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Lethe Hübner, [1819]
Treebrowns
Lethe Hübner, [1819]; Verz. bek. Schmett. (4): 56, TS: Papilio europa Fabricius
Tanaoptera Billberg, 1820; Enum. Ins. Mus. Billb. : 79, TS: Papilio europa Fabricius
Zophoessa Doubleday, [1849]; Gen. diurn. Lep. (1): pl. 61, f. 1, TS: Zophoessa sura Doubleday
Debis Doubleday, [1849]; Gen. diurn. Lep. (1): pl. 61, f. 3, TS: Debis samio Doubleday
Hanipha (Satyrinae)Moore, [1880]; Lepid. Ceylon 1 (1): 18, TS: Lethe sihala Moore
Tansima Moore, 1881; Trans. ent. Soc. Lond. 1881 (3) : 305, TS: Lethe satyrina Moore
Charma Doherty, 1886; J. Asiat. Soc. Bengal 55 Pt.II (2) : 117, TS: Zophoessa baladeva Moore
Rangbia Moore, [1892]; Lepidoptera Indica 1 : 232, TS: Debis scanda Moore
Nemetis Moore, [1892]; Lepidoptera Indica 1 : 237, TS: Papilio minerva Fabricius
Pegada Moore, [1892]; Lepidoptera Indica 1 : 224 nota, TS: Mycalesis oculatissima Poujade
Kirrodesa Moore, [1892]; Lepidoptera Indica 1 : 237 nota, TS: Debis sicelis Hewitson
Placilla Moore, [1892]; Lepidoptera Indica 1 : 253, TS: Lethe christophi Leech
Archondesa Moore, [1892]; Lepidoptera Indica 1 : 270, TS: Lethe lanaris Butler
Choranesa Moore, [1892]; Lepidoptera Indica 1 : 270, TS: Lethe trimacula Leech
Dionana Moore, [1892]; Lepidoptera Indica 1 : 271, TS: Lethe(?) margaritae Elwes
Sinchula Moore, [1892]; Lepidoptera Indica 1 : 275, TS: Debis sidonis Hewitson
Kerrata Moore, [1892]; Lepidoptera Indica 1 : 285, TS: Lethe tristigmata Elwes
Putlia Moore, [1892]; Lepidoptera Indica 1 : 287 (unn. repl. Charma Doherty, 1886), TS: Zophoessa baladeva Moore
Harima Moore, [1892]; Lepidoptera Indica 1 : 299, TS: Neope callipteris Butler
Magula Fruhstorfer, 1911; in Seitz, Gross-Schmett. Erde 9 : 313, TS: Zophoessa jalaurida de Niceville
Hermias Fruhstorfer, 1911; in Seitz, Gross-Schmett. Erde 9 : 324, TS: Satyrus verma Kollar
Lethe, NSG Voucher Specimen [Wahlberg]
Lethe (Satyrinae) ; Moore, [1880], Lepid. Ceylon 1 (1): 16
Lethe (Lethina) ; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 176; [Wahlberg]

The grouping from [BIR].


The 1 species-group

Lethe europa (Fabricius, 1775)
S.India, Dun - Assam, Burma, N.India, Japan, Hong Kong. See [About maps] Bamboo Tree-brown
Papilio europa Fabricius, 1775; Syst. Ent.: 500, no. 247, TL: S.India
Lethe europa, NSG Voucher Specimen [Wahlberg]
Lethe europa ; [BOW: pl. 167, f. 3]; [BOR, 416]; [MRS, 327]; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 176
[europa-1v.jpg] 512x341 (~27Kb) underside India: Guwahati, 16.11.2005, Photo © Kedar Tokekar

Larva on Bambusa multiplex, Microstegium ciliatum, Leleba multiplex [MRS], Bambusa, Dinochloa, Microstegium, Phyllostachys, Pleioblastus Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 176


Java, Bawean, Kangean, Bali Lethe europa europa ; [BOR, 416]
L. e. niladana Fruhstorfer, 1911
N.India - Burma, Thailand Lethe europa niladana Fruhstorfer, 1911; in Seitz, Gross-Schmett. Erde 9 : 315, TL: "Kuamon - Burma"
Lethe europa niladana, Butterflies in Indo-China [Yutaka Inayoshi]
Lethe europa niladana ; [BOR, 416]; [MRS, 327]
L. e. gada Fruhstorfer, 1911
S.Yunnan ?Lethe europa f. gada Fruhstorfer, 1911; in Seitz, Gross-Schmett. Erde 9 : 316, TL: Siam; Tonkin
Lethe europa gada ; Huang & Xue, 2004, Neue Ent. Nachr. 57: 140 (name)
L. e. malaya Corbet, 1941
Peninsular Malaya, Singapore Lethe europa malaya Corbet, 1941; Entomologist 74 (6): 133, TL: Malaysia
Lethe europa malaya, Butterflies in Indo-China [Yutaka Inayoshi]
Lethe europa malaya ; [BOR, 416]; [BMP: 124, pl. 15, f. 7-9]
[malaya-1fv.jpg] 1211x1031 (~343Kb) underside female Thailand, Phuket, a garden in the Nai Yang beach area. 16th February 2009, Photo © Oleg Kosterin

L. e. ragalva Fruhstorfer, 1911
S.India europa ragalva Fruhstorfer, 1911; in Seitz, Gross-Schmett. Erde 9 : 316, TL: Karwar
Lethe europa ragalva ; [BOR, 416]
L. e. nudgara Fruhstorfer, 1911
Andamans europa nudgara Fruhstorfer, 1911; in Seitz, Gross-Schmett. Erde 9 : 316, pl. 96 c, TL: Andamans
Lethe europa nudgara ; [BOR, 416]
L. e. tamuna de Nicéville, 1887
Nicobars Lethe tamuna de Nicéville, 1887; Proc. zool. Soc. Lond. 1887 (3) : 449, pl. 39, f. 6 ♀, TL: Little Nicobar
Lethe europa tamuna ; [BOR, 416]
L. e. kumara Fruhstorfer, 1911
Enggano europa kumara Fruhstorfer, 1911; in Seitz, Gross-Schmett. Erde 9 : 316, TL: Engano
Lethe europa kumara ; [BOR, 416]
L. e. mahamaya Fruhstorfer, 1911
Lombok europa mahamaya Fruhstorfer, 1911; in Seitz, Gross-Schmett. Erde 9 : 316, pl. 96 d, TL: Lombok
Lethe europa mahamaya ; [BOR, 417]
L. e. pavida Fruhstorfer, 1908
Japan, Taiwan Lethe europa pavida Fruhstorfer, 1908; Ent. Zs. 22 (35) : 141, TL: Kagi
Lethe europa pavida ; [BOR, 417]; [MRS, 327]

Larva on Phyllostachys makinoi, Bambusa multiplex, B. oldhamii [MRS]

L. e. alaca Fruhstorfer, 1911
Palawan europa alaca Fruhstorfer, 1911; in Seitz, Gross-Schmett. Erde 9 : 316
Lethe europa alaca ; [BOR, 417]
L. e. cevanna Fruhstorfer, 1911
Philippines (Mindanao) europa cevanna Fruhstorfer, 1911; in Seitz, Gross-Schmett. Erde 9 : 316, TL: Mindanao
Lethe europa cevanna ; [BOR, 417]
L. e. beroe (Cramer, [1775])
S.China, ?Hong Kong Papilio beroe Cramer, [1775]; Uitl. Kapellen 1 (1-7): pl. 79, f. C, D
Lethe europa beroe ; [BOR, 417]; [MRS, 327]

Lethe rohria (Fabricius, 1787)
Afghanistan, Ceylon, S.India - Pachmarhi, Kashmir - Assam, Burma. See [About maps] Common Treebrown
Papilio rohria Fabricius, 1787; Mantissa Insectorum 2: 45, TL: S.India
stenopoides Mell, 1942; Arch. Naturgesch. (N.F.) 11 : 272
= ; [NHM]
godona Fruhstorfer, 1911²; in Seitz, Gross-Schmett. Erde 9 : 315, TL: Java
= ; [NHM]
Lethe rohria borneensis Kalis, 1933; Tijdschr. Ent. 76 (1-2) : 69, TL: Borneo
= ; [NHM]
Lethe rohria, NSG Voucher Specimen [Wahlberg]
Lethe rohria rohria, Butterflies in Indo-China [Yutaka Inayoshi]
Lethe rohria ; [BOW: pl. 167, f. 13]; [BOR, 418]; [MRS, 327]
[rohria-1v.jpg] 600x400 (~55Kb) underside India: Pune, 18.12.2005 © Kedar Tokekar

Burma Debis dyrta C. & R. Felder, [1867]; Reise Fregatte Novara, Bd 2 (Abth. 2) (3): 497, TL: Bengalia
?dyrta Bingham, 1905²; Fauna Brit. Ind., Butts (ed. 1) 1: 80
= Lethe rohria ; [NHM]
Lethe rohria rohria ; [BOR, 418]
rohria r. dyrta ; [BIR, (Kashmir - Kumaon)]
rohria r. rohria ; [BIR, (Sikkim - Burma)]
L. r. neelgheriensis (Guérin-Méneville, 1843)
Ceylon, S.India - S.Bihar, Jubbulpore, Pachmarhi, Konkan Satyrus neelgheriensis Guérin-Méneville, 1843; in Delessert, Souvenirs Voy. Inde 2: 74, pl. 21, f. 1, 1a
= Lethe dyrta ; [NHM]
Debis neelgheriensis ; Hewitson, 1863, Ill. exot. Butts [4] (Debis II-III): [35], pl. [19], f. 6
Lethe neelgherriensis[sic, recte neelgheriensis] ; Moore, [1881], Lepid. Ceylon 1 (1): 16, pl. 7, f. 1, 1a
rohria r. nilgiriensis[sic, recte neelgheriensis] ; [BIR]
Lethe rohria neelgheriensis ; [BOR, 418]
L. r. yoga Fruhstorfer, 1911
Ceylon rohria yoga Fruhstorfer, 1911; in Seitz, Gross-Schmett. Erde 9 : 317, TL: Ceylon
= Lethe dyrta ; [NHM]
Lethe rohria yoga ; [BOR, 418]
L. r. permagnis Fruhstorfer, 1911
S.China rohria permagnis Fruhstorfer, 1911; in Seitz, Gross-Schmett. Erde 9 : 317
= Lether dyrta ; [NHM]
Lethe rohria permagnis ; [BOR, 418]; [MRS, 327]
L. r. anunda Fruhstorfer, 1911
Java, Bali rohria anunda Fruhstorfer, 1911; in Seitz, Gross-Schmett. Erde 9 : 317, TL: Bali, Java
= Lether dyrta ; [NHM]
Lethe rohria anunda ; [BOR, 418]
L. r. daemoniaca Fruhstorfer, 1908
Taiwan Lethe dyrta daemoniaca Fruhstorfer, 1908; Ent. Zs. 22 (35) : 141
= Lether dyrta ; [NHM]
Lethe rohria daemoniaca ; [BOR, 418]; [MRS, 327]

Larva on Miseanthus sinesis, M. flavidus, Brachiaria mutica, Imperata cylindrica [MRS]

L. r. sambaluna Fruhstorfer, 1911
Lombok rohria sambaluna Fruhstorfer, 1911; in Seitz, Gross-Schmett. Erde 9 : 317, pl. 97 b, TL: Lombok
= Lethe dyrta ; [NHM]
Lethe rohria sambaluna ; [BOR, 418]

Lethe drypetis (Hewitson, 1863)
Ceylon, S.India. See [About maps] Tamil Treebrown
Debis drypetis Hewitson, 1863; Ill. exot. Butts [4] (Debis II-III): [36], pl. [19], f. 11-12, TL: Ceylon
Debis embolina Butler, 1866; Ent. mon. Mag. 3 (28) : 77, TL: Ceylon
= ; [NHM]
Lethe todara Moore, 1881; Trans. ent. Soc. Lond. 1881 (3) : 305, TL: Nilgiris
Lethe drypetes[sic] ; Moore, [1880], Lepid. Ceylon 1 (1): 17, pl. 8, f. 1a-b
Lethe drypetis ; [BOW: pl. 167, f. 2]; [BOR, 418]

Lethe daretis (Hewitson, 1863)
Ceylon. See [About maps] Ceylon Treebrown
Debis daretis Hewitson, 1863; Ill. exot. Butts [4] (Debis II-III): [35], pl. [19], f. 7-8, TL: Ceylon
Lethe daretis ; Moore, [1880], Lepid. Ceylon 1 (1): 18, pl. 7, f. 2a-b; [BOW: pl. 167, f. 2 (text only)]; [BOR, 418]

Lethe confusa Aurivillius, 1897
Murree - Assam, Burma, Malaysia. See [About maps] Banded Treebrown
Lethe confusa Aurivillius, 1897; Ent. Tidskr. 18 (3/4) : 142, TL: [India]
Lethe confusa, NSG Voucher Specimen [Wahlberg]
Lethe confusa confusa, Butterflies in Indo-China [Yutaka Inayoshi]
Lethe confusa ; [EBW]; [BOW: pl. 166, f. 15-16]; [BOR, 420]; [MRS, 331]
[confusa-1v.jpg] 512x341 (~52Kb) underside India: Guwahati, 17.11.2005, Photo © Kedar Tokekar

Larva on Microstegium ciliatum, Bambusa multiplex [MRS]


N.India, Himalayas, Assam, Burma, Thailand, Indo-China, Malaya, SE.Tibet, S.China, Hainan, Yunnan Lethe confusa fuhaica Lee, 1962; Acta ent. sin. 11 (1) : 139, 144, TL: Yunnan
Lethe confusa confusa ; [BOR, 420]
Lethe confusa fuhaica ; Huang & Wu, 2003, Neue Ent. Nachr. 55: 126, pl. 10, f. 14
Lethe confusa confusa ; Huang, 2003, Neue Ent. Nachr. 55: 94 (note)
L. c. apara Fruhstorfer, 1911
S.Burma, Thailand - Indo China, Hong Kong, Hainan Lethe rohria var. apara Fruhstorfer, 1911; in Seitz, Gross-Schmett. Erde 9 : 318, TL: Tenasserim
Lethe confusa apara ; [BOR, 420]; [MRS, 331]
L. c. gambara Fruhstorfer, 1911
Sikkim, Assam Lethe rohria gambara Fruhstorfer, 1911; in Seitz, Gross-Schmett. Erde 9 : 315, TL: Assam
Lethe confusa gambara ; [BOR, 420]
L. c. enima Fruhstorfer, 1911
Sumatra, Peninsular Malaya Lehte enima Fruhstorfer, 1911; in Seitz, Gross-Schmett. Erde 9 : 315, pl. 97, f. a2 (80), (86)
Lethe confusa enima ; [BMP: 125, pl. 15, f. 19]; [BOR, 420]
L. c. godana Fruhstorfer²
Java Lethe confusa godana ; [BOR, 420]

Lethe dynsate (Hewitson, 1863)
Ceylon. See [About maps] Ceylon Forester
Debis dynsate Hewitson, 1863; Ill. exot. Butts [4] (Debis II-III): [36], pl. [19], f. 9-10, TL: Ceylon
Lethe sihala Moore, 1872; Proc. zool. Soc. Lond. 1872 (2) : 555
Lethe dynsate ; Moore, [1880], Lepid. Ceylon 1 (1): 20, pl. 9, f. 1, 1a; [BOR, 418]

Lethe mekara (Moore, [1858])
China, Assam - Burma, Malaya. See [About maps] Common Red Forester
Debis mekara Moore, [1858]; in Horsfield & Moore, Cat. lep. Ins. Mus. East India Coy 1 : 219, TL: Darjeeling
vajra Fruhstorfer, 1911; in Seitz, Gross-Schmett. Erde 9 : 319
= ; [NHM]
manis Corbet, 1942; Ann. Mag. nat. Hist. (11) 9: 609, TL: Mentawei I.
= ; [NHM]
Lethe mekara, NSG Voucher Specimen [Wahlberg]
Lethe mekara ; [BOW: pl. 167, f. 10]; [BOR, 422]; [MRS, 330]

Larva on Phyllostachys pubescens [MRS]

N.India, Sikkim Lethe mekara mekara ; [BOR, 422]
L. m. zuchara Fruhstorfer, 1911
Assam - Tenasserim, Indo China mekara zuchara Fruhstorfer, 1911; in Seitz, Gross-Schmett. Erde 9 : 319
Lethe mekara zuchara ; [BOR, 422]
L. m. crijnana Fruhstorfer, 1911
Thailand, Laos, N.Vietnam, S.Yunnan Lethe mekara crijnana Fruhstorfer, 1911; in Seitz, Gross-Schmett. Erde 9 : 319, pl. 98 c, TL: N.Vietnam
Lethe mekara crijnana, Butterflies in Indo-China [Yutaka Inayoshi]
Lethe mekara crijnana ; [MRS, 330]
L. m. gopaka Fruhstorfer, 1911
Peninsular Malaya Lethe mekara gopaka Fruhstorfer, 1911; in Seitz, Gross-Schmett. Erde 9 : 320, TL: Malaysia
Lethe mekara gopaka, Butterflies in Indo-China [Yutaka Inayoshi]
Lethe mekara gopaka ; [BOR, 422]; [BMP: 125, pl. 15, f. 14, 16, 18]
L. m. debata Fruhstorfer, 1911
Sumatra mekara debata Fruhstorfer, 1911; in Seitz, Gross-Schmett. Erde 9 : 320, pl. 98 d, TL: Celebes
Lethe mekara debata ; [BOR, 422]
L. m. sumati Fruhstorfer, 1911
Borneo mekara sumati Fruhstorfer, 1911; in Seitz, Gross-Schmett. Erde 9 : 319, pl. 98 c d, TL: N.Borneo
Lethe mekara sumati Fruhstorfer; [BOR, 422]

Lethe chandica (Moore, [1858])
W.China, Assam - Burma, Malaya. See [About maps] Angled Red Forester
Debis chandica Moore, [1858]; in Horsfield & Moore, Cat. lep. Ins. Mus. East India Coy 1 : 219, TL: Darjeeling, India
Debis isabella Butler, 1866; Proc. zool. Soc. Lond. 1866 (1) : 41, pl. 3, f. 4, TL: Philippines
obscura Mell, 1942; Arch. Naturgesch. (N.F.) 11 : 276
= ; [NHM]
rahula Fruhstorfer, 1911; in Seitz, Gross-Schmett. Erde 9 : 320, TL: Hainan
= ; [NHM]
Lethe chandica chandica, Butterflies in Indo-China [Yutaka Inayoshi]
Lethe chandica ; [BOW: pl. 166, f. 11]; [BOR, 422]; [MRS, 329]
[chandica-1v.jpg] 512x341 (~36Kb) underside India: Itanagar, 13.11.2005, Photo © Kedar Tokekar

Larva on Phyllostachys pubescens [MRS]


NW.India - Peninsular Malaya?, Sumatra?, Yunnan Lethe chandica chandica ; [BOR, 422]; Huang, 2003, Neue Ent. Nachr. 55: 94 (note)
L. c. namura Fruhstorfer, 1911
Peninsular Malaya, Sumatra? Lethe namura Fruhstorfer, 1911; in Seitz, Gross-Schmett. Erde 9 : 321
Lethe chandica namura ; [BMP: 125, pl. 15, f. 15, 17]
L. c. marga Fruhstorfer, 1911
Java chandica marga Fruhstorfer, 1911; in Seitz, Gross-Schmett. Erde 9 : 321, TL: W.Java
Lethe chandica marga ; [BOR, 422]
L. c. ratnacri Fruhstorfer, 1908
Taiwan Lethe chandica ratnacri Fruhstorfer, 1908; Insekten-Börse 25: 38
Lethe chandica ratnacri ; [BOR, 422]; [MRS, 329]
L. c. coelestis Leech, [1892]
W.China chandica coelestis Leech, [1892]; Butts China Japan Corea (1): 20, pl. 3, f. 7-8
Lethe chandica coelestis ; [BOR, 422]; [MRS, 329]
L. c. flanona Fruhstorfer, 1911
Lethe chandica flanona Fruhstorfer, 1911; in Seitz, Gross-Schmett. Erde 9 : 320, TL: Assam
Lethe chandica flanona ; [MRS, 329]
L. c. suvarna Fruhstorfer, 1908
Laos, N.Vietnam, Hainan Lethe chandica suvarna Fruhstorfer, 1908; Insekten-Börse 25: 38 [? not Fruhstorfer lit.]
?suvarna Fruhstorfer, 1911; in Seitz, Gross-Schmett. Erde 9 : 320 [?], TL: Chiem Hoa, N.Vietnam
Lethe chandica suvarna, Butterflies in Indo-China [Yutaka Inayoshi]
Lethe chandica suvarna ; [MRS, 329]
L. c. negrito (C. & R. Felder, 1863)
Philippines (Luzon) Debis negrito C. & R. Felder, 1863; Wien. ent. Monats. 7 (4) : 124, TL: Luzon
Lethe chandica negrito ; [BOR, 422]
L. c. sisapon Fruhstorfer, 1911
Philippines (Mindoro) chandica sisapon Fruhstorfer, 1911; in Seitz, Gross-Schmett. Erde 9 : 321, TL: Mindoro
Lethe chandica sisapon ; [BOR, 422]
L. c. byzaccus Fruhstorfer, 1911
Philippines (Mindanao) chandica byzaccus Fruhstorfer, 1911; in Seitz, Gross-Schmett. Erde 9 : 321, TL: Mindanao
Lethe chandica byzaccus ; [BOR, 422]
L. c. jomaria Fruhstorfer, 1911²
Sulu Is. chandica jomaria Fruhstorfer, 1911²; in Seitz, Gross-Schmett. Erde 9 : 321, TL: Jolo
Lethe chandica jomaria ; [BOR, 422]
L. c. ratnapandi Fruhstorfer, 1911²
Palawan chandica ratnapandi Fruhstorfer, 1911²; in Seitz, Gross-Schmett. Erde 9 : 321, TL: Palawan
?Lethe negrito var. ; Staudinger, 1889, Dt. ent. Z. Iris 2 (1) : 34
Lethe chandica ratnapandi ; [BOR, 422]

Lethe satyavati de Nicéville, 1880
Assam. See [About maps] Pallid Forester
Lethe satyavati de Nicéville, 1880; J. Asiat. Soc. Bengal 49 Pt.II (4) : 246

Lethe distans Butler, 1870
Bhutan, Sikkim, Assam - Burma, N.Thailand. See [About maps] Scarce Red Forester
Lethe distans Butler, 1870; Trans. ent. Soc. Lond. 1870 (4) : 488, TL: Darjeeling, India
Lethe distans, Butterflies in Indo-China [Yutaka Inayoshi]
Lethe distans ; [BOR, 424]

The 2 species-group

Lethe naga Doherty, 1889
Assam - Myanmar, Thailand, Laos. See [About maps] Naga Treebrown
Lethe naga Doherty, 1889; J. asiat. Soc. Bengal 58 Pt.II (1) : 123, pl. 10, f. 4, TL: Margherita
Lethe naga, Butterflies in Indo-China [Yutaka Inayoshi]
Lethe naga ; [BOR, 428]

Lethe margaritae (Elwes, 1882)
Bhutan, Sikkim. See [About maps] Bhutan Treebrown
Lethe(?) margaritae Elwes, 1882; Proc. zool. Soc. Lond. 1882 (4) : 405, pl. 25, f. 15
Lethe margaritae ; [BOR, 428]

Lethe insana (Kollar, [1844])
W.China, Chamba - Assam. See [About maps] Common Forester
Satyrus insana Kollar, [1844]; in Hügel, Kaschmir und das Reich der Siek, 4: 448, pl. 16, f. 3-4
hyrania (Kollar, [1844]) (Satyrus); in Hügel, Kaschmir und das Reich der Siek, 4: 449, pl. 17, f. 1-2
= ; [NHM]
caerulescens Mell, 1923²; Dt. ent. Z. 1923 (2) : 155
= ; [NHM]
Lethe insana insana, Butterflies in Indo-China [Yutaka Inayoshi]
Lethe isana[sic] ; [BOW: pl. 202, f. 2 (missp.)]
Lethe insana ; [BOR, 420]; [MRS, 331]; Yoshino, 2008, Futao (54): 9, pl. 2, f. 5-6

Larva on Arundinaria falcata [MRS]

NW.India, Bhutan, Sikkim, Nepal, Assam - Indo China Debis dinarbas Hewitson, 1863; Ill. exot. Butts [4] (Debis II-III): [37], pl. [20], f. 15, TL: North India
Lethe insana insana ; [BOR, 420]
L. i. procris Leech, 1891
N.Thailand, Laos, N.Vietnam ?Lethe procris Leech, 1891; Entomologist 24 (Suppl.) : 2, TL: Wa-Shan; Chang-Yang
= Lethe baucis ; [NHM]
L. i. formosana Fruhstorfer, 1908
Taiwan Lethe insana formosana Fruhstorfer, 1908; Ent. Zs. 22 (12): 48, TL: Formosa, Larachi
Lethe insana formosana ; [BOR, 420]; [MRS, 331]
L. i. brisanda de Nicéville, 1886
Bhutan, Assam, NE.Burma Lethe brisanda de Nicéville, 1886; J. Asiat. Soc. Bengal 55 Pt.II (3) : 249, pl. 11, f. 13 ♀, TL: Bhutan
Lethe brisanda ; de Nicéville, 1887, Proc. zool. Soc. Lond. 1887 (3) : 451, pl. 39, f. 8 ♂; [BOR, 421]; Yoshino, 2008, Futao (54): 9
Lethe insana brisanda ; [MRS, 331]
L. i. baucis Leech, 1891
W.China, C.China Lethe baucis Leech, 1891; Entomologist 24 (Suppl.) : 3, TL: Wa-Shan; Chia-Kou-Ho; Ta-Tsien-Lu
Lethe baucis ; [BOW: pl. 201, f. 8-9]
Lethe insana baucis ; [MRS, 331]

Lethe sadona Evans, 1932
NE.Burma (Sadon). See [About maps] sadona Evans, 1932; Indian Butterflies (edn. 2): 107, TL: Burma
Lethe sadona ; [BOR, 421]; Yoshino, 2008, Futao (54): 9

Lethe kazuichiroi Yoshino, 2008
N.Myanmar. See [About maps] Lethe kazuichiroi Yoshino, 2008; Futao (54): 9, pl. 2, f. 1-2, TL: Kaching state, N. Myanmar

Lethe kouleikouzana Yoshino, 2008
China(N.Yunnan). See [About maps] Lethe kouleikouzana Yoshino, 2008; Futao (54): 9, pl. 2, f. 3-4, TL: Mts. Gaolingonshan, 2500m, N. Yunnan

Lethe serbonis (Hewitson, 1876)
Bhutan, Sikkim, Assam - W.China. See [About maps] Brown Forester
Debis serbonis Hewitson, 1876; Ent. mon. Mag. 13 (7) : 151, TL: Darjeeling
Debis davidi Oberthür, 1881; Étud. d'Ent. 6 : 15, pl. 7, f. 5, TL: Mou-Pin
= ; [NHM]
flavofasciata Leech, [1892]²; Butts China Japan Corea (1): 30
= ; [NHM]
Lethe serbonis naganum Tytler, 1914; J. Bombay nat. Hist. Soc. 23 (2): 219, TL: Kabru Peak, Manipur; Naga Hills
teesta Talbot, 1947; Fauna Br. Ind., Butt. (2nd ed.) 2: 220, TL: Sikkim
bhutya Talbot, 1947; Fauna Br. Ind., Butt. (2nd ed.) 2: 221, TL: Bhutan
Lethe davidi f. angulata Seitz, 1907; Gross-Schmett. Erde 1 : 83
Lethe serbonis ; [BOW: pl. 167, f. 9]; [BOR, 424]; [MRS, 330]
?L. s. pallida Tytler, 1939
Burma Lethe serbonis pallida Tytler, 1939; J. Bombay nat. Hist. Soc. 41 (2): 245, TL: NE.Burma
Lethe serbonis pallida ; Huang, 2001

Lethe scanda (Moore, 1857)
Bhutan, Sikkim - Assam, Burma - W.China. See [About maps] Blue Forester
Debis scanda Moore, 1857; in Horsfield & Moore, Cat. lep. Ins. Mus. East India Coy 1 : 218, TL: Darjeeling
Debis nada Moore, 1857; in Horsfield & Moore, Cat. lep. Ins. Mus. East India Coy 1 : 218, TL: Darjeeling
= ; [NHM]
Zophoessa dirphia Druce, 1875; Cistula ent. 1 (12) : 357, TL: Darjeeling
= ; [NHM]
Lethe scanda ; [BOW: pl. 167, f. 14]

Lethe vindhya (C. & R. Felder, 1859)
Sikkim - Burma, Assam - Tenasserim. See [About maps] Black Forester
Debis vindhya C. & R. Felder, 1859; Wien. ent. Monats. 3 (12): 402, TL: Assam
Lethe alberta Butler, 1871; Ann. Mag. nat. Hist. (4) 8 (46) : 283, TL: "Benares"
Lethe dolopes Hewitson, 1872; Ent. mon. Mag. 9 : 85, TL: Darjeling
Lethe vindhya vindhya, Butterflies in Indo-China [Yutaka Inayoshi]
Lethe vindhya ; [BOR, 424]; [MRS, 339]

Larva on Bambusa spp. [MRS]

L. v. ladesta Fruhstorfer, 1911
Lethe vindhya ladesta Fruhstorfer, 1911; in Seitz, Gross-Schmett. Erde 9 : 319, TL: Tenasserim
Lethe vindhya ladesta ; [MRS, 339]
L. v. luaba Corbet, 1941
Lethe luaba Corbet, 1941; J. fed. Malay St. Mus. 18 (5): 807
Lethe vindhya luaba ; [BMP: 125, pl. 15, f. 11]

Lethe kansa (Moore, 1857)
Kumaon - Assam, Burma. See [About maps] Bamboo Forester
Debis kansa Moore, 1857; in Horsfield & Moore, Cat. lep. Ins. Mus. East India Coy 1 : 220, TL: Darjeeling
Lethe kansa kansa, Butterflies in Indo-China [Yutaka Inayoshi]
Lethe kansa ; [BOW: pl. 167, f. 7]; [BOR, 419]; [MRS, 324]
Sikkim Lethe kansa kansa ; [BOR, 419]
L. k. zeugitana Fruhstorfer, 1911
Assam, Manipur Lethe kansa zeugitana Fruhstorfer, 1911; in Seitz, Gross-Schmett. Erde 9 : 318, TL: Assam
Lethe kansa zeugitana ; [BOR, 419]
L. k. vaga Fruhstorfer, 1911
S.Burma, S.Thailand, S.Yunnan Lethe kansa vaga Fruhstorfer, 1911; in Seitz, Gross-Schmett. Erde 9 : 318-319, TL: Tenasserim
Lethe kansa vaga ; [BOR, 419]; [MRS, 324]

Lethe sinorix (Hewitson, 1863)
China, Sikkim - Assam, S.Burma, Malaya. See [About maps] Red Tailed Forester
Debis sinorix Hewitson, 1863; Ill. exot. Butts [4] (Debis II-III): [38], pl. [20], f. 19-20, TL: East India
kuatungensis Mell, 1923²; Dt. ent. Z. 1923 (2) : 154
= ; [NHM]
lotaoshanensis Tinkham, 1936; Lingman J. Sci. 15: 561
= ; [NHM]
Lethe sinorix sinorix, Butterflies in Indo-China [Yutaka Inayoshi]
sinoryx ; (missp.?)
Lethe sinorix ; [BOW: pl. 168, f. 2]; [BOR, 418]; [MRS, 324]
Sikkim, Bhutan, Assam, Burma, ?Thailand Lethe sinorix sinorix ; [BOR, 418]
L. s. vanda Corbet, 1941
Peninsular Malaya (Cameron Highlands) Lethe vanda Corbet, 1941; J. fed. Malay St. Mus. 18 (5): 807
Lehte sinorix vanda ; [BOR, 419]
Lethe sinorix vanda ; [BMP: 125, pl. 15, f. 10]
L. s. obscura Mell, 1942
sinorix obscura Mell, 1942; Arch. Naturgesch. (N.F.) 11 : 274
Lethe sinorix obscura ; [MRS, 324]

Lethe latiaris (Hewitson, 1862)
Sikkim - Assam, Karens. See [About maps] Pale Forester
Debis latiaris Hewitson, 1862; Ill. exot. Butts [4] (Debis I): [34], pl. [18], f. 4, TL: Sylhet
Lethe latialis[sic] hige Fujioka, 1970; Spec. Bull. Lepid. Soc. Japan (4) : 47, TL: Godavari (1600m), Kathmandu
= ; [NHM]
Lethe latiaris ; [BOW: pl. 167, f. 8]
Yunnan Lethe latiaris latiaris ; Huang, 2003, Neue Ent. Nachr. 55: 94 (note), f. 147-149
L. l. perimela Fruhstorfer, 1911
S.Burma, N.Thailand, Laos, N.Vietnam Lethe latiaris perimela Fruhstorfer, 1911; in Seitz, Gross-Schmett. Erde 9 : 323, pl. 97 c, TL: Tandong, Tenasserim
Lethe unistigma Lee, 1985; , TL: Shizong, E.Yunnan
= ?; Huang, 2003, Neue Ent. Nachr. 55: 94; Huang & Wu, 2003, Neue Ent. Nachr. 55: 125, pl. 10, f. 12-13
Lethe latiaris perimele[sic?] ; [BOR, 424]
L. l. lishadii Huang, 2002
Yunnan Lethe latiaris lishadii ; Huang, 2003, Neue Ent. Nachr. 55: 94 (note)

Lethe gulnihal de Nicéville, 1887
Bhutan - Assam, S.Burma. See [About maps] Dull Forester
Lethe gulnihal de Nicéville, 1887; Proc. zool. Soc. Lond. 1887 (3) : 450, pl. 39, f. 7, TL: Bhutan
?mandersi Elwes, 1891; Proc. zool. Soc. Lond. 1891 (2): 263 [ ?? ]
= ; [NHM]
?Lethe gulnihal var. ; Elwes, 1891, Proc. zool. Soc. Lond. 1891 (2): 263
Lethe gulnihal ; [BOR, 426]
Bhutan - N.Burma Lethe gulnihal gulnihal ; [BOR, 426]
L. g. peguana (Moore, [1891])
S.Burma, ?Thailand Rangbia peguana Moore, [1891]; Lepidoptera Indica 1 : 235, TL: Burma
?Lethe gulnihal peguana f. issa Fruhstorfer, 1911; in Seitz, Gross-Schmett. Erde 9 : 323, TL: Saipha, Upper Burma
= Lethe gulnihal ; [NHM]
Lethe gulnihal peguana, Butterflies in Indo-China [Yutaka Inayoshi]
Lethe gulnihal peguana ; [BOR, 426]

Lethe bhairava (Moore, 1857)
Bhutan - Assam. See [About maps] Rusty Forester
Debis bhairava Moore, 1857; in Horsfield & Moore, Cat. lep. Ins. Mus. East India Coy 1 : 217, TL: Darjeeling
Debis anysis Hewitson, 1862; Ill. exot. Butts [4] (Debis I): [33], pl. [18], f. 1, TL: East India
Lethe bhairava, Butterflies in Indo-China [Yutaka Inayoshi]
Lethe bhairava ; [BOW: pl. 166, f. 12]; [BOR, 426]
[bhairava-1v.jpg] 600x400 (~52Kb) underside India: Arunachal Pradesh, Taley Valley WLS, 11.11.2005, Photo © Kedar Tokekar


Lethe verma (Kollar, [1844])
Kashmir - Assam, Burma. See [About maps] Straight-banded Treebrown
Satyrus verma Kollar, [1844]; in Hügel, Kaschmir und das Reich der Siek, 4: 447, pl. 16, f. 1-2
Lethe verma ; [BOW: pl. 168, f. 5]; [BOR, 420]; [MRS, 332]
[verma-1u.jpg] 600x400 (~63Kb) upperside India: Arunachal Pradesh, Taley Valley WLS, 10.11.2005, Photo © Kedar Tokekar [verma-1v.jpg] 600x400 (~47Kb) underside India: Arunachal Pradesh, Taley Valley WLS, 10.11.2005, Photo © Kedar Tokekar

NW.India Lethe verma verma ; [BOR, 420]
L. v. sintica Fruhstorfer, 1911
Sikkim, Assam, Yunnan Lethe verma sintica Fruhstorfer, 1911; in Seitz, Gross-Schmett. Erde 9 : 324
Lethe verma sintica ; [BOR, 420]; Huang, 2001; Huang, 2003, Neue Ent. Nachr. 55: 94 (note)
L. v. stenopa Fruhstorfer, 1908
Burma, Thailand - Indo China, S.China, Hainan Lethe (Tausima) verma stenopa Fruhstorfer, 1908; Ent. Zs. 22 (31) : 127, TL: N.Vietnam
?Lethe (Tausima) verma stenopa f. laticincta Fruhstorfer, 1908; Ent. Zs. 22 (31) : 127
Lethe verma stenopa, Butterflies in Indo-China [Yutaka Inayoshi]
Lethe verma stenopa ; [BOR, 420]; [MRS, 332]
L. v. robinsoni Pendlebury, 1933
Peninsular Malaya (Cameron Highlands) Lethe robinsoni Pendlebury, 1933; J. Fed. Malay St. Mus. 17 (2): 386
Lethe verma robinsoni ; [BOR, 420]; [BMP: 125, pl. 15, f. 13]
L. v. satarnus Fruhstorfer, 1911²
W.China verma satarnus Fruhstorfer, 1911²; in Seitz, Gross-Schmett. Erde 9 : 324
Lethe verma satarnus ; [BOR, 420]; [MRS, 332]
L. v. cintamani Fruhstorfer, 1909
Taiwan Lethe verma cintamani Fruhstorfer, 1909; Ent. Zs. 23 (8) : 40, TL: Formosa
Lethe verma cintamani ; [BOR, 420]; [MRS, 332]

The 3 species-group

Lethe visrava (Moore, [1866])
Sikkim - Bhutan, Assam, Burma. See [About maps] White-edged Woodbrown
Debis visrava Moore, [1866]; Proc. zool. Soc. Lond. 1865 (3) : 768, pl. 41, f. 4
Debis deliades Hewitson, 1872; Ent. mon. Mag. 9 : 84, TL: Darjeeling
= ; [NHM]
Lethe visrava ; [BOR, 414]; [MRS, 344]

Lethe siderea Marshall, [1881]
Sikkim - W.China. See [About maps] Scarce Woodbrown
Lethe siderea Marshall, [1881]; J. Asiat. Soc. Bengal 49 Pt.II (4) : 246
Lethe siderea siderea, Butterflies in Indo-China [Yutaka Inayoshi]
Lethe siderea ; [BOR, 415]; [MRS, 343]
Lethe siderae[sic] ; [NHM]
Zophoessa siderea ;
L. s. kanoi Esaki & Nomura
Lethe siderea kanoi ; [MRS, 343]

Lethe sidonis (Hewitson, 1863)
Tibet, Kulu - Assam, S. Shan States. See [About maps] Common Woodbrown
Debis sidonis Hewitson, 1863; Ill. exot. Butts [4] (Debis II-III): [37], pl. [20], f. 16, TL: North India
?Lethe vaivarta Doherty, 1886; J. Asiat. Soc. Bengal 55 Pt.II (2) : 115, TL: Kumaon
?Lethe irma Evans, 1923; J. Bombay nat. Hist. Soc. 29: 531, TL: Gangtok
?Zophoessa nicevillei Evans, 1923; J. Bombay nat. Hist. Soc. 29: 531, TL: Bhutan, Garhwal
ranoi Esaki & Nomura, 1937; Zephyrus 7: 107, TL: Formosa
= ; [NHM]
Lethe sidonis sidonis, Butterflies in Indo-China [Yutaka Inayoshi]
Lethe sidonis ; [BOW: pl. 168, f. 1]; [BOR, 414]; [MRS, 333]
Zophoessa sidonis ; Huang, 2003, Neue Ent. Nachr. 55: 91 (note)
[sidonis-1v.jpg] 600x400 (~44Kb) underside India: Arunachal Pradesh, Taley Valley WLS, 10.11.2005, Photo © Kedar Tokekar


Lethe nicetella de Nicéville, 1887
Tibet, Sikkim. See [About maps] Small Woodbrown
Lethe nicetella de Nicéville, 1887; Proc. zool. Soc. Lond. 1887 (3) : 448, pl. 39, f. 5 ♂, TL: Sikkim
Lethe nicetella ; [BOR, 415]

Lethe maitrya de Nicéville, 1880
-. See [About maps] Barred Woodbrown
Lethe maitrya de Nicéville, 1880; J. Asiat. Soc. Bengal 49 Pt.II (4) : 245
Lethe maitrya ; [BOW: pl. 167, f. 15]; [BOR, 414]; [MRS, 335]
W.Himalayas, Nepal, Sikkim, Bhutan Lethe maitrya maitrya ; Huang, 2001, Neue Entomologische Nachrichten, 51: 96 (note)
L. m. thawgawa Tytler, 1939
NE.Burma, Nujiang Valley Lethe maitrya thawgawa Tytler, 1939; J. Bombay nat. Hist. Soc. 41 (2): 245, TL: Burma
Lethe maitrya thawgawa ; Huang, 2001, Neue Entomologische Nachrichten, 51: 96 (note), pl. 7, f. 56
Zophoessa maitrya thawgawa ; Huang, 2003, Neue Ent. Nachr. 55: 91 (note)
L. m. metokana Huang, 1998
SE.Tibet (Metok) Lethe maitrya metokana Huang, 1998; Neue Entomologische Nachrichten, 41: 223, pl. 4, f. 1a-c, 2a-c, TL: Metok
L. m. lijiangensis Huang, 2001
NW.Yunnan Lethe maitrya lijiangensis Huang, 2001; Neue Entomologische Nachrichten, 51: 96, TL: Lijiang, NW.Yunnan, 3500m
Misidentified/misapplied:
Lethe maitrya thawgawa; D'Abrera, 1993, 124 (Tsekou)
Lethe maitrya thawgawa; Huang, 1998, Neue Entomologische Nachrichten, 41: 251, pl. 4, f. 1d, 2d

Lethe nicetas (Hewitson, 1863)
Sikkim, Assam, Manipur. See [About maps] Yellow Woodbrown
Debis nicetas Hewitson, 1863; Ill. exot. Butts [4] (Debis II-III): [38], pl. [20], f. 17-18, TL: East India
Lethe nicetas nicetas, Butterflies in Indo-China [Yutaka Inayoshi]
Lethe nicetas ; [BOR, 414]
Zophoessa nicetas ; Huang, 2003, Neue Ent. Nachr. 55: 91 (note), f. 141

Lethe kabrua (Tytler, 1914)
Manipur (Kabru) Manipur Goldenfork
Zophoessa kabrua Tytler, 1914; J. Bombay nat. Hist. Soc. 23 (2): 222, 23 (3): pl. 1, f. 7, TL: Kabru Peak
Lethe kabrua ; [BOR, 416]

Lethe nosei (Koiwaya, 2000)
N.Burma. See [About maps] Lehte nosei ; Huang, 2001, Neue Entomologische Nachrichten, 51: 95 (note)

Lethe jalaurida (de Nicéville, 1880)
W.China, Kulu - Sikkim. See [About maps] Small Silverfork
Zophoessa jalaurida de Nicéville, 1880; J. Asiat. Soc. Bengal 49 Pt.II (4) : 245
plistia Fruhstorfer, 1911; in Seitz, Gross-Schmett. Erde 9 : 394
= ; [NHM]
Lethe jalaurida ; [BOW: pl. 167, f. 6]; [BOR, 416]; [MRS, 328]
W.Himalaya (foothills) Lethe jalaurida jalaurida ; [BOR, 416]
Lehte jalaurida jalaurida ; Huang, 2001, Neue Entomologische Nachrichten, 51: 94 (note)
L. j. elwesi (Moore, [1892])
Sikkim, Burma, W.China? Zophoessa elwesi Moore, [1892]; Lepidoptera Indica 1 : 298, pl. 92, f. 3
Lethe elwesi ; [BOW: pl. 167, f. 4]
Lethe jalaurida elwesi ; [BOR, 416]; Huang, 2001, Neue Entomologische Nachrichten, 51: 94 (note)
L. j. gelduba Fruhstorfer, 1911
Sichuan, Yunnan ?Lethe sidonis f. gelduba Fruhstorfer, 1911; in Seitz, Gross-Schmett. Erde 9 : 312, TL: Sikkim; Bhutan; Assam
= Lethe sidonis ; [NHM]
Lethe jalaurida gelduba ; [BOR, 416]; Huang, 2001, Neue Entomologische Nachrichten, 51: 94 (note)
L. j. nuolaensis Huang, 2001
Lethe jalaurida nuolaensis Huang, 2001; Neue Entomologische Nachrichten, 51: 94, pl. 7, f. 52, TL: 5km E of Nuola, Nujiang Valley, SE.Tibet

Lethe moelleri (Elwes, 1887)
Bhutan, Sikkim - Guanxi. See [About maps] Moeller's Silverfork
Zophoessa mölleri Elwes, 1887; Proc. zool. Soc. Lond. 1887 : 445
Lethe moelleri ; [BOR, 416]; [MRS, 328]
L. m. bruno Tytler, 1939
NE.Burma Lethe moelleri bruno Tytler, 1939; J. Bombay nat. Hist. Soc. 41 (2): 246, TL: Hthawgaw, NE Burma
Lethe moelleri bruno ; Huang, 2001, Neue Entomologische Nachrichten, 51: 95 (note)

Lethe bitaensis Yoshino, 1999
NW.Yunnan, SE.Tibet (Nujiang). See [About maps] Lethe moelleri bitaensis Yoshino, 1999; Neo Lepidoptera 4: 7, f. 49-50, TL: Bita Lake, Zhongdian, NW.Yunnan
= ?; Huang, 2001, Neue Entomologische Nachrichten, 51: 95
Lethe gesangdawai Huang, 2001; Neue Entomologische Nachrichten, 51: 95, f. 86, pl. 7, f. 55, TL: 10km E of Nuola, Nujiang Valley, SE.Tibet
= ; Yoshino, 2003, Futao (43): 8
Lethe bitaensis ; Yoshino, 2003, Futao (43): 8

Lethe goalpara (Moore, [1866])
Simla Hills, Sikkim, Assam. See [About maps] Large Goldenfork
Zophoessa goalpara Moore, [1866]; Proc. zool. Soc. Lond. 1865 (3) : 768
Lethe goalpara goalpara, Butterflies in Indo-China [Yutaka Inayoshi]
Lethe goalpara ; [BOW: pl. 167, f. 5]; [BOR, 414]
[goalpara-1u.jpg] 600x400 (~49Kb) upperside India: Arunachal Pradesh, Taley Valley WLS, 11.11.2005, Photo © Kedar Tokekar [goalpara-1v.jpg] 600x400 (~82Kb) underside India: Arunachal Pradesh, Taley Valley WLS, 11.11.2005, Photo © Kedar Tokekar

N.India - Assam ?narkunda Fruhstorfer, 1911; in Seitz, Gross-Schmett. Erde 9 : 314, TL: Simla
Lethe goalpara goalpara ; [BOR, 414]
L. g. kabruensis Tytler, 1939
Manipur (Kabru) goalpara kabruensis Tytler, 1939; J. Bombay nat. Hist. Soc. 41 (2): 246, TL: Burma
Lethe goalpara kabruensis ; [BOR, 414]
L. g. gana Talbot, 1947
Upper Burma goalpara gana Talbot, 1947; Fauna Br. Ind., Butt. (2nd ed.) 2: 186, TL: Sikkim
Lethe goalpara gana ; [BOR, 414]

Lethe sura (Doubleday, [1849])
Bhutan - Assam, Burma. See [About maps] Scarce Lilacfork
Zophoessa sura Doubleday, [1849]; Gen. diurn. Lep. (1): pl. 61, f. 1
Zophoessa sura Westwood, [1851]; Gen. diurn. Lep. (2): 362
Debis moupinensis Poujade, 1884; Bull. Soc. Ent. Fr. (6) 4 : cxl, TL: Mou-Pin
= ; [NHM]
Lethe sura, Butterflies in Indo-China [Yutaka Inayoshi]
Lethe (Zophoessa) sura ; de Nicéville, 1900, J. Bombay nat. Hist. Soc. 13 (1) : 163, pl. CC, f. 5-6
Lethe sura ; [BOW: pl. 168, f. 3]; [BOR, 412]; [MRS, 324]

Lethe baladeva (Moore, [1866])
Sikkim, Bhutan, Assam. See [About maps] Treble Silverstripe
Zophoessa baladeva Moore, [1866]; Proc. zool. Soc. Lond. 1865 (3) : 769, pl. 41, f. 5
aisa Fruhstorfer, 1911; in Seitz, Gross-Schmett. Erde 9 : 314
= ; [NHM]
Lethe baladeva ; [BOW: pl. 166, f. 14]; [BOR, 419]; [MRS, 336]

Lethe atkinsonia (Hewitson, 1876)
Sikkim - Bhutan. See [About maps] Small Goldenfork
Zophoessa atkinsonia Hewitson, 1876; Ent. mon. Mag. 13 (7) : 151, TL: Darjeeling
Lethe atkinsonia ; [BOR, 416]

Lethe ramadeva (de Nicéville, 1887)
Sikkim, Bhutan, Yunnan. See [About maps] Single Silverstripe
Zophoessa ramadeva de Nicéville, 1887; Proc. Asiat. Soc. Beng. 1887 : 147
Lethe ramadeva ; [BOR, 427]; Huang, 2003, Neue Ent. Nachr. 55: 91 (note)

Lethe violaceopicta (Poujade, 1884)
Tibet, W.China (Sichuan, Guizhou), Manipur. See [About maps] Manipur Woodbrown
Debis violaceopicta Poujade, 1884; Bull. Soc. Ent. Fr. (6) 4 : clviii
Lethe calisto Leech, 1891; Entomologist 24 (Suppl.) : 23, TL: Omei-Shan
Lethe violaceopicta, Butterflies in Indo-China [Yutaka Inayoshi]
Lethe violaceopicta ; [BOR, 414]; [MRS, 333]
Zophoessa violaceopicta ; Huang, 2003, Neue Ent. Nachr. 55: 91 (note), f. 143

Lethe kanjupkula Tytler, 1914
-. See [About maps] Lethe kanjupkula Tytler, 1914; J. Bombay nat. Hist. Soc. 23 (2): 220, 23 (3): pl. 1, f. 5-6, TL: Naga Hills
L. k. burmana Tytler, 1939
NE.Burma (Sadon), Manipur Lethe violaceopicta burmana Tytler, 1939; J. Bombay nat. Hist. Soc. 41 (2): 245, TL: NE.Burma
Lethe violaceopicta burmana ; [BOR, 414]
Zophoessa kanjupkula burmana ; Huang, 2003, Neue Ent. Nachr. 55: 91, f. 142

Lethe tristigmata Elwes, 1887
Sikkim, ?Nepal. See [About maps] Spotted Mystic
Lethe tristigmata Elwes, 1887; Proc. zool. Soc. Lond. 1887 : 444, TL: Sikkim
Lethe lyncus de Nicéville, 1897; J. Asiat. Soc. Bengal 66 Pt.II (3) : 544, pl. 1, f. 8, TL: Sikkim
= ; [NHM]
Lethe tristigmata ; de Nicéville, 1889, J. Bombay nat. Hist. Soc. 4 (3) : 163, pl. A, f. 4 ♀; [BOR, 414]
Zophoessa tristigmata ; Huang, Wu & Yuan, 2003, Neue Ent. Nachr. 55: 152 (note)

Lethe gemina Leech, 1891
W.China, Naga Hills. See [About maps] Tytler's Treebrown
Lethe gemina Leech, 1891; Entomologist 24 (Suppl.) : 24, TL: Moupin
Lethe gemina var. wilemani Matsumura, 1909; Ent. Zs. 23 (19) : 91, TL: Formosa
= ; [NHM]
gafuri Tytler, 1924²; J. Bombay nat. Hist. Soc. 29: 530, pl. 12, f. D 3, 1, TL: Naga Hills
= ; [NHM]
chekiangensis Mell, 1935; Mitt. Dtsch. Ent. Ges. 6 : 37
= ; [NHM]
Lethe gemina ; [BOR, 428]; [MRS, 342]
L. g. zaitha Fruhstorfer, 1914
Taiwan Lethe gemina zaitha Fruhstorfer, 1914; Ent. Rundschau 31 (5) : 25, TL: Formosa
Lethe gemina zaitha ; [MRS, 342]

The ? species-group

Lethe armandina (Oberthür, 1881)
W.China, Assam - N.Burma. See [About maps] Chinese Labyrinth
Debis armandina Oberthür, 1881; Étud. d'Ent. 6: 16, pl. 7, f. 6
Zophoessa armandina ; Huang, Wu & Yuan, 2003, Neue Ent. Nachr. 55: 151 (note)

Lethe albolineata (Poujade, 1884)
Yang-tes-kiang basin Debis albolineata Poujade, 1884; Bull. Soc. Ent. Fr. (6) 4 : clv
Lethe albolineata ; [BOW: pl. 201, f. 7]; [MRS, 335]

Lethe andersoni (Atkinson, 1871)
Assam (Cherrapunji), Upper Burma, W.China. See [About maps] Zophoessa andersoni Atkinson, 1871; Proc. zool. Soc. Lond. 1871 : 215, pl. 12, f. 3, TL: Yunnan
Lethe andersoni ; [BOR, 427]; [MRS, 336]; Huang, 2003, Neue Ent. Nachr. 55: 91 (note)

Lethe arete (Stoll, [1780])
-. See [About maps] Papilio arete Stoll, [1780]; in Cramer, Uitl. Kapellen 4 (26b-28): pl. 313, f. E, F, TL: Amboina
Papilio arete ; [BOR, 417]
Ambon, Serang Lethe arete arete ; [BAUR, 278]
L. a. advipa Fruhstorfer, 1911
Bachan, Halmahera, Ternate arete advipa Fruhstorfer, 1911; in Seitz, Gross-Schmett. Erde 9 : 316
Lethe arete advipa ; [BAUR, 277]
L. a. nagaraja Fruhstorfer, 1911
S.Sulawesi, Selajar arete nagaraja Fruhstorfer, 1911; in Seitz, Gross-Schmett. Erde 9 : 316
Lethe arete nagaraja ; [BOR, 417]
Lethe europa nagaraja ; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 176
L. a. arcuata Butler, 1868
N.Sulawesi, Lutungan I. Lethe arcuata Butler, 1868; Cat. diurn. Lep. Satyridae Brit. Mus. : 114, pl. 2, f. 3, TL: Celebes
Lethe arete arcuata ; [BOR, 417]
Lethe europa arcuata ; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 176
L. a. anatha Fruhstorfer, 1911
Sula Is. arete anatha Fruhstorfer, 1911; in Seitz, Gross-Schmett. Erde 9 : 316, TL: Sula I.
Lethe arete anatha ; [BOR, 417]
Lethe europa anatha ; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 176
L. a. velitra Fruhstorfer, 1911
Sanghie arete velitra Fruhstorfer, 1911; in Seitz, Gross-Schmett. Erde 9 : 316, TL: Sangir
Lethe arete velitra ; [BOR, 417]
Lether europa velitra ; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 176

Lethe argentata (Leech, 1891)
W.China. See [About maps] Zophoessa argentata Leech, 1891; Entomologist 24 (Suppl.) : 1, TL: Wa-Shan; Chia-Ting-Fu; Huang-Mu-Chang
Lethe argentata ; [MRS, 336]

?Lethe armanderia Oberthür²
Lethe armanderia ; [BOW: pl. 201, f. 14 (text only)]

A mispelling of Lethe armandina?


 

Lethe ocellata (Poujade, 1885)
Sichuan, Tibet, Sikkim, Manipur. See [About maps] Dismal Mystic
Debis ocellata Poujade, 1885; Bull. Soc. Ent. Fr. (6) 5 : x, TL: Mou-pin, Sichuan
Lethe simulans Leech, 1891; Entomologist 24 (Suppl.) : 23, TL: Omei-Shan
Lethe ocellata ; Leech, [1892], Butts China Japan Corea (1): 34-35; Seitz, [1909], Gross-Schmett. Erde 1 : 85, pl. 31c; Draeseke, 1925, Dt. ent. Z. Iris 39: 52; Lee & Zhu, 1992, Atlas of Chinesee butteflies: 93-94; [MRS, 333]
Zophoessa ocellata ; de Lesse, 1956, Ann. Soc. ent. Fr. 125 : 78; Huang, 2002, Atalanta 33 (3/4): 365, pl. 1, f. 4, 8; Huang, 2003, Neue Ent. Nachr. 55: 91 (note), f. 140; Huang, Wu & Yuan, 2003, Neue Ent. Nachr. 55: 145, pl. 12, f. 1
L. o. mon Yoshino, 2008
N.Vietnam Lethe ocellata mon Yoshino, 2008; Futao (54): 10, pl. 2, f. 11-12, TL: Sapa, 2000m, N.Vietnam
Lethe ocellata mon, Butterflies in Indo-China [Yutaka Inayoshi]

Lethe nigrifascia Leech, 1890
China (Hubei, Hunan, Henan, Shaanxi, Gansu, Sichuan). See [About maps] Lethe nigrifascia Leech, 1890; Entomologist 23 : 28, TL: Hubei, Chang-yang
Lethe nigrifascia ab. fasciata Seitz, [1909]; Gross-Schmett. Erde 1 : 86, pl. 31d, TL: Sichuan, Pu-tsu-fong
Lethe nigrifascia ; Leech, [1892], Butts China Japan Corea (1): 33; Seitz, [1909], Gross-Schmett. Erde 1 : 85-86, pl. 31e; Huang, 1999, Lambillionea XCIX (1): 129-131, f. 1b, 2b, 3b; Huang, 1998, Neue Entomologische Nachrichten, 41: 225 (note), pl. 4, f. 3c, 4c; [BOW: pl. 202, f. 6]; [MRS, 334]
Zophoessa nigrifascia ; de Lesse, 1956, Ann. Soc. ent. Fr. 125 : 79; Huang, 2003, Neue Ent. Nachr. 55: 91 (note), f. 138; Huang, Wu & Yuan, 2003, Neue Ent. Nachr. 55: 148, pl. 12, f. 2

Lethe baoshana (Huang, Wu & Yuan, 2003)
W.Yunnan (Bao-shan). See [About maps] Zophoessa baoshana Huang, Wu & Yuan, 2003; Neue Ent. Nachr. 55: 151, f. 1, pl. 12, f. 6, TL: Yi-wan-shui Forest Zon, Bao-shan, W.Yunnan, 2480m
Lethe baoshana? ; Yoshino, 2008, Futao (54): pl. f. 15-16

Lethe zhangi (Huang, Wu & Yuan, 2003)
W.Sichuan (Li-xian). See [About maps] Zophoessa zhangi Huang, Wu & Yuan, 2003; Neue Ent. Nachr. 55: 151, f. 2, pl. 12, f. 7-8, TL: Mi-ya-luo, Li-xian, W.Sichuan, 2780-3300m

Lethe baileyi South, 1913
China (NW.Yunnan, SE.Tibet). See [About maps] Lethe baileyi South, 1913; J. Bomb. nat. Hist. Soc. 22 (2): 346, TL: SE.Tibet, Loma
Lethe baileyi ; Huang, 1998, Neue Entomologische Nachrichten, 41: 251, pl. 4, f. 3d, 4d; Huang, 1999, Lambillionea XCIX (1): 131, f. 1d, 2d, 3d; Huang, 2000, Lambillionea, C (1): 154
Zophoessa baileyi ; de Lesse, 1956, Ann. Soc. ent. Fr. 125 : 79; Huang, 2002, Atalanta 33 (3/4): 365-366, pl. 21, f. 3, 7; Huang, 2003, Neue Ent. Nachr. 55: 91 (note), f 136; Huang, Wu & Yuan, 2003, Neue Ent. Nachr. 55: 148

Lethe neofasciata Lee, 1985
Yunnan. See [About maps] Lethe neofasciata Lee, 1985; Entomotaxonomia 7 (3): 193, pl. 2, f. 11-12, TL: Pianma, Gaoligongshan
Lethe nujiangensis Yoshino, 1997; Neo Lepidoptera 2: 5, f. 34-35, 58, 60, TL: Gaolongonshan, W slope of Nujiang Valley, Yunnan
= ?; Huang, 2001, Neue Entomologische Nachrichten, 51: 95
= ; Huang, Wu & Yuan, 2003, Neue Ent. Nachr. 55: 150
Lethe neofasciata ; Huang, 2001, Neue Entomologische Nachrichten, 51: 95 (note)
Zophoessa neofasciata ; Huang, 2002, Atalanta 33 (3/4): 365-366, pl. 21, f. 2, 6; Huang, 2003, Neue Ent. Nachr. 55: 91, f. 134-135; Huang, Wu & Yuan, 2003, Neue Ent. Nachr. 55: 150, pl. 12, f. 3-4

Lethe wui Huang, 1999
SE.Tibet (Metok). See [About maps] Lethe wui Huang, 1999; Lambillionea XCIX (1): 129-131, TL: SE.Tibet, Metok
Lethe wui ; Huang, 2000, Lambillionea, C (1): 155
Zophoessa wui ; Huang, 2003, Neue Ent. Nachr. 55: 91 (note), f. 139; Huang, Wu & Yuan, 2003, Neue Ent. Nachr. 55: 150, pl. 12, f. 5
Misidentified/misapplied:
Lethe nigrifasciata ssp.; Huang, 1998, Neue Entomologische Nachrichten, 41: 224-225, f. 8b, 251, pl. 4, f. 3a, 4a

Lethe lisuae (Huang, 2002)
NW.Yunnan (Gong-shan). See [About maps] Zophoessa lisuae Huang, 2002; Atalanta 33 (3/4): 365-366, pl. 21, f. 1, 5, TL: NW.Yunnan, Nujian valley, Gong-shan
Zophoessa lisuae ; Huang, 2003, Neue Ent. Nachr. 55: 91 (note), f. 137; Huang, Wu & Yuan, 2003, Neue Ent. Nachr. 55: 150

 

Lethe uemurai (Sugiyama, 1994)
Zophoessa uemurai Sugiyama, 1994
Zophoessa uemurai ; Huang, Wu & Yuan, 2003, Neue Ent. Nachr. 55: 151 (note)

Lethe bojonia Fruhstorfer, 1913
Taiwan Lethe (Kerrata) ocellata bojonia Fruhstorfer, 1913; Dt. ent. Z. Iris 27 (3) : 137, TL: Formosa, Jujimichi
Lethe ocellata bojonia ; [BOR, 414]; [MRS, 333]
Lethe bojonia ; Huang, Wu & Yuan, 2003, Neue Ent. Nachr. 55: 148 (note)

Lethe bipupilla Chou & Zhao, 1994
Sichua Lethe bipupilla Chou & Zhao, 1994; in Chou, Monographia Rhopalocerum Sinensium 1-2: 755, 341, f. 17, TL: Dayi (1000m), Sichuan

Lethe butleri Leech, 1889
China. See [About maps] Lethe butleri Leech, 1889; Trans. ent. Soc. Lond. 1889 (1) : 99, pl. 8, f. 3, TL: Kiukiang
Mycalesis turpilius Oberthür, 1890; Étud. d'Ent. 13 : 43, pl. 9, f. 101
= ; [NHM]
kuatunensis Mell, 1942; Arch. Naturgesch. (N.F.) 11 : 279
= ; [NHM]
Lethe butleri ; [BOR, 428]; [MRS, 342]
C.China, W.China Lethe butleri butleri ; [BOR, 428]
L. b. periscelis (Fruhstorfer)
Taiwan ?Mycalesis periscelis Fruhstorfer, 1908; Ent. Zs. 22 (35) : 141, TL: Formosa
marginalis Matsumura²
Lethe butleri periscelis ; [BOR, 428]; [MRS, 342]

Lethe callipteris (Butler, 1877)
Japan. See [About maps] Neope callipteris Butler, 1877; Ann. Mag. nat. Hist. (4) 19 (109) : 92
?suffusa Esaki, 1924²; Insec. Inscit. Mens. 12: 56
= ; [NHM]
diluta Esaki, 1924²; Insec. Inscit. Mens. 12: 56, pl. 3, f. 3-4, TL: Hokkaido
= ; [NHM]
Harima callipteris ; [BRU, 184]
Zophoessa callipteris ;
[callipteris-1.jpg] 720x670 (~52Kb) The Kojoro Lake (Kojoroga-ike) bank on Mt. Hira, 1060 m altitude, Katsuragawa Sakashita-cho, Ohtsu Cty, Shiga Prefecture, Honshu, Japan. 2nd August 2002, Photo © Oleg Kosterin [callipteris-1v.jpg] 636x636 (~58Kb) underside The Kojoro Lake (Kojoroga-ike) bank on Mt. Hira, 1060 m altitude, Katsuragawa Sakashita-cho, Ohtsu Cty, Shiga Prefecture, Honshu, Japan. 2nd August 2002, Photo © Oleg Kosterin

Larva on Sasa [BRU, Fukuda et al., 1983]


L. c. karafutonis Matsumura, 1919
Sakhalin callipteris karafutonis Matsumura, 1919; [?]
minima Esaki, 1924; Insec. Inscit. Mens. 12: 56, pl. 3, f. 8, TL: Sakhalin
?minima Esaki & Nakahara, 1924
L. c. obscura Nakahara, 1926
Kuriles callipteris obscura Nakahara, 1926; Ins. Insec. Mens 14: 50, TL: Kuriles

Lethe camilla Leech, 1891
W.China. See [About maps] Lethe camilla Leech, 1891; Entomologist 24 (Suppl.) : 3, TL: Wa-Shan; Chia-Kou-Ho; Chia-Ting-Fu
rufa Mell, 1939; Dt. Ent. Z. Iris 52 : 139, TL: Fukien
= ; [NHM]
Lethe camilla, Butterflies in Indo-China [Yutaka Inayoshi]

Lethe christophi Leech, 1891
C.China, W.China. See [About maps] Lethe christophi Leech, 1891; Entomologist 24 (Suppl.) : 67, TL: Omei-Shan, Chang-Yang
obscura Mell, 1942; Arch. Naturgesch. (N.F.) 11 : 276
= ; [NHM]
Lethe christophi ; [BOW: pl. 201, f. 10]; [BOR, 426]; [MRS, 337]
L. c. hanako Fruhstorfer, 1908
Taiwan Lethe christophi hanako Fruhstorfer, 1908; Insekten-Börse 25: 38, TL: Formosa
Lethe christophi hanako ; [BOR: 426]; [MRS, 337]

Lethe clarissa Murayama, 1982²
W.China, Szechwan. See [About maps] clarissa Murayama, 1982²; J. Ent. Soc. Shinshu 31 (4): 2

Lethe cybele Leech, 1894
Tibet, W.China. See [About maps] Lethe cybele Leech, 1894; Butts China Japan Corea (2): 643 (appendix)
Lethe cybele ; [MRS, 333]

Lethe cyrene Leech, 1890
W.China. See [About maps] Lethe cyrene Leech, 1890; Entomologist 23 : 27, TL: Chang Yang
Lethe cyrene ; [MRS, 337]

Lethe dakwania Tytler, 1939
N.India, Kashmir. See [About maps] dakwania Tytler, 1939; J. Bombay nat. Hist. Soc. 41 (2): 245, TL: Burma

Lethe darena (C. & R. Felder, [1867])
Java, Sumatra, Borneo. See [About maps] Debis darena C. & R. Felder, [1867]; Reise Fregatte Novara, Bd 2 (Abth. 2) (3): 498, pl. 68, f. 4-5, TL: Java
Lethe darena ; de Nicéville, 1893, J. Bombay Nat. Hist. Soc. 8 (1): 40, pl. K, f. 7; [BOW: pl. 167, f. 1]; [BOR, 424]
W.Java (Gunong Gede) Lethe darena darena ; [BOR, 424]
L. d. sumatrensis Staudinger, [1897]
Sumatra (Battak Mts) Lethe darena var. sumatrensis Staudinger, [1897]; Dt. Ent. Z. Iris 9 (2): 228
Lethe darena sumatrensis ; [BOR, 424]
L. d. borneensis Staudinger, [1897]
Borneo Lethe darena var. borneensis Staudinger, [1897]; Dt. Ent. Z. Iris 9 (2): 228, TL: Kina Balu
Lethe darena borneensis ; [BOR, 424]

Lethe dataensis (Semper, 1887)
Philippines (Luzon, Mindoro). See [About maps] dataensis (Semper, 1887) (Debis?); Reisen Philipp. (2): 57, pl. 11, f. 4, TL: NW.Luzon
= Lethe dura ; [NHM]
daatensis[sic, recte dataensis] ; Fruhstorfer, 1911, in Seitz, Gross-Schmett. Erde 9 : 315
Lethe dataensis ; [BOR, 415]
L. d. halcona (Yamamoto & Takei, 1993)
Philippines (Mindanao) Zophoessa dataensis halcona Yamamoto & Takei, 1993; Tyô to Ga 43 84) : 255, f. 1-2, TL: Mt. Halcon (1500m), N.Mindanao

Lethe delila Staudinger, [1897]
Borneo. See [About maps] Lethe delila Staudinger, [1897]; Dt. Ent. Z. Iris 9 (2): 225, pl. 5, f. 1, TL: Kina Balu
Lethe delila ; [BOR, 421]

Lethe diana (Butler, 1866)
E.China, S.Ussuri, Sakhalin, Kurile Is., Japan, Korea. See [About maps] Debis diana Butler, 1866; J. Linn. Soc. Zool. Lond. 9 (34) : 55, TL: Hakodate, Hokkaido, Japan
Lethe whitelyi Butler, 1867; Ann. Mag. nat. Hist. (3) 20 (120) : 403, pl. 9, f. 8, TL: Nagasaki (North Japan)
= ; [NHM]
fixseni Butler, 1881²; Ann. Mag. Nat. Hist. (5) 7 (38) : 133 [ ?? bad ref], TL: Korea
= ; [NHM]
Lethe consanguis Butler, 1881; Ann. Mag. Nat. Hist. (5) 7 (38) : 133, TL: Japan
= ; [NHM]
celeja Fruhstorfer, 1911; in Seitz, Gross-Schmett. Erde 9 : 322, TL: Hondo
= ; [NHM]
Lethe diana ; [BRU, 183]; [BOW: pl. 201, f. 11]; [MRS, 339]

Larva on Phragmites japonicus? [BRU, Fukuda et al, 1983]

L. d. sachalinensis Matsumura, 1925
S.Sakhalin diana sachalinensis Matsumura, 1925; J. Coll. Agric. Hokkaido imp. Univ. 15 : 92, pl. 9, f. 14
L. d. tomariope Bryk, 1942
Kurile Is. diana tomariope Bryk, 1942; Dt. Ent. Z. Iris 56 : 14
L. d. mikuraensis Shirozy, 1975
Japan (Honsu)

Lethe dora Staudinger, [1897]
Sumatra, Borneo. See [About maps] Lethe dora Staudinger, [1897]; Dt. Ent. Z. Iris 9 (2): 228, TL: SE.Borneo
cerama Shelford, 1904²; Straits asiat. Soc. 41: 96
= ; [NHM]
Lethe dora ; [BOR, 422]

Lethe dura (Marshall, 1882)
Himalays, W.China, Formosa. See [About maps] Zophoessa dura Marshall, 1882; J. Asiat. Soc. Bengal 51 Pt.II (2-3) : 38, pl. 4, f. 2, TL: Upper Tenasserim
Lethe dura dura, Butterflies in Indo-China [Yutaka Inayoshi]
Lethe dura ; [BOW: pl. 201, f. 13]; [BOR, 412]; [MRS, 322]
Assam, Burma, ?Thailand Lethe dura dura ; [BOR, 412]
Zophoessa dura dura ; Huang, 2003, Neue Ent. Nachr. 55: 91 (note)
L. d. gammiei (Moore, [1892])
NW.India, Bhutan, Sikkim dura gammiei (Moore, [1892]) (Zophoessa); Lepidoptera Indica 1 : 294, pl. 91, f. 3, TL: Bhutan
Lethe dura gammiei ; [BOR, 412]
L. d. mansonia Fruhstorfer, 1911
Indo China Lethe dura mansonia Fruhstorfer, 1911; in Seitz, Gross-Schmett. Erde 9 : 314, pl. 99 a, TL: Manson Mts., N.Vietnam
Lethe dura mansonia, Butterflies in Indo-China [Yutaka Inayoshi]
Lethe dura mansonia ; [BOR, 412]
L. d. moupiniensis (Poujade, 1884)
W.China Debis moupiniensis Poujade, 1884; Bull. Soc. Ent. Fr. (6) 4 : cxl
Zophoessa libitina Leech, 1891; Entomologist 24 (Suppl.) : 2, TL: Chia-Ting-Fu; Chang-Yang
Lethe dura moupinensis[sic?] ; [MRS, 322]; [BOR, 412]
L. d. neoclides Fruhstorfer, 1909
Taiwan dura neoclides Fruhstorfer, 1909; Soc. ent. 24 (16) : 121, TL: Formosa
Lethe dura neoclides ; [BOR, 412]; [MRS, 322]

Lethe gracilis (Oberthür, 1886)
Tibet. See [About maps] Pararge gracilis Oberthür, 1886; Étud. d'Ent. 11 : 23, pl. 4, f. 19, TL: Ta-Tsien-Lou

Lethe hecate Leech, 1891
W.China. See [About maps] Lethe hecate Leech, 1891; Entomologist 24 (Suppl.) : 3, TL: Wa-Shan; Chia-Ko-Hou; Huang-Mu-Chang
consobrina Watkins, 1928²; Ann. Mag. nat. Hist. (10) 1 (5) : 616
= ; [NHM]
L. h. zao Yoshino, 2008
Vietnam Lethe hecate zao Yoshino, 2008; Futao (54): 11, pl. 2, f. 17-18, TL: Sapa, 2000m, N.Vietnam
Lethe hecate zao, Butterflies in Indo-China [Yutaka Inayoshi]
L. h. haba Yoshino, 2008
N.Yunnan Lethe hecate haba Yoshino, 2008; Futao (54): 11, pl. 2, f. 19-20, TL: Zhondian County, 2000m, N.Vietnam

Lethe helena Leech, 1891
W.China. See [About maps] Lethe helena Leech, 1891; Entomologist 24 (Suppl.) : 3, TL: Chia-Ting-Fu
obscura Mell, 1942; Arch. Naturgesch. (N.F.) 11 : 279
= ; [NHM]
Lethe helena ; [MRS, 332]

Lethe helle (Leech, 1891)
W.China. See [About maps] Zophoessa helle Leech, 1891; Entomologist 24 (Suppl.) : 1, TL: Wa-Shan; Chia-Ting-Fu; Huang-Mu-Chang
Lethe helle ; [BOW: pl. 201, f. 14]; [MRS, 334]
Zophoessa helle ; Huang, Wu & Yuan, 2003, Neue Ent. Nachr. 55: 151 (note)
L. h. gregoryi Watkins, 1927
Yunnan Lethe helle gregoryi Watkins, 1927; Ann. Mag. nat. Hist. (9) 19 : 323, TL: Yunnan
Lethe helle gregoryi ; [MRS, 334]

Lethe kondoi Uemura, 1997
Laos. See [About maps] Lethe kondoi Uemura, 1997; Bull. Toyosato Mus. Ent. 6: 1-4, f. 1-9, TL: Vang Vieng, Laos
Lethe kondoi, Butterflies in Indo-China [Yutaka Inayoshi]

Lethe konkakini Monastyrskii & Devyatkin, 2000
Vietnam. See [About maps] Lethe konkakini Monastyrskii & Devyatkin, 2000; Atalanta 31 (3/4): 479, pl. XIX, f. 5-8, TL: C.Vietnam, Gia Lai, Kon Ka Kinh Nature Preserve
Lethe konkakini, Butterflies in Indo-China [Yutaka Inayoshi]
Lethe konkakini ; Huang & Wu, 2003, Neue Ent. Nachr. 55: 126

Lethe guansia Sugiyama, 1999
Guangxi, Guangdong. See [About maps] Lethe guansia ; Huang & Wu, 2003, Neue Ent. Nachr. 55: 126 (note)

Lethe labyrinthea Leech, 1890
W.China. See [About maps] Lethe labyrinthea Leech, 1890; Entomologist 23 : 28, TL: Chang Yang
Lethe labyrinthea ; [BOW: pl. 201, f. 15]; [MRS, 334]

Lethe lanaris Butler, 1877
W.China. See [About maps] Lethe lanaris Butler, 1877; Ann. Mag. nat. Hist. (4) 19 (109) : 95
conspicua Mell, 1942; Arch. Naturgesch. (N.F.) 11 : 279
= ; [NHM]
Lethe lanaris ; [BOW: pl. 202, f. 1]; [BOR, 428]; [MRS, 341]
L. l. exista Yoshino, 2008
N.Vietnam Lethe lanaris exista Yoshino, 2008; Futao (54): 10, pl. 2, f. 7-8, TL: Sapa, 2000m, N. Vietnam
Lethe lanaris exista, Butterflies in Indo-China [Yutaka Inayoshi]

Lethe laodamia Leech, 1891
W.China. See [About maps] Lethe laodamia Leech, 1891; Entomologist 24 (Suppl.) : 67, TL: Wa-Shan
Lethe laodamia ; [MRS, 338]

Lethe luteofasciata (Poujade, 1884)
China. See [About maps] Debis luteofasciata Poujade, 1884; Bull. Soc. Ent. Fr. (6) 4 : cliv
Lethe luteofasciata ; [MRS, 335]

Lethe manthara (C. & R. Felder, [1867])
Java, Bali. See [About maps] Debis manthara C. & R. Felder, [1867]; Reise Fregatte Novara, Bd 2 (Abth. 2) (3): 497, TL: Java
mangala Fruhstorfer, 1911; in Seitz, Gross-Schmett. Erde 9 : 320, pl. 98 e, TL: E.Java
= ; [NHM]
Lethe manthara ; Butler, 1867, Ann. Mag. nat. Hist. (3) 20 (120) : 403, pl. 9, f. 9; [BOR, 422]

Lethe manzorum (Poujade, 1884)
W.China. See [About maps] Satyrus manzorum Poujade, 1884; Bull. Soc. Ent. Fr. (6) 4 : cxxxiv
Lethe manzorum ; [BOW: pl. 202, f. 3]
Lethe manzora ; [MRS, 342]

Lethe marginalis Motschulsky, 1860
E.China, Amur, Korea, Japan. See [About maps] Lethe marginalis Motschulsky, 1860; Etud. Ent. 9: 29, TL: Hakodate, Hokkaido, Japan
Satyrus (Pararge) davidianus Poujade, 1885; Bull. Soc. Ent. Fr. (6) 5 : xciv, TL: Mou-Pin
= ; [NHM]
Lethe marginalis ; [BRU, 184]; [BOW: pl. 202, f. 4]; [MRS, 338]

Larva on Miscanthus sinensis, Spodiopogon sibiricus, Scirpus wichurae, Carex [BRU, Fukuda et al., 1983]

Lethe marginalis marginalis ; Huang, 2003, Neue Ent. Nachr. 55: 94 (note), f. 151
L. m. maacki (Bremer, 1861)
Amur, Ussuri marginalis maacki (Bremer, 1861) (Lasiommata); Bull. Acad. Imp. Sci. St. Petersb. 3 : 468
Yunnan Lethe marginalis obscurofasciata ; Huang, 2003, Neue Ent. Nachr. 55: 94 (note), f. 150

Lethe mataja Fruhstorfer, 1908
Taiwan Lethe mataja Fruhstorfer, 1908; Insekten-Börse 25: 38
Lethe mataja ; [BOW: pl. 202, f. 5]; [BOR, 428]; [MRS, 324]

Lethe minerva (Fabricius, 1775)
Malaya, Sumatra, Borneo, Java. See [About maps] Papilio minerva Fabricius, 1775; Syst. Ent.: 493, no. 216
Papilio arcadia Cramer, [1777]; Uitl. Kapellen 2 (9-16): pl. 116, f. E, F
= ; [NHM]
caumas (Godart, [1824]) (Satyrus); Encyclopédie Méthodique. 9 (2): 479
= ; [NHM]
Lethe minerva, NSG Voucher Specimen [Wahlberg]
Nemetis minerva ; [EBW]
Lethe minerva ; [BOW: pl. 167, f. 10]; [BOR, 421]
Java, Sumatra, Peninsular Malaya Lethe minerva minerva ; [BMP: 125, pl. 15, f. 12]; [BOR, 421]
L. m. tritogeneia Fruhstorfer, 1911
Burma, Thailand Lethe minerva tritogeneia Fruhstorfer, 1911; in Seitz, Gross-Schmett. Erde 9 : 318, TL: Tenasserim, Myanmar
Lethe minerva tritogeneia, Butterflies in Indo-China [Yutaka Inayoshi]
Lethe minerva tritogeneia ; [BOR, 421]

Lethe monilifera Oberthür, 1923
W.China. See [About maps] Lethe monilifera Oberthür, 1923; Etud. Lépid. Comp. 20: 199, f. 4866
Lethe monilifera ; [MRS, 342]

Lethe niitakana (Matsumura, 1906)
Taiwan Zophoessa insularis Fruhstorfer, 1908; Ent. Zs. 22 (12): 48, TL: Polisha
Lethe insularis ; [BOW: pl. 201, f. 12]
Lethe niitakana ; [BOR, 416]; [MRS, 335]

Larva on Arundinaria nitakayamensis [MRS]


Lethe oculatissima (Poujade, 1885)
W.China. See [About maps] Mycalesis oculatissima Poujade, 1885; Bull. Soc. Ent. Fr. (6) 5 : xxiv
Lethe occulta Leech, 1890; Entomologist 23 : 26, TL: Chang Yang
= ; [NHM]
Lethe oculatissima, NSG Voucher Specimen [Wahlberg]
Lethe oculatissima ; [BOW: pl. 202, f. 7]; [MRS, 332]

Lethe perimede Staudinger, [1897]
Borneo (Kina Balu). See [About maps] Lethe perimede Staudinger, [1897]; Dt. Ent. Z. Iris 9 (2): 226, TL: Kina Balu
Lethe perimede ; [BOR, 421]

Lethe philemon Fruhstorfer, 1911
N.Indo China, ?N.Burma, ?Assam. See [About maps] ?Lethe baucis philemon Fruhstorfer, 1902; Dt. ent. Z. Iris 14 (2) : 275, TL: Than-Moi, Tonkin
Lethe naga philemon Fruhstorfer, 1911; in Seitz, Gross-Schmett. Erde 9 : 318, pl. 97 b
Lethe philemon, Butterflies in Indo-China [Yutaka Inayoshi]
Lethe philemon ; [BOR, 426]

Lethe philesana Monastyrskii & Devyatkin, 2000
Lethe philesana Monastyrskii & Devyatkin, 2000; Atalanta 31 (3/4): 479, pl. XIX, f. 1-4, TL: N.Vietnam, Boc Can, Ba Be National Park
Lethe philesana, Butterflies in Indo-China [Yutaka Inayoshi]

Lethe privigna Leech, [1892]²
W.China. See [About maps] privigna Leech, [1892]²; Butts China Japan Corea (1): 32, pl. 5, f. 3-4
= Lethe camilla ; [NHM]

Lethe procne (Leech, 1891)
China (Yunnan, Sichuan, ...?). See [About maps] Zophoessa procne Leech, 1891; Entomologist 24 (Suppl.) : 2, TL: Wa-Shan; Huang-Mu-Chang; Ta-Tsien-Lu
Zophoessa procne ; Huang, Wu & Yuan, 2003, Neue Ent. Nachr. 55: 151 (note)

Lethe proxima Leech, [1892]
W.China. See [About maps] Lethe proxima Leech, [1892]; Butts China Japan Corea (1): 32, pl. 6, f. 8
Lethe proxima ; [BOW: pl. 202, f. 9]; [MRS, 341]

Lethe samio (Doubleday, [1849])
Java. See [About maps] Debis samio Doubleday, [1849]; Gen. diurn. Lep. (1): pl. 61, f. 3
Debis samio Westwood, [1851]; Gen. diurn. Lep. (2): 358
purana Felder²
= ; [NHM]
Lethe samio ; [BOR, 419]

Lethe satyrina Butler, 1871
W.China, Tibet - Shanghai. See [About maps] Lethe satyrina Butler, 1871; Trans. ent. Soc. Lond. 1871 (3) : 402
Lethe naias Leech, 1889; Trans. ent. Soc. Lond. 1889 (1) : 100, pl. 8, f. 4, TL: Kiukiang
= ; [NHM]
Mycalesis styppax Oberthür, 1890; Étud. d'Ent. 13 : 44, pl. 10, f. 110
= ; [NHM]
obscura Mell, 1942; Arch. Naturgesch. (N.F.) 11 : 280
= ; [NHM]
Lethe satyrina ; [BOW: pl. 202, f. 10]; [MRS, 343]

Lethe sicelides Grose-Smith, 1893
W.China. See [About maps] Lethe sicelides Grose-Smith, 1893; Ann. Mag. Nat. Hist. (6) 11 (63) : 218, TL: Omei-shan, North-west China
= Lethe hecate ; [NHM]
Lethe sicelides ; [MRS, 339]

Lethe sicelis (Hewitson, 1862)
Japan, China. See [About maps] Debis sicelis Hewitson, 1862; Ill. exot. Butts [4] (Debis I): [33], pl. [18], f. 3, TL: Japan
Lethe sicelis vanelia Fruhstorfer, 1909; Ent. Zs. 23 (25) : 118, TL: Kiu-shiu
= ; [NHM]
meridionalis Martin, 1921; Dt. ent. Z. Iris 35 (1/2) : 13, TL: Shikoku
= ; [NHM]

Lethe syrcis (Hewitson, 1863)
-. See [About maps] Debis syrcis Hewitson, 1863; Ill. exot. Butts [4] (Debis II-III): [37], pl. [20], f. 13-14, TL: North China
Debis syrcis ab. confluens Oberthür, 1913; Étud. Lépid. Comp. 7 : 669, pl. 186, f. 1820-1821, TL: Ta-tsien-lu
= ; [NHM]
sikiangensis Mell, 1942; Arch. Naturgesch. (N.F.) 11 : 277
= ; [NHM]
ochrescens Mell, 1942; Arch. Naturgesch. (N.F.) 11 : 277
= ; [NHM]
Lethe syrcis ; [BOW: pl. 168, f. 4]; [BOR, 427]; [MRS, 338]
China Lethe syrcis syrcis ; [BOR, 427]
L. s. diunaga Fruhstorfer, 1908
N.Vietnam Lethe syrcis diunaga Fruhstorfer, 1908; Ent. Zs. 22 (2) : 7, TL: Tonkin
?diunaga Fruhstorfer, 1911; in Seitz, Gross-Schmett. Erde 9 : 323, TL: Tonkin
Lethe syrcis diunaga, Butterflies in Indo-China [Yutaka Inayoshi]
Lethe syrcis diunaga ; [BOR, 427]

Lethe titania Leech, 1891
W.China (Sichuan). See [About maps] Lethe titania Leech, 1891; Entomologist 24 (Suppl.) : 67, TL: Pt-su-fong; Kia-ting-fu
Lethe titania ; [BOW: pl. 202, f. 11]; [MRS, 339]

Lethe trimacula Leech, 1890
W.China. See [About maps] Lethe trimacula Leech, 1890; Entomologist 23 : 27, TL: Chang Yang
pekiangensis Mell, 1935; Mitt. Dtsch. Ent. Ges. 6 : 37
= ; [NHM]
kuatunensis Mell, 1942; Arch. Naturgesch. (N.F.) 11 : 277
= ; [NHM]
Lethe trimacula ; [MRS, 341]

Lethe umedai Koiwaya, 1998
Sichuan. See [About maps] Lethe umedai Koiwaya, 1998; , TL: Sichuan
Lethe umedai, Butterflies in Indo-China [Yutaka Inayoshi]
L. u. albofasciata Huang, 2002
Yunnan Lethe umedai albofasciata ; Huang, 2003, Neue Ent. Nachr. 55: 93 (note), f. 144-145

Lethe liae Huang, 2002
Yunnan. See [About maps] Lethe liae ; Huang, 2003, Neue Ent. Nachr. 55: 93 (note), f. 146

Lethe violae Tsukada & Nishiyama, 1979
C.Sulawesi (Parado). See [About maps] Violet Tree-brown
Lethe violae Tsukada & Nishiyama, 1979; Mem. Tsukada Coll. 1: 13
Lethe violae ; [BOR, 418]; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 176

Lethe yantra Fruhstorfer, 1914
W.China. See [About maps] Lethe yantra Fruhstorfer, 1914; Ent. Rundschau 31 (5) : 25, TL: Yunnan
Lethe yantra ; [MRS, 334]
L. y. distincta Mell, 1923
Lethe yantra distincta Mell, 1923; Dt. ent. Z. 1923 (2) : 154
= Lethe callipteris ; [NHM]
Lethe yantra distincta ; [MRS, 334]

Lethe yunnana D'Abrera
Lethe yunnana ; [MRS, 336]

Lethe kinabalensis Okubo, 1979
N.Borneo. See [About maps] Lethe kinabalensis Okubo, 1979; Tyô to Ga 30 (1,2): 91, f. 1-2, TL: Mt. Kinabalu, 5800ft, Sabah
Lethe kinabalensis ; [NHM]


15.2.2013 (114)

References:
[About maps]
Warning! The maps are automatically generated from the textual information, and the process does not always produce acceptable result; See about maps for more info.
[BAUR]
D'Abrera, 1977; Butterflies of the Australian Region (2nd edn)
[BHL]
The; Biodiversity Heritage Library; and on; Internet Archive; Note: The linking to the content of Internet Archive is somewhat delicate -- notify me when links break!
[BIR]
Wynter-Blyth, 1957; Butterflies of the Indian Region; (1982 Reprint)
[BMP]
Eliot (Ed.), Corbet & Pendlebury, 1992; The Butterflies of The Malay Peninsula; 4th edition; ISBN 983-9681-05-2
[BOR]
D'Abrera, 1982, 1985, 1986; Butterflies of the Oriental Region, Part I-III
[BOW]
Lewis, 1974; Butterflies of the World; ISBN 0 245 52097 X
[BRU]
Tuzov, Bogdanov, Devyatkin, Kaabak, Korolev, Murzin, Samodurov, Tarasov, 1997; Guide to the Butterflies of Russia and adjacent territories; Hesperiidae, Papilionidae, Pieridae, Satyridae; Volume 1
[BnF]
Gallica / Bibliothèque nationale de France; Annales de la Société entomologique de France; Bulletin de la Sociéte entomologique de France
[CiNii]
http://ci.nii.ac.jp/
[EBW]
Smart, Paul, 1976; The Illustrated Encyclopedia of the Butterfly World; ISBN 0 600 31381 6
[HBN]
Francis Hemming, 1937; HÜBNER, A bibliographical and systematic account of the entomological works of Jacob Hübner and of the supplements thereto by Carl Geyer, Gottfried Franz von Frölich and Gottlieb August Wilhelm Herrich-Schäffer, Vol 1 & 2
[Kedar Tokekar]
India
[LAMAS-2005]
Gerardo Lamas; Entomofauna 26 (6): 57-100; A Bibliography of the Zoological Publications of Hans Fruhstorfer (1886* - 1922+)
[LAMAS-NEO]
Gerardo Lamas; Bibliography of butterflies; An Annocated Bibliography of the Neotropical Butterflies and Skipper (Lepididoptera: Papilionoidea and Hesperioidea); Revised Electronic Edition (2007)
[MRS]
Chou Io (Ed.); Monographia Rhopalocerum Sinensium, 1-2
[NHM]
Natural History Museum; The Global Lepidoptera Names Index
[Oleg Kosterin]
Institue of Cytology & Genetics; Siberian Division of Russian Academy of Sciences; Novosibirsk; e-mail: Oleg Kosterin; http://pisum.bionet.nsc.ru/kosterin/
[Wahlberg]
Niklas Wahlberg; The NSG's voucher specimen database; Nymphalidae Systematics Group
[Yutaka Inayoshi]
A Check List of Butterflies in Indo-China; Chiefly from THAILAND, LAOS & VIETNAM
[²]
This may require parentheses or not. I don't have the necessary information for this taxon.


Some related literature:
Atkinson, 1871
Descriptions of three New Species of Diurnal Lepidoptera from Western Yunan collected by Dr. Anderson in 1868 Proc. zool. Soc. Lond. 1871 : 215-216, pl. 12
Aurivillius, 1897
Bemerkungen zu den von. J. Chr. Fabricius aus danischen Sammlungen beschriebenen Lepidopteren Ent. Tidskr. 18 (3/4) : 139-174
Billberg, 1820
Enumeratio Insectorum Enum. Ins. Mus. Billb. : 1-138
Bingham, 1905
Butterflies. Vol. I. Fauna of British India Fauna Brit. Ind., Butts (ed. 1) 1: xxii, 511pp., 10pls
Bremer, 1861
Neue Lepidopteren aus Ost-Sibirien und dem Amur-Lande gesammelt von Radde und Maack, bescrieben von Otto Bremer Bull. Acad. Imp. Sci. St. Petersb. 3 : 461-496, Mélanges biol. St.-Pétersb. 3 (5-6): 538-589
Bryk, 1942
Zur Kenntnis der Großschmetterlinge der Kurilen Dt. Ent. Z. Iris 56 : 1-90, pl. 1-2
Butler, 1866
A list of the diurnal Lepidoptera recently collected by Mr. Whitely in Hakodadi (North Japan) J. Linn. Soc. Zool. Lond. 9 (34) : 50-58
Butler, 1866
Descriptions of some new exotic butterflies in the National Collection Proc. zool. Soc. Lond. 1866 (1) : 39-42, pl. 3
Butler, 1866
Descriptions of some new species of diurnal Lepidoptera in the collection of the British Museum Ent. mon. Mag. 3 (28) : 76-78
Butler, 1867
Description of new or little-known Species of Asiatic Lepidoptera Ann. Mag. nat. Hist. (3) 20 (120) : 399-404, pl. 8-9
Butler, 1868
A catalogue of diurnal Lepidoptera of the family Satyridae in the collection of the British Museum Cat. diurn. Lep. Satyridae Brit. Mus. : 1-211, pl. 1-5
Butler, 1870
Descriptions of some new diurnal Lepidoptera, chiefly Hesperiidae Trans. ent. Soc. Lond. 1870 (4) : 485-520
Butler, 1871
Descriptions of five new species, and a new genus , of diurnal Lepidoptera, from Shanghai Trans. ent. Soc. Lond. 1871 (3) : 401-403
Butler, 1871
Descriptions of some new Species of Lepidoptera chiefly from the Collection of Mr Wilson Saunders Ann. Mag. nat. Hist. (4) 8 (46) : 282-290
Butler, 1877
On Rhopalocera from Japan and Shanghai, with Descriptions of new Species Ann. Mag. nat. Hist. (4) 19 (109) : 91-97
Butler, 1881
On a Collection of Butterflies from Nikko, Central Japan Ann. Mag. Nat. Hist. (5) 7 (38) : 132-140
Corbet, 1941
A further contribution to a knowledge of the Rhopalocera of the Malay Peninsula J. fed. Malay St. Mus. 18 (5): 805-313
Corbet, 1941
Further revisional Notes on Malayan Rhopalocera Entomologist 74 (6): 133-135
Cramer, [1775]
Uitlandsche Kapellen (Papillons exotiques) Uitl. Kapellen 1 (1-7): 1-132, pl. 1-84 (1775), (8): 133-155, pl. 85-96 (1776)
Cramer, [1777]
Uitlandsche Kapellen (Papillons exotiques) Uitl. Kapellen 2 (9-16): 1-152, pl. 97-192 (1777)
de Nicéville, 1886
On some New Indian Butterflies J. Asiat. Soc. Bengal 55 Pt.II (3) : 249-256, pl. 11
de Nicéville, 1887
Descriptions of some new or little-known Butterflies from India, with some Notes on the seasonal Dimorphism obtaining in the Genus Melanitis Proc. zool. Soc. Lond. 1887 (3) : 448-467, pl. 39-40
de Nicéville, 1887
Description of a new Satyrid from India Proc. Asiat. Soc. Beng. 1887 : 147
de Nicéville, 1889
On new and little-known Butterflies from the Indian Region, with Revision of the Genus Plesioneura of Felder and of Authors J. Bombay nat. Hist. Soc. 4 (3) : 163-194, pl. A-B
de Nicéville, 1893
On new and little-known Butterflies from north-east Sumatra collected by Hofrath Dr. L. Martin J. Bombay Nat. Hist. Soc. 8 (1): 37-56, pl. K-M
de Nicéville, 1897
On new or little-known Butterflies from the Indo- and Austro-Malayan Regions J. Asiat. Soc. Bengal 66 Pt.II (3) : 543-577, pl. 1-4
de Nicéville, 1900
On new and little-known Butterflies, from Oriental region J. Bombay nat. Hist. Soc. 13 (1) : 157-176, pl. CC,DD,EE (1900)
Doherty, 1886
A list of Butterflies taken in Kumaon J. Asiat. Soc. Bengal 55 Pt.II (2) : 103-140
Doherty, 1889
Notes on Assam butterflies J. asiat. Soc. Bengal 58 Pt.II (1) : 118-134
Doubleday, [1849]; Westwood, [1851]
The genera of diurnal Lepidoptera, comprising their generic characters, a notice of their habitats and transformations, and a catalogue of the species of each genus; illustrated with 86 plates by W. C. Hewitson Gen. diurn. Lep. (1): 1-18, pl. A, 1-4 (1-2) (1846), (1): 19-132, pl. 4*, 5-25, 28 (3-14) (1847), (1): 133-200, pl. 1*, 26, 27, 29, 31-44 (15-23) (1848), (1): 201-242, pl. 30, 45-52, 56-58, 60-62, 64 (24-31) (1849), (1): 243-250 [?], (2): 243-326, pl. 53-55, 63, 65, 66 (32-38) (1850), (2): 327-466, pl. 59, 67-77 (39-50) (1851), (2): 467-534, 143, 144, pl. 78-80 (Index) (51-54) (1852)
Draeseke, 1925
Die Schmetterlinge der Stötznerschen Ausbeute (2,3,4) Dt. ent. Z. Iris 38: 1-8, 39: 48-57, 39: 211-231
Druce, 1875
Descriptions of new Species of diurnal Lepidoptera Cistula ent. 1 (12) : 357-363
Elwes, 1882
On a collection of Butterflies from Sikkim Proc. zool. Soc. Lond. 1882 (4) : 398-407, pl. 25
Elwes, 1887
Description of some new Lepidoptera from Sikkim Proc. zool. Soc. Lond. 1887 : 444-446
Elwes, 1891
On Butterflies collected by Mr. W. Doherty in the Naga and Karen Hills and in Perak. (1) and (2) Proc. zool. Soc. Lond. 1891 (2): 249-289, pl. 27, 1892 (4): 617-664, pl. 43-44
Evans, 1932
The Identification of Indian Butterflies (edn. 2) Indian Butterflies (edn. 2): x + 454pp, 9 figs, 32 pls
Fabricius, 1775
Systema Entomologiae, sistens Insectorum Classes, Ordines, Genera, Species, Adiectis Synonymis, Locis, Descriptionibus, Observationibus. Syst. Ent.: 832pp
C. & R. Felder, 1859
Lepidopterologische Fragmente. I-IV Wien. ent. Monats. 3 (6): 178-186, pl. 3-4, (9): 263-273, pl. 5, (11): 321-328, pl. 6, (12): 390-405, pl. 7-9
C. & R. Felder, 1863
Lepidoptera nova a Dr. Carolo Semper in Insulis Philippinis collecta diagnosibus. (3) Wien. ent. Monats. 7 (4) : 105-127
C. & R. Felder, [1867]
Reise der österreichischen Fregatte Novara um die Erde in den Jahren 1857, 1858, 1859 unter den Behilfen des Commodore B. von Wüllerstorf-Urbair. Zoologischer Theil. Band 2. Abtheilung 2. Lepidoptera. Rhopalocera Reise Fregatte Novara, Bd 2 (Abth. 2) (1): 1-136, pl. 1-21 (Rhop.) (C. & R. Felder, [1865]), (2): 137-378, pl. 22-47 (Rhop.) (C. & R. Felder, 1865), (3): 379-536, pl. 48-74 (Rhop.) (C. & R. Felder, 1867), (4): 537-548, pl. 75-120, (Inhalts-Verz.) 1-9 (pl. 1-74), (Felder & Rogenhofer, 1874) , (5): pl. 121-140, (erklärung) 1-10 (pl. 75-107), (Inhalts-Verz.) 1-20 (pl. 108-140), (Felder & Rogenhofer, 1875)
Fruhstorfer, 1902
Neue und seltene Lepidopteren aus Annam und Tonkin und dem malayischen Archipel Dt. ent. Z. Iris 14 (2) : 265-276, pl. 5 (1 March)
Fruhstorfer, 1908
Eine neue Lethe-Rasse aus Tonkin Ent. Zs. 22 (2) : 7 (11 April)
Fruhstorfer, 1908
Lepidopterologisches Pêle-Mêle. I. Neue ostasiatische Rhopaloceren Ent. Zs. 22 (11): 46-47 (13 June), (12): 48-49 (20 June)
Fruhstorfer, 1908
Neue ostasiatische Satyriden Ent. Zs. 22 (31) : 127 (31 October)
Fruhstorfer, 1908
Lepidopterologisches Pêle-Mêle. X. Neue Rhopaloceren von Formosa Ent. Zs. 22 (35) : 140-141 (28 November)
Fruhstorfer, 1909
Lepidopterologisches Pêle-Mêle. [XV]. Neue Rhopaloceren von Formosa Ent. Zs. 23 (8) : 40 (22 May)
Fruhstorfer, 1909
Neue japanische Satyriden Ent. Zs. 23 (25) : 117-118 (18 September)
Fruhstorfer, 1909
Neue Rhopaloceren von Formosa Soc. ent. 24 (16) : 121 (15 November)
Fruhstorfer, 1911
4. Familie: Satyridae in Seitz, Gross-Schmett. Erde 9 : 285-288 (14 February, 1911), : 289-296 (15 February, 1911), : 297-304 (22 February, 1911), : 305-312 (23 February, 1911), : 313-328 (before 25 April, 1911), : 329-336 (before 2 May, 1911), : 337-344 (before 9 May, 1911), : 345-352 (before 23 May, 1911), : 353-360 (before 30 May, 1911), : 361-368 (before 7 June, 1911), : 369-376 (before 13 June, 1911), : 377-384 (before 20 June, 1911), : 385-392 (before 27 June, 1911), : 393-400 (24 July), : 401 (15 August)
Fruhstorfer, 1913
Neue Indo-Australiche[sic] Rhopaloceren Dt. ent. Z. Iris 27 (3) : 130-139 ([14 October])
Fruhstorfer, 1914
Neue Satyriden Ent. Rundschau 31 (5) : 25-26 (14 March)
Fujioka, 1970
Butterflies collected by the Lepidopterological Research Expedition to Nepal Himalaya, 1963. Part 1. Papilionoidea Spec. Bull. Lepid. Soc. Japan (4) : 1-125
Godart, [1824]
in Latreille & Godart, Encyclopédie Méthodique. Histoire naturelle Entomologie, ou histoire naturelle des crustacés, des arachnides et des insectes Encyclopédie Méthodique. 9 (1): 3-328 (1819), (2): 329-828 ([1824])
Grose-Smith, 1893
Descriptions of Four new Species of Butterflies from Omei-shan, North-west China, in the Collection of H. Grose Smith Ann. Mag. Nat. Hist. (6) 11 (63) : 216-218
Hewitson, 1862; Hewitson, 1863
Illustrations of new species of exotic Butterflies selected chiefly from the collections of W. Wilson Saunders and William C. Hewitson Ill. exot. Butts [4] : (Systematic Index) IV, (Clerome): [1-2], pl. [1] (1863), (Drusilla & Hyantis): [3-4], pl. [2] (1862), (Dasyophthalma & Thaumantis): [5-6], pl. [3] (1862), (Morpho I): [7-8], pl. [4] (1856), (Pavonia I-II): [9-10], pl. [5-6] (1877), (Opsiphanes): [11-12], pl. [7] (1873), (Corades): [13-14], pl. [8] (1863), (Pronophila): [15-16], pl. [9] (1857), (Pronophila II): [17-18], pl. [10] (1860), (Pronophila III-IV): [19-22], pl. [11-12] (1868), (Pronophila V): [23-24], pl. [13] (1871), (Pronophila VI-VII): [25-28], pl. [14-15] (1872), (Pronophila VIII-IX): [29-32], pl. [16-17] (1874), (Debis I): [33-34], pl. [18] (1862), (Debis II-III): [35-38], pl. [19-20] (1863), (Haetera): [39-40], pl. [21] (1860), (Daedalma): [41-42], pl. [22] (1858), (Euptychia & Ragadia): [43-44], pl. [23] (1862), (Hypocista): [45-46], pl. [24] (1863), (Mycalesis I-IV): [47-54], pl. [25-28] (1862), (Mycalesis V-VI): [55-58], pl. [29-30] (1864), (Mycalesis VII-VIII): [59-62], pl. [31-32] (1866), (Mycalesis IX): [63-64], pl. [33] (1873), (Mycalesis & Idiomorphus): [65-66], pl. [34] (1877), (Idiomorphus): [67-68], pl. [35] (1865), (Melanitis): [69-70], pl. [36] (1863), (Taxila): [71-72], pl. [37] (1861), (Taxila II): [73-74], pl. [38] (1862), (Sospita): [75-76], pl. [39] (1861), (Sospita II): [77-78], pl. [40] (1862), (Dodona Sospita): [79-80], pl. [41] (1866), (Mesosemia I): [81-82], pl. [42] (1857), (Mesosemia II-III): [83-86], pl. [43-44] (1858), (Mesosemia IV-VI): [87-92], pl. [45-47] (1859), (Mesosemia VII-VIII): [93-96], pl. [48-49] (1860), (Mesosemia IX-X): [97-100], pl. [50-51] (1870), (Mesosemia XI): [101-102], pl. [52] (1871), (Mesosemia XII): [103-104], pl. [53] (1873), (Eurygona I-II): [105-108], pl. [54-55] (1853), (Eurygona III-IV): [109-112], pl. [56-57] (1855), (Eurygona V-VII): [113-118], pl. [58-60] (1856), (Eurygona VIII): [119-120], pl. [61] (1860), (Eurygona IX): [121-122], pl. [62] (1870), (Eurygona X): [123-124], pl. [63] (1872)
Hewitson, 1872
Descriptions of seven new exotic Rhopalocera Ent. mon. Mag. 9 : 83-86
Hewitson, 1876
Notes on Mr. Atkinson's collection of East Indian Lepidoptera, with descriptions of new species of Rhopalocera Ent. mon. Mag. 13 (7) : 149-152
Huang, 1998
Research on the butterflies of the Namjagbarwa Region, S. E. Tibet Neue Entomologische Nachrichten, 41: 207-264
Huang, 1999
Lethe wui sp. nov. from Metok, S.E. Tibet Lambillionea XCIX (1): 129-131
Huang, 2000
A list of butterflies collected from Tibet during 1993-1996, with descriptions, revisional notes and discussion on zoogeography - 1, part 1 Lambillionea, C (1): 141-158
Huang, 2001
Report of H. Huang's 2000 Expedition to SE. Tibet for Rhopalocera Neue Entomologische Nachrichten, 51: 65-152
Huang, 2002
Some new satyrids of the tribe Lethini from China Atalanta 33 (3/4): 361-372, pl. 19-23
Huang, 2003
A list of butterflies collected from Nujiang (Lou Tse Kiang) and Dulongiang, China with descriptions of new species, and revisional notes Neue Ent. Nachr. 55: 3-114
Huang & Wu, 2003
New and little known Chinese butteflies in the collection of the Institue of Zoology, Academia Sinica, Beijing - 1 Neue Ent. Nachr. 55: 115-143
Huang, Wu & Yuan, 2003
Zophoessa ocellata (Poujade, 1885) and its allies in China with the description of two new species. A review of the genera Lethe, Zophoessa and Neope in China - 1 Neue Ent. Nachr. 55: 145-158
Huang & Xue, 2004
A Contribution to the Butterfly Fauna of Southern Yunnan Neue Ent. Nachr. 57: 135-154, pl. 11
Hübner, [1819]
Verzeichniss bekannter Schmettlinge, 1816-[1826] Verz. bek. Schmett. (1): [1-3], 4-16 (1816), (2): 17-32 (1819), (3): 33-48 (1819), (4): 49-64 (1819), (5): 65-80 (1819), (6): 81-96 (1819), (7): 97-112 (1819), (8): 113-128 (1819), (9): 129-144 (1819), (10): 145-160 (1819), (11): 161-176 (1819), (12): 177-192 (1820), (13): 193-208 (1820), (14): 209-224 (1821), (15): 225-240 (1821), (16): 241-256 (1821), (17): 257-272 (1823), (18): 273-288 (1823), (19): 289-304 (1823), (20): 305-320 (1825), (21): 321-336 (1825), (22): 337-352 (1825), (23-27): 353-431 ([1825])
Kalis, 1933
Bijdrage tot de kennis van de Lepidoptera Rhopalocera van Nederlandsch-Indië Tijdschr. Ent. 76 (1-2) : 47-86
Lee, 1962
Some new species of Rhopalocera in China (2) Acta ent. sin. 11 (1) : 139-148, pl. 1-6
Lee, 1985
Some new species of Rhopalocera in China 6 Entomotaxonomia 7 (3): 191-195
Leech, 1889
On a collection of Lepidoptera from Kiukiang Trans. ent. Soc. Lond. 1889 (1) : 99-148, pl. 7-9
Leech, 1890
New species of Lepidoptera from China Entomologist 23 : 26-50, 81-83, : 109-114, : 187-192, pl. 1
Leech, 1891
New Species of Lepidoptera from China Entomologist 24 (Suppl.) : 1-6
Leech, 1891
New Species of Rhopalocera from North-west China Entomologist 24 (Suppl.) : 23-31
Leech, 1891
New Species of Rhopalocera from Western China Entomologist 24 (Suppl.) : 57-61, 66-68
Leech, [1892]; Leech, 1894
Butterflies from China, Japan, and Corea Butts China Japan Corea (1): 1-297, pl. 1-28 (1892-1894), (2): 297-681, pl. 29-43 (1893-1894)
de Lesse, 1956
Revision du genre Lethe (s.l.) (Lep. Nymphalidae Satyridae) Ann. Soc. ent. Fr. 125 : 75-95
Marshall, [1881]
Some new Species of rhopalocerous Lepidoptera from the Indian Region. J. Asiat. Soc. Bengal 49 Pt.II (4) : 245-248
Marshall, 1882
Some new or rare species of Rhopalocerous Lepidoptera from the Indian region J. Asiat. Soc. Bengal 51 Pt.II (2-3) : 37-43, pl. 4
Martin, 1921
Die Tagfalterfauna des Gefangenenlagers Matsuyama auf der japanischen Südinsel Shikoku Dt. ent. Z. Iris 35 (1/2) : 6-21
Matsumura, 1909
Die Danaiden und Satyriden Japans Ent. Zs. 23 (19) : 91-92
Matsumura, 1925
An ennumeration of the butterflies and moths from Saghalien, with descriptions of new species and subspecies J. Coll. Agric. Hokkaido imp. Univ. 15 : 83-196, pl. 8-11
Mell, 1923
Noch unbeschriebene Lepidopteren aus Südchina (2) Dt. ent. Z. 1923 (2) : 153-160
Mell, 1935
Noch unbeschriebene chinesische Lepidopteren. 4 Mitt. Dtsch. Ent. Ges. 6 : 36-38
Mell, 1939
Beiträge zur fauna Sinica. XVIII. Noch unbeschriebene Chinesische Lepidopteren (V) Dt. Ent. Z. Iris 52 : 135-152
Mell, 1942
Die Amathusiidae und Satyriden Süd (un Südost-) Chinas Arch. Naturgesch. (N.F.) 11 : 221-290, pl. 1-3
Monastyrskii & Devyatkin, 2000
New taxa and new records of butterflies from Vietnam Atalanta 31 (3/4): 471-492
Moore, 1857; Moore, [1858]
A Catalogue of the Lepidopterous Insects in the Museum of the Hon. East-India Company in Horsfield & Moore, Cat. lep. Ins. Mus. East India Coy 1: : 1-278, pl. 1-12,1a,2a,3a,4a,5a,6a
Moore, [1866]
On the Lepidopterous Insects of Bengal Proc. zool. Soc. Lond. 1865 (3) : 755-823, pl. 41-43
Moore, 1872
Descriptions of new Indian Lepidoptera Proc. zool. Soc. Lond. 1872 (2) : 555-583, pl. 32-34
Moore, 1881
Descriptions of new Asiatic diurnal Lepidoptera Trans. ent. Soc. Lond. 1881 (3) : 305-313
Moore, [1880]; Moore, [1881]
The Lepidoptera of Ceylon Lepid. Ceylon 1 (1): 1-40, pl. 1-18 (Dec 1880), (2): 41-80, pl. 19-36 (Jan 1881), (3): 81-136, pl. 37-54 (Jun 1881), (4): 137-190, pl. 55-71 (Dec 1881)
Motschulsky, 1860
Insectes du Japon Etud. Ent. 9: 4-39
Oberthür, 1886
Espèces Nouvelles de Lépidoptères du Thibet / Nouveaux Lépidoptères du Thibet Étud. d'Ent. 11 : 13-38, pl. 1-7
Oberthür, 1890
Lépidoptères de Chine Étud. d'Ent. 13 : 35-45, pl. 9-10
Okubo, 1979
A New Species of Lethe from North Borneo : Lepidoptera: Satyridae Tyô to Ga 30 (1,2): 91-92
Pendlebury, 1933
Notes and new Records of Butterflies from the Malay Peninsula J. Fed. Malay St. Mus. 17 (2): 377-401
Poujade, 1884
[no title] Bull. Soc. Ent. Fr. (6) 4 : cxxxiv-cxxxvi [134-136], : cxl-cxli [140-141], : cliv-clv [154-155], : clviii [158]
Poujade, 1885
[no title] Bull. Soc. Ent. Fr. (6) 5 : x-xi [10-11], : xxiv-xxv [24-25], : xli-xlii [41-42], : lxvi [61], : xciv-xcv [94-95], : cxliii [143], : cli [151], : cc [200], : ccvii-ccviii [207-208], : ccxv-ccxvi [215-216]
Seitz, 1907; Seitz, [1909]
Die Großschmetterlinge des Palaearktischen Faunengebietes. 1. Die Palaearktischen Tagfalter Gross-Schmett. Erde 1 : 1-379
Semper, 1887
Die Schmetterlinge der Philippinischen Inseln. Reisen im Archipel der Philippinen Reisen Philipp. (1): 1-46, pl. 1-8, A (1886), (2): 47-86, pl. 9-16 (1887), (3): 87-134, pl. 17-24 (1888), (4): 135-174, pl. 25-32 (1889), (5): 175-238, pl. 33-38, B (1890), (6): 239-270, pl. 39-46 (1891), (7): 271-380, pl. 47-49 [iv] (1892)
South, 1913
A list of butterflies collected by Captain F.M. Bailey in western China, south-eastern Tibet, and the Mishimi hills, 1911 J. Bomb. nat. Hist. Soc. 22 (2): 345-365, (3): 598-615
Staudinger, 1889
Lepidoptera der Insel Palawan Dt. ent. Z. Iris 2 (1) : 3-180, pl. 1-2
Staudinger, [1897]
Neue exotische Tagfalter Dt. Ent. Z. Iris 9 (1): 193-224, pl. 1-2 (July 1896), (2): 225-240, pl. 5 (Jan. 1897)
Stoll, [1780]
Uitlandsche Kapellen (Papillons exotiques) in Cramer, Uitl. Kapellen 4 (26b-28): 29-90, pl. 305-336 (1780), (29-31): 91-164, pl. 337-372 (1781), (32-32): 165-252, 1-29, pl. 373-400 (1782)
Talbot, 1947
Fauna of British India including Ceylon and Burma. (Danaidae, Satyridae, Amathusiidae, and Acraeidae) Fauna Br. Ind., Butt. (2nd ed.) 2: xv + 406pp, 2pls
Tsukada & Nishiyama, 1979
On some species of butterflies from Indonesia and Philippines with the description of new genus, new species and new subspecies Mem. Tsukada Coll. 1: 28pp
Tytler, 1914
Notes on some new and interesting butterflies from Manipur and the Naga Hills. Part 1-3 J. Bombay nat. Hist. Soc. 23 (2): 216-229 (1914), 23 (3): 502-515, pl. 1-2 (1915), 24 (1): 119-155, pl. 3-4 (1915)
Notes on some new and interesting butterflies from Manipur and the Naga Hills. Part 3
Tytler, 1939
Notes on some new and interesting butterflies chiefly from Burma 1 & 2 J. Bombay nat. Hist. Soc. 41 (2): 235-252, 42 (1): 109-123
Uemura, 1997
Description of a new species of the genus Lethe Hübner from Laos Bull. Toyosato Mus. Ent. 6: 1-4
Vane-Wright & de Jong, 2003
The butterflies of Sulawesi: annotated checklist for a critical island faunda Zool. Verh. Leiden 343: 3-268, pl. 1-16
Watkins, 1927
Butterflies from N.W. Yunnan Ann. Mag. nat. Hist. (9) 19 : 313-344, 512
Watkins, 1928
New Satyrid Butterflies Ann. Mag. nat. Hist. (10) 1 (5) : 615-618
Yamamoto & Takei, 1993
A new subspecies of Zophoessa dataensis (Semper) (Lepidoptera, Satyridae) from Mindoro, the Philippines Tyô to Ga 43 84) : 255-256
Yoshino, 1997
New Butterflies from China 2 Neo Lepidoptera 2: 1-8
Yoshino, 1999
New Butterflies from China 5 Neo Lepidoptera 4: 1-10
Yoshino, 2003
New butterflies from China 8 Futao (43): 6-19
Yoshino, 2008
New species and new subspecies of Lethe from Myanmar, China and Vietnam Futao (54): 9-19


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Keep in mind that the taxonomic information is copied from various sources, and may include many inaccuracies. Expert help is welcome.