wien. 79: 107
= ; [LCG, 41]
ab. flavoradiata Metschl, 1922; Mitt. munchn. ent. Ges. 12: 7
= ; [LCG, 41]
gartneri Skala, 1912; Ver. naturf. Ver. Brünn 50: 63
= ; [LCG, 41]
gavalgnii Stauder, 1921; Z. Wiss. InsekBiol. 16: 220
= ; [LCG, 41]
gloriciana Fritsch, 1918; Ent. Rundschau 35 : 6
= ; [LCG, 41]
grisandrocontata Lempke, 1954; Tijdschr. Ent. 97 : 311
= ; [LCG, 41]
griseoapicata Herzig, 1935; Int. ent. Z. 29: 365
= ; [LCG, 41]
heliceides de Selys de Longchamps, 1844; Mém. Soc. roy. Sci. Liége 2: 30
= ; [LCG, 41]
immaculata Derenne, 1925; Rev. Mens. Soc. ent. Namur 25: 92
= ; [LCG, 41]
impunctata Rocci, 1920; Atti Soc. Lig. 30 (4): 30
= ; [LCG, 41]
infrastriata Lempke, 1954; Tijdschr. Ent. 97 : 306
= ; [LCG, 41]
inornatura Derenne, 1923; Rev. Mens. Soc. ent. Namur 23: 39
= ; [LCG, 41]
intermedia Tutt, 1896; Brit. Butts: 253
= ; [LCG, 41]
inversa Alphéraky, 1881; Horae Soc. ent. Ross. 16 (3-4): 434
= ; [LCG, 41]
junior Geest, 1905; Z. wiss. InsekBiol. 1: 380
= ; [LCG, 41]
macropuncta Finke, 1934; Int. ent. Z. 28: 293, TL: Germany, Göttingen
= ; [LCG, 41]
magnoimaculata Lempke, 1954; Tijdschr. Ent. 97 : 314
= ; [LCG, 41]
melanina Verity, [1909]; Rhopalocera Palaearctica 1: 223
= ; [LCG, 41]
melaina Mück, 1919; Ent. Zeit. 33 (9): 35, TL: Bâle
= ; [LCG, 42]
mellaertsi Lambillion, 1913; Rev. Mesn. Soc. Ent. Namur 13: 126
= ; [LCG, 42]
metschli Gruber, 1932; Ent. Zeit. 46: 195, f. 15
= ; [LCG, 42]
minor van Mellaerts, 1926; Lambillionea 26: 84
= ; [LCG, 42]
myrmhyale Meeke, 1926; Int. ent. Z.: 173
= ; [LCG, 42]
nigriformis Finke, 1934; Int. ent. Z. 28: 394, TL: Germany
= ; [LCG, 42]
nigripuncta Caruel, 1955; Revue Française de Lépid. 1-2: 27
= ; [LCG, 42]
nigrofasciata Grum-Grshimailo, 1884; in Romanoff, Mém. Lép. 1 : 163-164, TL: Saratov
= ; [LCG, 42]
nigrosparsata Lempke, 1954; Tijdschr. Ent. 97 : 312
= ; [LCG, 42]
obsoleta Tutt, 1896; Brit. Butts: 259
= ; [LCG, 42]
ocellata Dufrane, 1947; Bull. Ann. Soc. ent. Belg. 83 (1/2) : 70
= ; [LCG, 42]
Colias hyale ♀ ab. ochro-cretacea Fritsch, 1918; Ent. Rundschau 35 : 6
= ; [LCG, 42]
Colias hyale ab. goriciana Fritsch, 1918; Ent. Rundschau 35 : 6
omnimarginata Hafner, 1912; Carniola: 221, pl. 3, f. 41
= ; [LCG, 42]
opposita Zusanek, 1918; Z. öst. EntVer. 3: 36
= ; [LCG, 42]
ab. pallescens Metschl, 1922; Mitt. munchn. ent. Ges. 12: 6
= ; [LCG, 42]
pallida Robson & Gardner, 1886; Young Naturalist, suppl.: 4
= ; [LCG, 42]
pallida Cockerell, 1889; Entomologist 22 : 4 [?]
= ; [LCG, 42]
pallidior Cockerell, 1912; Entomologist 45: 323
= ; [LCG, 42]
parisiensis Oberthür, 1936; Lambillionea 36: 90
= ; [LCG, 42]
parvipuncta Lempke, 1954; Tijdschr. Ent. 97 : 313
= ; [LCG, 42]
posticomunita Lempke, 1954; Tijdschr. Ent. 97 : 307
= ; [LCG, 42]
postrema Rocci, 1920; Atti Soc. Lig. 30 (4): 29
= ; [LCG, 42]
ab. pseudohelice Metschl, 1922; Mitt. munchn. ent. Ges. 12: 6
= ; [LCG, 42]
pupillata Lempke, 1936; Lambillionea 36: 103
= ; [LCG, 42]
pygmaea Lambillion, 1906; Rev. Mens. Soc. ent. namur. 6: 22
= ; [LCG, 42]
minor Vorbrodt, 1912; Die Schmett. Schweiz 1 : 30
= ; [LCG, 42]
radiata Geest, 1905; Z. wiss. InsekBiol. 1: 379
= ; [LCG, 42]
radiiformis Schultz, 1904; Nyt. Mag. naturvid. 42: 41
= ; [LCG, 42]
rufa Verity, [1909]; Rhopalocera Palaearctica 1: 222, pl. 41, f. 1
= ; [LCG, 42]
sagittata Schnaider, 1950; Polsk. Pismo. Ent. 19: 240
= ; [LCG, 42]
schonfeldi Mezger, 1926; Ent. Anz. 6: 18
= ; [LCG, 42]
sieversoides Verity, [1909]; Rhopalocera Palaearctica 1: 221, pl. 40, f. 34
= ; [LCG, 42]
simplex Neuburger, 1905; Soc. ent. 20 (6) : 42
= ; [LCG, 42]
striata Zusanek, 1918; Z. öst. EntVer. 3: 36
= ; [LCG, 42]
striata Romaniszyn, 1926; Polsk. Pismo ent. 5: 144
= ; [LCG, 42]
submarginata Finke, 1934; Int. ent. Z. 28: 394, TL: Germany
= ; [LCG, 42]
subobsoleta Pionneau, 1936; Echange 52: 23
= ; [LCG, 42]
sufflava Verity, 1947; Le Farfalle diurn. d'Italia 3: 262
= ; [LCG, 43]
Colias hyale var. supervacanea Krulikowsky, 1908; Soc. Ent. 23 (1): 3, TL: Wiatka; Kasan
= ; [LCG, 43]
trimaculata Steiner, 1935; Int. etn. Z. 29: 269
= ; [LCG, 43]
uber Verity, 1926; Ent. Rec. J. Var. 38 : 171
= ; [LCG, 43]
ubercalida Verity, 1947; Le Farfalle diurn. d'Italia 3: 266
= ; [LCG, 43]
uhli Kovats, 1899; Ent. Zeit. 12: 169
= ; [LCG, 43]
unimaculata Tutt, 1896; Brit. Butts: 253
= ; [LCG, 43]
venata Lempke, 1936; Lambillionea 36: 76
= ; [LCG, 43]
vernalis Verity, [1909]; Rhopalocera Palaearctica 1: 222, pl. 47, f. 32
= ; [LCG, 43]
viridescens Sloper, 1903; Wheeler. Bull. Switz.: 69
= ; [LCG, 43]
viridis van Mellaerts, 1926; Lambillionea 26: 84
= ; [LCG, 43]
Colias hyale hyale ; Winhard, 2000, Butterflies of the world, 10: 9, pl. 11, f. 1-2
Colias poliographus inversa ;
Colias hyale hyale ; [LCG]
C. h. altaica Verity, 1911
Altai, Sayan hyale altaica Verity, 1911; Rhopalocera Palaearctica 1: 348, pl. 69, f. 4-5
Colias hyale altaica ; [BRU, 172]
Colias hyale hyale f. altaica ; [LCG, 40]
C. h. alta Staudinger, 1886
Tian-Shan, Ghissar, Darvaz, Alai, Pamirs hyale alta Staudinger, 1886; Stett. Ent. Ztg. 47 (4-6): 200, TL: Kirgizia, Alai Mts.
Colias poliographus alta ; [BRU, 173]
Colias hyale alta ; [LCG, 43]
C. h. palidis Fruhstorfer, 1910
East Siberia hyale palidis Fruhstorfer, 1910; in Seitz, Gross-Schmett. Erde 9 : 164
= Colias hyale irkutskana ; [LCG, 44]
palides[sic, recte palidis] ; [LCG, 44]
Colias hyale palidis ; [BRU, 172]
C. h. irkutskana Stauder, 1924
Transbaikalian hyale irkutskana Stauder, 1924; Mitt. münch. ent. Ges. 14: 62, TL: Irkutsk, Ojek
f. fulvocoma Krulikovsky, 1907; Revue Russe Ent. 7: 29, TL: Viatka
= ; [LCG, 44]
Colias hyale irkutskana ; [BRU, 172]; [LCG, 43]
C. h. novasinensis Reissinger, 1989
Gansu Colias hyale novasinensis Reissinger, 1989; Neue Ent. Nachricten 26: 199, TL: Gansu, Xining
Colias hyale novasinensis ; [LCG]

Colias sareptensis Staudinger, 1871
SEU, CEU, S.Russia, Asia Minor, Caucasus, Transcaucasia - Middle Asia - China. See [About maps] Berger's Clouded Yellow Sydlig ljusgul höfjäril Szlaczkoń południowiec
Colias sareptensis Staudinger, 1871; in Staudinger & Wocke, Cat. Lepid. eur. Faunegeb. (Edn. 2) : 5, TL: Sarepta
Colias alfacariensis Ribbe, 1905; Soc. ent. 20 (18) : 137, TL: Andalusia
Colias australis Verity, 1911; Rhopalocera Palaearctica 1: 347, TL: Andalusia, Spain
calida Verity, 1916; Ent. Rec. J. Var. 28 (5): 99
= Colias alfacariensis ; [LCG, 45]
Colias alfacariensis ubercalida Reissinger, ?1972; 112[?]
= Colias alfacariensis ; [LCG, 45]
Colias alfacariensis senonica Reissinger, 1972; Atalanta 3: 359
= Colias alfacariensis ; [LCG, 45]
Colias alfacariensis paracalida Reissinger, 1972; Atalanta 3: 366
= Colias alfacariensis ; [LCG, 45]
Colias alfacariensis orthocalida Reissinger, 1974; Atalanta 5: (1-33)
= Colias alfacariensis ; [LCG, 45]
Colias alfacariensis kantaraica Reissinger, 1989; Neue Ent. Nachricten 26: 223
= Colias alfacariensis ; [LCG, 45]
Colias alfacariensis bergeri Reissinger, 1989; Neue Ent. Nachricten 26: 126
= Colias alfacariensis ; [LCG, 45]
Colias alfacariensis antebergeri Reissinger, 1989; Neue Ent. Nachricten 26: 126
= Colias alfacariensis ; [LCG, 45]
Colias alfacariensis postbergeri Reissinger, 1989; Neue Ent. Nachricten 26: 126
= Colias alfacariensis ; [LCG, 45]
anteorthocalida Reissinger, 1989; Neue Ent. Nachricten 26: 82
= Colias alfacariensis ; [LCG, 48]
postorthocalida Reissinger, 1989; Neue Ent. Nachricten 26: 82
= Colias alfacariensis ; [LCG, 48]
Colias alfacariensis rumilica Reissinger, 1989; Neue Ent. Nachricten 26: 129
= Colias alfacariensis ; [LCG, 48]
Colias alfacariensis slavonica Reissinger, 1989; Neue Ent. Nachricten 26: 154
= Colias alfacariensis ; [LCG, 50]
Colias alfacariensis magyarica Reissinger, 1989; Neue Ent. Nachricten 26: 162
= Colias alfacariensis ; [LCG, 46]
Colias alfacariensis metacalida Reissinger, 1989; Neue Ent. Nachricten 26: 170
= Colias alfacariensis ; [LCG, 47]
Colias alfacariensis hyrcanica Reissinger, 1989; Neue Ent. Nachricten 26: 133
= Colias alfacariensis ; [LCG, 46]
Colias alfacariensis mandarina Reissinger, 1989; Neue Ent. Nachricten 26: 189
= Colias alfacariensis ; [LCG, 47]
Colias alfacariensis, NSG Voucher Specimen [Wahlberg]
Colias alfacariensis, Larvae of North-European Lepidoptera [Kimmo Silvonen]
Colias alfacariensis, Moths and Butterflies of Europe [MBE]
Colias sareptensis, Extracts from [BRU] by [Ilya Osipov]
Colias australis, Europaische Schmetterlinge [Mario Maier]
Colias alfacariensis, Captain's European Butterfly Guide [Simon Coombes]
Colias australis ; [EBW]
Colias alfacariensis ; [BMAT: 101, pl. 32, f. 14-16]; Winhard, 2000, Butterflies of the world, 10: 9, pl. 11, f. 3-6; [LCG, 44]; [Otakar Kudrna]
Colias sareptensis ; [BRU, 172]

Larva on Coronilla varia, Hippocrepis comosa, Astragalus monspessulanus [LCG, (for alfacariensis)]

Colias alfacariensis ; [LCG, 49 (form)]
C. s. vihorlatensis Reissinger, 1989
Carpathians Colias alfacariensis vihorlatensis Reissinger, 1989; Neue Ent. Nachricten 26: 166
= Colias alfacariensis ; [LCG, 50]
C. s. remota Reissinger, 1989
SEU, Caucaus Colias alfacariensis remota Reissinger, 1989; Neue Ent. Nachricten 26: 149
= Colias alfacariensis ; [LCG, 48]
Colias sareptensis remota ; [BRU, 172]
C. s. fontainei Reissinger, 1989
Armenian Highland, Talysh, Kopet-Dagh Colias alfacariensis fontainei Reissinger, 1989; Neue Ent. Nachricten 26: 141
= Colias alfacariensis ; [LCG, 45]
C. s. saissanica Reissinger, 1989
SE.Kazakhstan Colias alfacariensis saissanica Reissinger, 1989
= Colias alfacariensis ; [LCG, 49]
Colias sareptensis saissanica ; [BRU, 172]

Colias erate (Esper, 1805)
-. See [About maps] Eastern Pale Clouded Yellow Szlaczkoń erate
Papilio erate Esper, 1805; Die Schmett., Th I, Suppl. Th. 2 (11): 13, pl. 119, f. 3, TL: Sarepta, S. Russia
Colias erate edusoides Krulikovsky, 1903; Rev. Russ. Ent. 3: 301, TL: Aksu
Colias erate, NSG Voucher Specimen [Wahlberg]
Colias erate, Moths and Butterflies of Europe [MBE]
Colias erate, Extracts from [BRU] by [Ilya Osipov]
Colias erate ; [BRU, 173]; [EBW]; [BOW: pl. 4, f. 1-2]; [MRS, 218]; [AfrL]; [Otakar Kudrna]
[erate-1mv.jpg] 951x624 (~56Kb) underside male Sukayu, Aomori, Japan, 8-90, Photo © S. Shuichi Haupt [erate-2v.jpg] 800x707 (~67Kb) underside Japan: Lake Kawaguchiko, Yamanashi Prefecture, 23.08.2007, Photo © Peter Payzant [erate-3fvU.jpg] identification uncertain 1127x1230 (~316Kb) underside female Ethiopia, Amhara, vicinity of Hayk city. 6th August 2012, Photo © Oleg Kosterin

Larva on Medicago, Trifolium, Onobrychis, Melilotus, (etc.) [BRU]


Ukraine, Turkey, Lebanon, Bulgaria, Romania, Macedonia, Greece, Hungary, Austria, Turkmenistan, Kazakhstan, Kirhizia, Uzbekistan, Tadzhikistan, Afghanistan Colias neriene Fischer von Waldheim, 1823; Entom. Imp. ross., 2: 251, pl. 11, f. 3-4
afghana Bang-Haas, 1927; Horae Macrolep. Palaearct. 1 : 42, TL: Afghanistan, Paghman Mts.
= ; [LCG, 52]
albida Rühl, [1893]; Palaearktischen Grosschmetterlinge: 157
= ; [LCG, 52]
androconiata Jachontov, 1910; Revue Russe Ent. 9 (4): 380
= ; [LCG, 52]
beckeri Gerhard, 1882; Berl. ent. Zs. 26 (1) : 125
= ; [LCG, 52]
Colias erate edusoides Verity, 1908; Rhopalocera Palaearctica 1: 219, pl. 47, f. 24
Colias erate benesignata Sheljuzhko, 1913; Dt. Ent. Z. Iris 27 (1) : 20 (repl. Colias erate edusoides Verity, 1908)
= ; [LCG, 52]
chryseis Röber, [1907]; in Seitz, Grossschmett. Erde 1 : 66, pl. 26b
= ; [LCG, 52]
chrysodona Boisduval, 1840; Genera Index Eur. Lepid. : 7
= ; [LCG, 52]
chrysohyaleoides Verity, [1908]; Rhopalocera Palaearctica 1: 219
chrysopallida Verity, [1908]; Rhopalocera Palaearctica 1: 221, pl. 57, f. 29
= ; [LCG, 52]
chrysothemoides Verity, [1908]; Rhopalocera Palaearctica 1: 219, pl. 47, f. 25
= ; [LCG, 52]
citronea Mezger, 1932; Lambillionea, 32: 4
= ; [LCG, 52]
conjuncta Verity, [1908]; Rhopalocera Palaearctica 1: 219
= ; [LCG, 52]
costaenigrata Mezger, 1932; Lambillionea, 32: 4
= ; [LCG, 52]
diffusa Verity, [1908]; Rhopalocera Palaearctica 1: 219, pl. 40, f. 30
= ; [LCG, 52]
flavomaculata Mezger, 1932; Lambillionea, 32: 4
= ; [LCG, 52]
garica Verity, 1911; Rhopalocera Palaearctica 1: 348, pl. 69, f. 6-7
= ; [LCG, 52]
helichta Lederer, 1853; Verh. zool.-bot. Ver. Wien 2 (Abh.) : 33 (spell.?)
= ; [LCG, 52 (as helichtha[sic])]
helichta Talbot, 1935; Lep. Cat. 66: 423
= Colias erate erate f. androconiata Jachontov, 1910; [LCG, 52]
hyaleoides Grum-Grshimailo, 1890; in Romanoff, Mém. Lép. 4 : 321, TL: Fergana
= ; [LCG, 52]
Colias erate ab. maculigena Avinoff, 1910; Horae Soc. Ent. Ross. 39 : 231, TL: Borokoro
= ; [LCG, 52]
pallida Staudinger, 1861; in Staudinger & Wocke, Cat. Lep. (ed. 1): 3
= ; [LCG, 52]
parva Rühl, [1893]; Palaearktischen Grosschmetterlinge: 157
= ; [LCG, 52]
semifaillaei Pionneau, 1928; Echange 44 (431): 3
= ; [LCG, 52]
unimaculata Pionneau, 1928; Echange 44 (431): 3
= ; [LCG, 52]
Colias erate erate f. chrysodona ; Winhard, 2000, Butterflies of the world, 10: 10, pl. 11, f. 10
Colias erate erate f. chrysopallida ; Winhard, 2000, Butterflies of the world, 10: 9, pl. 11, f. 9
Colias erate erate f. hyaleoides ; Winhard, 2000, Butterflies of the world, 10: 9, pl. 11, f. 8
Colias erate erate ; Winhard, 2000, Butterflies of the world, 10: 9, pl. 11, f. 7; [LCG, 51]
C. e. amdensis Verity, 1911
China (Qinghai, Gansu, Sichuan) erate amdensis Verity, 1911; Rhopalocera Palaearctica 1: 348, pl. 41, f. 8, TL: Amndo, Kukunor
Colias poliographus ab. chrysocoma Grum-Grshimailo, 1893; Horae Soc. ent. Ross. 27 : 379, TL: Sichuan
= ; [LCG, 52]
hera Grum-Grshimailo, 1893; Horae Soc. ent. Ross. 27 : 379, TL: Sichuan
= ; [LCG, 52]
Colias erate amdensis ; [LCG, 52]
C. e. marnoana Rogenhofer, 1884
Sudan, Ethiopia, SW.Arabia Colias marnoana Rogenhofer, 1884; Verh. zool.-bot. Ges. Wien 33 : 22, TL: Sudan
Colias electo ab. kostlani Strand, 1911; Ent. Rundsch. 28 : 140, TL: Ethiopia
= ; [AfrL]
Colias erate marnoana ♀ f. solilucis Berger, 1953; Entomologist 86 : 172
= ; [LCG, 54]
Colias erate marnoana f. androides Storace, 1956; Ann. Mus. Civ. Stor. nat. Genova 68 : 349
= ; [LCG, 54]; [LCG, 54]
Colias erate marnoana ; [BAFR, 54]; [LCG, 53]; [AfrL]
C. e. sinensis Verity, 1911
Yunnan?, Mandschuria, N.Korea Colias erate sinensis Verity, 1911; Rhopalocera Palaearctica 1: 349, pl. 69, f. 19-22, TL: Kioi-Kiang, Chang-Yang
Colias erate sinensis ; [MRS, 218]; [LCG, 55]
C. e. formosana Shirôzu, 1955
Taiwan Colias erate formosana Shirôzu, 1955; Sieboldia 1 (3): 229, pl. 30, f. 1-4
Colias erate formosana ; [MRS, 218]; [LCG, 52]

Larva on Trifolium pratense, T. repens, Medicago polymorpha, M. sativa, Lotus australis, Astragalus sinicus [MRS]

C. e. lativitta Moore, 1882
Nepal, N.India erate lativitta Moore, 1882; Proc. zool. Soc. Lond. 1882 (1) : 255, TL: Nepal
erate gigantea Verity, 1911; Rhopalocera Palaearctica 1: 347, pl. 72, f. 25 (preocc. Colias gigantea Strecker)
Colias erate tania Hemming, 1933; Entomologist 66: 278 (repl. erate gigantea Verity, 1911)
= ; [LCG, 53]
glicia Fruhstorfer, 1910; in Seitz, Gross-Schmett. Erde 9 : 164
= ; [LCG, 53]
kachgarica Verity, 1911; Rhopalocera Palaearctica 1: 348, pl. 69, f. 6-7
= ; [LCG, 53]
Colias erate lativitta ; [LCG, 53]
C. e. poliographus Motschulsky, 1860
Japan, Amur, Ussuri, Sakhalin, Tian Shan Colias poliographus Motschulsky, 1860; Etud. Ent. 9: 29, TL: "Khokodody", Japan
Colias simoda de l'Orza, 1869; Lep. Jap.: 16
= ; [LCG, 55]
immamis Verity, 1911; Rhopalocera Palaearctica 1: 349, pl. 41, f. 10
= ; [LCG, 55]
murina Fruhstorfer, 1910; in Seitz, Gross-Schmett. Erde 9 : 164
= ; [LCG, 55]
napata Fruhstorfer, 1910; in Seitz, Gross-Schmett. Erde 9 : 164
= ; [LCG, 55]
pyxagathus Fruhstorfer, 1910; in Seitz, Gross-Schmett. Erde 9 : 164
= ; [LCG, 55]
Colias hyale zuzuki Matsumura, 1908; Ent. mon. Mag. 2: 120, pl. 5, f. 8, TL: Kyoto
= ; [LCG, 55]
Colias erata poliographus f. sagakoe Yano, 1958; Tyô to Ga 9 (3): 36, TL: Senjo-ga-ha-ra, Nikko, Tochigi Pref., Japan
Colias poliographus, Extracts from [BRU] by [Ilya Osipov]
Colias poliographus ; [BRU, 173]
Colias erate poliographus ; Huang, 2001; Winhard, 2000, Butterflies of the world, 10: 10, pl. 11, f. 11-12; [LCG, 54]

Larva on Trifolium, Vicia, (etc.) Glycine max [BRU], Parochetus communis, Lucerne spp. [MRS]

C. e. tomarias Bryk, 1942
Kuriles Colias hyale tomarias Bryk, 1942; Dt. Ent. Z. Iris 56 : 12, TL: Kurile Is., Kumashir & Tomari
tokotana Bryk, 1942; Dt. Ent. Z. Iris 56 : 12
Colias poliographus tomarias ; [BRU]
Colias erate tomarias ; [LCG, 55]
C. e. naukratis Fruhstorfer, 1909
Altai, S.Siberia, Transbaikalia erate naukratis Fruhstorfer, 1909; in Seitz, Gross-Schmett. Erde 9 : 164
Colias poliographus naukratis ; [BRU]
Colias erate naukratis ; [LCG, 54]
C. e. nilagiriensis C. & R. Felder, 1859
S.India erate nilagiriensis C. & R. Felder, 1859; Wien. ent. Monats. 3 (12): 395, TL: S.India, Nilgiri Mts.
Colias erate nilagiriensis ; [LCG, 54]

Colias libanotica Lederer, 1858
-. See [About maps] Colias libanotica Lederer, 1858; Wien. ent. Monats. 2 (5) : 140, pl. 2, f. 1-2, TL: Syria [Lebanon]
Israel, Lebanon, Turkey hybrida Verity, [1909]; Rhopalocera Palaearctica 1: 259
= ; [LCG, 56]
offusa Verity, [1909]; Rhopalocera Palaearctica 1: 259
= ; [LCG, 56]
pulchior Verity, [1909]; Rhopalocera Palaearctica 1: 259
Colias libanotica libanotica ; [LCG, 56]
C. l. heldreichi Staudinger, 1862
Greece Colias libanotica heldreichi Staudinger, 1862; Stettin Ent. Ztg 23 (4-6) : 257, TL: Mt. Veluchi, Greece
fountainei Aigner-Abafi, 1901; Rovart. Lap. 8: 30, TL: Mt. Chelmos
= Colias libanotica heldreichii ; [LCG, 57]
griseoviridis Verity, [1909]; Rhopalocera Palaearctica 1: 260-274, pl. 45, f. 29, TL: Greece
= Colias libanotica heldreichii ; [LCG, 57]
Colias aurorina heldreichi ;
Colias libanotica heldreichii f. fountainei ; Winhard, 2000, Butterflies of the world, 10: 10, pl. 11, f. 14
Colias libanotica heldreichii ; Winhard, 2000, Butterflies of the world, 10: 10, pl. 11, f. 13; [LCG, 56]
C. l. sovarensis Blom, 1979
Iran Colias aurorina sovarensis Blom, 1979; Ent. Ber. Amst. 39 (1): (10-14), TL: Iran, 8km N-E Khosh-Yeilaq, Kuh-e-Sovar Mts, 2000-2500m
Colias libanotica sovarensis ; [LCG, 57]

Colias aurorina Herrich-Schäffer, 1850
-. See [About maps] Dawn Clouded Yellow
Colias aurorina Herrich-Schäffer, 1850; Syst. Bearb. Schmett. Europ. 1 (43): f. 453-456, TL: Armenia
Colias aurorina, NSG Voucher Specimen [Wahlberg]
Colias aurorina, Moths and Butterflies of Europe [MBE]
Colias aurorina, Extracts from [BRU] by [Ilya Osipov]
Colias aurorina, Captain's European Butterfly Guide [Simon Coombes]
Colias aurorina ; [BRU, 177]; [Otakar Kudrna]

Larva on Astragalus [H&R], Astracantha caucasica, A. denudata [BRU, Nekrutenko, 1990]

Armenia, Georgia, Azerbaijan, Talysh, N.Iran, Turkey alba Rühl, [1893]; Palaearktischen Grosschmetterlinge: 731
= ; [LCG, 58]
aurantiaca Rühl, [1893]; Palaearktischen Grosschmetterlinge: 730
= ; [LCG, 58]
deserticola Verity, [1909]; Rhopalocera Palaearctica 1: 258
= ; [LCG, 58]
Colias aurorina aurorina ; [LCG, 57]
C. a. daghestanica Verhulst, 1994
Daghestan, Armenia Colias aurorina daghestanica Verhulst, 1994; Lambillionea 94: 115, TL: Daghestan, Rutul
Colias aurorina daghestanica ; [LCG, 58]
C. a. kermana Eckweiler, 1979
Iran (Kerman) Colias aurorina kermana Eckweiler, 1979; Ent. Zs. Frankf. a. M. 89 (3): 29-32, TL: Iran, Kerman, Jiroft-Rayen, 2800-3000m
Colias auronina kermana ; [LCG, 59]

Larva on Astragalus sp. [LCG, 59]

C. a. rosei Gross & Ebert, 1975
N.Iran (Elburs) Colias aurorina rosei Gross & Ebert, 1975; J. ent. Soc. Iran, suppl. 1: 11, TL: Iran, Elburs, Damavand
Colias aurorina rosei ; [LCG, 59]
C. a. taurica Rebel, 1901
Turkey Colias aurorina f. taurica Rebel, 1901; Verh. zool.-bot. Ges. Wien 51 : 310, TL: Turkey, Taurus
leucothea Rebel, 1917; Sber. Akad. Wiss. Wien (126) 1: 251
= ; [LCG, 60]
Colias aurorina taurica ; [LCG, 59]
C. a. transcaspica Christoph, 1889
Turkmenistan (Kopet-Dagh, W.Transcaspia) Colias aurorina v. transcaspica Christoph, 1889; in Romanoff, Mém. Lépid. 5 : 4, pl. 1, f. 1, TL: W.Transcaspia, Nuchur
chrysocoma Eversmann, 1851; Bull. Soc. imp. Nat. Moscou 24 (2) : 622, TL: Caucasia
= ; [LCG, 60]
tamara Nordmann, 1851; Bull. Soc. imp. Nat. Moscou 24 (4): 413, TL: Caucasia
= ; [LCG, 60]
Colias aurorina transcaspica ; [LCG, 60]

Colias sagartia Lederer, 1869
N.Iran. See [About maps] Colias sagartia Lederer, 1869; Horae Soc. ent. ross. 6 (2) : 75, TL: Iran
daubii Geest, 1905; Z. wiss. InsekBiol. 1: (378-381)
= ; [LCG, 61]
lisa Geest, 1905; Z. wiss. InsekBiol. 1: (378-381)
= ; [LCG, 61]
pullata Neuburger, 1905; Soc. ent. 20 (6) : 42
= ; [LCG, 61]
Colias sagartia ; [LCG, 60]

Larva on Acantholimon, Astragalus spp. [LCG, 61]


Colias chlorocoma Christoph, 1888
Transcaucasia, Turkey, Iran. See [About maps] Colias chlorocoma Christoph, 1888; Horae Soc. ent. Ross. 22 (3-4) : 308, TL: Turkey
Colias chlorocoma, NSG Voucher Specimen [Wahlberg]
Colias chlorocoma, Extracts from [BRU] by [Ilya Osipov]
Colias chlorocoma ; [BRU, 177]

Larva on Onobrychis cornuta [BRU, Nekrutenko, 1990]

Turkey ochracea Verhulst, 1993; Lambillionea 93 (1): 44
= ; [LCG, 62]
Colias chlorocoma ♀ ab. sulphurea Miller, [1913]; Dt. ent. Z. Iris 26 (4) : 223, TL: Transcaucasia
= ; [LCG, 62]
Colias chlorocoma chlorocoma ; Winhard, 2000, Butterflies of the world, 10: 10, pl. 11, f. 15; [LCG, 61]

Larva on Astragalus aurens, A. lycioides [LCG, 62]

C. c. aladagensis Verhulst, 1993
Turkey Colias chlorocoma aladagensis Verhulst, 1993; Lambillionea 93 (1): 44, TL: Turkey, Nidge, Aladglari, Pinarbasi 200-2600m
Colias chlorocoma aladagensis ; [LCG, 62]

Larva on Astragalus angusitfolius? [LCG, 63]

C. c. tkatschukovi O. Bang-Haas, 1936
Armenia, Azerbaijan chlorocoma tkatschukovi O. Bang-Haas, 1936; Ent. Z. Frankf. a. M. 50: 287, TL: Armenia, Daralagaes 2000m, 45.30E, 39.25N
Colias chlorocoma tkatschukovi ; [LCG, 63]
C. c. wyatti Häuser & Schurian, 1978
Iran chlorocoma wyatti Häuser & Schurian, 1978; Atalanta, 9 (3): (240-244), TL: Iran, Dugijan
Colias chlorocoma wyatti ; [LCG, 63]

Colias caucasica Staudinger, 1871
W.Caucasus, NE.Turkey. See [About maps] Balkan Clouded Yellow
Colias caucasica Staudinger, 1871; in Staudinger & Wocke, Cat. Lepid. eur. Faunegeb. (Edn. 2) : 6, TL: vicinity of Akhaltsikhe, Georgia
Colias caucasica, Extracts from [BRU] by [Ilya Osipov]
Colias caucasica ; [BRU, 178]

Larva on Cytisus [BRU, Nekrutenko, 1990]

anna Gerhard, 1882; Berl. ent. Zs. 26 (1) : 125
= ; [LCG, 65]
olga Romanoff, 1882; Horae Soc. ent. ross. 17 : 127, pl. 4, f. 1-3
= ; [LCG, 65]
alba Rühl, [1893]; Palaearktischen Grosschmetterlinge: 162
= ; [LCG, 65 (= olga)]
Colias caucasica caucasica ; [LCG, 64]

Larva on Astragalus caucasicus [LCG, 65]


Colias balcanica Rebel, 1901
Bosnia, Yugoslavia, Bulgaria, Greece. See [About maps] Colias Myrmidone v. Balcanica Rebel, 1901; in Staudinger & Rebel, Cat. Lep. palaearct. Faunengeb. 1 : 250, TL: Bosnia and Herzegovina, Vucja Bara
anna Schawerda, 1913; Verh. zool.-bot. Ges. Wien 63 : 144
= ; [LCG, 66]
aphrodite Schawerda, 1908; Jber. Wien. ent. Ver. 19: 89
= ; [LCG, 66]
caltha Schawerda, 1922; Verh. zool.-bot. Ges. Wien 71 : 148
= ; [LCG, 66]
daphnis Schawerda, 1913; Verh. zool.-bot. Ges. Wien 63 : 144
= ; [LCG, 66]
isabellina Schawerda, 1922; Verh. zool.-bot. Ges. Wien 71 : 149
= ; [LCG, 66]
immaculata Schawerda, 1922; Verh. zool.-bot. Ges. Wien 71 : 148
= ; [LCG, 66]
leucorhorion Schawerda, 1922; Verh. zool.-bot. Ges. Wien 71 : 148
= ; [LCG, 66]
magnomaculata Schawerda, 1922; Verh. zool.-bot. Ges. Wien 71 : 148
= ; [LCG, 66]
micans Schawerda, 1922; Verh. zool.-bot. Ges. Wien 71 : 148
= ; [LCG, 66]
nicholli Schawerda, 1907; Verh. zool. bot. Ges. Wien 57: 219
= ; [LCG, 66]
nigrocuneata Schawerda, 1908; Jber. Wien. ent. Ver. 19: 89
= ; [LCG, 66]
nigrovittata Rebel, 1922; Verh. zool.-bot. Ges. Wien 71: 31
= ; [LCG, 67]
pseudobalcanica Pieszczek, 1917; Z. Öst. EntVer. 2 (9): 77
= ; [LCG, 67]
rebeli Schawerda, 1906; Verh. zool. bot. Ges. Wien 56 : 651
= ; [LCG, 67]
rosea Schawerda, 1919; Z. öst. EntVer. 4 (7): 67
= ; [LCG, 67]
scotina Schawerda, 1922; Verh. zool.-bot. Ges. Wien 71 : 148
= ; [LCG, 67]
semialba Wagner, 1907; Ent. Zeit. 21 (20) : 125
= ; [LCG, 67]
balcanica ; Abadjiev, 2002, Neue Ent. Nachr. 23: 16
Colias caucasica balcanica ; Winhard, 2000, Butterflies of the world, 10: 10, pl. 11, f. 16; [LCG, 65]
Colias balcanica ; [Otakar Kudrna]

Larva on Chamaecytisus hirsutus, Chamaecytisus absinthioides [LCG, 66]


Colias myrmidone (Esper, 1781)
-. See [About maps] Danube Clouded Yellow Orangeroter Heufalter Szlaczkoń szafraniec
Papilio myrmidone Esper, 1781; Die Schmett. Th. I, Bd. 2 (3): 88, pl. 65, f. 1-2, TL: Turnau, Hungary
Colias myrmidone, Larvae of North-European Lepidoptera [Kimmo Silvonen]
Colias myrmidone, Extracts from [BRU] by [Ilya Osipov]
Colias myrmidone, Europaische Schmetterlinge [Mario Maier]
Colias myrmidone ; [BRU, 178]; [Otakar Kudrna]

Larva on Cytisus [H&R]

Germany, Poland, Ukraine, Romania, Hungary, Slovakia, Austria, Bulgaria, Caucasus, SW.Russia aegra Verity, [1909]; Rhopalocera Palaearctica 1: 265
= ; [LCG, 67]
agnes Pieszczek, 1905; Verh. zool.-bot. Ges. Wien 55 : 416
= ; [LCG, 67]
alba Staudinger, 1871; in Staudinger & Wocke, Cat. Lepid. eur. Faunegeb. (Edn. 2) : 6
= ; [LCG, 67]
amicans Pieszczek, 1917; Z. Öst. EntVer. 2 (9): 73, pl. 1, f. 2
= ; [LCG, 67]
angustimarginata Metschl, 1925; Int. Ent. Zeit. 19: 25
= ; [LCG, 67]
antihygiaea Mayer, 1916; Ent. Zeit. 29 (26): 102
= ; [LCG, 67]
aurantiaca Pieszczek, 1917; Z. Öst. EntVer. 2 (9): 75, pl. 2, f. 18
= ; [LCG, 67]
bahri Skala, 1912; Int. ent. Z. 6 (3): 16
= ; [LCG, 68]
bicolor Metschl, 1922; Mitt. munchn. ent. Ges. 12: 12
= ; [LCG, 68]
Colias caliginosa Klos, 1914; Mitt. naturw. Ver. Steiermark 50: 214
= ; [LCG, 68]
callos Maurer, 1911; Jber. Wien ent. Ver. 22: 217
= ; [LCG, 68]
chrysothemides Verity, 1911; Rhopalocera Palaearctica 1: 358, pl. 72, f. 1-2
= ; [LCG, 68]
completa Pionneau, 1937; Echange 53: 10
= ; #
demarginata Metschl, 1922; Mitt. munchn. ent. Ges. 12: 8
= ; [LCG, 68 (= opposita)]
discolor Fritsch, 1912; Ent. Rundsch. 29 (21) : 143
= ; [LCG, 68]
edusaeformis Klemensiewicz, 1910; Spraw. Kom. fizyogr. 45: 50
= ; [LCG, 68]
edusoides Skala, 1912; Int. ent. Z. 6 (3): 16
= ; [LCG, 68]
pallida Zelezny, 1916; Z. öst. ent. Ver. 1: 20
= Colias myrmidone myrmidone f. edusoides ; [LCG, 68]
flavescens Garbowski, 1892; Sber. Akad. Wiss Wien.: 903
= ; [LCG, 68]
griseomarginata Berger, 1911; Jahrb. Wien. ent. Ver. 21: 77
= ; [LCG, 68]
depunctata Nitsche, 1912; Verh. zool.-bot. Ges. Wien 62: 109
= ; [LCG, 68 (= hartmanni Jouki, 1907)]
hartmanni Jouki, 1907; Cas. csl. Spol. ent. 4: 77
= ; [LCG, 68]
helma Geest, 1905; Z. wiss. InsekBiol. 1: 379
= ; [LCG, 68]
impunctata Verity, [1909]; Rhopalocera Palaearctica 1: 264
= ; [LCG, 68]
inspersa Metschl, 1922; Mitt. munchn. ent. Ges. 12: 11
= ; [LCG, 68]
intermedia Maurer, 1905; Verh. zool.-bot. Ges. Wien: 148
= ; [LCG, 68]
inumbrata Schultz, 1904; Nyt. Mag. naturvid. 42: 43, TL: Austria
= ; [LCG, 68]
lutea Zelezny, 1915; Ent. Rdsch. 32: 59
= ; [LCG, 58]
mediocarnea Metschl, 1922; Mitt. munchn. ent. Ges. 12: 11
= ; [LCG, 58]
micans Röber, [1907]; in Seitz, Grossschmett. Erde 1 : 69
= ; [LCG, 68]
nana Mayer, 1910; Int. ent. Z. 4: 182, TL: Graz
= ; [LCG, 68]
myrmidonides Ksienschopolski, 1911; Trudy obsch. Izsl. Volyni Zitomir 8: 30
= ; [LCG, 68 (nana Mayer, 1910)]
nigerrima Pieszczek, 1905; Verh. zool.-bot. Ges. Wien 55 : 420
= ; [LCG, 68]
nigrovenata Pieszczek, 1917; Z. Öst. EntVer. 2 (9): 75, pl. 2, f. 13, TL: Graz
= ; [LCG, 68]
obscura Skala, 1912; Ver. naturf. Ver. Brünn 50: 63
= ; [LCG, 68]
opposita Metschl, 1922; Mitt. munchn. ent. Ges. 12: 8
= ; [LCG, 68]
orcus Ronnicke, 1917; Z. Öst. EntVer. 2 (9): 77, pl. 3, f. 28
orcusides Strand, 1928; Arch. Naturg. 92 (A) 8: 51 (repl. orcus Ronnicke, 1917)
pallescens Fritsch, 1912; Ent. Rundsch. 29 (21) : 143
= ; [LCG, 68]
pallida Pieszczek, 1912; Jber. Wien Ent. Ver. 22: 223
= ; [LCG, 68]
pallidomaculata Metschl, 1922; Mitt. munchn. ent. Ges. 12: 8
= ; [LCG, 68]
permaculata Pieszczek, 1917; Z. Öst. EntVer. 2 (9): 76, pl. 2, f. 20
= ; [LCG, 68]
pieszezeki Predota, 1909; Jber. Wien ent. Ver. 20: 141
= ; [LCG, 68]
pseudomaureri Metschl, 1925; Int. Ent. Zeit. 19: 25
= ; [LCG, 68]
Colias pseudorebeli Klos, 1906; Verh. zool.-bot. Ges. Wien 56 ??: 651
= ; [LCG, 68]
pupillata Pieszczek, 1917; Z. Öst. EntVer. 2 (9): 71
= ; [LCG, 68]
rima Metschl, 1922; Mitt. munchn. ent. Ges. 12: 11
= ; [LCG, 68]
rubroflammea Zelezny, 1916; Z. öst. ent. Ver. 1: 20
= ; [LCG, 68]
rubroisabellina Fritsch, 1912; Ent. Rundsch. 29 (21) : 142
= ; [LCG, 68]
rudolfi Ronnicke, 1927; Z. Öst. EntVer. 12 (8): 76
= ; [LCG, 69]
schwabi Pieszczek, 1912; Jber. Wien Ent. Ver. 22: 232
= ; [LCG, 69]
striata Pieszczek, 1917; Z. Öst. EntVer. 2 (9): 71
= ; [LCG, 69]
striata Metschl, 1922; Mitt. munchn. ent. Ges. 12: 8 (preocc. striata Pieszczek, 1917)
unipuncta Pieszczek, ?1922; ?8
= ; [LCG, 69]
verna Verity, 1911; Rhopalocera Palaearctica 1: 358
= ; [LCG, 69]
virescalis Stöckl, 1929; Polsk. Pismo ent. 7: 65
= ; [LCG, 69]
Colias myrmidone myrmidone ; [LCG, 67]; [LCG, 68]

Larva on Cytisus ratisbonensis, C. capitatus, Chamaecytisus ruthenicus [LCG, 67 (Gorbunov & Korshunov, 1995)]

C. m. ermak Grum-Grshimailo, 1890
S.Urals, SW.Siberia, NW.Kazakhstan myrmidone ermak Grum-Grshimailo, 1890; in Romanoff, Mém. Lép. 4 : 301, TL: SE.Ural
Colias myrmidone ab. stellata Krulikovsky, 1908; Soc. Ent. 23 (1): 3, TL: Wiatka; Kasan
= ; [LCG, 69]
Colias myrmidone ermak ab. ♀ irinae Krulikovsky, 1909; Rev. russ. ent. 9 (1-2) : 110
= ; [LCG, 69]
Colias myrmidone ermak ab. ♀ sophiae Krulikovsky, 1909; Rev. russ. ent. 9 (1-2) : 110
= ; [LCG, 69]
vernalis Ksienschopolski, 1911; Trudy obsch. Izsl. Volyni Zitomir 8: 30
= ; [LCG, 69]
Colias myrmidone ermak ; [LCG, 69]

Colias nastes Boisduval, 1832
Altai, Sayan - Chukot Peninsula, Alaska, Greenland, Labrador, Greenland. See [About maps] Labrador Sulphur
Colias nastes Boisduval, 1832; Icon. hist. Lépid. Europe 1 (23-24): 245, (3-4): pl. 8, f. 4-5
Colias nastes, E.H. Strickland Ent. Mus, U of [ALBERTA]
Colias nastes, Lepidoptera collection [SZMN]
Colias nastes, NSG Voucher Specimen [Wahlberg]
Colias nastes, Butterflies and Moths of North America [BAMONA]
Colias nastes, Norges Sommerfugler [NORLEP]
Colias nastes, Extracts from [BRU] by [Ilya Osipov]
Eurymus nastes ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 11
Colias nastes ; [NACL, #4216]; [BRU, 170]; [Opler]
?Colias nastes ; [EBW, (or = tyche?)]
Colias nastes ; [BOW: pl. 4, f. 6 (or = tyche?)]

Larva on Astragalus alpinus, Oxytropis, Hedysarium [BRU, Scott, 1986]

Canada (Labrador, Northwest Territories, Baffin Island, Southampton Island) gueneei Avinoff, 1935; in Holland & Avinoff, Mem. Carn. Mus. 12 (2): 13
= ; [LCG, 70]
Colias nastes rossii Guenée, 1864; Ann. Soc. Ent. Fr. (4) 4 : 199
= ; [LCG, 70]
Eurymus nastes rossii ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 11
Colias nastes rossii ; [NACL, #4216a]
Colias nastes nastes ; [LCG, 69]

Larva on Astragalus alpinum [LCG, 70]

C. n. aliaska Bang-Haas, 1927
Alaska, Canada (Yukon, Northwest Territories, Victoria Island, Banks Island) Colias nastes aliaska Bang-Haas, 1927; Horae Macrolep. Palaearct. 1 : 41, pl. 5, f. 24-25, TL: Alaska, Ramport
subarctica McDunnough, 1928; Can. ent. 60 (11): 270, TL: Bernard Harbour, N. W. T.
= ; [LCG, 71]
alaskae ; McDunnough, 1938, Mem. South. Calif. Acad. Sci. 1: 8 (missp.)
Colias nastes aliaska ; [NACL, #4216e]; Winhard, 2000, Butterflies of the world, 10: 10, pl. 11, f. 17-18; [LCG, 70]
C. n. dezhnevi Korshunov, 1996
NE.Siberia (Magadan, Chukotka, Bilibino) Colias nastes dezhnevi Korshunov, 1996; Dopolneniya i ispravleniya k knige "Dnevnye babochki aziatskoi chasti Rossii": 18, TL: Siberia, Verchoiansk, Adytch River
Colias nastes dezhnevi, Lepidoptera collection [SZMN]
Colias tyche dezhnevi ; Grieshuber, 1998
Colias nastes dezhnevi ; [LCG, 71]
C. n. dioni Verhulst, 1999
Canada (Alberta) Colias nastes dioni Verhulst, 1999; Lambillionea 99 (4): 624, TL: Canada, Alberta, Kananaskis Co., Plateau Mtn 2500m
Colias nastes dioni ; [LCG, 71]
C. n. jakutica Kurentzov, 1970
Far East (Yakutia) Colias nastes jacutica Kurentzov, 1970; Butterflies of the USSR Far East: ? (spell.?)
?Colias nastes jakutica Kurentzov, 1970; (spell.?)
Colias jacuticola Weiss & Mracek, 1989; Nachr. ent. Ver. Apollo 10 (4): 281 (spell.?)
= ; [LCG, 72]
?Colias nastes jakuticola Weiss & Mracek, 1989; (spell. ?)
Colias nastes jacutica, Lepidoptera collection [SZMN]
Colias nastes jakutica ; [LCG, 72]
C. n. moina Strecker, 1880
Canada (Northwest Territories, Manitoba) moina Strecker, 1880; Bull. Brooklyn ent. Soc. 3 (5) : 34, TL: Canada, N.Manitoba, Churchill
harperi (Gunder, 1932); Can. Ent. 64 (2): 278, TL: Ft. Churchill, Manitoba
Colias nastes moina ab. harperi ; [LCG, 73]
Eurymus moina ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 11
Colias nastes moina ; [NACL, #4216b]; [LCG, 73]
C. n. streckeri Grum-Grshimailo, 1895
Canada (Alberta, British Columbia), N.Washinginton, Montana streckeri Grum-Grshimailo, 1895; Horae Soc. ent. Ross. 29 : 290, TL: Alberta, Laggan, alke Louise
sibirica Kurentzov, 1970; (nom. praeocc.)
obscurata Verity, 1911; Rhopalocera Palaearctica 1: 355, pl. 61, f. 6, TL: Alberta, Lake Louise
= ; [LCG, 74 (ab.)]
palliflava McDunnough, 1927; Can. ent. 59 : 154
= ; [LCG, 74 (ab.)]
Eurymus nastes streckeri ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 11
Colias nastes streckeri ; [NACL, #4216c]; [LCG, 74]
C. n. zemblica Verity, 1911
Novaja Zemlja nastes zemblica Verity, 1911; Rhopalocera Palaearctica 1: 355, pl. 71, f. 10, TL: Siberia, Novaja Zemlja
?zembalica Verity, 1911; (spell.?)
Colias tyche zemblica ; Grieshuber, 1998
Colias nastes zemblica ; [LCG, 74]
C. n. cocandicides Verity, 1911
Colias nastes cocandicides ; [NACL, #4216d]
C. n. ferrisi Verhulst, 2004
Alaska Colias nastes ferrisi Verhulst, 2004; Lambillionea 104 (1): 129, TL: Alaska

fiColias tyche (Böber, 1812)
. See [About maps] Tunturikeltaperhonen Pale Arctic Clouded Yellow Fjällfjäril
tyche (Böber, 1812); Mém. Soc. Imp. Nat. Moscou 3: 21, ?72, pl. 3, f. 3-6 [??ref, see melinos]
Colias melinos Eversmann, 1847; Bull. Soc. Imp. Nat. Moscou 20 (3) : 72, pl. 3, f. 3-6 [??ref, see tyche }
Colias tyche, NSG Voucher Specimen [Wahlberg]
Colias tyche, Butterflies and Moths of North America [BAMONA]
Colias tyche, Larvae of North-European Lepidoptera [Kimmo Silvonen]
Colias tyche, Svenska fjärilar [SVF]
Colias tyche, Moths and Butterflies of Europe [MBE]
Colias tyche, Extracts from [BRU] by [Ilya Osipov]
Colias tyche ; [BRU, 170]; [Opler]
[tyche-1fv.jpg] 533x400 (~46Kb) underside female Open pine forest on a hill just NE of the Chita City, Transbaikalia, Siberia, Russia. 17th June 1995 (ssp. deckerti Verity, 1909), Photo © Oleg Kosterin

Larva on Astragalus frigidus [SPRK, given as C. nastes]

Larva on Astragalus alpinus, Oxytropis, Caragana [BRU]


N.Siberia, SE.Siberia, Altai, Sayan, Yakutia, Transbaikalia, Baikal, Amur, Mongolia deckerti Verity, [1909]; Rhopalocera Palaearctica 1: 236, pl. 42, f. 27-28, TL: SE.Siberia, Mt. Apfel
= ; [LCG, 75]
herzi Staudinger, 1901; in Staudinger & Rebel, Cat. Lepid. palaearct. Faunengeb., 3: 16, TL: Viluy
= ; [LCG, 76]
jeholensis Matsumura, 1939; Bull. biogeogr. Soc. Japan 9 (20): 348, TL: Mongolia, Re He, Jehol, Kingan Mtn
= ; [LCG, 76]
ludmilla Hemming, 1933; Entomologist 66: 278 (repl. chryseis Verity, 1911
= ; [LCG, 76]
chryseis Verity, 1911; (praeocc. Colias erate chryseis Röber, 1907)
vitimensis Austaut, 1899; Le Naturaliste 21: 284
= ; [LCG, 76]
magna Rühl, [1893]; Palaearktischen Grosschmetterlinge: 153, TL: ?Lake Baikal
= ; [LCG, 75]
melinoides Verity, 1926; Ent. Rec. J. Var. 38 : 171
= ; [LCG, 75]
montana Verity, 1911; Rhopalocera Palaearctica 1: 354, pl. 70, f. 34-38
= ; [LCG, 75]
viridis Bang-Haas, 1915; Dt. ent. Z. Iris 29 (2/3) : 99, TL: Mts Nan Shan, China
= ; [LCG, 75]
Colias tyche montana ;
Colias tyche tyche ; [LCG, 74]
C. t. relicta Kurentzov, 1970
Chukot Peninsula, Far East Colias melinos relicta Kurentzov, 1970; Butterflies of the USSR Far East: ?, TL: NE.Siberia, R. Okhota
Colias tyche relicta ; [LCG, 77]
fiC. t. werdandi Zetterstedt, 1839
NEU tyche werdandi Zetterstedt, 1839; Insecta Lapponica : 908, TL: Torne-Lappland
alba Schulte, 1953; Ent. Zeit. 62: 150
= ; [LCG, 78]
anastigma Röber, [1907]; in Seitz, Grossschmett. Erde 1 : 63
= ; [LCG, 78]
bruneomaculata Schulte, 1953; Ent. Zeit. 62: 151
= ; [LCG, 78]
castaneapunctata Sheldon, 1912; Entomologist 45: 27
= ; [LCG, 78]
christiernssoni Lampa, 1885; Ent. Tidskr. 6 (1) : 11
= ; [LCG, 78]
connexa Sheldon, 1912; Entomologist 45: 27
= ; [LCG, 78]
ellingeri Schulte, 1953; Ent. Zeit. 62: 151
= ; [LCG, 78]
flavopunctata Sheldon, 1912; Entomologist 45: 27
= ; [LCG, 78]
grisea Schulte, 1953; Ent. Zeit. 62: 151
= ; [LCG, 78]
immaculata Lampa, 1885; Ent. Tidskr. 6 (1) : 11
= ; [LCG, 78]
ab. insignata Thurau, 1903; Berl. ent. Z. 48 : 113
= ; [LCG, 78]
mariannae Schulte, 1953; Ent. Zeit. 62: 152
= ; [LCG, 78]
nigrita Schulte, 1953; Ent. Zeit. 62: 152
= ; [LCG, 78]
palaeonides Schulte, 1953; Ent. Zeit. 62: 151
= ; [LCG, 78]
phicomonides Verity, 1911; Rhopalocera Palaearctica 1: 355, TL: Norway
= ; [LCG, 78]
ab. radiata Thurau, 1903; Berl. ent. Z. 48 : 113
= ; [LCG, 78]
sulphurea Lampa, 1885; Ent. Tidskr. 6 (1) : 10
= ; [LCG, 78]
torneoensis Sheldon, 1912; Entomologist 45: 27
= ; [LCG, 78]
Colias tyche werdandi ; [LCG, 77]
Colias werdandi ; [Otakar Kudrna]

Larva on Astragalus alpinus, Vaccinium myrtillus, Oxytropis lappona [LCG, 77]

C. t. magadanica Churkin, Grieshuber, Bogdanov & Zamolodchikov, 2001
Magadan Colias tyche magadanica Churkin, Grieshuber, Bogdanov & Zamolodchikov, 2001; Helios, Coll. lep. articles II: (103-115)
C. t. flinti Churkin, Grieshuber, Bogdanov & Zamolodchikov, 2001
Wrangel Is. Colias tyche flinti Churkin, Grieshuber, Bogdanov & Zamolodchikov, 2001; Helios, Coll. lep. articles II: (103-115)
C. t. kolosovae Churkin, Grieshuber, Bogdanov & Zamolodchikov, 2001
E.Chukotka Colias tyche kolosovae Churkin, Grieshuber, Bogdanov & Zamolodchikov, 2001; Atalanta 32 (1/2): 262 (repl. Colias tyche olga Churkin, Grieshuber, Bogdanov & Zamolodchikov, 2001)
Colias tyche olga Churkin, Grieshuber, Bogdanov & Zamolodchikov, 2001; Helios, Coll. lep. articles II: (103-115) (preocc. Colias olga Romanoff, 1882)

Colias phicomone (Esper, 1780)
-. See [About maps] Mountain Clouded Yellow Grünlicher Heufalter
Papilio phicomone Esper, 1780; Die Schmett. Th. I, Bd. 2 (1): 32, pl. 56, f. 1-2, TL: Styria
Colias phicomone, NSG Voucher Specimen [Wahlberg]
Colias phicomone, Moths and Butterflies of Europe [MBE]
Colias phicomone, Europaische Schmetterlinge [Mario Maier]
Colias phicomone, Captain's European Butterfly Guide [Simon Coombes]
Colias phicomone ; [EBW]; [BOW: pl. 4, f. 8]; [Otakar Kudrna]
Alps, N.Carpathians, N.Italy alpiumnitida Verity, 1926; Ent. Rec. J. Var. 38 : 171
= ; [LCG, 79]
anticopupillata Verity, 1947; Le Farfalle diurn. d'Italia 3: 269
= ; [LCG, 79]
bipupillata Mezger, 1932; Lambillionea, 32: 210, TL: Simplon
= ; [LCG, 79]
blachieri Culot, 1909; Bull. Soc. Lép. Genève 1: 265
= ; [LCG, 79]
derosea Wagner, 1926; Z. Öst. Ent.-Ver. Wien 11: 25
= ; [LCG, 79]
elegans Schultz, 1904; Nyt. Mag. naturvid. 42: 42
= ; [LCG, 79]
fasciata Ronnicke, 1920; Ent. Zeit. 34: 70
= ; [LCG, 79]
flavida Osthelder, 1913; Mitt. Münch. ent. Ges. 4: 28
= ; [LCG, 79]
flava Osthelder, 1925; Mitt. Münch. ent. Ges. 15 (suppl.): 67
= ; [LCG, 78 (= flavida Osthelder, 1913)]
Colias phicomone f. fleischmanni Stauder, 1924; Int. ent. Z. 18: 37
= ; [LCG, 79]
geesti Neuburger, 1905; Soc. ent. 20 (6) : 42
= ; [LCG, 79]
gynomorpha Verity, 1911; Rhopalocera Palaearctica 1: 353, pl. 70, f. 22
= ; [LCG, 79]
albidior Ronnicke, 1920; Ent. Zeit. 34: 71
= ; [LCG, 79 (= gynomorpha Verity, 1911)]
hyalides Verity, 1911; Rhopalocera Palaearctica 1: 353, pl. 70, f. 24
= ; [LCG, 79]
incompleta Mezger, 1934; Lambillionea, 34: 54, TL: Switzerland, Valais, Rieffelalp
= ; [LCG, 79 (ab.)]
lanceolata Verity, 1947; Le Farfalle diurn. d'Italia 3: 270, TL: Dolomites, Alpi Agordine, Falcade, Ponte de la Schita 2000m
= ; [LCG, 79]
obscura Ronnicke, 1920; Ent. Zeit. 34: 71
= ; [LCG, 79]
pallida Hoffmann & Klos, 1914; Mitt. nat. Ver. Steierm. 50: 208
= ; [LCG, 79]
passa Verity, 1911; Rhopalocera Palaearctica 1: 353, pl. 70, f. 23
= ; [LCG, 79]
Colias phicomone periphaes Fruhstorfer, 1908; Ent. Zs. 22 (12) : 51, TL: Dachstein
= ; [LCG, 79]
phaedra Schawerda, 1924; Mitt. münchn. ent. Ges. 14: 79
= ; [LCG, 79]
porrecta Heinrich, 1930; Ent. Zeit. 43: 295, TL: Zürich
= ; [LCG, 79]
privata Ronnicke, 1920; Ent. Zeit. 34: 71
= ; [LCG, 79]
pulverulenta Verity, 1926; Ent. Rec. J. Var. 38 : 171, TL: Cottian Alps
= ; [LCG, 79]
pupillata Refhous, 1909; Bull. Soc. Lép. Genève 1: 26, pl. 8
= ; [LCG, 79]
pupillata Ronnicke, 1920; Ent. Zeit. 34: 71
= ; [LCG, 79 (= pupillata Refhous, 1909)]
unimaculata Pionneau, 1924; Echange 39: 23
= ; [LCG, 79 (= pupillata Refhous, 1909)]
Colias phicomone var. saturata Austaut, 1905; Ent. Zs. 18 (36) : 143
= ; [LCG, 79]
theia Schawerda, 1919; Z. öst. EntVer. 4 (7): 67
= ; [LCG, 79]

Larva on Vicia sativa, Vicia pirenaica, V. angustifolia, V. narbonensis, Medicago sativa, Coronilla sp., Lotus corniculatus, Oxytropis montana, Hippocrepis comosa [LCG, 78]

C. p. juliana Hospital, 1948
Cantabria phicomone juliana Hospital, 1948; Trab. Mus. Cienc. nat. Barcelona 1 (2): 11, TL: Cordillera Cantabrica, Picos de Europa
impunctata Verity, [1909]; Rhopalocera Palaearctica 1: 230, pl. 62, f. 3, TL: Asturites, Picos de Europa, 2000m
= ; [LCG, 80]
Colias phicomone juliana ; [LCG, 79]
C. p. oberthueri Verity, [1909]
Pyrenees phicomone oberthueri Verity, [1909]; Rhopalocera Palaearctica 1: 231, pl. 42, f. 5-6, TL: Pyrenees, Vernet-les-Bains, Gèdre
Colias phicomone oberthueri ; [LCG, 80]
C. p. phila Fruhstorfer, 1903
Kashmir Colias phicomone phila Fruhstorfer, 1903; Dt. ent. Z. Iris 16 (1) : 48, TL: Kashmir

fiColias palaeno (Linnaeus, 1761)
. See [About maps] Suokeltaperhonen Moorland Clouded Yellow Palaeno Sulphur Arctic Sulfur Svavelgul höfjäril Svavelfjäril Hochmoorgelbling Szlaczkoń torfowiec
Papilio palaeno Linnaeus, 1761; Fauna Suecica (Edn 2): 272, TL: Uppsala, Sweden
Colias palaeno, Lepidoptera collection [SZMN]
Colias palaeno, NSG Voucher Specimen [Wahlberg]
Colias palaeno, Butterflies and Moths of North America [BAMONA]
Colias palaeno, Larvae of North-European Lepidoptera [Kimmo Silvonen]
Colias palaeno, Svenska fjärilar [SVF]
Colias palaeno, Norges Sommerfugler [NORLEP]
Colias palaeno, Moths and Butterflies of Europe [MBE]
Colias palaeno, Extracts from [BRU] by [Ilya Osipov]
Colias palaeno, Europaische Schmetterlinge [Mario Maier]
Eurymus palaeno ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 10
Colias palaeno ; [NACL, #4218]; [BRU, 179]; [EBW]; [BOW: pl. 4, f. 7]; [MRS, 219]; [Otakar Kudrna]
[palaeno-1vf.jpg] 600x645 (~34Kb) underside female FINLAND: Li: Utsjoki Ailigas, 775:50, 14.7.1998, Photo © Markku Savela [palaeno-2.jpg] 750x499 (~34Kb) Sipoo 670:40, 18.06.1980, Photo © Pertti Pakkanen

Larva on Vaccinium uliginosum [SPRK] [MRS], Vaccinium uliginosum, V. caespitosum [NAB]


Finland, Sweden, Norway, Estonia lapponica Staudinger, 1871; in Staudinger & Wocke, Cat. Lepid. eur. Faunegeb. (Edn. 2) : 5
philomene Hübner, 1803; Samml. eur. Schmett. [1] : f. 602-603
avinoffi Verity, 1911; Rhopalocera Palaearctica 1: 346, TL: Finland
= ; [LCG, 81]
cretacea Schilde, 1884; Ent. Nachr. : (335-346)
= ; [LCG, 81]
nordströmi Bryk, 1923; Ent. Tidskr. 44: 109
= ; [LCG, 81]
octava Bryk, 1923; Ent. Tidskr. 44: 108
= ; [LCG, 81]
parva Hoyningen-Huene, 1901; Sber. Naturf. Ges. Dorpat (Jurjew) 12: 460, TL: Estonia
= ; [LCG, 81]
schildei Staudinger, 1892; Dt. Ent. Z. Iris 5 (2) : 310
= ; [LCG, 81]
Colias palaeno palaeno ; [LCG, 81]
Misidentified/misapplied:
?Eurymus palaeno werdandi Herrich-Schäffer, 1848; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 10
werdandi Herrich-Schäffer, 1848; Schmett. Eur. 1: 41-42, f. 403-404

Larva on Vaccinium uliginosum, V. myrtillus [LCG, 81]

C. p. aias Fruhstorfer, 1903
Japan Colias palaeno aias Fruhstorfer, 1903; Dt. ent. Z. Iris 16 (1) : 48, TL: Japan
sugitanii Esaki, 1929; Zephyrus 1 (2): 42, f. 3-4
= ; [LCG, 82]
nekkana Matsumura, 1939; Bull. biogeogr. Soc. Japan 9 (20): 348
= ; [LCG, 82]
Colias palaeno aias ; Winhard, 2000, Butterflies of the world, 10: 10, pl. 11, f. 19; [LCG, 81]
C. p. europome (Esper, 1778)
Germany, Belgium, Slovakia, Romania, Ukraine palaeno europome (Esper, 1778); Die Schmett. Th. I, Bd. 1 (7): pl. 42, f. 1-2, TL: Saxony
?europome (Esper, 1789); Die Schmett., Suppl. Th 1 (3-4): pl. 100, f. 5
atavista Stephan, 1923; Dt. Ent. Z. Iris 37: 30
= ; [LCG, 82]
aurantiaca Hackray, 1934; Lambillionea 34: 193
= ; [LCG, 82]
belga Derenne, 1928; Lambillionea 28: 23
= ; [LCG, 82]
bimaculata Schröder, 1923; Mitt. Bad. ent. Ver. 1: 13
= ; [LCG, 82]
binotata Cabeau, 1925; Rev. mens. Soc. ent. Namur 25: 39
= ; [LCG, 82]
cafflischi Talbot, 1935; Lep. Cat. 66: 413
= ; [LCG, 82]
cavroi Dufrane, 1947; Bull. Ann. Soc. ent. Belg. 83 (1/2) : 69
= ; [LCG, 82]
cretacea Schilde, 1884; Ent. Nachr. 10 (22): 339
= ; [LCG, 82]
dentata Derenne, 1934; Lambillionea 34: 191
= ; [LCG, 82]
illgneri Rühl, 1890; Soc. ent. 5: 89
= ; [LCG, 82]
illgneriana Vorbrodt, 1912; Die Schmett. Schweiz 1 : 28
= ; [LCG, 82]
illgneri-reducta Schröder, 1923; Mitt. Bad. ent. Ver. 1: 13
= ; [LCG, 82]
posterodimidiata Derenne, 1934; Lambillionea 34: 191
= ; [LCG, 82]
pruefferi Krzywicki, 1967; Ann. zool. Warsz. 25: 106
= ; [LCG, 82]
pupillata Pieszczek, 1917; Z. Öst. EntVer. 2 (9): 71
= ; [LCG, 83]
roseomaculata Loritz, 1953; Bull. Soc. ent. Mulhouse: 8, TL: Alpes-Maritimes
= ; [LCG, 83]
rubeopunctata Loritz, 1953; Bull. Soc. ent. Mulhouse: 8, TL: Alpes-Maritimes
= ; [LCG, 83]
schroderi Hommel, 1923; Mitt. Bad. ent. Ver. 1: 13
= ; [LCG, 83]
sitowskii Biezanko, 1924; Archiv Naturg. (A) 90 (5): 241
= ; [LCG, 83]
striata van Mellaerts, 1929; Lambillionea 29: 123
= ; [LCG, 83]
subobsoleta Pionneau, 1936; Echange 51: 18
= ; [LCG, 83]
valeria Sievers, 1859; Motschulsky Etudes Ent. 8: 146, TL: St. Petersburg
= ; [LCG, 83]
viridis Pionneau, 1936; Echange 51: 17
= ; [LCG, 83]
Colias palaeno europome ; [LCG, 82]
C. p. europomene Ochsenheimer, 1808
Alps palaeno europomene Ochsenheimer, 1808; Schmett. Europa 1 (2) : 184
altotineenis Loritz, 1953; Bull. Soc. ent. Mulhouse: 7
= ; [LCG, 83]
cafflishi Caradja, 1893; Soc. Ent. 8 : 26
= ; [LCG, 83]
cenisia Rocca, 1944; Mem. Soc. ent. Ital. 23: 21
= ; [LCG, 83]
deprunneri Rocca, 1944; Mem. Soc. ent. Ital. 23: 21
= ; [LCG, 83]
flavoinspersa Heinrich, 1919; Int. Ent. Z. 13: 132
= ; [LCG, 83]
flavoradiata Wheeler, 1903; Butts Switzerland: 68
= ; [LCG, 83]
herrichi Staudinger, 1892; Dt. Ent. Z. Iris 5 (2) : 311
= ; [LCG, 83]
herrichina Geest, 1905; Z. wiss. InsekBiol. 1: 381
= ; [LCG, 83]
jurassica Verity, [1908]; Rhopalocera Palaearctica 1: 216
= ; [LCG, 83]
obliterata Verity, 1911; Rhopalocera Palaearctica 1: 346, pl. 68, f. 42
= ; [LCG, 83]
ochracea Geest, 1905; Z. wiss. InsekBiol. 1: 381
= ; [LCG, 83]
Colias palaeno europomene ; [LCG, 83]
C. p. orientalis Staudinger, 1892
Far East, Kamchatka, Transbaikalia, Amur palaeno orientalis Staudinger, 1892; Dt. Ent. Z. Iris 5 (2) : 311, TL: Kentei, Amur region
arctica Verity, [1908]; Rhopalocera Palaearctica 1: 216, pl. 49, f. 11, TL: Siberia, Vitim
karaganica Kurentzov, 1970
Colias palaeno arctica, Lepidoptera collection [SZMN]
Colias palaeno arctica ; [LCG, 84]
Colias palaeno orientalis ; [MRS, 219]; [LCG, 84]
[orientalis-1fv.jpg] 300x400 (~33Kb) underside female A tundrous coastal terrace in the vicinity of the mouth of the Burgauli River on the south (Ochot Sea) coast of the Koni Peninsula, Ola District, S Magadan Province, Russia. 17th July, 1989, Photo © Oleg Kosterin

C. p. poktusani O. Bang-Haas, 1934
N.Korea palaeno poktusani O. Bang-Haas, 1934; Ent. Z. 48 : 183, TL: Korea, Paektusen Mts.
Colias palaeno poktusani ; [LCG, 84]
C. p. sachalinensis Matsumura, 1919
Sakhalin palaeno sachalinensis Matsumura, 1919; Thous. Ins. Japan. Addit. 3 : 725, TL: Sakhalin, Tonniacha, Shiraraka
Colias palaeno sachalinensis ; [LCG, 84]
C. p. synonyma Bryk, 1923
Sweden, Denmark palaeno synonyma Bryk, 1923; Ent. Tidskr. 44: 108, TL: Sweden
Colias palaeno synonyma ; [LCG, 84]

Colias chippewa Edwards, 1872
-. See [About maps] Colias chippewa Edwards, 1872; Synopsis N. Amer. Butts.: , TL: W. and Great Slave L, Northwest Territories
Colias chippewa, E.H. Strickland Ent. Mus, U of [ALBERTA]
Colias chippewa, Extracts from [BRU] by [Ilya Osipov]
Colias chippewa ; [BRU, 179]
Colias palaeno chippewa ; [NACL, #4218a]; [Opler]

Larva on Vaccinium uliginosum [BRU, Scott, 1986]

Canada (Alberta, British Columbia, Manitoba, Northwest Territories, Ontario, Saskatchewan, Yukon), Alaska helena Edwards, 1863; Proc. ent. Soc. Philad. 2 (2): 80 (preocc. Herrich-Schäffer, 1843)
= ; [LCG, 86]
kohlsaati Gunder, 1931; Bull. South. Calif. Acad. Sci. 30 (2): 45
= ; [LCG, 86]
pelidneides Staudinger, 1901; in Staudinger & Rebel, Cat. Lepid. palaearct. Faunengeb., 3: 15
= ; [LCG, 86]
Colias chippewa chippewa ; [LCG, 85]
C. c. baffinensis Ebner & Ferris, 1978
Northwest Territories (Baffin Island, ...) chippewa baffinensis Ebner & Ferris, 1978; J. Res. Lep. 16 (3): 155(?)
Colias chippewa baffinensis ; [LCG, 87]
C. c. gomojunovae Korshunov, 1996
Far East (tundras, Chukot, Magadan) chippewa gomojunovae Korshunov, 1996; Dopolneniya i ispravleniya k knige "Dnevnye babochki aziatskoi chasti Rossii": 18, TL: Magadan, Bilibinski district
Colias chippewa gomojunovae ; [LCG, 87]

Colias hyperborea Grum-Grshimailo, 1899
-. See [About maps] Colias hyperborea Grum-Grshimailo, 1899; Ann. Mus. Zool. Imp. Acad. Sci. St.-Petersb. 4 (4) : 455, TL: Lena Valley, Yakutia
Colias hyperborea, NSG Voucher Specimen [Wahlberg]
Colias hyperborea, Extracts from [BRU] by [Ilya Osipov]
Colias hyperborea ; [BRU, 176]
[hyperborea-1mv.jpg] 556x400 (~48Kb) underside male Photographed on a fruticulose tundra on the southern coastal terrase of the Koni Peninsula protruding into the Ochot Sea south of Magadan. The plants seen are Arctous alpinus (broad leaves) and Phyllodoce coerulea L. (narrow leaves) 17th of July 1988., Photo © Oleg Kosterin

Larva on Astragalus spp. [BRU, Gorbunov, Korshunov, 1995]


NE.Siberia, Chukot Peninsula alba Sheljuzhko, 1923; Ann. Soc. ent. Fr. 91 (4) : 282
= ; [LCG, 88]
viluiensoides Verity, [1909]; Rhopalocera Palaearctica 1: 250
= ; [LCG, 88]
Colias hyperborea hyperborea ; [LCG, 88]

Larva on Astragalus alpinus, Oxytropis czukotica [LCG, 88]

C. h. kurnakovi Kurentzov, 1970
Magadan Colias hyperborea kurnakovi Kurentzov, 1970; Butterflies of the USSR Far East: ?, TL: Magadan, Omsuktchansk
Colias hyperborea kurnakovi ; [LCG, 89]
C. h. tunkuna Austaut, 1912
Sayan Colias tunkuna Austaut, 1912; Int. Ent. Zs 6 (13) : 88, f. 2, TL: Sayan Mts.
Colias hyperborea tunkuna ; [LCG, 88]
C. h. puella Churkin & Grieshuber, 2001
E.Chukotka Colias hyperborea puella Churkin & Grieshuber, 2001; Helios, Coll. lep. articles II: (1174-189), TL: E.Chukotka, Anadyr
C. h. paradoxa Churkin & Grieshuber, 2001
Colias hyperborea paradoxa Churkin & Grieshuber, 2001; Helios, Coll. lep. articles II: (1174-189), TL: Putorana plateau

Colias meadii Edwards, 1871
-. See [About maps] Mead's Sulphur
Colias meadii Edwards, 1871; Trans. amer. ent. Soc. 3 : 267, TL: Colorado, Lake Park Co., Mosquito Pass
Colias meadii, E.H. Strickland Ent. Mus, U of [ALBERTA]
Colias meadii meadii, Butterflies of America [BOA]
Colias meadii, Butterflies and Moths of North America [BAMONA]
Colias meadi ; [BOW: pl. 17, f. 10 (text only)]
Eurymus meadii ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 9
Colias meadii ; [NACL, #4213]; [EBW]; [Opler]
[meadii-1v.jpg] 800x1031 (~154Kb) underside USA: Colorado, Clear Creek Co., Summit Lake, Mt. Evans, 23.7.2012 (39°35'52"N 105°38'23"W), Photo © Markku Savela

Larva on Trifolium, T. parryi, Trifolium sasyphyllum


Colorado, Utah, New Mexico, Wyoming, Montana medi (Gunder, 1934); Can. Ent. 66 (6): 125
= ; [LCG, 90]
Colias meadii meadii ; Winhard, 2000, Butterflies of the world, 10: 10, pl. 11, f. 20; [LCG, 89]

Larva on Tripholium dasyphyllum, T. nanum, T. parryi, Astragalus alpinus, Oxytropis deflexa, Vicia americana [LCG, 90]

C. m. elis Strecker, 1885
Alberta, British Columbia Colias elis Strecker, 1885; Proc. Acad. Nat. Sci. Philad. 37 : 24, TL: Alberta, Kicking Horse Pass, Rocky Mts.
alberta Bowman, 1942; (unavail. hybrid. name)
pallidissima Bowman, 1942; Can. Ent. 74 (2): 25 (unavail. hybrid. name)
= ; [LCG, 91]
lambillioni Dufrane, 1947; Bull. Ann. Soc. ent. Belg. 83 (1/2) : 70
= ; [LCG, 91]
Eurymus meadii elis ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 9
Colias elis ; [BOW: pl. 17, f. 10]
Colias meadii elis ; [NACL, #4213a]; [LCG, 90]
C. m. lemhiensis Curtis & Ferris, 1985
Idaho, Montana meadii lemhiensis Curtis & Ferris, 1985; Bull. Allyn Mus. 91 (2): (1-9), TL: Idaho, Lemhi Co., Meadow Creek Lake
Colias meadii lemhiensis ; [LCG, 91]

Larva on Astragalus miser [LCG, 92]


Colias johanseni Troubridge & Philip, 1990
Northwest Territories (Mackenzie, Bernard Harbour). See [About maps] Colias johanseni Troubridge & Philip, 1990; Can. Ent. 122 (1-2): (15-20), TL: Northwest Territories, Mackenzie, Bernard Harbour
Colias johanseni ; [LCG, 93]; [Opler]

Larva on Hedysarum mackenzii [LCG, 93]


Colias aquilonaris Grum-Grshimailo, 1899
Chukot region. See [About maps] aquilonaris Grum-Grshimailo, 1899; Ann. Mus. Zool. Imp. Acad. Sci. St.-Petersb. 4 (4) : 459, TL: Olenek River Valey, 67,5 lat
Colias aquilonaris, NSG Voucher Specimen [Wahlberg]
Colias viluiensis aquilonaris ; [BRU]
Colias aquilonaris ; [LCG, 93 (= hecla?)]

fiColias hecla Lefèbvre, 1836
. See [About maps] Lapinkeltaperhonen Northern Clouded Yellow Greenland Sulphur Häcklefjäril
Colias hecla Lefèbvre, 1836; Ann. Soc. ent. Fr. 5 : 383, pl. 9, f. 3-6, TL: "Island" [Greenland]
groenlandica Lampa, 1893
Colias hecla, Butterflies and Moths of North America [BAMONA]
Colias hecla, Larvae of North-European Lepidoptera [Kimmo Silvonen]
Colias hecla, Svenska fjärilar [SVF]
Colias hecla, Norges Sommerfugler [NORLEP]
Colias hecla, Moths and Butterflies of Europe [MBE]
Colias hecla, Extracts from [BRU] by [Ilya Osipov]
Eurymus hecla ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 9
Colias hecla ; [NACL, #4214]; [BRU, 176]; [EBW]; [BOW: pl. 4, f. 3]; [Opler]; [Otakar Kudrna]

Larva on Astragalus frigidus [SPRK], Astragalus alpinus, Amoria repens, Arctostaphylos, Vaccinium, Hedysarium sp. [BRU]

Greenland Colias hecla var. pallida Skinner & Mengel, 1892; Proc. Acad. Nat. Sci. Philad. 44 : 156, Ent. News 3 (3): pl. 2, f. 4
palamedes Hemming, 1934; Stylops 3 : 98 (repl. pallida Skinner & Mengel, 1892)
palmedes Talbot, 1935; Lep. Cat. 66: 447 (repl. pallida Skinner & Mengel, 1892)
groenlandica Rühl, 1895; in Heyne, Palaearkt. Grossschmett. 1 : 158
= ; [LCG, 95]
Eurymus hecla pallida ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 9
Colias hecla hecla f. pallida ; [LCG, 94]
Colias hecla hecla ; [LCG, 94]
C. h. glacialis McLachlan, 1878
Alaska, Canada (Northwest Territories, Yukon, Quebec, Labrador) glacialis McLachlan, 1878; J. Linn. Soc. Zool. Lond. 14 : 108, TL: Northwest Territories, 81.45N. Lat (Grinnel Land, Ellesmere Is.)
Eurymus hecla glacialis ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 9
Colias hecla glacialis ; [NACL, #4214b]; [LCG, 95]
C. h. hela Strecker, 1880
Canada (Manitoba, Northwest Territories) hela Strecker, 1880; Bull. Brooklyn ent. Soc. 3 (5) : 33, TL: Manitoba, 3 miles N. Fort Churchill, W. Hudson Bay
chrysothemoides Verity, 1911
Eurymus hecla hela ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 9
Colias hecla hela ; [NACL, #4214a]; [LCG, 95]
C. h. orientis Wnukowsky, 1929
Chukot region, Far East hecla orientis Wnukowsky, 1929; Zool. Anz. 83 (9/10): 221 (repl. orientalis Grum-Grshimailo, 1899), TL: Chukot
orientalis Grum-Grshimailo, 1899; (nom. praeocc.), TL: Chukot
Colias hecla orientis ; [LCG, 96]
fiC. h. sulitelma Aurivillius, 1890
N.Scandinavia, NW.Siberia (Arctic Ural) hecla sulitelma Aurivillius, 1890; Bihang K. svenska VetensAkad. Handl. 15 (4) (1) : 12, pl. 2, f. 10, TL: Mt. Sulitelma, Sweden
carnea Mezger, 1932; Lambillionea, 32: 210, TL: Lappmark
= ; [LCG, 97 (ab.)]
citrina Staudinger, 1901; in Staudinger & Rebel, Cat. Lepid. palaearct. Faunengeb., 3: 18
= ; [LCG, 97]
distincta Sheldon, 1912; Entomologist 45: 340
= ; [LCG, 97]
holmbomi Bang-Haas, 1915; Dt. ent. Z. Iris 29 (4) : 193
= ; [LCG, 97]
jora Rosa, 1921; Entomologist 54: 265
= Colias hecla sulitelma f. citrina ; [LCG, 97]
nigropunctata Sheldon, 1912; Entomologist 45: 340
= ; [LCG, 97]
obscura Sheldon, 1912; Entomologist 45: 340
= ; [LCG, 97]
ochreasuffusa Sheldon, 1912; Entomologist 45: 340
= ; [LCG, 97]
radiata Mezger, 1935; Lambillionea, 35: 188
= ; [LCG, 97]
rosea Sheldon, 1912; Entomologist 45: 340
= ; [LCG, 97]
sandahli Lampa, 1885; Ent. Tidskr. 6 (1) : 11
= ; [LCG, 97]
sulphurea Röber, [1907]; in Seitz, Grossschmett. Erde 1 : 67
= ; [LCG, 97]
supresa Sheldon, 1912; Entomologist 45: 340
= ; [LCG, 97]
Colias sulitelma, NSG Voucher Specimen [Wahlberg]
Colias sulitelma ;
Colias hecla sulitelma ; [LCG]

Larva on Astragalus alpinus, A. eucosmus, Hedysarum mackenziei, H. alpinum, Lupinus arcticus [LCG, 97]

C. h. zamolodchikovi Churkin & Grieshuber, 2001
E.Chukotka Colias hecla zamolodchikovi Churkin & Grieshuber, 2001; Helios, Coll. lep. articles II: (174-189)

Colias canadensis Ferris, 1982
British Columbia, Alberta, Yukon, Alaska. See [About maps] Colias canadensis Ferris, 1982; Bull. Allyn Mus. 71: 11, TL: British Columbia, Mile 208, Alaska Hwy
Colias canadensis, E.H. Strickland Ent. Mus, U of [ALBERTA]
Colias canadensis, Butterflies and Moths of North America [BAMONA]
Colias canadensis ; [LCG, 97]; [Opler]

Colias boothi Curtis, 1835
Northwest Territories, Yukon, Baffin I., Victoria I., Banks I., Alaska. See [About maps] Arctic Green Sulfur
Colias boothii Curtis, 1835; in Ross, Nar. 2nd Voy. N.W. Passage, App.: 62 [65?]
chione Curtis, 1835; in Ross, Nar. 2nd Voy. N.W. Passage, App.: 62
Eurymus boothii chione ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 9
Colias boothi ab. chione ; [LCG, 98]
Eurymus boothii ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 9
Colias tyche boothii ;
Colias boothii ; [EBW]; [NACL, #4215]; [BOW: pl. 17, f. 8]
Colias boothi ; [LCG, 98]

Larva on Astragalus [EOB]


Colias thula Hovanitz, 1955
Northwest Territories, Victoria I., Banks I., Baffin I., Southampton I., Yukon, Alaska. See [About maps] Thula Sulphur
thula Hovanitz, 1955; Wasmann J. of Biol. 13 (1): (1-8), f. 1-2, TL: Alaska, Meade River S. Pt. Barrow (70.45N, 156.30W)
Colias nastes thula ;
Colias tyche thula ;
Colias thula ; [NACL, #4217]; [LCG, 99]

Larva on Lupinus arcticus [NAB], (unknown) [LCG, 99]


Colias behrii Edwards, 1866
California. See [About maps] Behr's Sulphur Sierra Green Sulfur
Colias behrii Edwards, 1866; Proc. ent. Soc. Philad. 6 (4) : 201, TL: Sierra Nevada, Yosemite Mts.
canescens (Comstock, 1925); Bull. Soc. Calif. Acad. Sci. 24 (1): 3
Colias behrii, NSG Voucher Specimen [Wahlberg]
Colias behrii f. canescens ; [LCG, 101]
Eurymus behrii ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 11
Colias behrii ; [NACL, #4219]; [EBW]; Winhard, 2000, Butterflies of the world, 10: 10, pl. 11, f. 21; [LCG, 101]; [Opler]

Larva on Vaccinium caespitosum, Vaccinium nivictum, Gentiana newberry [LCG, 101]


Colias alexandra Edwards, 1863
-. See [About maps] Queen Alexandra's Sulphur Alexandra Sulphur Ultraviolet Sulfur
Colias alexandra Edwards, 1863; Proc. ent. Soc. Philad. 2 (1): 14, pl. 11, f. 1-3, TL: Colorado, W. Denver, Pike's Peak
alba Strecker, 1878
pallida Cockerell, 1889; (not Cockerell 1888)
Colias alexandra, E.H. Strickland Ent. Mus, U of [ALBERTA]
Colias alexandra, Butterflies and Moths of North America [BAMONA]
Eurymus alexandra ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 10
Colias alexandra ; [NACL, #4211]; [EBW]; [BOW: pl. 17, f. 7]; [Opler]

Larva on Thermopsis, Astragalus, Lathyrus, Oxytropis, Lupinus [NAB]

Utah, Wyoming, Colorado, Montana, South Dakota, Nebraska, Canada (Alberta, Saskatchewan) alba Strecker, 1878; Synon. Cat. Macroplep.: 81
= ; [LCG, 102]
pallida Cockerell, 1889; Entomologist 22 : 128
= ; [LCG, 102]
Colias alexandra alexandra ; [LCG, 101]

Larva on Astragalus miser, A. lentiginosus, Astragalus diphysus, A. araneosus, A. salinus, A. eremiticus, A. canadensis, A. serotinus, A. alpinus, A. douglasii, A. parishii, A. antiselli, A. bisulcatus var. haydenianus, Hedysarum boreale, Hedysarum mackenzii, H. alpinum var. americanum, Lathyrus leucanthus, Lotus scoparins, Lupinus, Medicago sativa, Oxytropis lambertii, O. splendens, Thermopsis divaricarpa, T. pinetorum, T. montana, T. rhombifolia, Trifolium pratense, T. repens, Vicia americana [LCG, 102]

C. a. apache Ferris, 1988
Arizona, New Mexico Colias alexandra apache Ferris, 1988; Bull. Allyn Mus. 116: (1-28), TL: Arizona, Apache Co., 13 miles E. Mc Nary
Colias alexandra apache, Butterflies of America [BOA]
Colias alexandra apache ; [LCG, 102]

Larva on Thermopsis sp., Astragalus sp., Oxytropis sp., Lathyrus sp. [LCG, 103]

C. a. columbiensis Ferris, 1973
British Columbia, Alberta, Washington, Idaho, Montana Colias alexandra columbiensis Ferris, 1973; J. Lep. Soc. 27 (1): 68, f. 6, TL: British Columbia, Anderson Lake/D'Arcy
Colias alexandra columbiensis ; [NACL, #4211c]; [LCG, 103]
C. a. edwardsii Edwards, 1870
Nevada, California, Oregon, Idaho, Washington Colias alexandra edwardsii Edwards, 1870; Trans. amer. ent. Soc. 3 (1) : 11, TL: Nevada, Storey Co., Virginia City
?edwardsii Behr, 1870; in Edwards, Butts. N. Amer. 1: 55
emilia Edwards, 1870; Trans. amer. ent. Soc. 3 (1) : 12
= ; [LCG, 105]
hatui (Barnes & Benjamin, 1926); Bull. South. Calif. Acad. Sci. 25 (3): 89, TL: Stockton, Utah
= ; [LCG, 105]
Colias alexandra edwardsii, Butterflies of America [BOA]
Eurymus alexandra edwardsii ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 10
Eurymus alexandra emilia ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 10
Colias alexandra edwardsii ; [NACL, #4211a]; [LCG]

Larva on Astragalus sp., Medicago sp., Thermopsis sp., Trifolium sp. [LCG, 105]

?C. a. alberta Bowman, 1942

Colias christina Edwards, 1863
-. See [About maps] Colias christina Edwards, 1863; Proc. ent. Soc. Philad. 2 (2): 79, TL: Northwest Territories, Slave River Crossing
?pallida Lyman, 1884; Can. Ent. 16 (1) : 6
pallida Cockerell, 1889; (not Cockerell, 1988)
Eurymus christina pallida Lyman, 1884
Colias christina, E.H. Strickland Ent. Mus, U of [ALBERTA]
Colias christina, Butterflies and Moths of North America [BAMONA]
Eurymus christina ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 10
Eurymus christina pallida ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 10
Colias alexandra christina ; [NACL, #4211e]
Colias christina ; [BOW: pl. 17, f. 12]; [Opler]
Montana, Washington, British Columbia, Alberta, Manitoba, Northwest Territories Colias christina christina ; [LCG, 105]

Larva on Trifolium, Hedysarum sp. [LCG, 107]

C. c. astraea Edwards, 1872
Montana, Wyoming, Utah Colias christina astraea Edwards, 1872; Trans. amer. ent. Soc. 4 : 61, TL: Wyoming, Yellowstone Lake, Yellowstone N. P.
?Colias scudderii astraea ;
Colias alexandra astraea ; [NACL, #4211d]
Colias christina astraea ; [LCG, 107]

Colias pseudochristina Ferris, 1989
Utah, Idaho, Washington (Asotin Co.). See [About maps] Colias pseudochristina Ferris, 1989; Bull. Allyn Mus. 128: (1-11), TL: Utah, Summit Co., along North Fork of the Provo River
Colias pseudochristina, NSG Voucher Specimen [Wahlberg]
Colias christina pseudochristina ;
Colias pseudochristina ; [LCG, 107]; Verhulst, 2002, Lambillionea 102 (2): 123-126, f. 1-6

Larva on Vicia americana [LCG, 108] Verhulst, 2002, Lambillionea 102 (2): 126


Colias krauthii Klots, 1935
-. See [About maps] krauthii Klots, 1935; Amer. Mus. Novit. 767: 1, TL: South Dakota, Custer Co., Black Hills, 12mi W Custer
Colias alexandra krauthii ; [NACL, #4211f]
Colias christina krauthii ;
Colias krauthii ;
South Dakota, Wyoming Colias krauthii krauthii ; [LCG, 108]

Larva on Astragalus sp. [LCG, 109]

C. k. kluanensis Ferris, 1981
Yukon, Alaska, British Columbia Colias alexandra kluanensis Ferris, 1981; Bull. Allyn Mus. 63: 3, TL: Yukon, Alaska Hwy, E. Haines Junction
Colias christina kluanensis ; [LCG, 109]

Colias scudderii Reakirt, 1865
Rocky Mountains. See [About maps] Willow Sulphur
Colias scudderii Reakirt, 1865; Proc. ent. Soc. Philad. 4 (2) : 217, TL: Colorado, Rocky Mountains [?d'Empire, Clear Creek Co.]
Colias scudderi, Butterflies of America [BOA]
Colias scudderi, NSG Voucher Specimen [Wahlberg]
Colias scudderi, Butterflies and Moths of North America [BAMONA]
Eurymus scudderi ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 10
Colias scudderii ; [NACL, #4223]; [Opler]
Colias scudderi ; [BOW: pl. 17, f. 30]

Larva on Salix [NAB]

Colorado, Utah, New Mexico, Wyoming flavotincta Cockerell, 1901; Psyche 9: 186, TL: Lake Co., Colorado
= Colias scudderi scudderi ; [LCG, 119]
?flavotincta Klots, 1937
Eurymus scudderi flavotincta ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 10
Colias scudderi scudderi ; [LCG, 110]

Larva on Salix reticulata, S. lutea, S. planifolia [LCG, 111]

C. s. ruckesi Klots, 1937
New Mexico scudderii ruckesi Klots, 1937; J. N.Y. Ent. Soc. 45 (3/4) : 324, TL: New Mexico, Windsor Creek Canyon, W of Cowles
Colias scudderii ruckesi ; [NACL, #4223a]
Colias scudderi ruckesi ; [LCG, 111]

Colias pelidne Boisduval & Le Conte, [1829]
-. See [About maps] Blueberry Sulphur
Colias pelidne Boisduval & Le Conte, [1829]; Hist. Lép. Am. Sept. (7/8): 62-66, pl. 21, f. 4-5, TL: Canada, Arctic region [Labrador?]
Colias pelidne, E.H. Strickland Ent. Mus, U of [ALBERTA]
Colias pelidne, NSG Voucher Specimen [Wahlberg]
Colias pelidne, Butterflies and Moths of North America [BAMONA]
Eurymus pelidne ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 10
Colias pelidne ; [NACL, #4221]; [EBW]; [BOW: pl. 17, f. 19]; [Opler]

Larva on Vaccinium, Gaultheria humifusa [NAB]

Manitoba, Labrador, New Foundland, Ontario, Quebec, Saskatchewan, Yukon Colias labradorensis Scudder, 1862; Proc. Boston Soc. Nat. Hist. 9 : 107
= ; [LCG, 113]; Verhulst, 2002, Lambillionea 102 (4, I): 515 (form); Dyar, 1903, Bull. U.S. nat. Mus. 52 : 10
Colias minisni Bean, 1895; Psyche 7 : 228, TL: Alberta
= ; [LCG, 113]; Verhulst, 2002, Lambillionea 102 (4, I): 515 (form)
mira Verity, 1911; Rhopalocera Palaearctica 1: 347, TL: Labrador
= ; [LCG, 113]; Verhulst, 2002, Lambillionea 102 (4, I): 515 (form)
Colias pelidne ab. moeschleri Grum-Grshimailo, 1893; Horae Soc. ent. Ross. 27 : 379
= ; [LCG, 113]; Verhulst, 2002, Lambillionea 102 (4, I): 515 (form)
standfussi Röber, [1909]; in Seitz, Grossschmett. Erde 5 : 91
= pelidne × nastes ; [LCG, 113]; Verhulst, 2002, Lambillionea 102 (4, I): 515 (hybdrid)
isni (Barnes & Benjamin, 1926)
menisme ; Verity, 1908, (missp.)
Colias pelidne labradorensis ;
Colias pelidne minisini[sic?] ; [NACL, #4221a]
Colias pelidne pelidne ; [LCG, 112]; Verhulst, 2002, Lambillionea 102 (4, I): 514
Misidentified/misapplied:
Colias anthyale; Möschler, 1870, Stettin Ent. Ztg 31 (1-3): 113
= ; [LCG, 113]; Verhulst, 2002, Lambillionea 102 (4, I): 515 (form)
C. p. skinneri Barnes, 1897
Alberta, British Columbia, Idaho, Montana, Wyoming skinneri Barnes, 1897; Can. Ent. 29 (2) : 41-42, TL: Wyoming, Yellowstone N. P.
neri Barnes & Benjamin, 1926; (female)
Eurymus pelidne skinneri ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 10
Colias pelidne skinneri ; [NACL, #4221b]; [LCG, 113]; Verhulst, 2002, Lambillionea 102 (4, I): 517

Colias gigantea Strecker, 1900
-. See [About maps] Great Northern Sulphur
Colias pelidne var. gigantea Strecker, 1900; Lep. Rhopal. Het., Suppl. 3 : 19, TL: W. coast Hundson Bay, N. Fort York
?gigantea Verity, 1911²; Rhopalocera Palaearctica 1: 347
Colias gigantea, E.H. Strickland Ent. Mus, U of [ALBERTA]
Colias gigantea, NSG Voucher Specimen [Wahlberg]
Colias gigantea, Butterflies and Moths of North America [BAMONA]
?Colias scudderii gigantea ;
Eurymus pelidne gigantea ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 10
Colias gigantea ; [EBW]; [NACL, #4222]; [BOW: pl. 17, f. 15]; [Opler]

Larva on Salix reticulata, Salix [NAB]

Alaska, Alberta, British Columbia, Manitoba, Saskatchewan, Ontario, Yukon, Northwest Territories marjorie F. & R. Chermock, 1940; Can. Ent. 72 (4): 81
= ; [LCG, 115 (form)]
Colias gigantea gigantea ; [LCG, 114]
C. g. mayi F. & R. Chermock, 1940
Colias christina mayi F. & R. Chermock, 1940; Can. Ent. 72 (4): 82
Colias gigantea mayi ; [NACL, #4222a]
C. g. harroweri Klots, 1940
Wyoming, Montana, Oregon Colias gigantea harroweri Klots, 1940; Amer. Mus. Novit. 1054: 4, TL: Wyoming, Sublette Co., Clear Creek env./vic. Lower Green River Lake 2560m
Colias scudderii harroweri ;
Colias gigantea harroweri ; [NACL, #4222b]; [LCG, 115]

Colias occidentalis Scudder, 1862
-. See [About maps] Western Sulphur Golden Sulfur
Colias occidentalis Scudder, 1862; Proc. Boston Soc. Nat. Hist. 9 : 109, TL: British Columbia, Mackenzie River, Georgia Golf
Colias occidentalis occidentalis, Butterflies of America [BOA]
Colias occidentalis, NSG Voucher Specimen [Wahlberg]
Colias occidentalis, Butterflies and Moths of North America [BAMONA]
Eurymus occidentalis ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 10
Colias occidentalis ; [NACL, #4212]; [EBW]; [Opler]

Larva on Lathyrus spp., Thermopsis spp. Hammond & McCorkle, 2003, J. Lep. Soc. 57 (4): 276

British Columbia, Washington, Oregon Colias occidentalis occidentalis ; [LCG, 115]
C. o. chrysomelas H. Edwards, 1877
California, Oregon Colias occidentalis chrysomelas H. Edwards, 1877; Proc. Cal. Acad. Sc.: 8, TL: California, Napa Co.
shastae (Barnes & Benjamin, 1926); Bull. South. Calif. Acad. Sci. 25 (3): 88
= ; [LCG, 117 (form)]
Colias occidentalis chrysomelas, Butterflies of America [BOA]
chryomelas ; (missp.)
chrysomelas ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 10 (missp.?)
Colias chrysomelas ; [BOW: pl. 17, f. 9 (text only)]
Colias occidentalis chrysomelaena ; [NACL, #4212a (missp.?)]
Colias occidentalis chrysomelas ; [LCG, 116]
C. o. sullivani Hammond & McCorkle, 2003
Oregon Colias occidentalis sullivani Hammond & McCorkle, 2003; J. Lep. Soc. 57 (4): 275, f. 1, TL: Oregon, Harney Co., Alvord Desert Rd at north end of Steens Mts

Larva on Lathyrus rigidus Hammond & McCorkle, 2003, J. Lep. Soc. 57 (4): 276


Colias interior Scudder, 1862
Canada, New Foundland - Michigan. See [About maps] Pink-edged Sulphur
Colias interior Scudder, 1862; Proc. Boston Soc. Nat. Hist. 9 : 108, TL: N. shore of Lake Superior
Colias interior, E.H. Strickland Ent. Mus, U of [ALBERTA]
Colias interior, Butterflies of America [BOA]
Colias interior, NSG Voucher Specimen [Wahlberg]
Colias interior, Butterflies and Moths of North America [BAMONA]
interier Kirby, 1877; (missp.)
Eurymus interior ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 10
Colias interior ; [NACL, #4220]; [EBW]; [BOW: pl. 17, f. 18]; [Opler]
[interior-1vU.jpg] identification uncertain 600x574 (~40Kb) underside USA: Gnat Flat, Wenatchee National Forest, (47°08N 120°58W), Kittitas Co., WA, 29.7.1999, Photo © Markku Savela

Larva on Vaccinium myrtilloides, Vaccinium canadensis [NAB]


Alberta, British Columbia, Labrador, Brunswick, New Foundland, Northwest Territories, Nova Scotia, Ontario, Prince Edward Is., Quebec, Saskatchewan, Idaho, Maine, Michigan, Minnesota, Montana, Oregon, Washington, Virginia, Wisconsin nepi Barnes & Benjamin, 1926; Bull. South. Calif. Acad. Sci. 25 (3): 89
= ; [LCG, 119 (form)]
solivaga Edwards, 1877; Trans. amer. ent. Soc. 6 : 17
= ; [LCG, 119 (form)]
Colias interior interior ; [LCG, 117]; Verhulst, 2002, Lambillionea 102 (3, I): 291, f. 1-8

Larva on Vaccinium myrtilloides, V. corymbosum, V. caespitosum, Vaccinium angustifolium, V. vacillans Verhulst, 2002, Lambillionea 102 (3, I): 294

C. i. laurentina (Scudder, 1875)
Nova Scotia (Cape Breton) Eurymus philodice[?] var. laurentina Scudder, 1875; Proc. Boston Soc. nat. Hist. 18: 188, TL: Canada, Nova Scotia, Cape Breton Island, Baddeck (neotype)
raritus Gunder, 1928
Colias interior laurentina ; [NACL, #4220b]; [LCG, 119]; Verhulst, 2002, Lambillionea 102 (3, I): 294

Larva on Vaccinium myrtilloides, V. caespitosum, V. corymbosum Verhulst, 2002, Lambillionea 102 (3, I): 295

C. i. vividior Berger, 1945
Wisconsin Colias interior vividior Berger, 1945; Lambillionea 45 (6-8): 42, TL: Wisconsin, Oneida, Eagle River
Colias interior vividior ; [NACL, #4220a]; [LCG, 120]; Verhulst, 2002, Lambillionea 102 (3, I): 295

Colias harfordii H. Edwards, 1877
California, Baja California. See [About maps] Colias harfordii H. Edwards, 1877; Proc. Cal. Acad. Sc.: 9, TL: California, Kern Co. & Contra Costa Co., Havilnh
Colias barbara H. Edwards, 1877; Proc. Cal. Acad. Sc.: 7, TL: California
= ; [LCG, 120 (form)]; [NL4A, #56]
Eurymus hartfordii[sic] ab. weaverae Gunder, 1924; Ent. News 35 (5) : 156, TL: California
= ; [LCG, 120 (form)]; [NL4A, #56]
Eurymus harfordii f. martini Gunder, 1931; Bull. South. Calif. Acad. Sci. 30 (2): 45, TL: California
= ; [LCG, 120 (form)]; [NL4A, #56]
Colias harfordii, Listing of Mexican Butterflies [ILMB]
Colias harfordii, Butterflies of America [BOA]
Colias harfordii, NSG Voucher Specimen [Wahlberg]
Colias harfordii, Butterflies and Moths of North America [BAMONA]
Colias alexandra harfordii ; [NACL, #4211b]
Eurymus hartfordii ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 10 (missp.)
Colias harfordi ; [EBW]
Colias harfordii ; [LCG, 120]; [Opler]; [NL4A, #56]

Larva on Astragalus douglasii, Astragalus antiselli, Lotus scoparius, L. crassifolius [LCG, 120]


Colias philodice Godart, 1819
-. See [About maps] Common Sulphur
Colias philodice Godart, 1819; Encyclopédie Méthodique. 9 (1): 100, TL: "Amérique sept."
europome Haworth, 1803; (not Esper, 1778)
dorippe Godart, 1819
europome Stephens, 1828; (not Esper, 1778)
suffusa Cockerell, 1889
Colias philodice, E.H. Strickland Ent. Mus, U of [ALBERTA]
Colias philodice, NSG Voucher Specimen [Wahlberg]
Colias philodice, Butterflies and Moths of North America [BAMONA]
Eurymus philodice ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 10
Colias philodice ; Godman & Salvin, [1889], Biol. centr.-amer., Lep. Rhop. 2: 152; [EBW]; [NACL, #4209]; [BOW: pl. 17, f. 9]; [Opler]; [NL4A, #58]
[philodice-1v.jpg] 600x745 (~66Kb) underside USA: Alabama, Jeff. Co., 7.9.1998, Photo © Vitaly Charny [philodice-2v.jpg] 900x817 (~85Kb) underside USA: Alabama, Jackson Co., 26.5.2001, Photo © Vitaly Charny [philodice-3v.jpg] 1000x801 (~133Kb) underside USA: Alabama, 27.3.2004, Photo © Vitaly Charny

Canada (Quebec, Alberta), Montana, Colorado, California, Virginia, NA.Georgia, Massachusetts, Illinois, Michigan, Pennsylvania, New York, Utah, Mexico Papilio ?notatus Megerle, 1803; (nom. dub.), TL: Georgia
= ; [NL4A, #58a]
Colias dorippe Godart, 1819; , TL: "Cape of Good Hope"
= ; [NL4A, #58a]
Zerene anthyale Hübner, 1823; Zuträge Samml. exot. Schmett. 2 : f. 307-308 (spr. f.), TL: Pennsylvania
= ; [LCG, 121 (form)]; [NL4A, #58a]
enegthenu Boisduval, 1852; (missp.)
= ; [LCG, 121 (form)]
santes Fitch, 1854; Trans. N.Y. St. agric. Soc. 13: 378
= ; [LCG, 122 (form)]
flava Strecker, 1878; Synon. Cat. Macroplep.: 82
= ; [LCG, 121 (form)]
philodin Edwards, 1872; in Hayden, Geol. Surv. Montana: 466
= ; [LCG, 122 (form)]
ab. alba Strecker, 1878; Synon. Cat. Macroplep.: 82, TL: USA
= ; [LCG, 121 (form)]; [NL4A, #58a]
ab. nig Strecker, 1878; Synon. Cat. Macroplep.: 82, TL: New York,Canada
= ; [LCG, 122 (form)]; [NL4A, #58a]
ab. virida Strecker, 1878; Synon. Cat. Macroplep.: 82, TL: Canada
= ; [LCG, 122 (form)]; [NL4A, #58a]
alba Pilate, 1888; Papilio (2) 5[?]: 65
= ; [LCG, 121 (form, = alba Strecker, 1878)]
= f. alba (Strecker, 1878); Verhulst, 2003, Lambillionea 103 (1): 73
maria Edwards, 1883; Papilio 3 (7-10) : 171 [?] (?1885)
= ; [LCG, 122 (form)]; [NL4A, #58a]
hagenii Edwards, 1883; Papilio 3 (7-10) : 160 (Canada)
= ; [LCG, 121 (form)]
autumnalis Cockerell, 1888; W. Amer. Sci (4) 33: 42 (fall f.)
= ; [LCG, 121 (form)]
Colias philodice suffusa Cockerell, 1889
= ; [NL4A, #58a]
Eurymus philodice nigridice Scudder, 1889; Butts eastern U.S. Canada 2: 1114, TL: New York
= ; [LCG, 122 (form)]; [NL4A, #58a]
Eurymus philodice miscidice Scudder, 1889; Butts eastern U.S. Canada 2: 1115, TL: Massachusetts
= ; [LCG, 122 (form)]; [NL4A, #58a]
Eurymus philodice pallidice Scudder, 1889; Butts eastern U.S. Canada 2: 1115, TL: USA
= ; [LCG, 122 (form)]; [NL4A, #58a]
Colias philodice luteitincta Wolcott, 1893; Can. Ent. 25: 104, TL: Illinois; Michigan
= ; [LCG, 121 (form)]; [NL4A, #58a]
var. albinic Skinner, 1898; , TL: USA
= ; [NL4A, #58a]
var. melanic Skinner, 1898; Cat. North Am. Rhopalocera: 68, TL: USA
= ; [NL4A, #58a]
ab. nigrina Strecker, 1900; Lep. Rhopal. Het., Suppl. 3 : 19 (emend.)
= ; [LCG, 122 (form)]; [NL4A, #58a]
ab. alba Röber, 1909; , TL: Pennsylvania
= ; [NL4A, #58a]
alba Röber, [1910]; in Seitz, Grossschmett. Erde 5 : 91
= ; [LCG, 121 (form, = alba Strecker, 1878)]
= f. alba (Strecker, 1878); Verhulst, 2003, Lambillionea 103 (1): 73
Colias philodice f. nigrofasciata Reiff, 1917; Massachusetts 1 (3) : 22-23, pl. 3 (preocc. Grum-Grshimailo, 1884), TL: Massachusetts
= ; [NL4A, #58a]
kootenai Cockle, 1910; Can. Ent. 42 (6) : 203 (altid. f.)
= ?Colias philodice eriphyle ;
Colias philodice f. rothkei Reiff, 1917; Lepidopterist 1 (11) : 84, pl. 7, TL: Pennsylvania
= ; [NL4A, #58a]
Colias philodice pallidice f. nigrofasciata Reiff, 1918; (not Reiff, 1917), TL: Massachusetts
= ; [NL4A, #58a]
f. plicaduta Nakahara, 1926; Bull. Brooklyn ent. Soc. 20 (5): 222, TL: New York
= ; [LCG, 122 (form)]; [NL4A, #58a]
ab. inversata Nakahara, 1926; Bull. Brooklyn ent. Soc. 20 (5): 223, TL: Pennsylvania
= ; [LCG, 121 (form)]; [NL4A, #58a]
f. plicaduta ab. minor Chermock, 1927; Bull. Brooklyn ent. Soc. 22 (2): 118, TL: Pennsylvania
= ; [LCG, 122 (form)]; [NL4A, #58a]
f. plicaduta ab. ehrmanni Chermock, 1927; Bull. Brooklyn ent. Soc. 22 (2): 118, TL: Pennsylvania
= ; [LCG, 121 (form)]; [NL4A, #58a]
f. plicaduta var. alba Chermock, 1927; Bull. Brooklyn ent. Soc. 22 (2): 119, TL: Pennsylvania
= ; [NL4A, #58a]
f. plicaduta var. albida (Chermock, 1928) (Eurymus); , TL: Pennsylvania
= ; [LCG, 121 (form)]; [NL4A, #58a]
tr.f. raritus (Gunder, 1928) (Eurymus); Can. Ent. 60 : 163, TL: Pennsylvania
= ; [LCG, 122 (form)]; [NL4A, #58a]
var. serrata (Chermock, 1929) (Eurymus); Bull. Brooklyn ent. Soc. 24 (1): 21, TL: Pennsylvania
= ; [LCG, 122 (form)]; [NL4A, #58a]
laurae Chermock, 1929; Bull. Brooklyn ent. Soc. 24 (1): 21
= ; [LCG, 121 (form)]
eriphyle var. nigricosta Chermock, 1929; Bull. Brooklyn ent. Soc. 24 (1): 21
= ; [LCG, 122 (form)]
notatus Clark & Clark, 1941; Proc. Ent. Soc. Wash. 43 (4): 83
= ; [LCG, 122 (form)]
pallidissima Bowman, 1942; Can. Ent. 74 (2): 25
= ; [LCG, 122 (form)]
ab. minor Dufrane, 1947; , TL: [Canada]
= ; [NL4A, #58a]
f. anthyale ab. reducta Dufrane, 1947; Bull. Ann. Soc. ent. Belg. 83 (1/2) : 69, TL: Pennsylvania
= ; [LCG, 122 (form)]; [NL4A, #58a]
Eurymus philodice anthyale ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 10
Colias hageni ; [BOW: pl. 17, f. 17]
Eurymus philodice nigridice ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 10
miscidice ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 10
Eurymus philodice luteitincta ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 10
Colias philodice philodice ; Winhard, 2000, Butterflies of the world, 10: 10, pl. 11, f. 22; [LCG]; Verhulst, 2003, Lambillionea 103 (1): 72; [NL4A, #58a]

Larva on Astragalus miser, A. adsurgens, Astragalus decumbens, A. crassicarpus, A. plattensis, Baptisia tinctoria, Caragana, Cytisus, Hedysarum boreale, Lathyrus leucanthus, Lotus, Lupinus perennis, Medicago sativa, M. hispida, Melilotus alba, Pisum sativum, Robinia pseudoacacia, Thermopsis divaricarpa, Trifolium repens, T. pratense, T. agrarium, T. hybridum, T. longipes, Vicia cracca, V. americana Verhulst, 2003, Lambillionea 103 (1): 74

C. p. eriphyle Edwards, 1876
British Columbia, Colorado, Wyoming, Montana Colias eriphyle Edwards, 1876; Trans. amer. ent. Soc. 5 : 202, TL: British Columbia, Lake Labache
?Colias eriphyle Edwards, 1878; Bull. Geol. Surv. Terr. 4 (2) : 514, TL: Milk River
Colias philodice eriphyle, Listing of Mexican Butterflies [ILMB]
Colias philodice eriphyle, Butterflies of America [BOA]
Colias philodice eriphyle, Butterflies of Southeastern Arizon [Bruce Walsh]
Eurymus eurytheme eriphyle ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 9
Colias philodice eriphyle ; [LCG, 122]; Verhulst, 2003, Lambillionea 103 (1): 75
C. p. guatemalena Röber, [1909]
Guatemala philodice guatemalena Röber, [1909]; in Seitz, Grossschmett. Erde 5: (?) , TL: Guatemala
= Colias philodice philodice ; [LCG, 121 (form)]
Colias philodice guatemalena, Butterflies of America [BOA]
Colias philodice guatemalena ; [NL4A, #58b]
C. p. vitabunda Hovanitz, 1943
Alaska, Canada (British Columbia, Yukon, Northwest Territories) Colias philodice vitabunda Hovanitz, 1943; Amer. Mus. Novit. 1240: 3, TL: Alaska, McKinley National Park
Colias philodice vitabunda ; [NACL, #4209a]; [LCG, 123]

Colias eurytheme Boisduval, 1852
New Mexico, California, Oregon, Washington, Colorado, Nevada, Arkansas, Tenessee, New York, Missouri, Illinois, Mexico. See [About maps] Orange Sulphur
Colias eurytheme Boisduval, 1852; Ann. Soc. ent. Fr. (2) 10 : 286, TL: California
Colias amphidusa Boisduval, 1852; Ann. Soc. ent. Fr. (2) 10 : 286, TL: California
= ; [LCG, 125 (form)]; [NL4A, #53]
Colias edusa var. californiana Ménétriés, 1855; Cat. lep. Petersb. 2: 80, TL: California
= ; [LCG, 125 (form)]; [NL4A, #53]
Colias keewaydin Edwards, 1869; Butts. N. Amer. 1: 49, TL: Missouri
= ; [LCG, 125 (form)]; [NL4A, #53]
Colias ariadne Edwards, 1870; Trans. amer. ent. Soc. 3 (1) : 12, TL: California
= ; [LCG, 125 (form)]; [NL4A, #53]
Colias philodice ab. hybrida Strecker, 1878; , TL: Illinois
= Colias eurytheme × philodice pallida ; [LCG, 125 (hybrid)]
= ; [NL4A, #53]
Colias chrysotheme var. eurytheme ab. alba Strecker, 1878; , TL: USA
= ?Colias philodice philodice ; [LCG, 125 (form)]
= ?; [LCG, 125 (form)]
= ; [NL4A, #53]
Colias chrysotheme var. keewaydin ab. alba Strecker, 1878; , TL: California; Texas
= ; [NL4A, #53]
f. intermedia Cockerell, 1888; W. Amer. Sci (4) 33: 41, TL: Colorado
= ; [LCG, 125 (form)]; [NL4A, #53]
Colias eurytheme intermedia f. pallida Cockerell, 1888; W. Amer. Sci (4) 33: [?] 41 , TL: Colorado
= ; [LCG, 125 (form)]; [NL4A, #53]
f. pallida Cockerell, 1888; , TL: Colorado
= ; [NL4A, #53]
autumnalis Cockerell, 1888; W. Amer. Sci (4) 33: 42 (fall f.)
= ; [LCG, 125 (form)]
typica Cockerell, 1888; W. Amer. Sci (4) 33: 41
= ; [LCG, 125 (form)]
ab. fumosa Strecker, 1900; Lep. Rhopal. Het., Suppl. 3 : 19, TL: Colorado
= ; [LCG, 125 (form)]; [NL4A, #53]
ab. albina Röber, [1909]; in Seitz, Grossschmett. Erde 5: [??] 93, TL: [Nevada]
= ; [LCG, 125 (form, = alba)]; [NL4A, #53]
ab. alba Röber, 1909; (nom. nud.)
= ; [NL4A, #53]
f. amphidusa ab. unicitrina (Gunder, 1924) (Eurymus); Ent. News 35 (5) : 158, TL: California
= ; [LCG, 125 (form)]; [NL4A, #53]
tr.f. rudkini (Gunder, 1932) (Eurymus); Can. Ent. 64 (2): 277, TL: California
= ; [LCG, 125 (form)]; [NL4A, #53]
Colias eurytheme alberta Bowman, 1942; Can. Ent. 74 (2): 25, TL: Canada
= ; [LCG, 125 (form)]; [NL4A, #53]
Colias eurytheme, E.H. Strickland Ent. Mus, U of [ALBERTA]
Colias eurytheme, Listing of Mexican Butterflies [ILMB]
Colias eurytheme, Butterflies of America [BOA]
Colias eurytheme, NSG Voucher Specimen [Wahlberg]
Colias eurytheme, Butterflies and Moths of North America [BAMONA]
Colias eurytheme, Butterflies of Southeastern Arizon [Bruce Walsh]

Colias eurytheme keewaydin ;
amphidona Edwards, 1869; (missp.)
enegthenu Edwards, 1872; (missp.)
Eurymus eurytheme ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 9
Eurymus eurytheme ariadne ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 9
Eurymus eurytheme fumosa ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 9
Colias eurytheme f. ariadne ; Winhard, 2000, Butterflies of the world, 10: 10, pl. 11, f. 23
Colias eurytheme ; [NACL, #4210]; [EBW]; [BOW: pl. 17, f. 16]; Winhard, 2000, Butterflies of the world, 10: 10, pl. 11, f. 24; [LCG, 124]; [Opler]; [NL4A, #53]
Misidentified/misapplied:
?Colias chrysotheme; Godman & Salvin, [1889], Biol. centr.-amer., Lep. Rhop. 2: 151
[eurytheme-1v.jpg] 500x470 (~35Kb) underside USA, Texas, Irving, 3-14.6.1996, Photo © Tero Piirainen [eurytheme-2v.jpg] 967x985 (~103Kb) underside USA: Alabama, 23.3.2001, Photo © Vitaly Charny [eurytheme-3o.jpg] 1300x975 (~156Kb) ova USA: Alabama, 11.5.2004, Photo © Vitaly Charny
[eurytheme-4v.jpg] 1071x1218 (~120Kb) underside USA: Alabama, 24.9.2004, Photo © Vitaly Charny [eurytheme-5v.jpg] 800x1012 (~123Kb) underside USA: Alabama, 24.9.2004, Photo © Vitaly Charny [eurytheme-6fvU.jpg] identification uncertain 1600x1320 (~214Kb) underside female USA: Colorado, Clear Creek Co., Summit Lake, Mt. Evans, 23.7.2012 (39°35'52"N 105°38'23"W), Photo © Markku Savela
[eurytheme-6mu.jpg] 700x798 (~75Kb) upperside male USA: Colorado, Clear Creek Co., roadside on 103 (Squaw Pass Rd) from Mt Evans to I-70, 23.7.2012, Photo © Markku Savela [eurytheme-6mv.jpg] 1000x846 (~103Kb) underside male USA: Colorado, Clear Creek Co., roadside on 103 (Squaw Pass Rd) from Mt Evans to I-70, 23.7.2012, Photo © Markku Savela [eurytheme-7fu.jpg] 1200x1123 (~164Kb) upperside female USA: Colorado, Aurora, Plains Conservation Center, 24.7.2012, Photo © Markku Savela

Larva on Astragalus drummondii, A. flexuosus, A. racemosus, A. bisulcatus, A. crassicarpus, A. plattensis, A. crotalariae, Astragalus antiselli, A. whitneyi, A. alpinus, Baptisia, Cassia, Coronilla, Glycine max, Glycyrrhiza lepidota, Lotus scoparius, L. purshianus, L. grandiflorus, L. crassifolius, L. subpinnatus, L. strigosus, Lathyrus leucanthus, L. jepsoni ssp. californicus, Lespedeza, Lupinus minimus, L. perennis, L. succulentus, L. bicolor, Medicago sativa, M. hispida, M. lupulina, Melilotus officinalis, M. alba, Phaseolus, Pisum sativum, Psoralea, Sesbania exalata, Thermopsis montana, Trifolium repens, T. nanum, T. pratense, T. reflexum, T. stoloniferum, T. tridentatum, T. longipes, T. wormskjoldii, Vicia americana, V. cracca, V. angustifolia, V. sativa [LCG, 124]



Colias lada Grum-Grshimailo, 1891
Tibet, China (Gansu). See [About maps] lada Grum-Grshimailo, 1891; Horae Soc. ent. Ross. 25 (3-4) : 447, TL: China, Amdo
Colias lada ab. oreas Grum-Grshimailo, 1893; Horae Soc. ent. Ross. 27 : 383
= ; [LCG, 126 (form)]
Colias lada ; [LCG, 125]

Colias marcopolo Grum-Grshimailo, 1888
-. See [About maps] Colias marco-polo Grum-Grshimailo, 1888; Horae Soc. ent. Ross. 22 (3-4) : 304, TL: Kunzhut Mts., E. Hindu Kush
Colias marcopolo, NSG Voucher Specimen [Wahlberg]
Colias marcopolo, Extracts from [BRU] by [Ilya Osipov]
Colias marcopolo ; [BOW: pl. 158, f. 20]; [BRU, 179]

Larva on Astragalus spp. [BRU]

Pamirs, Hindukush diana Schulte, 1989; Nacr. Ent. Ver. Apollo Frankf. N. F. 10 (4): (291-297)
= ; [LCG, 126 (form)]
nicopolo Röber, [1907]; in Seitz, Grossschmett. Erde 1 : 62
= ; [LCG, 126 (form)]
Colias marcopolo marcopolo ; [LCG, 126]
C. m. afganipolo Schulte, 1977
Afghanistan marcopolo afganipolo Schulte, 1977; Entomops, Nice 42: 35
Colias marcopolo afganipolo ; [LCG, 126]
C. m. kushana Wyatt & Omoto, 1966
Afghanistan Colias marcopolo kushana ; [LCG, 126]

Colias christophi Grum-Grshimailo, 1885
-. See [About maps] Colias christophi Grum-Grshimailo, 1885; in Romanoff, Mém. Lép. 2 : 220, TL: Archa-Bashi N. of Pass Kara-Kasyk, Alaisky Mts, Kirghizia
Colias christophi, NSG Voucher Specimen [Wahlberg]
Colias christophi, Extracts from [BRU] by [Ilya Osipov]
Colias christophi ; [BRU, 171]
Colias cristophi ; [EBW, (missp.?)]

Larva on Astragalus spp. [BRU]

Kirghizia aufderheidei Kotzsch, 1929; Ent. Zeit. 43: 205
= ; [LCG, 127 (= novosiltzovi)]
heidei Kotzsch, 1929; Ent. Zeit. 43: 205
= ; [LCG, 127]
bang-haasi Bollow, 1930; in Seitz, Macrolep. du Globe, 1: 110
= ; [LCG, 127 (form)]
Colias christophi ab. novosiltzovi Avinoff, 1910; Horae Soc. Ent. Ross. 39 : 248, pl. 14, f. 5
= ; [LCG, 127 (form)]
simplicissima Avinoff, 1910; Horae Soc. Ent. Ross. 39 : 248, pl. 14, f. 5 [?]
= ; [LCG, 127 (= novosiltzovi)]
Colias christophi helialaica Schulte, 1988; Nacr. Ent. Ver. Apollo 9 (3): 149[?], TL: Kirgizia, Alai Mts., ca. 100 km SE Fergana, between Schachimardan and Uchkurgan, Kaindy Mt., Isfara
= ; [LCG, 127]
Colias christophi christophi ; [LCG, 127]
C. c. kali Korb, 1999
Tadzhikistan Colias christophi kali Korb, 1999; Alexanor 20 (8): 452[?], TL: Tadzhikistan, Turkestansky Ridge, Chakhristan
Colias christophi kali ; Winhard, 2000, Butterflies of the world, 10: 10, pl. 12, f. 1-2; [LCG, 128]

Colias shahfuladi Clench & Shoumatoff, 1956
Afghanistan. See [About maps] shahfuladi Clench & Shoumatoff, 1956; Vidensk. Medd. Dansk Naturh. Forening 118 : 173, TL: Afghanistan, Takatu 4500m
Colias shahfuladi ; [LCG, 128]

Colias alpherakii Staudinger, 1882
-. See [About maps] Colias alpherakii Staudinger, 1882; in Staudinger & Bang-Haas, Berl. ent. Z. 26 (1) : 164, TL: Northern slopes of the Alaisky Mts., Kirghizia
Colias alpherakyi, NSG Voucher Specimen [Wahlberg]
Colias alpherakyi, Extracts from [BRU] by [Ilya Osipov]
Colias alpherakyi ; [BRU, 171 (missp.)]
Colias alpherakii ; [BOW: pl. 158, f. 14]

Larva on Onobrychis echidna [BRU]

Kirghizia, Tadzhikistan Colias alpherakii ab. ♀ leucophryna Grum-Grshimailo, 1893; Horae Soc. ent. Ross. 27 : 381
= ; [LCG, 130 (form)]
versa Verity, 1911; Rhopalocera Palaearctica 1: 349
= ?Colias alpherakii roschana ;
= ; [LCG, 130 (form)]
Colias alpherakii alpherakii ; Winhard, 2000, Butterflies of the world, 10: 10, pl. 12, f. 3-4; [LCG, 129]

Larva on Astragalus sp. [LCG, 130]

C. a. chitralensis Verity, 1911
Pakistan (Chitral, ...) alpherakii chitralensis Verity, 1911; Rhopalocera Palaearctica 1: 349, pl. 69, f. 31-32, TL: Pakistan, Chitral, Shandur Lake, 3600m
Colias alpherakii chitralensis ; [LCG, 130]
C. a. kohibaba Wyatt & Omoto, 1966
Afghanistan, Tadzhikistan (W.Pamir, ...) alpherakii kohibaba Wyatt & Omoto, 1966; Entomops 5: 146, TL: Afghanistan, Panjaw
Colias alpherakyi[sic] kohabiba[sic, recte kohibaba] ; [NHM]
Colias alpherakii kohibaba ; [LCG, 131]
C. a. roschana Grum-Grshimailo, 1893
W.Pamirs Colias alpherakii var. roschana Grum-Grshimailo, 1893; Horae Soc. ent. Ross. 27 : 381, TL: Tadzhikistan, Koudara
Colias alpherakii roschana ; [LCG, 131]
C. a. tashkurgonica Kesküla, 1997
Uzbekistan Colias alpherakii tashkurgonica Kesküla, 1997; Atalanta 28 (1/2): 33[?], TL: Uzbekistan, Kashkadarya prov., Chakchar Range, Tashkurgon Pass
Colias alpherakii tashkurgonica ; [LCG, 132]
C. a. usmatica Shchetkin, 1990
Uzbekistan (W.Turkestan Range) Colias alpherakii usmatica Shchetkin, 1990; Bull. Soc. nat. Moscou 95: (58-60), TL: Uzbekistan, W.Turkestan Range, Tchoukmartaou Mts., env. village Ousmat, 2750-2800m
Colias alpherakii usmatica ; [LCG, 132]

Colias montium Oberthür, 1886
-. See [About maps] Colias montium Oberthür, 1886; Étud. d'Ent. 11 : 16, pl. 6, f. 1, TL: Sinkiang, Ta-Tsien-Lou
Colias montium, NSG Voucher Specimen [Wahlberg]
Colias montium ; [BOW: pl. 4, f. 8 (text only)]; [MRS, 221]
Tibet, Sichuan Colias montium montium ; [LCG, 133]
C. m. viridis Bang-Haas, 1915
Qinghai montium viridis Bang-Haas, 1915; Dt. ent. Z. Iris 29 (2/3) : 99, TL: Chotan
Colias montium viridis ; [LCG, 134]
?C. m. longto Evans, 1924
S.Tibet Colias montium longto Evans, 1924; J. Bombay nat. Hist. Soc. 29 (4): 973, TL: Tibet, Gyangze
?Colias montium longto ; [LCG, 134 (stat. uncert.)]
?C. m. fasciata Kocman, 1999
Qinghai Colias montium fasciata Kocman, 1999; Wallace 5: 55[?], TL: S.Qinghai, Chiehku, 2300m
?Colias montium fasciata ; [LCG, 135 (= thrasibulus?)]

Colias sieversi Grum-Grshimailo, 1887
Tadzhikistan (Darvaz?, Peter I Mts.). See [About maps] Colias sieversi Grum-Grshimailo, 1887; in Romanoff, Mém. Lép. 3 : 397, TL: Kara-Shura River valley near Pass Gardani-Kaftar, Tuptchek Plataeu, Peter 1 Mts, Tajikistan
Colias sieversi, NSG Voucher Specimen [Wahlberg]
Colias sieversi, Extracts from [BRU] by [Ilya Osipov]
Colias sieversi ; [BRU, 171]; [BOW: pl. 158, f. 21]; [MRS, 222]; [LCG, 135]

Colias sifanica Grum-Grshimailo, 1891
-. See [About maps] Colias sifanica Grum-Grshimailo, 1891; Horae Soc. ent. Ross. 25 (3-4) : 447, TL: China, Amdo, Kukunor
Colias sifanica, NSG Voucher Specimen [Wahlberg]
Colias sifanica ; [MRS, 222]
Colias sifanica sifanica ; Winhard, 2000, Butterflies of the world, 10: 10, pl. 12, f. 5-6
Amdo, Qinghai, Gansu tancrei Röber, [1907]; in Seitz, Grossschmett. Erde 1 : 64, pl. 25
= ; [LCG, 137 (form)]
Colias sifanica sifanica ; [LCG, 136]

Larva on Caragana sp. [LCG, 137]

C. s. herculeana Bollow, 1930
Gansu sifanica herculeana Bollow, 1930; in Seitz, Macrolep. du Globe, 1: 113, suppl. pl. 7, f, TL: China, Richthofen Mts.
Colias sifanica herculeana ; [LCG, 137]

Colias aegidii Verhulst, 1990
Gansu, Qinghai. See [About maps] Colias aegidii Verhulst, 1990; Lambillionea 90 (3): 42[?], TL: Gansu, Xhiahe, 3000-3300m
Colias aegidii ; [LCG, 137]

Colias nebulosa Oberthür, 1894
-. See [About maps] Colias nebulosa Oberthür, 1894; Étud. d'Ent. 19 : 8, pl. 8, f. 65, TL: China, Tchang-Kou
Colias nebulosa, NSG Voucher Specimen [Wahlberg]
Colias sifanica nebulosa ;
Sichuan Colias nebulosa nebulosa ; [LCG, 138]
C. n. karoensis Hoshiai & Rose, 1998
S.Tibet nebulosa karoensis Hoshiai & Rose, 1998; Butterflies 20: (58-61, f. 1-8, TL: S.Tibet, Karo La, 4700-5000m, 60km E of Gyangze
Colias nebulosa karoensis ; [LCG, 138]

Larva on Astragalus epineux [LCG, 139]

C. n. niveata Verity, [1909]
Qinghai, ...? nebulosa niveata Verity, [1909]; Rhopalocera Palaearctica 1: 228, pl. 41, f. 28, TL: Kukunor Lake
melanina Bollow, 1930; (repl. melanitica Bang-Haas, 1915)
f. melanitica Bang-Haas, 1915; Dt. ent. Z. Iris 29 (2/3) : 99
= ; [LCG, 139 (form)]
Colias sifanica niveata ;
Colias nebulosa niveata ; [LCG, 139]
[niveata-1mu.jpg] 1024x768 (~111Kb) upperside male China, Qinghai prov, W.Qinghai lake, Nan Shan mts, 3500m, 28.6.2005, Photo © A. Timchenko [niveata-1mv.jpg] 1024x768 (~132Kb) underside male China, Qinghai prov, W.Qinghai lake, Nan Shan mts, 3500m, 28.6.2005, Photo © A. Timchenko [niveata-2m.jpg] 1500x1473 (~374Kb) male China, Qinghai prov, 20 km S Tianjun (3900m), Photo © A. Timchenko leg.

Larva on Astragalus epineux [LCG, 139]


C. n. pugo Evans, 1924
SE.TIbet nebulosa pugo Evans, 1924; J. Bombay nat. Hist. Soc. 29 (4): 972, TL: Pugo, SE.Tibet
Colias nebulosa pugo ; [LCG, 139]
C. n. richthofeni Bang-Haas, 1927
Richthofen Mts., Kukunor Mts., ...? Colias richthofeni Bang-Haas, 1927; Horae Macrolep. Palaearct. 1 : 42, 110, pl. 6, f. 1-2, TL: China, W. Liang-Tchou
Colias richthofeni ; [MRS, 221]
Colias nebulosa richthofeni ; [LCG, 140]
C. n. sungpani Bang-Haas, 1927
Sichuan, Gansu nebulosa sungpani Bang-Haas, 1927; Horae Macrolep. Palaearct. 1 : 109, TL: Sichuan, Songpan
Colias nebulosa sungpani ; [LCG, 140]

Colias cocandica Erschoff, 1874
-. See [About maps] Colias cocandica Erschoff, 1874; in Fedschenko, Travels in Turkestan. 2 (5) : 6, pl. 1, f. 3, TL: Upper reaches of Sokh River near the Stschurowski Glacier, Turestansky Mts.
Colias cocandica, NSG Voucher Specimen [Wahlberg]
Colias cocandica, Extracts from [BRU] by [Ilya Osipov]
Colias cocandica ; [BRU, 170]; [MRS, 223]
Tadzhikistan (Turkestan Range), Kirgizia (Fergana, E.Turkestan Range, Aksu Valley, Alai Mts.) Colias cocandica ab. ♀ galba Grum-Grshimailo, 1893; Horae Soc. ent. Ross. 27 : 381
= ; [LCG, 141 (form)]
Colias eogene ab. hybrida Grum-Grshimailo, 1893; Horae Soc. ent. Ross. 27 : 381
= ; [LCG, 141 (form)]
aurantiacopunctata Verity, [1909]; Rhopalocera Palaearctica 1: 274
= ; [LCG, 141 (form)]
Colias cocandica var. tatarica Bang-Haas, 1915; Dt. ent. Z. Iris 29 (2/3) : 98, TL: Chotan
= ; [LCG, 141 (form)]
Colias cocandica taratarica f. aurantiacomaculata Bang-Haas, 1915; Dt. ent. Z. Iris 29 (2/3) : 98
= ; [LCG, 141 (form)]
Colias cocandica tatarica f. brunneo-viridis Bang-Haas, 1915; Dt. ent. Z. Iris 29 (2/3) : 99, TL: Chotan
= ; [LCG, 141 (form)]
Colias cocandica tatarica f. griseo viridis Bang-Haas, 1915; Dt. ent. Z. Iris 29 (2/3) : 99
= ; [LCG, 141 (form)]
Colias cocandica tatarica f. immaculata Bang-Haas, 1915; Dt. ent. Z. Iris 29 (2/3) : 99
= ; [LCG, 141 (form, = integra Verity, 1911)]
Colias cocandica tatarica f. melanitica Bang-Haas, 1915; Dt. ent. Z. Iris 29 (2/3) : 99, TL: Chotan
= ; [LCG, 141 (form)]
Colias cocandica tatarica f. minor Bang-Haas, 1915; Dt. ent. Z. Iris 29 (2/3) : 99, TL: Chotan
= ; [LCG, 141 (form)]
Colias cocandica tatarica f. circumiens Bang-Haas, 1915; Dt. ent. Z. Iris 29 (2/3) : 99, TL: Chotan
= ; [LCG, 141 (form)]
Colias cocandica tatarica f. impunctatas Bang-Haas, 1915; Dt. ent. Z. Iris 29 (2/3) : 99, TL: Chotan
= ; [LCG, 141 (form)]
Colias cocandica tatarica f. radiata Bang-Haas, 1915; Dt. ent. Z. Iris 29 (2/3) : 99, TL: Chotan
= ; [LCG, 141 (form)]
Colias cocandica tatarica f. viridis Bang-Haas, 1915; Dt. ent. Z. Iris 29 (2/3) : 99, TL: Chotan
= ; [LCG, 141 (form)]
integra Verity, 1911; Rhopalocera Palaearctica 1: 353
= ; [LCG, 141 (form)]
pupillata Stauder, 1924; Mitt. münch. ent. Ges. 14: 64
= ; [LCG, 141 (form)]
xantha Kotzsch, 1936; Ent. Rundsch. 54: 45
= ; [LCG, 141 (form)]
Colias cocandica mraceki Verhulst, 1999; Lambillionea 99 (2): 268, TL: Tadzhikistan, Turkestan Range, Kumbel Pass 2500-3000m
= ; [LCG, 141]
Colias cocandica cocandica ; Winhard, 2000, Butterflies of the world, 10: 10, pl. 12, f. 7-8; [LCG, 140]
C. c. hinducucica Verity, 1911
Afghanistan (Hindu Kush), Tadzhikistan (Pamirs) cocandica hinducucica Verity, 1911; Rhopalocera Palaearctica 1: 353, pl. 70, f. 25-27, TL: E.Hindu Kush, Beik, Pamir
culminicola Kotzsch, 1936; Ent. Rundsch. 54: 45
= ; [LCG, 143 (form)]
Colias cocandica hinducucica ; [LCG, 142]

Larva on Oxytropis sp., Echinospermum sp. [LCG, 142]

C. c. kunjerabi Verhulst, 1999
Pakistan (Karakorum) cocandica kunjerabi Verhulst, 1999; Lambillionea 99 (2): 267, TL: Pakistan, Karakoram Hwy, Kunjerab Pass
Colias cocandica kunjerabi ; [LCG, 143]
C. c. maja Grum-Grshimailo, 1891
Tian-Shan cocandica maja Grum-Grshimailo, 1891; Horae Soc. ent. Ross. 25 (3-4) : 447, TL: China, Borohoro Mts
obscura Austaut, 1898; Le Naturaliste 20: 202
= ; [LCG, 144 (form)]
Colias cocandica maja ; [LCG, 143]
C. c. nastoides Verity, 1911
Alaisky Mts. cocandica nastoides Verity, 1911; Rhopalocera Palaearctica 1: 354, pl. 72, f. 26, TL: Kirgizia, W. Transalai, Aram Kungey
Colias cocandica nastoides ; [LCG, 144]
C. c. pljushtchi Verhulst, 2000
Kirgizia (E.Tian Shan), Kazakhstan (Transili Ala Tau) Colias cocandica pljushtchi Verhulst, 2000; [LCG, 144], TL: Kirgizia, Dolon Pass 3000-3200m

Colias tamerlana Staudinger, 1897
-. See [About maps] Colias tamerlana Staudinger, 1897; Dt. Ent. Z. Iris, 10 (1) : 152, pl. 5, f. 1-3, TL: Xinjiang, E. Tian Shan, N. Chamyl
Colias tamerlana, NSG Voucher Specimen [Wahlberg]
Colias cocandica tamerlana ; [MRS, 223]
Xinjiang, Kirgizia (E.Issyk Kul) Colias tamerlana tamerlana ; [LCG, 145]
C. t. mongola Alphéraky, 1897
Altai, Sayan Colias mongola Alphéraky, 1897; in Romanoff, Mém. Lép. 9 : 188, TL: Ourga?
Colias nastes mongola ;
Colias coccandica mongola ;
Colias tamerlana mongola ; [LCG, 146]
[mongola-1fv.jpg] 468x350 (~29Kb) underside female A tundro-steppe landscapes at Lake Tarkhattinskoe, 2350 m above sea level, S of the junction of Yuzhno-Chuiiskii and Sailyugem mountain ranges, Kosh-Agach District, Altai Republic, West Siberia, Russia. 9th July 1998 (on Oxytropis recognita), Photo © Oleg Kosterin [mongola-1v.jpg] 700x656 (~92Kb) underside Tundrosteppes on the southerm principle slope of the Yuzhno- Chuiskii mountain range E of the Chikty Rivulet, 2400 m above sea level, the upper Dzhazator River basin, Kosh- Agach District, Altai Republic, West Siberia, Russia. 13th July 1998, Photo © Oleg Kosterin

Larva on Oxytropis oligantha [LCG, 147]


C. t. sidonia Weiss, 1968
N.Mongolia Colias cocandica sidonia Weiss, 1968; Sb. faun. Prac. ent. Odd. nat. Mus. Praze 13: 112, TL: N.Mongolia, Rincinlchumbe, 2000m
?sidonia Weiss, 1968; J. Res. Lep. 16: 208
Colias tamerlana sidonia ; [LCG, 147]

Colias tibetana Riley, 1923
Colias cocandica tibetana Riley, 1923; Trans. ent. Soc. Lond. 1922 (3-4) : 465, pl. 36, f. 9, TL: Nyenyam, 13000ft
Colias tibetana ; [LCG]

Colias grumi Alpheraky
-. See [About maps] Colias grumi ; [MRS, 222]
Colias grumi grumi ; [LCG]
C. g. burchana Mrácek & Schulte, 1991
Colias grumi burchana Mrácek & Schulte, 1991; Nachr. ent. Ver. Apollo N.F. 12: (164-165)
Colias grumi burchana ; [MRS, 222]; [LCG]
C. g. aljinshana Huang & Murayama, 1992
Xinjiang Colias alpherakyi aljinshana Huang & Murayama, 1992; Tyô to Ga 43 (1) : 5, f. 11, TL: Aljinshan, 4300m
Colias grumi aljinshana ; Hoshiai, 1993, Tyô to Ga 44 (1): 16; [MRS, 222]; [LCG]
C. g. dvoraki Kocman, 1994
Colias grumi dvoraki ; [LCG]

Colias thrasibulus Fruhstorfer, 1908
Colias thrasibulus ; [LCG]
C. t. zanskarensis Hoshiaei, 1996
Colias thrasibulus zanskarensis ; [LCG]

Colias ladakensis C. & R. Felder, 1865
Himalayas, Kashmir, Tibet. See [About maps] Colias ladakensis C. & R. Felder, 1865; Reise Fregatte Novara, Bd 2 (Abth. 2) (2): 197, pl. 27, f. 8-9
Colias shipkee Moore, 1865; Proc. zool. Soc. Lond. 1865 (2) : 492, pl. 31, f. 13
Colias ladakensis ; [MRS, 223]
Colias ladakensis ladakensis ; [LCG]
C. l. fatulaensis Weiss, 2000
Colias ladakensis fatulaensis ; [LCG]
C. l. seitzi Bollow, 1939
Colias ladakensis seitzi ; [LCG]

Colias nina Fawcett, 1904
Colias ladakensis nina ; [MRS, 223]
Colias nina nina ; [LCG]
C. n. hingstoni Riley, 1923
Colias nina hingstoni ; [LCG]
C. n. tsurpuana Grieshuber, 1996
Colias nina tsurpuana ; [LCG]
[tsurpuana-1afu.jpg] 1500x969 (~195Kb) upperside female China, C.Tibet, Dangxiong vill. (4500m), 11.7.2010, Photo © A. Timchenko leg. [tsurpuana-1afv.jpg] 1400x1256 (~328Kb) underside female China, C.Tibet, Dangxiong vill. (4500m), 11.7.2010, Photo © A. Timchenko leg. [tsurpuana-1amu.jpg] 1500x918 (~234Kb) upperside male China, C.Tibet, Dangxiong vill. (4500m), 11.7.2010, Photo © A. Timchenko leg.
[tsurpuana-1bfu.jpg] 1500x1121 (~334Kb) upperside female China, C.Tibet, Dangxiong vill. (4500m), 11.7.2010, Photo © A. Timchenko leg. [tsurpuana-1bfv.jpg] 1400x1294 (~245Kb) underside female China, C.Tibet, Dangxiong vill. (4500m), 11.7.2010, Photo © A. Timchenko leg. [tsurpuana-1bmu.jpg] 1500x1033 (~238Kb) upperside male China, C.Tibet, Dangxiong vill. (4500m), 11.7.2010, Photo © A. Timchenko leg.


Colias berylla Fawcett, 1904
Sikkim, Tibet. See [About maps] Colias berylla Fawcett, 1904; Proc. zool. Soc. 2: 139
Colias berylla, NSG Voucher Specimen [Wahlberg]
Colias berylla ; [MRS, 221]
Colias berylla berylla ; [LCG]
C. b. bergeriana Verhulst, 1992
Colias berylla bergeriana ; [LCG]

Colias adelaidae Verhulst, 1991
Colias adelaidae ; [LCG]
C. a. karmalana Grieshuber, 1999
Colias adelaidae karmalana ; [LCG]; Huang, 2001

Colias arida Alphéraky, 1889
Tibet, W.China. See [About maps] Colias eogene var. arida Alphéraky, 1889; in Romanoff, Mém. lép. 5 : 76
?Colias eogene ab. auritheme Grum-Grshimailo, 1893; Horae Soc. ent. Ross. 27 : 383
?Colias cocandica irma Evans, 1924; J. Bombay nat. Hist. Soc. 29 (4): 972, TL: SE.Tibet, Loutsang-Po
= ?; [LCG, 134]
= ?Colias adelaidae Verhulst, 1991; [LCG, 134]
?Colias montium irma ;
Colias arida ; [MRS, 221]
Colias arida arida ; [LCG]
C. a. cakana Rose & Schulte, 1992
Colias arida cakana ; [LCG]
C. a. muetingi Rose & Schulte, 1992
Colias arida muetingi ; [LCG]
C. a. wanda Grum-Grshimailo, 1893
Colias arida var. wanda Grum-Grshimailo, 1893; Horae Soc. ent. Ross. 27 : 383
Colias arida wanda ; [MRS, 221]

Colias baeckeri Kotzsch, 1930
Colias baeckeri baeckeri ; [LCG]
C. b. paskoi Kocman, 1999
Colias baeckeri paskoi ; [LCG]

Colias stoliczkana Moore, 1878
Sikkim, Kashmir, Tibet. See [About maps] Colias stoliczkana Moore, 1878; Ann. Mag. nat. Hist. (5) 1 (3) : 229
Colias stoliczkana ; [MRS, 223]
Colias stoliczkana stoliczkana ; [LCG]
C. s. miranda Fruhstorfer, 1903
Colias stoliczkana miranda ; [LCG]
C. s. cathleenae Epstein, 1979
Colias stoliczkana cathleenae ; [LCG]
C. s. yangguifei Huang & Murayama, 1992
Xinjiang Colias chrysotheme yangguifei Huang & Murayama, 1992; Tyô to Ga 43 (1) : 4, f. 7-8, TL: Aqikfu, Aljin, East Kunlun, 4350-4400
Colias stoliczkana yangguifei ; Hoshiai, 1993, Tyô to Ga 44 (1): 15

Colias dubia Elwes, 1906
Sikkim. See [About maps] Colias dubia Elwes, 1906; Proc. zool. Soc. Lond. 1906 : 481, pl. 36, f. 8-9
Colias dubia ; [LCG]

Colias staudingeri Alphéraky, 1881
Turkestan, Kadzhikistan, Uzbekistan. See [About maps] Colias staudingeri Alphéraky, 1881; Horae Soc. ent. Ross. 16 (3-4): 368, TL: upper reaches of Kunghes River, Boro-Khoro Mts, China
Colias staudingeri, NSG Voucher Specimen [Wahlberg]
Colias staudingeri, Extracts from [BRU] by [Ilya Osipov]
Colias staudingeri ; [BRU, 174]; [EBW]
Colias staudingeri staudingeri ; Winhard, 2000, Butterflies of the world, 10: 10, pl. 12, f. 9; [LCG]
C. s. maureri Staudinger, 1901
Alaisky Mts.
C. s. emivittata Verity, 1911
C.Tien-Shan Colias staudingeri emivittata ; [LCG]
C. s. pamira Grum-Grshimailo, 1890
Zaalaisky Mts. Colias staudingeri var. pamira ab. verityi Avinoff, 1910; Horae Soc. Ent. Ross. 39 : 248
Colias staudingeri pamira ; Winhard, 2000, Butterflies of the world, 10: 10, pl. 12, f. 10; [LCG]

Colias felderi Grum-Grshimailo, 1891
Tibet, China. See [About maps] felderi Grum-Grshimailo, 1891; Hor. Soc. ent. Ross. 25: 448
Colias felderi ; [LCG]

Colias eogene C. & R. Felder, 1865
E.Hindukush, Afghanistan, Kashmir, Himalaya. See [About maps] Colias eogene C. & R. Felder, 1865; Reise Fregatte Novara, Bd 2 (Abth. 2) (2): 196, pl. 27, f. 7, TL: "Himalaya"
?Colias eogene ab. cana Grum-Grshimailo, 1890; in Romanoff, Mém. Lép. 4 : 298, 333
?Colias eogene ab. oshanini Avinoff, 1910; Horae Soc. Ent. Ross. 39 : 248, pl. 14, f. 4
Colias eogene, NSG Voucher Specimen [Wahlberg]
Colias eogene ; [BRU, 175]; [EBW]; [BOW: pl. 158, f. 13, 15]; [MRS, 222]

Larva on Astragalus spp. [BRU]

Colias eogene eogene ; [LCG]
C. e. erythas Grum-Grshimailo, 1890
Pamirs Colias eogene var. erythas Grum-Grshimailo, 1890; in Romanoff, Mém. Lép. 4 : 298, 332
Colias eogene erythas ; [LCG]
C. e. elissa Grum-Grshimailo, 1890
Alai Colias eogene var. elissa Grum-Grshimailo, 1890; in Romanoff, Mém. Lép. 4 : 298, 332
Colias eogene elissa ; Winhard, 2000, Butterflies of the world, 10: 10, pl. 12, f. 11-12; [LCG]
C. e. francesca Watkins, 1927
Colias eogene francesca ; [LCG]
C. e. shandura Evans, 1926
Colias eogene shandura ; [LCG]

Colias leechi Grum-Grshimailo, 1893
Colias eogene var. leechi Grum-Grshimailo, 1893; Horae Soc. ent. Ross. 27 : 382
Colias leechi ; [LCG]

Colias regia Grum-Grshimailo, 1887
Alai, Tien-Shan. See [About maps] Colias regia Grum-Grshimailo, 1887; in Romanoff, Mém. Lép., 3: 398, TL: Aram-Kungei
Colias regia, NSG Voucher Specimen [Wahlberg]
Colias regia, Extracts from [BRU] by [Ilya Osipov]
Colias regia ; [BRU, 174]; [LCG]

Colias romanovi Grum-Grshimailo, 1885
Ghissar, Alai, Tian-Shan. See [About maps] Colias romanovi Grum-Grshimailo, 1885; in Romanoff, Mém. Lép. 2 : 229 nota
Colias romanovi Grum-Grshimailo, 1887; in Romanoff, Mém. Lép. 3 : 398
Colias romanovi, NSG Voucher Specimen [Wahlberg]
Colias romanovi, Extracts from [BRU] by [Ilya Osipov]
Colias romanovi ; [BRU, 174]; [EBW]; Winhard, 2000, Butterflies of the world, 10: 10, pl. 12, f. 13
Colias romanovi romanovi ; [LCG]
C. r. seravschana Lukhtanov, 1999
Colias romanovi seravschana Lukhtanov, 1999; Atalanta 30 (1/4): 138, TL: Tadjikistan, Seravschanksi, Dascht, 3200-3500m
Colias romanovi seravschana ; [LCG]

Colias heos (Herbst, 1792)
Altai - S.Siberia, Mongolia - Ussuri, SE.China. See [About maps] heos (Herbst, 1792); in Jablonsky, Naturs. Schmett. 5: 213, pl. 144 [or 114?], f. 5-6, TL: "Siberia"
Papilio aurora Esper, 1783; Die Schmett. Th. I, Bd. 2 (8): 161, pl. 83, f. 3 (preocc.)
Colias aurora ab. obscura Moltrecht, 1909; Ent. Zs. 22 (44) : 185
?chloe Eversmann, 1847; Bull. Soc. Imp. Nat. Moscou 20 (3) : 73, pl. 4, f. 3-4
sibirica Lederer, 1852
kenteana Rühl, 1895
Colias heos, NSG Voucher Specimen [Wahlberg]
Colias heos, Extracts from [BRU] by [Ilya Osipov]
Colias heos ; [BRU, 176]; [MRS, 220]
[heos-1fv.jpg] 549x505 (~53Kb) underside female A steppefied meadow on a high right bank of the Onon River 5 km upstream of the village Nizhnii Tsasuchei, Onon District, S Chita Province (Dahuria), S Transbaikalia, Siberia, Russia. 30th June 1996, Photo © Oleg Kosterin

Larva on Vicia, Astragalus, Trifolium lucanicum [BRU, Kurentzov, 1970, Danchenko]


Colias heos heos ; [LCG]
C. h. alpina Verity, 1911
Altai, Sayan
C. h. semenovi Shtandel, 1960
Colias semenovi Shtandel, 1960
Colias heos semenovi ; Winhard, 2000, Butterflies of the world, 10: 10, pl. 12, f. 14; [LCG]
C. h. vespera Bang-Haas, 1929
Colias heos vespera ; [LCG]

Colias diva Grum-Grshimailo, 1891
Tibet, W.China. See [About maps] diva Grum-Grshimailo, 1891; Hor. Ent. Soc. Ross. 25: 449
Colias diva ; [LCG]

Colias viluiensis Ménétriés, 1859
Transbaikalia - Chukot Peninsula. See [About maps] Colias viluiensis Ménétriés, 1859; in Schrenck, Reise Forschungen Amur-Lande 2 (1) : 18, TL: "Vilui", SE.Siberia
chilkana Austaut, 1898
Colias viluiensis, Extracts from [BRU] by [Ilya Osipov]
Colias viluiensis ; [BRU, 175]
Colias viluiensis viluiensis ; [LCG]
C. v. dahurica Austaut, 1899
Transbaikalia Colias viliuensis dahurica ; [LCG]
C. v. heliophora Churkin & Grieshuber, 2001
Chukotka Colias viluiensis heliophora Churkin & Grieshuber, 2001; Helios, Coll. lep. articles II: (174-189)

Colias erschoffi Alphéraky, 1881
Tien-Shan. See [About maps] Colias erschoffi Alphéraky, 1881; Horae Soc. ent. Ross. 16 (3-4): 362, t. 13, f. 1-2, TL: E. Tian-Shan, Kuldzha
Colias erschoffi ; [BRU, 174]; [LCG]

Larva on Astragalus [BRU]


Colias wiskotti Staudinger, 1882
Turkestan, Uzbekistan, Afghanistan, Kadzikhistan, Kashmir. See [About maps] Colias wiskotti Staudinger, 1882; in Staudinger & Bang-Haas, Berl. ent. Z. 26 (1) : 166, pl. 2, f. 9-10, TL: Alai Mts.
Colias wiskotti, Extracts from [BRU] by [Ilya Osipov]
Colias wiskotti ; [BRU, 177]; [EBW]

Larva on Acantolimon, Oxytropis, Onobrychis echidna [BRU]

C. w. wiskotti Staudinger, 1882
Ghissar (west) leuca Staudinger, 1882
palaenoides Verity, 1909
Colias wiskotti wiskotti ; Winhard, 2000, Butterflies of the world, 10: 10, pl. 12, f. 15-16; [LCG]
C. w. sagina Austaut, 1891
Ghissar (east), Darvaz
C. w. separata Grum-Grshimailo, 1888
Alaisky Mts. Colias wiskotti var. separata Grum-Grshimailo, 1888; Horae Soc. ent. Ross. 22 (3-4) : 305, TL: Montes Alaienses
leucotheme Grum-Grshimailo, 1890
Colias wiskotti var. (ab.?) aurantiaca Staudinger, [1892]; Dt. Ent. Z. Iris 4 (2) : 230
Colias wiskotti separata ; [LCG]
C. w. aurea Kotzsch, 1937
W.Pamirs Colias wiskotti aurea, NSG Voucher Specimen [Wahlberg]
Colias wiskotti aurea ; [LCG]
C. w. chrysoptera Grum-Grshimailo, 1888
E.Pamirs Colias wiskotti var. chrysoptera Grum-Grshimailo, 1888; Horae Soc. ent. Ross. 22 (3-4) : 305, TL: Afghanistan (Roschan); Transalai; Sarykol
Colias wiskotti chrysoptera ; [LCG]
C. w. draconis Grum-Grshimailo, 1891
W.Tian-Shan Colias draconis Grum-Grshimailo, 1891; Horae Soc. ent. Ross. 25 (3-4) : 465, TL: Kara-Kul
Colias draconis, NSG Voucher Specimen [Wahlberg]
Colias draconis, Extracts from [BRU] by [Ilya Osipov]
Colias draconis ; [BRU, 178]
Colias wiskotti draconis ; [LCG]
C. w. sweadneri Clench & Shoumatoff, 1956
Afghanistan Colias wiskotti sweadneri Clench & Shoumatoff, 1956; Vidensk. Medd. Dansk Naturh. Forening 118: (141-192) , TL: btw Surtu and the top of Mt. Shah Fuladi, nr. 5000m
Colias wiskotti sweadneri ; [LCG]

Colias chrysotheme (Esper, 1781)
EEU, Asia Minor, N.Iran, W.Siberia, Altai, S.Russia - Transbaikalia, Mongolia, NE.China. See [About maps] Lesser Clouded Yellow Hellorangegrüner Heufalter
Papilio chrysotheme Esper, 1781; Die Schmett. Th. I, Bd. 2 (3): 89, pl. 65 f. 3-4, TL: "Kremnitz" [Slovakia]
Colias chrysotheme, NSG Voucher Specimen [Wahlberg]
Colias chrysotheme, Extracts from [BRU] by [Ilya Osipov]
Colias chrysotheme ; [BOW: pl. 3, f. 26]; [MRS, 220]; [Otakar Kudrna]

Larva on Vicia hirsuta [H&R], Vicia [BRU, Nekrutenko, 1990]

Colias chrysotheme chrysotheme ; [LCG]
C. c. caspicus Stichel, 1911
Colias chrysotheme caspicus ; [LCG]
C. c. elena P. Gorbunov, 1995
Altai Colias chrysotheme elena, Lepidoptera collection [SZMN]

Colias audre Hemming, 1933
Colias chrysotheme var. sibirica Grum-Grshimailo, 1893; Horae Soc. ent. Ross. 27 : 380 (praeocc.)
?Colias chrysotheme sibirica ab. ♀ malmyzhensis Krulikovsky, 1909; Rev. russ. ent. 9 (3) : 296
Colias chrysotheme audre ; [MRS, 220]
Colias audre ; [LCG]

Colias thisoa Ménétriés, 1832
Turkestan, Uzbekistan, Kadzhikistan - Altai, Tuva, E.Sayan, China. See [About maps] Colias thisoa Ménétriés, 1832; Cat. Rais.: 244, TL: Mt. Shakhdagh, Daghestan
eos Herrich-Schäffer, 1852
Colias thisoa, NSG Voucher Specimen [Wahlberg]
Colias thisoa, Extracts from [BRU] by [Ilya Osipov]
Colias thisoa ; [BRU, 175]
[thisoa-1vf.jpg] 475x450 (~37Kb) underside female A steppe on the Dzhazator River right bank in its upper flow, between the Chikty and Akbul River, 2000 m above sea level, Kosh-Agach District, Altai Republic, West Siberia, Russia. 12th July 1998(on Taraxacum), Photo © Oleg Kosterin [thisoa-1vm.jpg] 640x636 (~78Kb) underside male A steppe on the Dzhazator River right bank in its upper flow, between the Chikty and Akbul River, 2000 m above sea level, Kosh-Agach District, Altai Republic, West Siberia, Russia. 12th July 1998, Photo © Oleg Kosterin

Larva on Astragalus spp. [BRU, Nekrutenko, 1990]


Colias thisoa thisoa ; [LCG]
C. t. strandiana Sheljuzhko, 1935
Caucasus Minor, Armenian Highland Colias thisoa strandiana ; [LCG]
C. t. shakuhensis Sheljuzhko, 1935
Talysh, Kopet-Dagh Colias thisoa shakuhensis ; [LCG]
C. t. aeolides Grum-Grshimailo, 1890
N. and Inner Tian-Shan, Ghissar, Darvaz, Alai Colias thisoa aeolides ; [LCG]
C. t. urumtsiensis Verity, 1909
C.Tian-Shan Colias thisoa urumtsiensis ; [LCG]
C. t. irtyschensis Lukhtanov, 1999
SW.Altai Colias thisoa irtyschensis Lukhtanov, 1999; Atalanta 30 (1/4): 137, TL: Kasakstan, Ust-Kamenogorsk, Altai, Kalba, Tainty
Colias thisoa irtyschensis ; [LCG]
C. t. nikolaevi Korshunov, 1998
Colias thisoa nikolaevi ; [LCG]

Colias fieldii Ménétriés, 1855
S.Iran - India - S.China, Ussuri. See [About maps] Colias fieldii Ménétriés, 1855; Cat. lep. Petersb. 2: 79, 1: pl. 1, f. 5, TL: "Monts Himalaya"
Colias fieldii, NSG Voucher Specimen [Wahlberg]
Colias fieldii, Extracts from [BRU] by [Ilya Osipov]
Colias fieldii ; [BRU, 176]; [MRS, 219]
Colias fieldi ; [EOB, 87]; [EBW]; [BOR, 182]; Huang, 2001
[fieldii-1mu.jpg] 1024x768 (~129Kb) upperside male China, Qinghai prov, ~30 km SW Xining town, 3600m, 6.7.2005, Photo © A. Timchenko [fieldii-1mv.jpg] 1024x768 (~113Kb) underside male China, Qinghai prov, ~30 km SW Xining town, 3600m, 6.7.2005, Photo © A. Timchenko [fieldii-2f.jpg] 1000x931 (~92Kb) female China, S.Tibet, 10 km N. Nyalam town (4300m), 7.7.2010, Photo © A. Timchenko leg.
[fieldii-2mu.jpg] 1400x945 (~305Kb) upperside male China, S.Tibet, 10 km N. Nyalam town (4300m), 7.7.2010, Photo © A. Timchenko leg. [fieldii-2mv.jpg] 1400x1050 (~216Kb) underside male China, S.Tibet, 10 km N. Nyalam town (4300m), 7.7.2010, Photo © A. Timchenko leg.

Larva on Leguminosae [EOB]


Yunnan Colias fieldii fieldii ; [LCG]
C. f. chinensis Verity, 1909
S.Ussuri Colias fieldii chinensis ; [MRS, 219]; [LCG]

fiColias croceus (Geoffroy, 1785)
NAF, Fezzan, Cyrenaica, SEU, CEU, Asia Minor, Iran, W.Siberia. See [About maps] Etelänkeltaperhonen Clouded Yellow Rödgul höfjäril Saffransfjäril Postillon Orangeroter Kleefalter Le Souci Szlaczkoń sylwetnik
Papilio croceus Geoffroy, 1785; in Fourcroy, Ent. Paris. 2: 250, TL: Paris
Papilio edusa Fabricius, 1787; Mantissa Insectorum 2: 23 (preocc. Papilio edusa Fabricius, 1777)
Colias croceus f. helice
Colias croceus f. helicina Oberthür, 1880; Bull. Soc. Ent. Fr.: 145, TL: Italy
Colias croceus f. deserticola Verity, 1909; Rhopalocera Palaearctica 1: 268, TL: Algeria
Colias croceus ab. cremonae Bang-Haas, 1912; Dt. ent. Z. Iris 26 (2): 103, TL: Lebanon
Colias croceus f. mediterranea Stauder, 1913; Boll. Soc. adriat. Sci. nat. 27 (1): 147, pl. 2, f. 5, TL: Italy
Colias edusa f. obsoleta Grüber, 1929; Ent. Z. Frankf. a. M. 43: 21, pl. f. 10
Colias crocea, NSG Voucher Specimen [Wahlberg]
Colias croceus, Larvae of North-European Lepidoptera [Kimmo Silvonen]
Colias croceus, Svenska fjärilar [SVF]
Colias croceus, Norges Sommerfugler [NORLEP]
Colias crocea, Moths and Butterflies of Europe [MBE]
Colias crocea, Extracts from [BRU] by [Ilya Osipov]
Colias crocea, Europaische Schmetterlinge [Mario Maier]
Colias crocea, Captain's European Butterfly Guide [Simon Coombes]
Colias crocea ; [BRU, 179]; [EBW]; [BOW: pl. 3, f. 27-28]; [LCG]
Colias croceus ; [BMAT: 18, pl. 7, f. 15-24]; [AfrL]; [Otakar Kudrna]
[croceus-1.jpg] 500x401 (~57Kb) Clouded yellow Turunc, SW Turkey 14/04/95, Photo © Simon Coombes [croceus-2fv.jpg] 500x690 (~31Kb) underside female France, Sophia Antipolis, 5.10.2000, Photo © Markku Savela [croceus-3vmf.jpg] 600x435 (~41Kb) underside male female Morocco, Djebel Lekst, Anti-Atlas , Photo © Michel Tarrier
[croceus-4v.jpg] 1200x900 (~178Kb) underside Italy, Ravenna, August 2002, Photo © Maurizio Elia

Larva on Trifolium pratense, Medicago, Trifolium [SPRK], Vicia, Lotus, Onobrychis, Astragalus [BRU, Nekrutenko, 1990, Gorbunov, Korshunov, 1995], Colutea arborescens, Medicago sativa, Acanthyllis tragacanthoides, A. numidica, Anthyllis tetraphylla, Erophaca baetica, Hippocrepis sp., Medicago lappacea, M. hispida, M. polymorpha, M. sulcata [BMAT]



Colias electo (Linnaeus, 1763)
-. See [About maps] African Clouded Yellow
Papilio electo Linnaeus, 1763; Amoenitates Acad., 6: 405
Colias xenodice C. & R. Felder, 1859; Wien. ent. Monats. 3 (12): 394
= ; [NHM]
?Colias edusina C. & R. Felder, 1860; Wien. ent. Monats. 4 (4): 100
Colias electo, NSG Voucher Specimen [Wahlberg]
Colias electo ; [EBW]; [AfrL]

Larva on Vicia sativa, Trifolium, Robinia pseudacacia [PBSA]

South Africa, S.Mozambique, Namibia, Rhodesia, Zambia Papilio electra Linnaeus, 1767; Syst. Nat. (Edn 12) 1 (2): 764
= ; [AfrL]
Papilio palaeno Stoll, [1781]; in Cramer, Uitl. Kapellen 4 (29-31): 96
= ; [AfrL]
Colias aurivillius Keferstein, 1883; Verh. zool.-bot. Ges. Wien 32 : 457
= ; [AfrL]
Colias electo electoe ♀ f. overlaeti Berger, 1940; Expl. Parc. nat. Albert Mission G. F. de Witte 30 : 34
= ; [AfrL]
Colias electo ♀ millari Stoneham, 1957; Bull. Stoneham Mus. (70) : [2]
= ; [AfrL]
Colias electo ab. eremna Vári, 1976; Ann. Transv. Mus. 30 (13) : 125, TL: South Africa
= ; [AfrL]
Colias electo electo ♂ f. elysium Kroon, 1985; Ent. Rec. J. Var. 97 (1-2) : 6, TL: Vaal R., Sasolburg, South Africa
= ; [AfrL]
Colias electo electo ab. capensis McLeod & McLeod, 2002; Ent. Rec. J. Var. 114 (5) : 224, TL: Cape
= ; [AfrL]
Colias electo electo ab. inconstantis McLeod, 2004; Ent. Rec. J. Var. 116 (2): 85, TL: Cape
= ; [AfrL]
Colias electo electo ab. memorabilis McLeod, 2004; Ent. Rec. J. Var. 116 (2): 86, TL: Cape
= ; [AfrL]
Colias electo electo ab. pallidula McLeod, 2004; Ent. Rec. J. Var. 116 (2): 86, TL: Cape
= ; [AfrL]
Colias electo electo ; [BAFR, 54]; [LCG]; [AfrL]
C. e. hecate Strecker, 1900
Angola, S.Zaire, N.Zambia, NW.Malawi Colias hecate Strecker, 1900; in Weeks, Proc. New England Zool. Club. 2 : 7, pl. 3, TL: Ovim-Bunda, Congo, West Africa
Colias electo hecate f. katangae Berger, 1940; Expl. Parc. nat. Albert Mission G. F. de Witte 30 : 38
= ; [AfrL]
Colias electo hecate ♀ f. bunda Berger, 1940; Expl. Parc. nat. Albert Mission G. F. de Witte 30 : 38
= ; [AfrL]
Colias electo hecate ♀ f. elisabethae Berger, 1940; Expl. Parc. nat. Albert Mission G. F. de Witte 30 : 39
= ; [AfrL]
Colias hecate ; Strecker, 1900, Lep. Rhopal. Het., Suppl. 3 : 19
Colias electo hecate ; [AfrL]; [BAFR, 54 (text)]
C. e. pseudohecate Berger, 1940
Malawi, Tanzania, Kenya, Uganda, E.Zaire, Rwanda, Burundi, S.Sudan, S.Ethiopia, N.Somalia Colias electo pseudohecate Berger, 1940; Expl. Parc. nat. Albert Mission G. F. de Witte 30 : 40
Colias electo pseudohecate ♀ f. licina Berger, 1940; Expl. Parc. nat. Albert Mission G. F. de Witte 30 : 42
= ; [AfrL]
Colias electo pseudohecate ♀ f. lecerfi Berger, 1940; Expl. Parc. nat. Albert Mission G. F. de Witte 30 : 43
= ; [AfrL]
Colias electe pseudohecate ♀ f. splendens Berger, 1940; Expl. Parc. nat. Albert Mission G. F. de Witte 30 : 43
= ; [AfrL]
Colias electo pseudohecate ab. dormonti Dufrane, 1947; Bull. Ann. Soc. ent. Belg. 83 (1/2) : 72
= ; [AfrL]
Colias electo ♀ f. affricana Stoneham, 1957; Bull. Stoneham Mus. (70) : [2], TL: Kenya
= ; [AfrL]
Colias electo ♀ f. ambreana Stoneham, 1957; Bull. Stoneham Mus. (70) : [3], TL: Kenya
= ; [AfrL]
Colias electo pseudohecate ; [BAFR, 54]; [LCG]; [AfrL]
C. e. meneliki Berger, 1940
C.Ethiopia, N.Ethiopia, Eritrea, S.Arabia, W.Arabia Colias electo meneliki Berger, 1940; Expl. Parc. nat. Albert Mission G. F. de Witte 30 : 44
Colias electo meneliki ♀ f. bafanae Berger, 1940; Expl. Parc. nat. Albert Mission G. F. de Witte 30 : 45
= ; [AfrL]
Colias electo philbyi Berger, 1953; Entomologist 86 : 171, TL: Saudi Arabia
= ; [AfrL]
Colias electo philbyi ♀ f. bafanae Berger, 1953; Entomologist 86 : 172
= ; [AfrL]
Colias electo philbyi ♂ f. pauper Berger, 1953; Entomologist 86 : 172
= ; [AfrL]
Colias electo philbyi ; [BAFR, 54 (text)]; [LCG]
Colias electo meneliki ; [BAFR, 54 (text)]; [LCG]; [AfrL]
Cameroon Colias electo manengoubensis Darge, 1968; Bull. Soc. Ent. Mulhouse 1968 : 15, TL: Cameroon
Colias electo manengoubensis ; [LCG]; [AfrL]

Colias mukana Berger, 1981
-. See [About maps] Colias electo mukana Berger, 1981; Les Papillons du Zaire: 81
Colias electo mukana ; [LCG]
Colias mukana ; [AfrL]
Colias mukana mukana ; [AfrL]
C. m. jolyi Verhulst, 2006
Malawi Colias mukana jolyi Verhulst, 2006; Lambillionea 106 (2 II) : 288, TL: Malawi
Colias mukana jolyi ; [AfrL]

Colias ponteni Wallengren, 1860
Chile. See [About maps] Colias ponteni Wallengren, 1860; Wien ent. Monats. 4: 33, TL: Honolulu
Colias imperialis Butler, 1871; Proc. zool. Soc. Lond. 1871 : 250, pl. 19, f. 2, TL: [Chile]
= ; Shapiro, 1991, J. Res. Lepid. 28 (3): 180; [NL4A, #59]
Colias imperialis ; Staudinger, 1899, Naturhist. Mus. Hamburg 2 (6) : 27
Colias ponteni ; Shapiro, 1991, J. Res. Lepid. 28 (3): 180; [LCG]; [NL4A, #59]

Colias lesbia (Fabricius, 1775)
S.Brazil. See [About maps] Papilio lesbia Fabricius, 1775; , TL: "Patagoniae littoribus"
Colias lesbia, NSG Voucher Specimen [Wahlberg]
Colias lesbia ; [EBW]; [BOW: pl. 49, f. 18]; [NL4A, #57]
Tierra del Fuego, Argentina, Brazil(Rio Grande do Sul), Uruguay Colotis pyrrhothea Hübner, 1823; , TL: Argentina
= ; [NL4A, #57a]
Colias pyrrhothea var. heliceoides Capronnier, 1874; , TL: Argentina
= ; [NL4A, #57a]
Colias aquilo Austaut, 1912; Int. Ent. Zs 6 (13) : 88, f. 3, TL: "Lappland"
= ; [NL4A, #57a]
f. obscura Köhler, 1923; , TL: Argentina
= ; [NL4A, #57a]
f. micans Köhler, 1923; , TL: Argentina
= ; [NL4A, #57a]
f. flaveola Köhler, 1923; (preocc. Blanchard, 1852), TL: Argentina
= ; [NL4A, #57a]
f. maculata Köhler, 1923; , TL: Argentina
= ; [NL4A, #57a]
Colias lesbia eurinice Martin, [1923]; (nom. nud.), TL: Argentina
= ; [NL4A, #57a]
ab. rufa Köhler, 1928; (nom. nud.), TL: Argentina
= ; [NL4A, #57a]
ab. citrina Breyer, 1930; , TL: Argentina
= ; [NL4A, #57a]
ab. privata Breyer, 1939; , TL: Argentina
= ; [NL4A, #57a]
ab. angustimargo Breyer, 1939; , TL: Argentina
= ; [NL4A, #57a]
ab. koehleri Breyer, 1939; , TL: Argentina
= ; [NL4A, #57a]
ab. nigromarginata Breyer, 1939; , TL: Argentina
= ; [NL4A, #57a]
ab. punctata Breyer, 1939; , TL: Argentina
= ; [NL4A, #57a]
ab. citronigra Breyer, 1939; , TL: Argentina
= ; [NL4A, #57a]
Colias lesbia pyrrhothea ab. almeidai Biezanko, 1949; , TL: Brazil(Rio Grande do Sul)
= ; [NL4A, #57a]
Colias lesbia pyrrhothea f. heliceoides ab. lutzi Biezanko, 1949; , TL: Brazil(Rio Grande do Sul)
= ; [NL4A, #57a]
Colias lesbia pyrrhothea f. heliceoides ab. travassosi Biezanko, 1949; , TL: Brazil(Rio Grande do Sul)
= ; [NL4A, #57a]
Colias lesbia pyrrhothea ab. strandi Biezanko, 1949; , TL: Brazil(Rio Grande do Sul)
= ; [NL4A, #57a]
Colias lesbia pyrrhothea ab. costa-limai Biezanko, 1949; , TL: Brazil(Rio Grande do Sul)
= ; [NL4A, #57a]
Colias lesbia pyrrhothea f. heliceoides ab. schweizeri Biezanko, 1949; , TL: Brazil(Rio Grande do Sul)
= ; [NL4A, #57a]
Colias lesbia pyrrhothea ab. williamsi Biezanko, 1949; , TL: Brazil(Rio Grande do Sul)
= ; [NL4A, #57a]
Colias lesbia pyrrhothea ab. forbesi Biezanko, 1949; , TL: Uruguay
= ; [NL4A, #57a]
Colias lesbia pyrrothea[sic?] ; [LCG]
Colias lesbia lesbia ; [LCG]; [NL4A, #57a]
C. l. andina Staudinger, 1894
Bolivia Colias lesbia var. andina Staudinger, 1894; Dt. Ent. Z. Iris 7 (1) : 65, TL: Bolivia
Colias lesbia andina ; [LCG]; [NL4A, #57b]
C. l. dinora Kirby, 1881
Ecuador, Colombia Colias dinora Kirby, 1881; Trans. ent. Soc. Lond. 1881 (3) : 358, TL: Chimborazo, Ecuador
Colias lesbia meieri Berger, 1983; , TL: Ibarra, Baños, Ecuador
= ; [EcuL]; [NL4A, #57c]
Colias lesbia roberti Salazar, 2002; , TL: Colombia
= ; [NL4A, #57c]
Colias lesbia meieri ; Winhard, 2000, Butterflies of the world, 10: 11, pl. 12, f. 19; [LCG]
Colias dinora ; [BOW: pl. 49, f. 15]
Colias lesbia dinora ; [EcuL]; [NL4A, #57c]
C. l. mineira Zikán, 1940
Brazil(Minas Gerais) lesbia mineira Zikán, 1940; , TL: Brazil, Minas Gerais
Colias lesbia mineira ; [NL4A, #57d]
C. l. vauthierii Guérin-Méneville, [1830]
Chile, Argentina lesbia vauthierii Guérin-Méneville, [1830]; , TL: Chile
Colias rutilans Boisduval, 1836; , TL: Chile
= ; [NL4A, #57e]
Colias cunninghami Butler, 1881; , TL: Chile
= ; [NL4A, #57e]
Colias minuscula Butler, 1881; , TL: Chile
= ; [NL4A, #57e]
Colias lesbia var. (ab.)? antarctica Staudinger, 1899; Naturhist. Mus. Hamburg 2 (6) : 23, TL: Argentina
= ; [NL4A, #57e]
Colias lesbia var.(?) arena Staudinger, 1899; Naturhist. Mus. Hamburg 2 (6) : 23, TL: Chile
= ; [NL4A, #57e]
Colias vautieri[sic] ab. micans Breyer, 1939; , TL: Argentina
= ; [NL4A, #57e]
Colias vauthierii ljungneri Bryk, 1944; , TL: Argentina
= ; [NL4A, #57e]
Colias antarctica ; [BOW: pl. 49, f. 12]
Colias vautieri[sic, recte vauthierii] ; [BOW: pl. 49, f. 19]
Colias vautieri vautieri[sic, recte vauthierii] ; Winhard, 2000, Butterflies of the world, 10: 11, pl. 12, f. 23-24; [LCG]
Colias vautieri cunninghami ; [LCG]
Colias vauthierii ; Shapiro, 1991, J. Res. Lepid. 28 (3): 180
Colias lesbia vauthierii ; [NL4A, #57e]
C. l. verhulsti Berger, 1983
Peru Colias verhulsti Berger, 1983; , TL: Peru, Cuzco 2800-3400m
Colias verhulsti f. aurantiaca Berger, 1983; , TL: Peru
= ; [NL4A, #57f]
Colias verhulsti ; Winhard, 2000, Butterflies of the world, 10: 11, pl. 12, f. 20; [LCG]
Colias lesbia verhulsti ; [NL4A, #57f]

Colias euxanthe C. & R. Felder, 1865
-. See [About maps] Colias euxanthe C. & R. Felder, 1865; Reise Fregatte Novara, Bd 2 (Abth. 2) (2): 196, TL: Peru
Colias euxanthe, NSG Voucher Specimen [Wahlberg]
Colias euxanthe ; [NL4A, #54]
Peru Colias euxanthe euxanthe ; Winhard, 2000, Butterflies of the world, 10: 11, pl. 12, f. 21; [LCG]; [NL4A, #54a]
C. e. alticola Godman & Salvin, 1891
Ecuador euxanthe alticola Godman & Salvin, 1891; , TL: Pichincha; Tortorillas, Chimborazo; Cayambe; Chimborazo W side; Antisana W side
Colias euxanthe alticola ; [NL4A, #54b]; [EcuL]
C. e. hermina (Butler, 1871)
Peru, Bolivia Scalidoneura hermina Butler, 1871; Proc. zool. Soc. Lond. 1871 : 251, pl. 19, f. 5, TL: Peru
Colias lesbia f. puna Fruhstorfer, 1907; Stettin ent. Ztg 68 (2): 284, TL: Peru
= ; [NL4A, #54c]
Colias flaveola ab. saturata Röber, 1909; , TL: Bolivia
= ; [NL4A, #54c]
Colias flaveola ab. alba Röber, 1909; , TL: [Bolivia]
= ; [NL4A, #54c]
Colias euxanthe ab. dimorpha Hemming, 1925; , TL: Peru
= ; [NL4A, #54c]
Colias euxanthe euxanthe f. gynomorpha Berger, 1981; , TL: Bolivia
= ; [NL4A, #54c]
Colias euxanthe euxanthe f. lutea Berger, 1981; , TL: Bolivia
= ; [NL4A, #54c]
Colias euxanthe hermina ; [NL4A, #54c]
C. e. stuebeli Reissinger, 1972
Peru Colias euxanthe stuebeli Reissinger, 1972; Atalanta 4: (60-64), TL: NW Peru, Cajamarca, Cutervo
Colias euxanthe coeneni Berger, 1983; , TL: Peru
= ; [NL4A, #54d]
Colias euxanthe coeneni ; [LCG]
Colias euxanthe stuebeli ; [EcuL]; [LCG]; [NL4A, #54d]

Colias flaveola Blanchard, 1852
-. See [About maps] Colias flaveola Blanchard, 1852; in Gay, Hist. Chile, (Zool.) 7 : 19, pl. 1, f. 6a-b, TL: Chile
Colias flaveola ; [BOW: pl. 49, f. 17]; [LCG]; Shapiro, 1991, J. Res. Lepid. 28 (3): 180; [NL4A, #55]
Chile Colias flaveola flaveola ; [NL4A, #55a]
C. f. weberbaueri Strand, 1912
Peru, Bolivia Colias weberbaueri Strand, 1912; Archiv Naturg. 78 A (9) : 185, TL: Peru
Colias euxanthe alticola ab. virescens Rothschild, 1913; Novit. Zool. 20 (1): (187-188) , TL: Peru
= ; [NL4A, #55f]
Colias euxanthe nigerrima Fassl, 1915; Dt. Ent. Z. Iris 29 (2/3) : 180, pl. 6, f. 8a-b, TL: Bolivia,Peru
= ; [NL4A, #55f]
Colias weberbaueri ; Strand, 1916, Lepidoptera Niepeltiana (2) : 14, pl. 16, f. 5; [LCG]
Colias weberbaueri weberbaueri ; [LCG]
Colias flaveola weberbaueri ; [NL4A, #55f]
C. f. mossi Rothschild, 1913
Peru flaveola mossi Rothschild, 1913; , TL: Peru
Colias mossi Rothschild, 1913; Novit. Zool. 20 (1): (187-188) , TL: Peru
Colias mossi ; [LCG]
Colias flaveola mossi ; [NL4A, #55e]
C. f. blameyi Jörgensen, 1916
Argentina Colias blameyi Jörgensen, 1916; , TL: Argentina
Colias blameyi ; Shapiro, 1991, J. Res. Lepid. 28 (3): 182
Colias weberbaueri blameyi ; [LCG]
Colias flaveola blameyi ; [NL4A, #55b]
C. f. mendozina Breyer, 1939
Argentina Colias mendozina Breyer, 1939; , TL: Argentina
Colias weberbaueri mendozina ; [LCG]
Colias flaveola mendozina ; [NL4A, #55d]
C. f. erika Lamas, 1981
Peru flaveola erika Lamas, 1981; , TL: Peru
Colias weberbaueri erika ; [LCG]
Colias flaveola erika ; [NL4A, #55c]

Colias dimera Doubleday, 1847
Venezuela, Colombia, Peru, Ecuador. See [About maps] Colias dimera Doubleday, 1847; , TL: [Colombia]
Colias erythrogrammus Kollar, 1850; , TL: Nova Granada
= ; [EcuL]; [NL4A, #52]
Colias erythrogramma Kollar, 1850
ab. semperi Strecker, 1873; , TL: Colombia
= ; [NL4A, #52]
ab. meridensis Neuburger, 1905; , TL: Venezuela
= ; [NL4A, #52]
Colias dimera mariae Apolinar, 1914; , TL: Colombia
= ; [NL4A, #52]
var. fassli Apolinar, 1914; , TL: Colombia
= ; [NL4A, #52]
Colias tolima Fassl, 1915; Dt. Ent. Z. Iris 29 (2/3) : 180, pl. 6, f. 9a-b, TL: Colombia
= ; [NL4A, #52]
f. andromorpha Fernández, 1928; , TL: Colombia
= ; [NL4A, #52]
f. superba Berger, 1937; , TL: Colombia
= ; [NL4A, #52]
f. alba Berger, 1937; , TL: Colombia
= ; [NL4A, #52]
Colias minuscula f. peruviensis Dufrane, 1947; , TL: "Peru"
= ; [NL4A, #52]
Colias dimera, NSG Voucher Specimen [Wahlberg]
Colias erythrorammus ; Kollar, 1850, (missp.)
Colias dimera ; [EBW]; [BOW: pl. 49, f. 14, 16]; Winhard, 2000, Butterflies of the world, 10: 11, pl. 12, f. 22; [LCG]; [EcuL]; [NL4A, #52]

The ??? species-group

Colias hofmannorum Eckweiler, 2000
Iran. See [About maps] Colias hofmannorum Eckweiler, 2000; , TL: Iran, Khorasan, Birjand, 20km N Sar Bisheh, 2200-2500m
Colias hofmannorum ; Winhard, 2000, Butterflies of the world, 10: 10, pl. 12, f. 17 (holotype), 18 (paratype)


Unknown or unplaced taxa

Colias dinora goryna Martin, [1923]; (nom. nud.), TL: Peru
dinora goryna ; [NL4A, 103]

Colias elwesii Butler, 1881; Ann. Mag. Nat. Hist. (5) 7 (38) : 135, TL: Japan

Colias subaurata Butler, 1881; Ann. Mag. Nat. Hist. (5) 7 (38) : 138, TL: Japan

Colias hyale poliographus ab. daisetsuzana Matsumura, 1928; Insecta Matsumurana 2 (4) : 198, TL: Hokkaido

Colias poliographus ab. ♀ hera Grum-Grshimailo, 1893; Horae Soc. ent. Ross. 27 : 380

Colias chrysotheme ab. ♀ standfussi Grum-Grshimailo, 1893; Horae Soc. ent. Ross. 27 : 380

Colias cocandica ab. hybrida Grum-Grshimailo, 1893; Horae Soc. ent. Ross. 27 : 380

Colias edusa var. pyrenaica Grum-Grshimailo, 1893; Horae Soc. ent. Ross. 27 : 383


9.7.2013 (83)

References:
[A. Timchenko]
Andrew Timchenko; Beijing, China (ex-Moscow, Russia)
[AJA]
African Journal Archive
[ALBERTA]
University of Alberta, E.H. Strickland Entomological Museum; Browse Entomology Collection
[About maps]
Warning! The maps are automatically generated from the textual information, and the process does not always produce acceptable result; See about maps for more info.
[AfrL]
Mark C. Williams; Checklist of Afrotropical Papilionoidea and Hesperoidea; Compiled by Mark C. Williams, 7th ed. (2008) (April 2007); Afrototropical Butterflies; e-mail: Mark C. Williams
[BAFR]
D'Abrera, 1980; Butterflies of the Afrotropical Region
[BAMONA]
Opler, Pavulaan, Stanford & Pogue; Butterflies and Moths of North America; Mountain Prairie Information Node
[BCA]
electronic; BIOLOGIA CENTRALI-AMERICANA
[BHL]
The; Biodiversity Heritage Library; and on; Internet Archive; Note: The linking to the content of Internet Archive is somewhat delicate -- notify me when links break!
[BMAT]
Tennent, 1996; The butterflies of Morocco, Algeria and Tunisia; ISBN 0 906802 05 9
[BOA]
Interactive Listing of American Butterflies; Listado Interactivo de la Mariposas Americanas; Butterflies of America
[BOR]
D'Abrera, 1982, 1985, 1986; Butterflies of the Oriental Region, Part I-III
[BOW]
Lewis, 1974; Butterflies of the World; ISBN 0 245 52097 X
[BRU]
Tuzov, Bogdanov, Devyatkin, Kaabak, Korolev, Murzin, Samodurov, Tarasov, 1997; Guide to the Butterflies of Russia and adjacent territories; Hesperiidae, Papilionidae, Pieridae, Satyridae; Volume 1
[Bruce Walsh]
Bruce Walsh; e-mail: Bruce Walsh; The Moths of Southeastern Arizona; Butterflies of Southeastern Arizona
[CiNii]
http://ci.nii.ac.jp/
[D. Smirnov]
Russia, Moscow; e-mail: D. Smirnov
[EBW]
Smart, Paul, 1976; The Illustrated Encyclopedia of the Butterfly World; ISBN 0 600 31381 6
[EOB]
Feltwell; The Encyclopedia of Butterflies; Prentice Hall, 1993; ISBN 0-671-86828-4
[ESP]
Heppner, 1981; The dates of E. J. C. Esper's Die Schmetterlinge in Abblidungen... 1776-[1830]; Archives of Natural History (1981) 10 (2): 251-254
[EcuL]
Tommaso & Racheli, 2001; Fragmenta entomologica 33 (2): 213-380; An annotated list of Ecuadorian Butterflies (Lepidoptera: Papilionidae, Pieridae, Nymphalidae)
[Erich Mangl]
http://www.mangl.at
[H&R]
Higgins, L.G, Riley, N.D, 1970; A Field Guide to the Butterflies of Britain and Europe
[HBN]
Francis Hemming, 1937; HÜBNER, A bibliographical and systematic account of the entomological works of Jacob Hübner and of the supplements thereto by Carl Geyer, Gottfried Franz von Frölich and Gottlieb August Wilhelm Herrich-Schäffer, Vol 1 & 2
[Heiner Ziegler]
Switzerland / Schweiz; www.pieris.ch
[ILMB]
Interactive Listing of Mexican Butterflies
[Ilya Osipov]
The insects from former USSR for collectors Web site; e-mail: Ilya Osipov; http://www.osipov.org/insects/
[JLS]
Journal of the Lepidopterists' Society; the original J. Lep. Soc. articles
[JRL]
The Journal of Research on the Lepidoptera; JRL Archive
[Kimmo Silvonen]
Espoo, Finland; Kimmo's Lep Site
[LAMAS-2005]
Gerardo Lamas; Entomofauna 26 (6): 57-100; A Bibliography of the Zoological Publications of Hans Fruhstorfer (1886* - 1922+)
[LAMAS-NEO]
Gerardo Lamas; Bibliography of butterflies; An Annocated Bibliography of the Neotropical Butterflies and Skipper (Lepididoptera: Papilionoidea and Hesperioidea); Revised Electronic Edition (2007)
[LANDESMUSEUM]
Oberösterreichisches Landesmuseum; Digitale Literatur
[LCG]
Verhulst, 2000; Les Colias du Globe, 1 & 2: (1) 262pp (Texte), (2) pl. 1-183; Monograph of the genus Colias
[MBE]
Moths and Butteflies of Europe; Paolo Mazzei, Diego Reggianti and Ilaria Pimpinelli; http://www.leps.it/
[MRS]
Chou Io (Ed.); Monographia Rhopalocerum Sinensium, 1-2
[Mario Maier]
Europäische Schmetterlinge; e-mail: Mario Maier
[Maurizio Elia]
Ravenna, Italy; e-mail: Maurizio Elia
[Michel Tarrier]
Michel R. Tarrier; Malaga, Spain; http://www.tarrier.org
[NAB]
Pyle, 1981; Field Guide to North American Butterflies, National Audubon Society; ISBN 0-394-51914-0
[NACL]
Hodges, 1983; Check List of the Lepidoptera of America North of Mexico
[NHM]
Natural History Museum; The Global Lepidoptera Names Index
[NL4A]
Gerardo Lamas, 2004; Atlas of Neotropical Lepidoptera; Checklist: Part 4A; Hesperioidea - Papilionoidea
[NORLEP]
Norges Sommerfugler
[Oleg Kosterin]
Institue of Cytology & Genetics; Siberian Division of Russian Academy of Sciences; Novosibirsk; e-mail: Oleg Kosterin; http://pisum.bionet.nsc.ru/kosterin/
[Opler]
Opler & Warren, 2003 (April); Butterflies of North America. 2. Scientific Names List for Butterfly Species of North America, north of Mexico.
[PBSA]
Dickson, C.G.C (ed.), Kroon, D.M.; Pennington's Butterflies of Southern Africa; AD. DONKER 1978
[Pertti Pakkanen]
e-mail: Pertti Pakkanen; Butterfly and moth observations from Finland
[Peter Payzant]
Canada; http://www.pbase.com/ppayzant/profile
[S. Shuichi Haupt]
e-mail: S. Shuichi Haupt; http://www.tu-berlin.de/fb7/ioeb/biologie/stefan.htm
[SPRK]
Seppänen, E. J, 1970; Suomen suurperhostoukkien ravintokasvit, Animalia Fennica 14
[SVF]
Svenska fjärilar; Naturhistoriska riksmuseet; (Sweden)
[SZMN]
Lepidoptera; Siberian Zoological Museum
[Simon Coombes]
1 Park Street, Plymouth, PL3 4BL, UK; 01752 607854; e-mail: Simon Coombes; http://www.butterfly-guide.co.uk
[Tero Piirainen]
Tampere, Finland; e-mail: Tero Piirainen
[Vitaly Charny]
Birmingham, Alabama, USA
[Wahlberg]
Niklas Wahlberg; The NSG's voucher specimen database; Nymphalidae Systematics Group
[²]
This may require parentheses or not. I don't have the necessary information for this taxon.


Some related literature:
Abadjiev, 2002
Types of Balkan butterflies in the collection of The Natural History Museum, London Neue Ent. Nachr. 23: 3-53
Alphéraky, 1881
Lépidoptères du district de Kouldjà et des montagnes environnantes. (1-3) Horae Soc. ent. Ross. 16 (3-4): 334-435, pl. 14-15 (1881), 17 (1-2): 15-103, pl. 1-3 (1882), 17 (3-4): 156-227, pl. 8-9 (1883)
Alphéraky, 1889
Lépidoptères rapportés du Thibet par le Général N. M. Przewalsky de son voyage de 1884-1885 in Romanoff, Mém. lép. 5 : 59-89, pl. 4
Alphéraky, 1897
Mémoire sur différents Lépidoptéres, tant nouveaux que peu connus, de la faune paléarctique in Romanoff, Mém. Lép. 9 : 185-227
Aurivillius, 1890
Gölands Insektfauna. I. Lepidoptera. Hymenoptera Bihang K. svenska VetensAkad. Handl. 15 (4) (1) : 1-33, pl. 1-3
Austaut, 1905
Notice sur quelques lépidoptères nouveaux Ent. Zs. 18 (36) : 143
Austaut, 1912
Lépidoptères asiatiques nouveaux Int. Ent. Zs 6 (13) : 87-89
Avinoff, 1910
[Contribution à la faune des Rhopaloceres du Pamir] (in Russian) Horae Soc. Ent. Ross. 39 : 225-246, pl. 14
Avinoff, 1910
[New forms of Rhopalocera from Fergana] (in Russian) Horae Soc. Ent. Ross. 39 : 247-250, pl. 14
Avinoff, 1935
Lepidoptera collected by G. M. Sutton on Southampton Island. Rhopalocera in Holland & Avinoff, Mem. Carn. Mus. 12 (2): 13...?
Bang-Haas, 1912
Neue oder wenig bekannte palaearktische Makrolepidopteren. (IV-VI) Dt. ent. Z. Iris 26 (2): 103-110 (1912), (3): 139-162 (1912), (4): 229-230, pl. 6 (1913)
Bang-Haas, 1915
Rhopalocera der Chotan-Ausbeite 1914 Dt. ent. Z. Iris 29 (2/3) : 92-100
Bang-Haas, 1915
Einige seltene Pieriden-Aberrationen Dt. ent. Z. Iris 29 (4) : 193-194 (1916)
Bang-Haas, 1927
Horae Macrolepidopterologicae Regionis Palaearcticae Horae Macrolep. Palaearct. 1 : 1-128, pl. 1-11
O. Bang-Haas, 1934
Neubeschreibungen und Berichtigungen der Palaearktischenk Macrolepidopterenfauna. VIII, XIII, XIV-XVII Ent. Z. 48 : 7-8, (14): 108-110, (16): 128, : 135, : 152 (1935), : 183-184 (1935)
Barnes, 1897
Some new species and varieties of Lepidoptera from the Western U. S. Can. Ent. 29 (2) : 39-42
Barnes & Benjamin, 1926
Notes on diurnal Lepidoptera, with addition and corrections to the recent "List of Diurnal Lepidoptera" Bull. South. Calif. Acad. Sci. 25 (3): 88-98
Bean, 1895
A comparison of Colias hecla with Colias meadii and Colias elis Psyche 7 : 219-229
Berger, 1940
Lepidoptera-Rhopalocera. Exploration du Parc National Albert. Mission G.F. de Witte (1933-1935) Expl. Parc. nat. Albert Mission G. F. de Witte 30 : 1-55
Berger, 1945
Une nouvelle sous-espèce de Colias interior Scud. Lambillionea 45 (6-8): 42
Berger, 1953
New forms of Colias (Lep. Pieridae) collected in Arabia by Major H. St. J.B. Philby, C.I.E Entomologist 86 : 171-172
Blom, 1979
Descriptions of new butterflies form Iran Ent. Ber. Amst. 39 (1): 10-14
Boisduval, 1832
Icones historique des Lépidoptères nouveaux ou peu connus. Collection, avec Figures coloritées, des Papillons d'Europe nouvellement découverts; ouvrage formant le complément de tous les auteurs iconographes Icon. hist. Lépid. Europe 1 (1-2): 1-24, pl. 1-4 (1832), (3-4): 25-40, pl. 5-8 (1832), (5-6): 41-56, pl. 9-12 (1832), (7-8): 57-72, pl. 13-16 (1832), (9-10): 73-104, pl. 17-20 (1833), (11-12): 105-120, pl. 21-24 (1833), (13-14): 121-144, pl. 25-28 (1833), (15-16): 145-160, pl. 29-32 (1833), (17-18): 161-184, pl. 33-36 (1833), (19-20): 185-208, pl. 37-40 (1834), (21-22): 209-224, pl. 41-44 (1834), (23-24): 225-248, pl. 45-47 (1834), (25-28): 249-252 (1834)
Boisduval, 1840
Genera et Index Methodicus Europaeorum Lepidopterorum Genera Index Eur. Lepid.: 238pp : 1-238
Boisduval, 1852
Lépidoptères de la Californie Ann. Soc. ent. Fr. (2) 10 : 275-324
Boisduval & Le Conte, [1829]
Histoire générale et iconographie des lépidoptères et des chenilles de l'Amérique septentrionale Hist. Lép. Am. Sept. (1): 1-16. pl. 1-3 (1829), (2): 17-24, pl. 4-6 (1829), (3/4): 25-40, pl. 7-12 ([1830]), (5/6): 41-56, pl. 13-18 ([1830]), (7/8): 57-80, pl. 19-24 ([1830]), (9/10): 81-100, pl. 25-30 (1833), (11-22): 101-196, pl. 31-65 ([1835]), (23-26): 197-228, pl. 66-78 ([1837])
Bryk, 1942
Zur Kenntnis der Großschmetterlinge der Kurilen Dt. Ent. Z. Iris 56 : 1-90, pl. 1-2
Butler, 1866
A list of the diurnal Lepidoptera recently collected by Mr. Whitely in Hakodadi (North Japan) J. Linn. Soc. Zool. Lond. 9 (34) : 50-58
Butler, 1871
Descriptions of some new Species and a new Genus of Pierinae, with a Monographic Lists of the Species of Ixias Proc. zool. Soc. Lond. 1871 : 250-254, pl. 19
Butler, 1881
On a Collection of Butterflies from Nikko, Central Japan Ann. Mag. Nat. Hist. (5) 7 (38) : 132-140
Böber, 1812
Contribution de la description de quelques nouvelles espèces de papillons découverts en Sibérie ... Mém. Soc. Imp. Nat. Moscou 3: 20-21,?72
Caradja, 1893
Colias Palaeno L. var. Caflischi Soc. Ent. 8 : 26-27
Le Charles, 1927
Descriptions de formes nouvelles de Lépidoptères Bull. Soc. ent. Fr. 1927 (8) : 137-139, pl. 2
Chermock, 1927
New forms of Lepidoptera Bull. Brooklyn ent. Soc. 22 (2): 118-119
Chermock, 1929
Notes on North American Lepidoptera Bull. Brooklyn ent. Soc. 24 (1): 20-21
Christoph, 1888
Diagnosen zu einigen neuen Lepidopteren des palaearktischen Faunengebietes Horae Soc. ent. Ross. 22 (3-4) : 308-314
Christoph, 1889
Lepidoptera aus dem Achal-Tekke-Gebiete. Vierter Theil in Romanoff, Mém. Lépid. 5 : 1-58, pl. 1-3
Churkin & Grieshuber, 2001
Taxonomic notes on Colias Fabricius, 1807 from North Russia with description of new subspecies Helios, Coll. lep. articles II: 174-189
Churkin, Grieshuber, Bogdanov & Zamolodchikov, 2001
Taxonomic notes on Colias tyche Böber, 1812 and Colias nastes Boisduval, 1832 from Russian Far East with the description of new subscpecies Helios, Coll. lep. articles II: 103-115
Churkin, Grieshuber, Bogdanov & Zamolodchikov, 2001
Ein Ersatzname für eine kürzlich beschriebende Unterart von Colias tyche Böber, 1812 Atalanta 32 (1/2): 262-263
Clench & Shoumatoff, 1956
Lepidoptera Rhopalocera (Insecta) from Afghanistan. The 3rd Danish Expedition to Central Asia (Zoological Results 21) Vidensk. Medd. Dansk Naturh. Forening 118 : 141-192, 23 fgs
Cockerell, 1889
On the variation of insects Entomologist 22 : 1-6, 26-29, 54-56, 98-100, 125-130, 147-149, 176-178, 198-202, 226-229, 243-245
Cockle, 1910
Notes on a few butterflies found at Kaslo and in northern British Columbia Can. Ent. 42 (6) : 203-204
Curtis & Ferris, 1985
A review of Colias meadii W.H. Edwards with a description of a new subspecies from Idaho (Pieridae: Coliadinae) Bull. Allyn Mus. 91 (2): 1-9
Darge, 1968
Description d'une forme nouvelle de Colias electo L.: Colias electo mamengoubensis subsp. nov. Bull. Soc. Ent. Mulhouse 1968 : 15-16
Dufrane, 1947
Pieridae Bull. Ann. Soc. ent. Belg. 83 (1/2) : 46-73
Dyar, 1903
A List of North American Lepidoptera and Key to the Literature of this Order of Insects Bull. U.S. nat. Mus. 52 : 1-723
Edwards, 1863
Descriptions of certain species of Diurnal Lepidoptera, found within the limits of the United States and British America. (No. 1-3) Proc. ent. Soc. Philad. 2 (1): 14-22, pl. 1, 11 (1863), (2): 78-82 (1863), (4): 501-507 (1864)
Edwards, 1866
Descriptions of certain species of Diurnal Lepidoptera found within the limits of the United States and British America. No. 5 Proc. ent. Soc. Philad. 6 (4) : 200-208
Behr in Edwards, 1870; Edwards, 1869
The Butteflies of North America. Vol. I. Butts. N. Amer. 1: 218pp, 50pls
Edwards, 1870
Descriptions of new species of diurnal Lepidoptera found within the United States Trans. amer. ent. Soc. 3 (1) : 10-22
Edwards, 1871
Descriptions of new species of North American butterflies Trans. amer. ent. Soc. 3 : 266-277
Edwards, 1872
Descriptions of new species of diurnal Lepidoptera found within the United States Trans. amer. ent. Soc. 4 : 61-70
Edwards, 1872
List of species of butterflies collected by Campbell Carrington and Wm. B. Logan, of the expedition [to Montana], in 1871. Preliminary report [5th Ann. rep.] U. S. Geological Survey of Montana, by F. V. Hayden in Hayden, Geol. Surv. Montana: 466-467
Edwards, 1872
Synopsis of North American butterflies Synopsis N. Amer. Butts.: 52pp
Edwards, 1876
Description of new species of diurnal Lepidoptera found within the United States and British North America Trans. amer. ent. Soc. 5 : 202-208
Edwards, 1877
Catalogue of the Diurnal Lepidoptera of America North of Mexico Trans. amer. ent. Soc. 6 : 1-68
Edwards, 1878
On the Lepidoptera collected by Dr. Elliott Coues, U. S. A., in Montana, during 1874 Bull. Geol. Surv. Terr. 4 (2) : 513-517
Edwards, 1883
Notes upon a small collection of butterflies, made in Judith Mountains. Montana, in 1883, by Wm. M. Courtis, M. E. Papilio 3 (7-10) : 157-164
Edwards, 1883
Comments on a paper by Dr. H. A. Hagen entitled "The Genus Colias" Papilio 3 (7-10) : 169-185
H. Edwards, 1877
Pacific Coast Lepidoptera. No. 24 - Notes on the Genus Colias, with descriptions of some apparently new forms Proc. Cal. Acad. Sc.: 1-11 (Feb. 5)
Elwes, 1906
On the Lepidoptera collected by the Officer on the recent Tibet Frontier Commission. Butterflies Proc. zool. Soc. Lond. 1906 : 479-485 (Butterflies, Elwes), pl. 36, f. 8-15, : 485-496 (Lepidoptera-Phalaenae, Hampson), : 496-498 (Tineina, Durrant)
Esper, 1778
Die Schmetterlinge in Abbildungen nach der Natur mit Beschreibungen. Theil I. Die Tagschmetterlinge. Band 1 Die Schmett. Th. I, Bd. 1 (1): 1-40, pl. 1-6 (1776), (2): 41-72, pl. 7-12 (1776), (3): 73-104, pl. 13-18 (1777), (4): 105-120, pl. 19-24 (1777), (5): 121-144, pl. 25-30 (1777), (6): 145-176, pl. 31-36 (1777), (7): 177-200, pl. 37-46 (1778), (8): 201-216, pl. 43-48 (1778), (9): 217-388, pl. 49-50 (1779)
Esper, 1780; Esper, 1781; Esper, 1783
Die Schmetterlinge in Abbildungen nach der Natur mit Beschreibungen. Theil I. Die Tagschmetterlinge. Fortsetzung. Band 2 Die Schmett. Th. I, Bd. 2 (1): 1-36, pl. 51-56 (1780), (2): 37-68, pl. 57-62 (1781), (3): 69-100, pl. 63-68 (1781), (4): 101-124, pl. 69-74 (1781), (5): 125-140, pl. 75-80 (1782), (7): 141-156, pl. 81-82 (1783), (8): 157-172, pl. 83-88 (1783), (9): 173-184, pl. 89-92 (1784), (28): 185-190, pl. 93 (1786)
Esper, 1789
Die Schmetterlinge in Abbildungen nach der Natur mit Beschreibungen. Theil I. Die Tagschmetterlinge. Supplement Theil 1. Abschnitt 1 Die Schmett., Suppl. Th 1 (1-2): 1-8, pl. 94 (1789), (3-4): 9-60, pl. 95-101 (1789), (5-6): 61-68, pl. 102-106 (1794), ([7]): [69-88], pl. [107-109] (1798), (8-9): 89-104, pl. 110-112 (1800), (10): 105-120, pl. 113-116 ([1803-1804])
Esper, 1805
Die Schmetterlinge in Abbildungen nach der Natur mit Beschreibungen. Theil I. Die Tagschmetterlinge. Supplement Theil 2 Die Schmett., Th I, Suppl. Th. 2 (11): 1-24, pl. 117-122 (1805), (12): 25-48, pl. 123-126 [1830]
Erschoff, 1874
Travels in Turkestan. Volume 2. Zoogeographical Investigations. Lepidoptera [in Russian] in Fedschenko, Travels in Turkestan. 2 (5) : 1-128, pl. 1-6
Eversmann, 1847
Lepidoptera quaedam nova Rossiae et Sibiriae indigena descripsit et delineavit Bull. Soc. Imp. Nat. Moscou 20 (3) : 66-83, 6 pls.
Eversmann, 1851
Description de quelques nouvelles espéce de Lépidoptéres de la Russie Bull. Soc. imp. Nat. Moscou 24 (2) : 610-644
Fabricius, 1807
Systema Glossatorum. Die neueste Gattungs-Eintheilung der Schmetterlinge aus den Linnéischen Gattungen Papilio und Sphinx Magazin f. Insektenk. (Illiger) 6: 277-295
Fassl, 1915
Neue Pierieden aus Süd-Amerika Dt. Ent. Z. Iris 29 (2/3) : 176-181, pl. 6 (1915)
C. & R. Felder, 1859
Lepidopterologische Fragmente. I-IV Wien. ent. Monats. 3 (6): 178-186, pl. 3-4, (9): 263-273, pl. 5, (11): 321-328, pl. 6, (12): 390-405, pl. 7-9
C. & R. Felder, 1860
Lepidopterologische Fragmente. V-VI Wien. ent. Monats. 4 (4): 97-112, pl. 1-2, (8): 225-251, pl. 3-4
C. & R. Felder, 1865
Reise der österreichischen Fregatte Novara um die Erde in den Jahren 1857, 1858, 1859 unter den Behilfen des Commodore B. von Wüllerstorf-Urbair. Zoologischer Theil. Band 2. Abtheilung 2. Lepidoptera. Rhopalocera Reise Fregatte Novara, Bd 2 (Abth. 2) (1): 1-136, pl. 1-21 (Rhop.) (C. & R. Felder, [1865]), (2): 137-378, pl. 22-47 (Rhop.) (C. & R. Felder, 1865), (3): 379-536, pl. 48-74 (Rhop.) (C. & R. Felder, 1867), (4): 537-548, pl. 75-120, (Inhalts-Verz.) 1-9 (pl. 1-74), (Felder & Rogenhofer, 1874) , (5): pl. 121-140, (erklärung) 1-10 (pl. 75-107), (Inhalts-Verz.) 1-20 (pl. 108-140), (Felder & Rogenhofer, 1875)
Ferris, 1973
A revision of the Colias alexandra complex (Pieridae) aided by ultraviolet reflectance photography with designation of a new subspecies J. Lep. Soc. 27 (1): 57-73
Ferris, 1981
A new subspecies of Colias alexandra W. H. Edwards and notes on Colias hecla Lefebvre (Pieridae: Coliadinae) Bull. Allyn Mus. 63: 1-12
Ferris, 1982
Revision of the North American Colias hecla Lefebvre (Pieridae: Coliadinae) Bull. Allyn Mus. 71: 1-19
Ferris, 1988
Revision on several North American leguminosae-feeding Colias spec., with description of a new ssp. Bull. Allyn Mus. 116: 1-28
Ferris, 1989
A new species of Colias from Utah Bull. Allyn Mus. 128: 1-11
Fitch, 1854
Insects of Algiers, from the Museum of Natural History of Paris Trans. N.Y. St. agric. Soc. 13: 234,378
Geoffroy in Fourcroy, 1785
Entomologia parisiensis, sive catalogus Insectorum, etc. Ent. Paris. 2: 233-544
Fritsch, 1912
Neue Spielarten Ent. Rundsch. 29 (21) : 142-144
Fritsch, 1913
Lepidopterologische Beobachtungen Ent. Rundschau 30 : 46-47
Fritsch, 1918
Neue Schmetterlingsformen Ent. Rundschau 35 : 5-6
Fruhstorfer, 1903
Neue Pieriden aus Ost- und Süd-Asien Dt. ent. Z. Iris 16 (1) : 47-50, pl. 1, f. 3-4 (early October)
Fruhstorfer, 1907
Verzeichniss der von Herrn Dr. Theodor Koch-Grünberg am oberen Waupes 1903-1905 gesammelten Rhopaloceren mit Besprechung verwandter Arten Stettin ent. Ztg 68 (1): 117-164 (May), (2): 207-309, pl. 1 (July)
Fruhstorfer, 1908
Neue paläarktische Pieriden Ent. Zs. 22 (12) : 50-51 (20 June)
Fruhstorfer, 1909; Fruhstorfer, 1910
2. Familie: Pieridae, Weisslinge in Seitz, Gross-Schmett. Erde 9 : 119-120 (1909), : 121-144 (14 January, 1910), : 145-152 (11 February, 1910), : 153-168 (2 April, 1910), : 169-176 (4 April, 1910), : 177-190 (20 May, 1910)
Gerhard, 1882
Lepidopterologisches Berl. ent. Zs. 26 (1) : 125-128
Godart, 1819
in Latreille & Godart, Encyclopédie Méthodique. Histoire naturelle Entomologie, ou histoire naturelle des crustacés, des arachnides et des insectes Encyclopédie Méthodique. 9 (1): 3-328 (1819), (2): 329-828 ([1824])
Godman & Salvin, [1889]
Biologia Centrali-Americana. Rhopalocera. (1887-1901) Biol. centr.-amer., Lep. Rhop. 2: 1-782, 3: pl. 1-112
Gross & Ebert, 1975
Beitrag zur Kenntnis der Rhopaloceren Irans. 2. Beitrag: Neue Taxa der Pieridae, Satyridae und Nymphalidae J. ent. Soc. Iran, suppl. 1: 8-45
Grum-Grshimailo, 1884
Lepidopterologische Mittheilungen in Romanoff, Mém. Lép. 1 : 162-173
Grum-Grshimailo, 1885
Bericht über meine Reise in das Alai Gebiet (Correspondenz) in Romanoff, Mém. Lép. 2 : 212-247
Grum-Grshimailo, 1887
Bericht über meine Reise in das östliche Buchara (Correspondez; mit 1 Karte). Nebst Anghang: Dianosen einger neuen Species in Romanoff, Mém. Lép. 3 : 357-402
Grum-Grshimailo, 1888
Novae species et varietates Rhopalocerum e Pamir Horae Soc. ent. Ross. 22 (3-4) : 303-307
Grum-Grshimailo, 1890
Le Pamir et sa faune lépidoptérologique in Romanoff, Mém. Lép. 4 : 1-576, pl. A,1-21
Grum-Grshimailo, 1891
Lepidoptera nova in Asia centrali novissime lecta et descripta Horae Soc. ent. Ross. 25 (3-4) : 445-465
Grum-Grshimailo, 1893
Lepidoptera Palearctica Nova Horae Soc. ent. Ross. 27 (1-2): 127-129, : 379-386
Grum-Grshimailo, 1895
Lepidoptera Palearctica Nova (3) Horae Soc. ent. Ross. 29 : 290-293
Grum-Grshimailo, 1899
Lepidoptera nova vei[sic] rarum cognita regionis palaearcticae. I Ann. Mus. Zool. Imp. Acad. Sci. St.-Petersb. 4 (4) : 455-472
Guenée, 1864
Note sur Quelques espècs du genre Colias Ann. Soc. Ent. Fr. (4) 4 : 197-200
Gunder, 1924
Several New Aberrant Lepidoptera (Rhopalocera) from California Ent. News 35 (5) : 153-158
Gunder, 1928
Additional transitional forms Can. Ent. 60 : 162-168, 2pls
Gunder, 1932
New Rhopalocera Can. Ent. 64 (2): 276-284
Hammond & McCorkle, 2003
A new desert subspecies of Colias occidentalis (Pieridae) from southeastern Oregon J. Lep. Soc. 57 (4): 274-275
Hemming, 1934
Some notes in the nomenclature of Palaearctic and African Rhopalocera Stylops 3 : 97-99
Herrich-Schäffer, 1850
Systematische Bearbeitung der Schmetterlinge von Europa, zugleich als Text, Revision und Supplement zu J. Hübner's Sammlung europäischer Schmetterlinge, 1843 (-1855), Die Tagfalter Syst. Bearb. Schmett. Europ. 1 (9): Title ("1843"), (1): [1-3], 4-16, (ii) pl. 1-5, f. 1-29 (1843), (2): 17-40, (ii) pl. 29-34, f. 131-156 (1843), (3): 41-56, (ii) pl. 6-12, f. 30-60, pl. 35-36, f. 157-164 (1844), (4): 57-80, (ii) pl. 13-22, f. 61-107 (1844), (5): 81-90, (ii) pl. 23-28, f. 108-130, pl. 37-41, f. 165-189 (1844), (6): 91-98, (ii) pl. 42-44, f. 190-203 (1844), (7): 99-114, (ii) pl. 45-52, f. 204-248 (1844), (9): 115-122 (1844), (10): 123-146, (ii) pl. 53-57, f. 249-274 (1845), (11): 147-162 (1845), (12): (iii) pl. 1-3, f. 1-17 (1845), (13): 163-164, (ii) pl. 58-65, f. 275-315 (1846), (14): (ii) pl. 66-70, f. 316-338, (iii) pl. 4-5, f. 18-30 (1846), (26): (ii) pl. 71-77, f. 339-376 (1847), (32): (ii) pl. 78-82, f. 377-400 (1848), (36): (ii) pl. 83-89, f. 401-426, (iii) pl. 6, f. 31-36 (1848), (37): (ii) pl. 90-91, f. 427-437 (1849), (43): (ii) pl. 92-94, f. 438-450, 452-453 (1850), (45): (ii) pl. 95-102, f. 453-489 (1850), (47): (ii) pl. 103, f. 490-495 (1851), (49): (ii) pl. 104-109, f. 496-528 (1851), (51): (ii) pl. 110-118, f. 529-570 (1851), (54): (ii) pl. 119-123, f. 571-596 (1852), (56): (ii) pl. 124-127, f. 597-618, pl. 130-132, f. 628-639, pl. 134, f. 646-651 (1852), (59): (ii) pl. 128-129, f. 619-627, pl. 133, f. 640-645 (1853), (65): (iii) pl. 7, f. 37-42, (index) [1], 2-20 (1854), (66): (index) 21-24 (1855)
Horsfield, [1829]
Descriptive Catalogue of the Lepidopterous Insects contained in the Museum of the Horourable East-India Company, illustrated by coloured figures of new species Descr. Cat. lep. Ins. Mus. East India Coy (1): 1-80, pl. 1-4 (1828), (2): 81-144, pl. 5-8, (expl.) [1-4] (1829)
Hoshiai, 1993
On two subspecies of Colias (Lepidoptera, Pieridae) recently described from China Tyô to Ga 44 (1): 15-17
Hoshiai & Rose, 1998
A description of a new subspecies of Colias nebulosa Oberthur, 1894 (Lepidoptera, Pieridae) Butterflies 20 : 58-61
Hovanitz, 1943
The nomenclature of the Colias chrysotheme complex in North America Amer. Mus. Novit. 1240: 1-4
Huang & Murayama, 1992
Butterflies of Xinjiang province, China Tyô to Ga 43 (1) : 1-22
Hübner, 1803
Sammlung europäischer Schmetterlinge. I. Papiliones - Falter ("Erste Band") Samml. eur. Schmett. [1] : 3-74, : pl. 1-88, f. 1-446 (1799-1800), : pl. 89-96, f. 447-490 (1800-1803), : pl. 97-114, f. 491-588 (1803-1804), : pl. 115-119, f. 589-612 (1805), : pl. 120-124, f. 613-636 (1805-1806), : pl. 125-128, f. 637-652 (1806-1808), : pl. 129-144, f. 653-727 (1808-1813), : pl. 145-147, f. 728-741 (1814-1816), : pl. 148-150, f. 742-756 (1816-1817), : pl. 151, f. 757-762 (1818-1819), : pl. 152-154, f. 757-774 (1819-1822), : pl. 155, f. 775-778 (1822-1823), : pl. 156-161, f. 779-806 (1823), : pl. 162-175, f. 807-874 (1823-1824), : pl. 176, f. 869-874 (1824-1825), : pl. 177-181, f. 875-895 (1825-1826), : pl. 182-187, f. 896-981 (1827-1828), : pl. 188-195, f. 922-968 (1828-1832), : pl. 196, f. 969-974 (1832-1833), : pl. 197-201, f. 975-997 (1834-1836), : pl. 202-203, f. 998-1009 (1836-1838), : pl. 204-207, f. 1010-1029 (1837-1838)
Hübner, 1823
Zuträge zur Sammlung exotischer Schmettlinge, Vol. 2 [1819-] 1823 Zuträge Samml. exot. Schmett. 2 : [1-3], 4-6, [7], 8-32, [33-40] (1823), : pl. [36-38], f. 201-218 (1819), : pl. [39-66], f. 219-382 (1819 - 1821), : pl. [67-69], f. 383-400 (1821 - 1822)
Keferstein, 1883
Ueber die Tagschmetterlings-Gattung Colias F. Verh. zool.-bot. Ges. Wien 32 : 449-458
Kirby, 1881
Notes on new or interesting species of Papilionidae and Pieridae Trans. ent. Soc. Lond. 1881 (3) : 351-358
Klots, 1935
A new Colias from South Dakota (Lepidoptera, Pieridae) Amer. Mus. Novit. 767: 1-2
Klots, 1937
Some notes on Colias and Brenthis (Lepidoptera, Pieridae and Nymphalidae) J. N.Y. Ent. Soc. 45 (3/4) : 311-333
Klots, 1940
New butterfly subspecies from Wyoming (Nymphalidae, Pieridae) Amer. Mus. Novit. 1054: 1-6
Korshunov, 1996
[Additions and corrections to the book "Butterflies of the Asian part of Russia."] Dopolneniya i ispravleniya k knige "Dnevnye babochki aziatskoi chasti Rossii": 66pp
Kroon, 1985
A new form of the common pierid butterfly Colias electo eclecto (L.) Ent. Rec. J. Var. 97 (1-2) : 6
Krulikovsky, 1907
Petites notices lepidopterologique Revue Russe Ent. 7: 27-34
Krulikovsky, 1908; Krulikowsky, 1908
Einige neue Varietäten und Aberrationen der Lepidopteren des östlichen Russlands Soc. Ent. 23 (1): 2-3, (2): 11-12, (3): 18
Krulikovsky, 1909
[russian] Rev. russ. ent. 9 (1-2) : 109-114
Krulikovsky, 1909
[russian] Rev. russ. ent. 9 (3) : 292-323
Lampa, 1885
Förteckning öfver Skandinaviens och Finlands Macrolepidoptera Ent. Tidskr. 6 (1) : 1-137
Lederer, 1853
Versuch die europäischen Lepidopteren (einschlissig der ihrem Habitus nach noch zure europäischen Fauna gehörigen Arten Labradors, der asiatischen Türkei un des asiatischen Russlands) in möglichst natürliche Reihenfolge zu stellen, nebst Bemerkungen zu eineg Familien und Arten. Rhopaloceren & Heteroceren Verh. zool.-bot. Ver. Wien 2 (Abh.) : 14-54 (Rhopaloceren), : 65-126 (Heteroceren)
Lederer, 1858
Noch einige syrische Schmetterlinge Wien. ent. Monats. 2 (5) : 135-152, pl. 2-4
Lederer, 1869
Verzeichniss der von Herrn Jos. Haberhauer bei Astrabad in Persion gesammelten Schmetterlinge Horae Soc. ent. ross. 6 (2) : 73-93, pl. 4-5
Lefèbvre, 1836
Description d'une nouvelle espèce de coliade Ann. Soc. ent. Fr. 5 : 383-387, pl. 9
Lempke, 1954
Catalogus der Nederlandse Macrolepidoptera (Tweede Supplement) Tijdschr. Ent. 97 : 301-345
Linnaeus, 1758
Systema Naturae per Regna Tria Naturae, Secundum Clases, Ordines, Genera, Species, cum Characteribus, Differentiis, Symonymis, Locis. Tomis I. 10th Edition Syst. Nat. (Edn 10) 1 : 1-338, : 339-824
Linnaeus, 1763
in Johansson (Thesis), Centuria Insectorum Amoenitates Acad., 6: -,(121) 384-415
Linnaeus, 1767
Systema Naturae per Regna tria Naturae, secundum Classes, Ordines, .... Editio Duocecima Reformata. Tom. 1. Part II. Syst. Nat. (Edn 12) 1 (2): -,533-1327
Lukhtanov, 1999
Neue Taxa und Synonyma zentalasiatischer Tagfalter Atalanta 30 (1/4): 135-150
Lyman, 1884
Notes on Colias christina Can. Ent. 16 (1) : 5-9
Matsumura, 1919
Thousand Insects of Japan. Additamenta 3 [Shin Nihon senchu zukai] Thous. Ins. Japan. Addit. 3 : 475-742, pl. 26-53
Matsumura, 1928
New Butterflies especially from the Kuriles Insecta Matsumurana 2 (4) : 191-201
Matsumura, 1929
New butterflies from Japan, Korea and Formosa Insecta Matsumurana 3 (2/3) : 87-107, pl. 4
Matsumura, 1939
The butterflies from Jehol (Nekka) Manchoukuo, collected by Marqis Y. Yamashina Bull. biogeogr. Soc. Japan 9 (20): 343-359, pl. XIII-XIV
McDunnough, 1927
The Lepidoptera of the Seton Lake Region, British Columbia Can. ent. 59 : 152-162,193-199,207-214,239-246,266-277
McDunnough, 1928
Notes on Canadian diurnal Lepidoptera Can. ent. 60 (11): 266-275
McDunnough, 1938
Check list of the Lepidoptera of Canada and the United States of America. Part I. Macrolepidoptera Mem. South. Calif. Acad. Sci. 1: 175pp
McLachlan, 1878
Report on Insecta (including Arachnida) collected by Captain Feilden and Mr. Hart between Parallels of 78° and 83° North Latitude, during the recent Arctic Expedition J. Linn. Soc. Zool. Lond. 14 : 98-122
McLeod, 2004
More aberrations of Colias electo electo L. (Lep.: Pieridae) from the Cape, South Africa Ent. Rec. J. Var. 116 (2): 85-89
McLeod & McLeod, 2002
Colias electo electo L. ab. capensis ab. nov. (Lep.: Pieridae): A new aberration of the African Clouded Yellow from the Cape, South Africa Ent. Rec. J. Var. 114 (5) : 221-234
Ménétriés, 1832
Catalogue raisonne des objects de zoologie recuelis dans un voyage au Caucase et jusqu'aux frontières de la Perse Cat. Rais.: -
Ménétriés, 1855
Enumeratio corporum animalium Musei Imperialis Academiae Scientiarum Petropilitanae. Classis Insectorum, Ordo Lepidopterorum Cat. lep. Petersb. 1: 1-66, pl. 1-6, (supplement) 67-112 (1855), 2: 67-97,99-144, pl. 7-14 (1857), 3: 145-161, pl. 15-18 (1863)
Ménétriés, 1859
Lépidoptères de la Sibérie orientale et en particulier des rives de l'Amour in Schrenck, Reise Forschungen Amur-Lande 2 (1) : 1-75, pl. 1-5
Metschl, 1922
Colias hyale L., C. edusa F. und C. myrmidone E., ihr Formenkreis und einige neue Aberrationen aus dem Faunenebiete Regensburgs Mitt. munchn. ent. Ges. 12: 6-12
Metschl, 1925
Einige interessante Schmetterlingsformen aus dem Faunengebiet von Regensburg Int. Ent. Zeit. 19: 25
Miller, [1913]
Neue Rhopalocera aus Transkaukasien Dt. ent. Z. Iris 26 (4) : 220-223 ([1913])
Moltrecht, 1909
Beitrag zur Kenntnis der ostpaläarktischen Lepidopterenfauna Ent. Zs. 22 (44) : 184-185
Moore, 1865
List of diurnal Lepidoptera collected by Capt. A.M. Lang in the N. W. Himalayas Proc. zool. Soc. Lond. 1865 (2) : 486-509, pl. 30-31
Moore, 1878
Descriptions of Lepidopterous Insects collected the late Dr. F. Stoliczka during the Indian-Government Mission to Yarkund in 1873 Ann. Mag. nat. Hist. (5) 1 (3) : 227-237
Moore, 1882
List of the Lepidoptera collected by the Rev. J.H. Hocking, chiefly in the Kangra Discrict, N.W Hiamalaya; with Descriptions of new Genera and Species Proc. zool. Soc. Lond. 1882 (1) : 234-263, pl. 11-12
Motschulsky, 1860
Insectes du Japon Etud. Ent. 9: 4-39
Mrácek & Schulte, 1991
Colias grumi burhcana nov. ssp., eine neue Unterart von Colias grumi Alpherakyi[sic] aus Provinz Qinghai (China) Nachr. ent. Ver. Apollo N.F. 12: 164-165
Möschler, 1870
Beiträge zur Schmetterlingsfauna von Labrador Stettin Ent. Ztg 31 (1-3): 113-125, (4-6): 251-254, (7-9): 265-272
Nakahara, 1926
Some Rhopalocera of eastern US, including three new Forms Bull. Brooklyn ent. Soc. 20 (5): 221-224
Neuburger, 1905
Üeber einige noch unbenannte Coliasabarten Soc. ent. 20 (6) : 42
Nordmann, 1851
Die im Gebiete der Fauna Taurico-Caucasica beobachteten Schmetterlinge Bull. Soc. imp. Nat. Moscou 24 (4): 395-428, 6 pls
Oberthür, 1886
Espèces Nouvelles de Lépidoptères du Thibet / Nouveaux Lépidoptères du Thibet Étud. d'Ent. 11 : 13-38, pl. 1-7
Oberthür, 1894
Lépidoptères d'Europe, d'Algérie, d'Asie et d'Océanie Étud. d'Ent. 19 : 1-41, pl. 1-8
Ochsenheimer, 1808
Die Schmetterlinge von Europa. Erster Theil, Zweyte Abtheilung. Falter, oder Tagschmetterlinge Schmett. Europa 1 (2) : 1-240
Pieszczek, 1905
Über die Variabilität von Colias myrmidone Esp. Verh. zool.-bot. Ges. Wien 55 : 401-423, pl. 1
Pieszczek, 1912
Über die Verbreitung der Colias myrmidone Esp, in Österreich - Ungarn und deren Variabilität Jber. Wien Ent. Ver. 22: 199-225
Pieszczek, 1917
Die Zucht der Colias myrmidone Esper und ab. alba aus dem Ei, sowie deren Variabilität Z. Öst. EntVer. 2 (9): 69-79
Pilate, 1888
List of Lepidoptera taken in and around Dayton Papilio (2) 5[?]: 65-71
Reakirt, 1865
Observations upon some American Pierinae Proc. ent. Soc. Philad. 4 (2) : 216-222
Rebel, 1901
[no title] Verh. zool.-bot. Ges. Wien 51 : 310-311
Nitsche in Rebel, 1913
Bericht der Sektion für Lepidopterologie Verh. zool.-bot. Ges. Wien 63 : (11) -(37),(49)-(62)
Reiff, 1917
Colias philodice f. n. nigrofasciata Massachusetts 1 (3) : 22-23, pl. 3
Reiff, 1917
Colias philodice f. n. rothkei Lepidopterist 1 (11) : 84, pl. 7
Reissinger, 1972
Die geographisch-subspecifische Gliederung von Colias alfacariensis Ribbe unter Berücksichtigun der Migrationsverhältnisse Atalanta 3: 349-376
Reissinger, 1972
Eine neue Colias-unterart aus Peru Atalanta 4: 60-64
Reissinger, 1974
Die geographisch-subspecifische Gliederung von Colias alfacariensis Ribbe unter Berücksichtigun der Migrationsverhältnisse Atalanta 5: 1-33
Reissinger, 1989
Die geographisch-subspecifische Gliederung von Colias alfacariensis Ribbe unter Berücksichtigun der Migrationsverhältnisse Neue Ent. Nachricten 26: 1-351
Reutti, 1853
Uebersicht der Lepidoperen-Fauna des Grossherzogthum's Baden Beitr. Rhein. Naturg. 3 : 1-216
Ribbe, 1905
Einige neue Formen von Schmetterlingen aus Andalusien Soc. ent. 20 (18) : 137-138
Riley, 1923
The Rhopalocera of the Mt. Everest 1921 Expedition Trans. ent. Soc. Lond. 1922 (3-4) : 461-483, pl. 36-37
Romanoff, 1882
Une nouvelle Colias du Caucase Horae Soc. ent. ross. 17 : 127-134, pl. 4-5
Rothschild, 1913
Descriptions of two new Colias and some African Syntomidae Novit. Zool. 20 (1) : 187-188
Röber, [1909]; Röber, [1910]
2. Familie: Pieridae, Weisslinge in Seitz, Grossschmett. Erde 5 : 53-56 (25 May 1908), : 57-64 (26 Sep 1908), : 65-72 (26 Nov 1908), : 73-80 (16 Feb 1909), : 81-88 (19 Feb 1909), : 89-96 (26 Mar 1909), : 97-104 (26 Apr 1909), : 105-111 (21 May 1909)
Schawerda, 1906
Materialvorweisung über Colias myrmidone - balcanica - caucasica Verh. zool. bot. Ges. Wien 56 : 651-652
Schawerda, 1907
Colias myrmidone balcanica variation, male genitalia Verh. zool. bot. Ges. Wien 57: 216-220
Schawerda, 1908
Bericht über lepidopterologische Sammelreisen in Bosnien und in der Herzegowina Jber. Wien. ent. Ver. 19: 85-126 (?1909)
Schawerda, 1913
Siebenter Nachtrag zur Lepidopterenfauna Bosniens und der Herzegovina Verh. zool.-bot. Ges. Wien 63 : 141-178
Schawerda, 1919
Neue Formen aus meiner Sammlung Z. öst. EntVer. 4 (7): 67-68
Schawerda, 1922
Zwölfter Nachtrag zur Lepidoperenfauna Bosniens und der Herzegowina Verh. zool.-bot. Ges. Wien 71 : 145-170, pl., f. 1-26
Schawerda, 1924
Faunula dolomitana Mitt. münchn. ent. Ges. 14: 73-119
Schilde, 1884
Beiträge und Unterhaltungen zur Schmetterlingskunde Ent. Nachr. 10 (22): 333-346 (1884), 11 (9): 133-137 (1885)
Schnaider, 1950
Formes de lépidoptères nouvelles ou plus rares Polsk. Pismo. Ent. 19: 237-255
Scudder, 1862
On the genus Colias in North America Proc. Boston Soc. Nat. Hist. 9 : 103-111
Scudder, 1889
The Butterflies of the Eastern United States and Canada, with special reference to New England Butts eastern U.S. Canada 1: 1-766, 2: 767-1774, 3: 1775-1958, pl. 1-89
Shapiro, 1991
The zoogeography and systematics of the Argentine Andean and Patagonian pierid fauna J. Res. Lepid. 28 (3): 137-238 ("1989")
Shchetkin, 1990
The new subspecies Colias alpherakyi from Uzbekistan Bull. Soc. nat. Moscou 95: 58-60
Sheljuzhko, 1913
Lepidopterologische Notizen Dt. Ent. Z. Iris 27 (1) : 13-22
Sheljuzhko, 1923
Lépidoptères nouveaux ou peu connus de la Sibérie Ann. Soc. ent. Fr. 91 (4) : 281-290
Skala, 1912
Einige Formen der Colias myrmidone Esper Int. ent. Z. 6 (3): 16
Skala, 1912
Das zweibändige, ausgezeichnet besprochene Werk: "Die Lepidopterenfauna Mährens" Ver. naturf. Ver. Brünn 50: 63-241
Skinner, 1898
A synonymic catalaogue of the North American Rhopalocera Cat. North Am. Rhopalocera: 215pp
Skinner & Mengel, 1892
Greenland Lepidoptera Proc. Acad. Nat. Sci. Philad. 44 : 156-159, Ent. News 3 (3): pl. 2
Stauder, 1913
Makrolepidopteren-fauna de Adriatischen Küstengebiete Boll. Soc. adriat. Sci. nat. 27 (1): 105-166, 3pls
Stauder, 1922
Neue Palaearktenformen. I Mitt. münch. ent. Ges. 12 (1-6) : 17-25
Stauder, 1924
Neue Palaearktenformen. II Mitt. münch. ent. Ges. 14: 59-66
Staudinger, 1861
Catalog der Lepidopteren Europas und der angrendzenden Länder in Staudinger & Wocke, Cat. Lep. (ed. 1): 1-192
Staudinger, 1862
Ueber einige neue griechische Lepidopteren Stettin Ent. Ztg 23 (4-6) : 257-271
Staudinger, 1886
Centralasiatische Lepidopteren Stett. Ent. Ztg. 47 (4-6): 193-215, (7-9): 225-256
Staudinger, [1892]
Neue Arten und Varietäten von Lepidopteren des paäarktischen Faunengebiets Dt. Ent. Z. Iris 4 (2) : 224-339, pl. 3-4 (1892)
Staudinger, 1892
Lepidopteren des Kentei-Gebirges Dt. Ent. Z. Iris 5 (2) : 300-393
Staudinger, 1894
Hochlandine Lepidoptera Dt. Ent. Z. Iris 7 (1) : 43-100, pl. 1-2
Staudinger, 1897
Drei neue paläarktische Lepidopteren Dt. Ent. Z. Iris, 10 (1) : 152-156
Staudinger, 1899
Lepidopteren. Hamburger Magalhaensische Sammelreise Naturhist. Mus. Hamburg 2 (6) : 1-117, pl., f. 1-27
Staudinger in Staudinger & Bang-Haas, 1882
Ueber einige neue Parnassius- und andere Tagfalter-Arten Central-Asiens Berl. ent. Z. 26 (1) : 161-177, pl. 1-2
Rebel in Staudinger & Rebel, 1901
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Staudinger in Staudinger & Wocke, 1871
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Storace, 1956
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Stoll, [1781]
Uitlandsche Kapellen (Papillons exotiques) in Cramer, Uitl. Kapellen 4 (26b-28): 29-90, pl. 305-336 (1780), (29-31): 91-164, pl. 337-372 (1781), (32-32): 165-252, 1-29, pl. 373-400 (1782)
Stoneham, 1957
Two new generic names, and new subspecies and forms of Pieridae Bull. Stoneham Mus. (70) : 1-3
Strand, 1911
Neue Gross-Schmetterlinge aus Abyssinien, etc. Ent. Rundsch. 28 : 137-141
Strand, 1912
Über einige exotische Lepidoptera aus der Sammlung des Herrn W. Niepelt in Zirlau Archiv Naturg. 78 A (9) : 178-186
Strand, 1916
Abbildungen und Beschreibungen neuer und wenig bekannter Lepidoptera aus der Sammlung W. Niepelt Lepidoptera Niepeltiana (2) : 1-26, pl. 13-17
Strand, 1918
Nachtrag zum zweiten Teil meiner "Lepidoptera Niepeltiana" Soc. ent. 33 (1): 2-3, (3): 11-12, (5): 19-20, (7): 26-28, (8): 30-31, (10): 38-39, (11): 43-44, (12): 46-47
Strecker, 1878
Butterflies and Moths of North America Synon. Cat. Macroplep.: -
Strecker, 1880
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Strecker, 1885
Description of a new Colias from the Rocky Mountains, and of an Example of Polymelianism in Samia Cecropia Proc. Acad. Nat. Sci. Philad. 37 : 24-27
Strecker, 1900
Lepidoptera, Rhopaloceres and Heteroceres, indigenous and exotic Lep. Rhopal. Het., Suppl. 3 : 13-37,38
Thurau, 1903
Colias nastes Bsd. var. werdandi Zett. und ihre Aberrationen Berl. ent. Z. 48 : 113-116
Troubridge & Philip, 1990
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Vári, 1976
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Verhulst, 1999
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Verhulst, 2002
Fiches specifiques des Colias Fabricius Lambillionea 102 (2): 123-126
Verhulst, 2002
Fiches specifiques des Colias Fabricius Lambillionea 102 (3, I): 291-297
Verhulst, 2002
Fiches specifiques des Colias Fabricius Lambillionea 102 (4, I): 514-520
Verhulst, 2003
Fisches specifiques des Colias Fabricius Lambillionea 103 (1): 72-80
Verhulst, 2006
Colias mukana Berger, 1981 stat. nov. et description de Colias mukana jolyi ssp. nov. (Lepidoptera, Pieridae, Coliadinae) Lambillionea 106 (2 II) : 286-292
Verity, [1908]; Verity, 1909; Verity, 1911
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Verity, 1916
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Verity, 1926
Zygaenae, Grypocera and Rhopalocera of the Cottian Alps compared with other races Ent. Rec. J. Var. 38 : 101-106,120-126,170-176
Vorbrodt, 1912
Die Schmetterlinge der Schweiz. I. Band Die Schmett. Schweiz 1 : 1-489
Wagner, 1907
Colias myrmidone balcanicus Rbl., from nov. ♀ semialba Ent. Zeit. 21 (20) : 125
Wallengren, 1860
Lepidopterologische Mittheilungen (1-3) Wien ent. Monats. 4: 33-46,161-176 (1860), 7: 137-151 (1863)
Watson, 1895
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Strecker in Weeks, 1900
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Winhard, 2000
Pieridae I Butterflies of the world, 10: (1) 40pp, (2) 48pls
Wnukowsky, 1929
Einige Nomeklatur-Notizen über die paläarktischen Lepidopteren Zool. Anz. 83 (9/10): 221-224
Wyatt & Omoto, 1966
New Lepidoptera from Afghanistan Entomops 5: 138-167, 6: 170-200
Yano, 1958
A new form of Colias erate poliographus Motschulsky from Japan (Peridae) Tyô to Ga 9 (3): 36
Zetterstedt, 1839
Insecta Lapponica Insecta Lapponica : 1-1014 ([1839]), : 1015-1140 (1840)


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Keep in mind that the taxonomic information is copied from various sources, and may include many inaccuracies. Expert help is welcome.