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Clossiana Reuss, 1920
Clossiana Reuss, 1920; Ent. Mitt. 9 : 192 nota, TS: Papilio selene Denis & Schiffermüller
= Boloria ; [Wahlberg]
Clossiana (Nymphalidae) ; [BRU2, 43]


[sp-1.jpg] 700x504 (~51Kb) USA: Corral Pass region (47°00'15N 121°29'02W), Pierce Co., WA, 18.7.2000, Photo © Markku Savela

The exact identification of this species is still unknown, but tentatively assumed to belong into this group.



fiClossiana selene ([Schiffermüller], 1775)
. See [About maps] Niittyhopeatäplä Small Pearl-bordered Fritillary Silver-bordered Fritillary Silver Meadow Fritillary Brunfläckig pärlemorfjäril Ängspärlefjäril Braunfleckiger Perlmutterfalter Le Petit Collier Argenté Perlada castańa Dostojka selene
Papilio selene [Schiffermüller], 1775; Ankündung eines systematischen Werkes von den Schmetterlingen der Wienergegend: 321, TL: Austria
euphrosyne (Bergsträsser, 1779)
euphrasia (Lewin, 1795)
silene (Haworth, 1803)
Boloria selene, E.H. Strickland Ent. Mus, U of [ALBERTA]
Boloria selene, NSG Voucher Specimen [Wahlberg]
Boloria selene, Butterflies and Moths of North America [BAMONA]
Boloria selene, Larvae of North-European Lepidoptera [Kimmo Silvonen]
Boloria selene, Svenska fjärilar [SVF]
Boloria selene, Norges Sommerfugler [NORLEP]
Clossiana selene, Moths and Butterflies of Europe [MBE]
Clossiana selene, Europaische Schmetterlinge [Mario Maier]
Clossiana selene, Captain's European Butterfly Guide [Simon Coombes]
Clossiana selene ; [NACL, #4464]; [BOW: pl. 2, f. 2-3]; [EBW]; [BARU, #188]; [BRU2: 45, pl. 32, f. 1-6]
Boloria selene ; [BNA: 318, #239, pl. 3, 5, 24, 26]; [BOC, 281]; [Opler]; [Otakar Kudrna]
[selene-1u.jpg] 450x697 (~67Kb) upperside [selene-1v.jpg] 458x713 (~67Kb) underside [selene-2u.jpg] 425x374 (~35Kb) upperside
[selene-2v.jpg] 401x353 (~34Kb) underside [selene-3u.jpg] 500x391 (~75Kb) upperside Haldon woods, Devon, UK 21/05/95, Photo © Simon Coombes [selene-3v.jpg] 500x389 (~57Kb) underside Haldon woods, Devon, UK 21/05/95, Photo © Simon Coombes
[selene-4v.jpg] 837x1028 (~120Kb) underside Finland, Kuhmalahti, Mattilankylä, Setälänlahti, 23.6.2005, Photo © Harri Arkkio [selene-4vx.jpg] 1371x992 (~199Kb) underside Finland, Kuhmalahti, Mattilankylä, Setälänlahti, 23.6.2005, Photo © Harri Arkkio [selene-5fu.jpg] 852x639 (~123Kb) upperside female Russia, Moscow area (36°25'E, 56°00'N), 14.6.2009, Photo © D. Smirnov
[selene-5mu.jpg] 711x624 (~103Kb) upperside male Russia, Moscow area, 14.6.2010 (36°25'E, 56°00'N), Photo © D. Smirnov [selene-5v.jpg] 824x689 (~114Kb) underside Russia, Moscow area, 10.6.2010 (36°25'E, 56°00'N), Photo © D. Smirnov [selene-6mu.jpg] 918x694 (~122Kb) upperside male Russia, Moscow area, 3.7.2010 (36°25'E, 56°00'N), Photo © D. Smirnov
[selene-6mv.jpg] 726x732 (~106Kb) underside male Russia, Moscow area, 3.7.2010 (36°25'E, 56°00'N), Photo © D. Smirnov

Larva on Viola, V. palustris, V. canina, V. riviniana [SPRK], Fragaria, Vaccinium uliginosum [BRU2, 46], Viola nephrophylla, V. glabella [BNA, 318]


EU, W.Siberia, N.Kazakhstan, Altai, Sayan, Transbaikalia Clossiana selene selene ; [BRU2, 45]
C. s. thalia (Hübner, 1790)
Arctic Europe, N.Siberia ?Papilio thalia Hübner, [1799-1800]; Samml. eur. Schmett. [1] : pl. 11, f. 57-58
hela (Staudinger, 1861)
obscurum (Sedych, 1977)
Clossiana selene thalia ; [BARU]; [BRU2, 45]
C. s. dilutior (Fixsen, 1887)
E.Amur, Ussuri sugitanii (Seok, 1938)
Clossiana selene dilutior ; [BARU]; [BRU2, 45]
C. s. chibiana (Matsumura, 1927)
Sakhalin, Kunashir Argynnis selene chibiana Matsumura, 1927; Ins. Matsumurana 2 (2) : 116, pl. 3, f. 4, TL: Kuriles
Clossiana selene chibiana ; [BARU]; [BRU2, 45]
C. s. myrina (Cramer, [1777])
Papilio myrina Cramer, [1777]; Uitl. Kapellen 2 (9-16): 141, pl. 189, f. B, C
Papilio myrinus Herbst, 1798
Argynnis myrissa Godart, 1819; 253
Brenthis myrina nubes Scudder, 1889; 595, TL: Sanbornton, New Hampshire
ab. nivea Gunder, 1928; Can. Ent. 60: (albino) , TL: Staten Island, New York
Brenthis selene marilandica Clark, 1941; 384, TL: Beltsville, Maryland
Boloria selene myrina, Butterflies of America [BOA]

Argynnis myrina ; Scudder, 1863, 166; Scudder, 1868, 379; Edwards, 1876, 161
Brenthis myrina ab. nubes ; McDunnough, 1938, 16
Brenthis myrina ; Dyar, 1900, 487; Cary, 1901, 307; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 15; Cary, 1906, 439; Elrod, 1906, 69; Franzen, 1914, 366; McDunnough, 1928, 274; Holland, 1931, The Butterfly Book (1931 edn): 105; Clark, 1932, 102; dos Passos & Grey, 1934, 190; McDunnough, 1938, 15; Davenport & Dethier, 1938, 158; Leussler, 1938, 216; Rysgaard, 1939, 193; Comstock, 1940, 53; Brooks, 1942, 34
Brenthis selene myrina ; Clark, 1941, 384
Boloria myrina ; dos Passos & Grey, 1945, 4
Boloria selene marilandica ; Klots, 1951, 89; dos Passos, 1964, 89
Clossiana selene myrina ; [NACL, #4464a]; [BNA, #239]
Boloria selene myrina ; Klots, 1951, 89; dos Passos, 1964, 89; Shepard, 1975, in Howe, 246; Kohler, 1977, J. Lep. Soc. 31 (4): 245; [Jonathan Pelham]
Boloria myrina ; [Joseph Belicek]

There is split of opinions whether 'myrina' is considered as own species or as ssp. of 'selene'. I have no idea of the "correct" ranking, but the setup here requires me to choose either one or the other, the choice is random, tentative, and with no taxonomic statement assocated with it! -- msa

C. s. nebraskensis (Holland, 1928)
E.Nebraska - Ohio Brenthis myrina var. nebraskensis Holland, 1928; 36, TL: Dodge Co., Nebraska
Boloria selene nebraskensis, Butterflies of America [BOA]

Brenthis myrina var. nebraskensis ; Holland, 1931, The Butterfly Book (1931 edn): 105, pl. LV, f. 8 (type)
Brenthis myrina r. nebraskensis ; Leussler, 1938, 216
Brenthis myrina nebraskensis ; McDunnough, 1938, 16
Brenthis selene nebraskensis ; Clark, 1941, 384
Clossiana selene nebraskensis ; [NACL, #4464b]
Boloria selene nebrascensis ; Forbes, 1960, 160 (missp.)
Boloria selene nebraskensis ; Klots, 1951, 89; Brown, 1957, 64; dos Passos, 1964, 89; Shepard, 1975, in Howe, 246; Kohler, 1977, J. Lep. Soc. 31 (4): 248, f. 2, 6; [BNA, #239]
C. s. sabulocollis (Kohler, 1977)
Nebraska, South Dakota, North Dakota, Colorado Boloria selene sabulocollis Kohler, 1977; J. Lep. Soc. 31 (4): 252, f. 3, 6-7, TL: Smith Lake, Sheridan Co., Nebraska
Boloria selene sabulocollis, Butterflies of America [BOA]
Clossiana selene sabulicollis[sic?] ; [NACL, #4464c]
C. s. tollandensis (Barnes & Benjamin, 1925)
Rocky Mountains (Colorado, Idaho, Montana, New Mexico, Utah, Wyoming), British Columbia Brenthis myrina tollandensis Barnes & Benjamin, 1925; Ent. News 36 : 44, TL: Tolland, Gilpin Co., Colorado
ab. serratimarginata (Gunder, 1926); Ent. News 37 (1): (1-9) , TL: Vernon, British Columbia
Boloria selene tollandensis, Butterflies of America [BOA]
Brenthis myrina tollandensis ; Holland, 1928, 37; Klots, 1937, 328; McDunnough, 1938, 16
Brenthis myrina var. tollandensis ; Holland, 1931, The Butterfly Book (1931 edn): 106
Brenthis selene tollandensis ; Clark, 1941, 384
Boloria myrina tollandensis ; Brown, 1954, 64
Boloria selene tollandensis ; Remington, 1952, 67; Brown, 1957, 64; Albright, 1960, 158; Newcomer & Rogers, 1963, 171; dos Passos, 1964, 89; Shepard, 1975, in Howe, 246; Kohler, 1977, J. Lep. Soc. 31 (4): 253
Clossiana selene tollandensis ; [NACL, #4464d]
C. s. albequina (Holland, 1928)
Yukon, SW.Alaska Brenthis albequina Holland, 1928; 40, f. 4-6, TL: White Horse Pass, Yukon Territory
Brenthis albeguina var. baxteri Holland, 1928; 42
Brenthis albequina ; Holland, 1931, The Butterfly Book (1931 edn): 107, pl. LV, f. 19-21 (types); McDunnough, 1932, 269
Brenthis myrina atrocostalis ab. baxteri ; McDunnough, 1938, 16
Boloria selene albequina ab. baxteri ; dos Passos, 1964, 89
Boloria selene albequina ; dos Passos, 1964, 89; Kohler, 1977, J. Lep. Soc. 31 (4): 255, f. 4
Clossiana selene albequina ; [NACL, #4464e]
C. s. atrocostalis (Huard, 1927)
Nova Scotia, Quebec, Saskatchewan, Manitoba, Maine, Vermont, New Hampshire, New York, Michinag, Wisconsin, Minnesota, North Dakota Argynnis atrocostalis Huard, 1927; 313 (1926), TL: Chicoutini, Quebec
Brenthis myrina ab. jenningase Holland, 1928; 36
Brenthis myrina var. jenningsae Holland, 1931; The Butterfly Book (1931 edn): 105, pl. LVI, f. 9 (type)
Boloria selene atrocostalis, Butterflies of America [BOA]

Brenthis myrina atrocostalis ; McDunnough, 1930, 107; McDunnough, 1938, 16; dos Passos & Grey, 1934, 190
Brenthis myrina atrocostalis ab. jenningsae ; McDunnough, 1932, 269; McDunnough, 1938, 16
Boloria selene atrocostalis ab. jenningsae ; dos Passos, 1964, 89
Boloria selene atrocostalis ; Clark, 1941, 384; Klots, 1951, 89; dos Passos, 1954, 89; Shepard, 1975, in Howe, 246
Bororia selene atrocostalis ; Kohler, 1977, J. Lep. Soc. 31 (4): 258, f. 5, 8
Boloria selene atrocostalis ; [BNA, 318]
Clossiana selene atrocostalis ; [NACL, #4464f]
C. s. terraenovae (Holland, 1928)
Newfoundland Brenthis myrina terrae-novae Holland, 1928; 36, TL: Newfoundland
Brenthis myrina terrae-novae ; dos Passos, 1935, 87; McDunnough, 1938, 16
Brenthis myrina var. terrae-novae ; Holland, 1931, The Butterfly Book (1931 edn): 105, pl. LV, f. 13 (m.type)
Brenthis selene terrae-novae ; Clark, 1941, 384
Boloria selene terrae-novae ; Klots, 1951, 89
Boloria selene terraenovae ; dos Passos, 1964, 89; Shepard, 1975, in Howe, 264; Kohler, 1977, JLS#: 362, f. 5
Clossiana selene terraenovae ; [NACL]

Clossiana perryi (Butler, 1882)
S.Ussuri, N.Korea, Amur?. See [About maps] Argynnis perryi Butler, 1882; Ann. mag. nat. Hist. (5) 9 (49) : 16
Clossiana selene perryi ; [BARU]
Clossiana perryi ; [BRU2: 46, pl. 32, f. 7-9]

Clossiana erubescens (Staudinger, 1901)
-. See [About maps] Argynnis erubescens Staudinger, 1901; in Staudinger & Rebel, Cat. Lepid. palaearct. Faunengeb., 1: 35
?Argynnis hegemone Alphéraky, 1881; Horae Soc. ent. Ross. 16 (3-4): 407, pl. 15, f. 16-17, TL: Kuldja?
Argynnis hegemone Staudinger, 1881; Stettin ent. Ztg 42 (7-9): 292, TL: "Ala Tau" [Dzhungarsky Alatau Mts., Kazakhstan]
Clossiana hegemone; [BRU2: 46, pl. 33, f. 25-30]
Clossiana hegemone ; [BOW: pl. 194, f. 12]
Clossiana erubescens ; [Oleg Kosterin]
C. e. tienschanica (Wagner, 1913)
Tian-Shan Argynynnis hegemone tienschanica Wagner, 1913; Ent. Mitt. 2 (4): 114, f. 9, TL: Juldus
= Clossiana hegemone hegemone ; [BRU2, 46]
darjana (Seitz, [1909])
= Clossiana hegemone hegemone ; [BRU2, 46]
Clossiana erubescens tienschanica ; [Oleg Kosterin]
[tienschanica-1v.jpg] 400x533 (~49Kb) underside An alpine meadow on the right bank terrace of the fourth tributary of the Sargir River (the Usek River basin), altitude 3200 m, 47 km NEN of Zharkent, 21 km NEN of village Sarybel', the mountain massif Tyshkantau, the Southern Dzhungarian Alatau mountain chain (NE part of Tien Shan), Zharkent District, Taldy-Kurgan Province, SE Kazakhstan. 27th of July 1994, Photo © Oleg Kosterin

C. e. houri (Wyatt, 1961)
W.Tian-Shan, Ghissar-Darvaz, Pamirs-Alai Boloria hegemone houri Wyatt, 1961; J. Lep. Soc. 15 (1): 8, TL: Bala Quran, Anjuman Valley, Hindu Kush Mts., 14300ft
Clossiana hegemone houri ; [BRU2, 46]
?C. e. chotana (O. Bang-Haas, 1915)
Argynnis erubescens chotana O. Bang-Haas, 1915; Dt. ent. Z. Iris 29 (2/3) : 92-100, TL: Kun-Lun Mts., China
?Clossiana hegemone chotana ; [BRU2, 46 (?)]

Clossiana selenis (Eversmann, 1837)
-. See [About maps] Argynnis selenis Eversmann, 1837; Bull. Soc. imp. Nat. Moscou 1837 (1): 10, TL: "Kasan" [Kazan, Tatarstan, Russia]
?Clossiana speranda Grosser, 1979; , TL: NW.Mongolia
= ?; [BRU2, 44 (?)]
Argynnis selenis takamukuella Matsumura, 1929; Insecta Matsumurana 3 (4) : 155, TL: Korea
Boloria selenis, NSG Voucher Specimen [Wahlberg]
Clossiana selenis ; [BARU, #189]; [BOW: pl. 194, f. 15]; [MRS, 478]; [BRU2: 43, pl. 31, f. 25-30]
Boloria selenis ; [Otakar Kudrna]

Larva on Viola [BRU2, 44]

S.Urals, Volga Valley Clossiana selenis selenis ; [BRU2, 43]
C. s. sibirica (Erschoff, 1870)
Siberia, Amur festiva (Krulikovsky, 1893)
Argynnis (Brenthis) selenis meinhardi Sheljuzhko, 1929; Mitt. Münch. Ent. Ges. 19 : 359, TL: Tomsk
Clossiana selenis sibirica ; [Oleg Kosterin]; [BRU2, 44]
[sibirica-1v.jpg] 532x326 (~51Kb) underside A southern steppefied slope of the mountain on the Chuya River right bank between the villages Kurai and Chagan-Uzun, Altai Mts., West SIberia, Russia. 7th July 1988, Photo © Oleg Kosterin

C. s. samkoi (Sheljuzhko, 1931)
Polar Urals, Yamal Peninsula Clossiana angarensis samkoi ; [BARU]
Clossiana selenis samkoi ; [BRU2, 44]
C. s. kononovi (Kurentzov, 1970)
NE.Siberia perunovi Korb, 1999
Clossiana selenis kononovi ; [BRU2, 44]
C. s. onorensis (Matsumura, 1925)
Sakhalin Clossiana selenis onorensis ; [BRU2, 44]
C. s. chosensis (Matsumura, 1927)
Ussuri, Korea Argynnis selenis chosensis Matsumura, 1927; Insecta Matsumurana 1 (4) : 165, pl. 5, f. 6, TL: Corea, Shuotsu
Clossiana selenis chosensis ; [BRU2, 44]

Clossiana angarensis (Erschoff, 1870)
-. See [About maps] Argynnis angarensis Erschoff, 1870; Bull. Soc. Imp. Nat. Moscou 43 (1) : 112, TL: "the Irkutsk Province"
Boloria angarensis, NSG Voucher Specimen [Wahlberg]
Clossiana angarensis ; [BARU, #193]; [BOW: pl. 194, f. 10]; [BRU2: 43, pl. 30, f. 7-19]
Boloria angarensis ; [Otakar Kudrna]
[angarensis-1fu.jpg] 533x400 (~45Kb) upperside female An open larch (Larix gmelinii (Rupr.) Rupr.)/birch (Betula pendula Roth. and B. davurica Pallas) forest above the left bluff of the Budyumkan River valley 5 km of its confluence with the Argun' River 6 km S of the village Uryupino, Gazimurskozavodskoi District, E Chita Province, E Transbaikalia, Siberia, Russia. 2nd August 1997, Photo © Oleg Kosterin [angarensis-1fv.jpg] 352x299 (~22Kb) underside female A road in a larch (Larix gmelinii (Rupr.) Rupr.) forest not far from the Argun' River left bank between the village Uryupino and the Gazimur River, Gazimurskozavodskoi District, E Chita Province, E Transbaikalia, Siberia, Russia. 31th July 1997, Photo © Oleg Kosterin

Larva on Vaccinium [BRU2, 43]


Transbaikalia, S.Siberia Clossiana angarensis angarensis ; [BRU2, 43]
C. a. herzi (Wnukowsky, 1927)
N.Transbaikalia, Far East Yakutia Clossiana angarensis herzi ; [BARU]; [BRU2, 43]
C. a. sedychi (Weiss, 1964)
Polar Urals, Yamal Peninsula Clossiana angarensis sedychi ; [BRU2, 43]
C. a. alticola (Sushkin & Tschetverikov, 1907)
Sayan, Tuva (mountains) shelkovnikovi Korb, 1997
Clossiana angarensis alticola ; [BARU]; [BRU2, 43]
C. a. hakutozana (Matsumura, 1927)
Amur, Ussuri, N.Korea, NE.China Argynnis hakutozana Matsumura, 1927; Insecta Matsumurana 1 (4) : 165, pl. 5, f. 13, TL: Korea
Clossiana hakutozana ; [BARU, #195]
Clossiana hakutosana[sic, recte hakutozana] ; [EBW, (name only)]
Clossiana angarensis hakutozana ; [BRU2, 43]
C. a. miakei (Matsumura, 1919)
Sakhalin Clossiana angarensis miakei ; [BARU]; [BRU2, 43]

Clossiana oscarus (Eversmann, 1844)
-. See [About maps] Argynnis oscarus Eversmann, 1844; Bull. Soc. Imp. Nat. Moscou 17 (3) : 588, pl. 14, f. 1 a,b, TL: Altai [Irkutsk region?]
Boloria oscarus, NSG Voucher Specimen [Wahlberg]
Clossiana oscarus ; [BARU, #192]; [BOW: pl. 194, f. 14]; [BRU2: 44, pl. 34, f. 1-6]
[oscarus-1mv.jpg] 488x400 (~38Kb) underside male A bushy meadow on the second right bank terrace of the Koen River at the Volchikha Rivulet just upstream of its fall into the Koen River near the village Nizhniy Koen, Iskitim District, Novosibirsk Province, West Siberia, Russia. 18th June, 1989, Photo © Oleg Kosterin

Larva on Viola [BARU]


S.Siberia, E.Amur Clossiana oscarus oscarus ; [BRU2, 44]
C. o. australis (Graeser, 1888)
S.Amur, Ussuri, Sakhalin maxima (Fixsen, 1887); (preocc.)
major (Graeser, 1888)
Clossiana oscarus australis ; [BARU]; [BRU2, 44]
C. o. oscaroides (Ménétriés, 1859)
E.Yakutia Clossiana oscarus oscaroides ; [BRU2, 44]

fiClossiana freija (Thunberg, 1791)
Japan. See [About maps] Muurainhopeatäplä Freija's Fritillary Freya's Fritillary Zigzag Fritillary Frejas pärlemorfjäril Myrpärlefjäril
Papilio freija Thunberg, 1791; Diss. Ent. sistens Insecta Suecica (2): 34, TL: Vestrobothnia & Lapponica
?Papilio freija Beclin, 1791
freya (Godart, 1819)
gunderi (Harper, 1933)
Boloria freija, E.H. Strickland Ent. Mus, U of [ALBERTA]
Boloria freija freija, Butterflies of America [BOA]
Boloria freija, NSG Voucher Specimen [Wahlberg]
Boloria freija, Butterflies and Moths of North America [BAMONA]
Boloria freija, Larvae of North-European Lepidoptera [Kimmo Silvonen]
Boloria freija, Svenska fjärilar [SVF]
Boloria freija, Norges Sommerfugler [NORLEP]
Clossiana freija, Moths and Butterflies of Europe [MBE]
Brenthis freija ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 16
Clossiana freija ; [BARU, #201]; [NACL, #4471]; [MRS, 477]; [BRU2: 48, pl. 32, f. 24-29]
Boloria freija ; [BNA: 323, #249, pl. 26]; [BOC, 284]; [Opler]; [Otakar Kudrna]

Larva on Rubus chamaemorus, Vaccinium uliginosum [SPRK], Vaccinium vitis-idaeus, Rhododendron auerum [BARU], Empetrum nigrum [MRS], Vaccinium caespitosum, V. uliginosum, Arctostaphylos uva-ursi, Empetrum nigrum [NAB], Arctous alpina, (Scandinavia) Sieversia, Rhododendron, (Japan) Dryas integrifolia, (N.America) [BRU2, 48]

Fennoscandia, W.Siberia, C.Siberia, Far East, Transbaikiali, Chukotka, Amur, Ussuri, Canada, Alaska Boloria freija freija
freija (Quensel, 1791); (preocc.)
?Papilio lapponica Esper, 1789; Die Schmett., Suppl. Th 1 (3-4): 21, pl. 97, f. 3
?Papilio lapponica Esper, 1798; Die Schmett., Suppl. Th 1 ([7]): 83, pl. 108, f. 5
Boloria freija freija ; [BNA, #249]
Clossiana freija freija ; [BRU2, 48]
C. f. pallida (Elwes, 1899)
Altai, Sayan, Tuva calais (Fruhstorfer, 1916)
Clossiana freija pallida ; [BARU]
Clossiana freija calais ; [BARU, (Tuva)]
Clossiana freija pallida ; [Oleg Kosterin]; [BRU2, 48]
[pallida-1fv.jpg] 533x400 (~67Kb) underside female A dwarf birch (Betula rotundifolia Spach) montane tundra at 2200 m above sea level in the Argem (Direntai) Stream basin, 15 km W of the village Dzhazatr, on the eastern spurs of the Katunskiy Mt. Range, Kosh-Agach District, Central Altai Mts., West Siberia, Russia. 13th July, 1988, Photo © Oleg Kosterin [pallida-1mu.jpg] 508x444 (~37Kb) upperside male A meadowy-tundrous SW slope of a ridge between the headwaters of the Chikty Rivulet, 2600 m above sea level, the southern principle slope of the Yuzhno-Chuiskii mountain range, the upper Dzhazator River basin, Kosh-Agach District, Altai Republic, West Siberia, Russia. 10th July 1998 (on Aster alpinum L.), Photo © Oleg Kosterin

C. f. jakutensis (Wnukowsky, 1927)
NE.Yakutia Clossiana freija jakutensis ; [BARU]; [BRU2, 48]
C. f. tarquinius (Curtis, 1835)
freija tarquinius (Curtis, 1835); in Ross, Nar. 2nd Voy. N.W. Passage, App.: 68
Brenthis freija tarquinius ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 16
Clossiana freija tarquinius ; [NACL, #4471a]
C. f. browni (Higgins, 1953)
Rocky Mountains Boloria freija browni, Butterflies of America [BOA]
Clossiana freija browni ; [NACL, #4471d]
Boloria freija browni ; [BNA, #249]

Clossiana natazhati (Gibson, 1920)
Yukon, Victoria I.. See [About maps] Pleistocene Fritillary
Brenthis natazhati Gibson, 1920; Rep. Can. arctic exped. 1913-18. 3 (1): 21i-22i, TL: Mt. Natazhat, St. Elias Mts., Yukon Territory, 2470-2620m
Boloria freija nabokovi Stallings & Turner, 1947; Can. Ent. 78 (7-8): (134-135), TL: Alaska Military Highway, 102 miles N of Summit 2, 1830m
= ; Troubridge & Wood, 1990, J. Lep. Soc. 44 (3): 181
Boloria natazhati natazhati, Butterflies of America [BOA]
Boloria natazhati, Butterflies and Moths of North America [BAMONA]
Boloria freija nabokovi ; dos Passos, 1964; Howe, 1975
Clossiana freija nabokovi ; Miller & Brown, 1981; [NACL, #4471c]; Tilden & Smith, 1986
Boloria freija natazhati ; dos Passos, 1964; Howe, 1975
Clossiana freija natazhati ; Miller & Brown, 1981; Ferris, dos Passos, Ebner & Lafontaine, 1983; [NACL, #4471b]; Tilden & Smith, 1968
Boloria natazhati ; [BNA: 323, #250, pl. 24]; Troubridge & Wood, 1990, J. Lep. Soc. 44 (3): 181, f. 1-2; [Opler]

Larva on Dryas integrifolia? Troubridge & Wood, 1990, J. Lep. Soc. 44 (3): 185


fiClossiana polaris (Boisduval, [1828])
Arctic Europe, Arctic Asia, Arctic America. See [About maps] Pohjanhopeatäplä Polar Fritillary Polaris Fritillary Polarpärlefjäril
polaris (Boisduval, [1828]); Eur. Lepid. Index Meth. 1: 15, TL: Nordkap Peninsula [Norway]
groenlandica (Skinner, 1892)
americana (Strand, 1905)
Boloria polaris polaris, Butterflies of America [BOA]
Boloria polaris, Butterflies and Moths of North America [BAMONA]
Boloria polaris, Larvae of North-European Lepidoptera [Kimmo Silvonen]
Boloria polaris, Svenska fjärilar [SVF]
Boloria polaris, Norges Sommerfugler [NORLEP]
Brenthis polaris ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 16
Clossiana polaris ; [NACL, #4470]; [BOW: pl. 2, f. 1]; [BARU, #202]; [BRU2: 48, pl. 33, f. 1-3]
Boloria polaris ; [BNA: 323, #248, pl. 26]; [Opler]; [Otakar Kudrna]
[polaris-1fu.jpg] 660x498 (~61Kb) upperside female FINLAND: Li: Utsjoki Puollamoaivi, 775:50, 8.7.1997, Photo © Markku Savela

Larva on Cassiope tetragona, Vaccinium uliginosum, Dryas octopetala, D. punctata [BRU2, 48]


C. p. stellata (Masters, 1972)
Clossiana polaris stellata ; [NACL, #4470a]

fiClossiana thore (Hübner, [1803-1804])
Japan. See [About maps] Purohopeatäplä Thore's Fritillary Gråkantad pärlemorfjäril Bäckpärlefjäril Alpen-Perlmutterfalter
Papilio thore Hübner, [1803-1804]; Samml. eur. Schmett. [1] : pl. 111, f. 571-573, TL: Alps of Tirol
Boloria thore, NSG Voucher Specimen [Wahlberg]
Boloria thore, Larvae of North-European Lepidoptera [Kimmo Silvonen]
Boloria thore, Svenska fjärilar [SVF]
Boloria thore, Norges Sommerfugler [NORLEP]
Clossiana thore, Moths and Butterflies of Europe [MBE]
Clossiana thore, Europaische Schmetterlinge [Mario Maier]
Clossiana thore, Captain's European Butterfly Guide [Simon Coombes]
Clossiana thore ; [BOW: pl. 2, f. 4]; [BARU, #194]; [MRS, 478]
Boloria thore ; [BRU2: 46, pl. 29, f. 22-24]; [Otakar Kudrna]
[thore-1v.jpg] 374x506 (~29Kb) underside Among the dead branches of the Siberian dwarf pine (Pinus pumila Ledeb.) A forest/tundra on the Koni Peninsula, S Magadan Region, Russia. 11th of July 1989, Photo © Oleg Kosterin [thore-u.jpg] 494x684 (~70Kb) upperside [thore-v.jpg] 470x733 (~69Kb) underside

Larva on Viola bifolia, (Finland) Viola altaica, (Urals) [BRU2, 47] Viola selkirkii, Viola grypoceras [MRS]


Alps Clossiana thore thore ; [BRU2, 46]
C. t. exellens (Krulikovsky, 1893)
splendida (Krulikovsky, 1895)
Clossiana thore exellens ; [BRU2, 47]
C. t. borealis (Staudinger, 1861)
N.Scandinavia, N.Siberia, Far East, Kamchatka Clossiana thore borealis ; [BRU2, 47]
C. t. transuralensis (Sheljuzhko, 1931)
Ural, W.Siberia Clossiana thore transuralensis ; [BARU]; [BRU2, 47]
C. t. hyperlampra (Fruhstorfer, 1907)
Pyrenees Argynnis thore hyperlampra Fruhstorfer, 1907; Ent. Zs. 21 (27) : 164, TL: Pyrenees
C. t. hypercala (Fruhstorfer, 1907)
Altai, Sayan, Transbaikalia Argynnis thore hypercala Fruhstorfer, 1907; Ent. Zs. 21 (27) : 164
lacrimosa (Fruhstorfer, 1907)
C. t. herculea (Fruhstorfer, 1907)
? herculea Fruhstorfer, 1907; , TL: Altai
Clossiana thore herculea ; [BARU]
C. t. hyperusia (Fruhstorfer, 1907)
Amur, Ussuri Argynnis thore hyperusia Fruhstorfer, 1907; Ent. Zs. 21 (27) : 164, TL: Amur
meridionalis (Kardakov, 1928)
= ; [BRU2, 47]
Clossiana brenda Hemming, 1933
= ; [BRU2, 47]
Clossiana thore meridionalis ; [BARU]
Clossiana thore hyperusia ; [BARU]; [MRS, 478]; [BRU2, 47]
C. t. excellens Krulikovsky²
Argynnis thore excellens ; Fruhstorfer, 1907, Ent. Zs. 21 (27) : 164
C. t. karafutonis (Matsumura, 1919)
Sakhalin Clossiana thore karafutonis ; [BRU2, 47]

fiClossiana frigga (Thunberg, 1791)
. See [About maps] Rahkahopeatäplä Frigga's Fritillary Willow-Bog Fritillary Friggas pärlemorfjäril Hjortronpärlefjäril
Papilio frigga Thunberg, 1791; Diss. Ent. sistens Insecta Suecica (2): 33, TL: Lappland
?Papilio frigga Becklin, 1791
Boloria frigga, E.H. Strickland Ent. Mus, U of [ALBERTA]
Boloria frigga, NSG Voucher Specimen [Wahlberg]
Boloria frigga, Butterflies and Moths of North America [BAMONA]
Boloria frigga, Larvae of North-European Lepidoptera [Kimmo Silvonen]
Boloria frigga, Svenska fjärilar [SVF]
Boloria frigga, Norges Sommerfugler [NORLEP]
Clossiana frigga, Moths and Butterflies of Europe [MBE]
Brenthis frigga ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 16
Clossiana frigga ; [NACL, #4466]; [BARU, #196]; [BOW: pl. 1, f. 27]; [BRU2: 47, pl. 32, f. 13-23]
Boloria frigga ; [BNA: 321, #244, pl. 26]; [Opler]; [Otakar Kudrna]
[frigga-u.jpg] 461x381 (~37Kb) upperside [frigga-v.jpg] 447x373 (~36Kb) underside

Larva on Rubus chamaemorus [SPRK], Salix, Betula, Dryas integrifolia [BNA, 321]


N.Scandinavia helvenacius (Sedykh, 1977)
Clossiana frigga frigga ; [BRU2, 47]
C. f. alpestris (Elwes, 1899)
Altai, Sayan, Transbaikalia Clossiana frigga alpestris ; [BARU]; [BRU2, 47]
[alpestris-1fu.jpg] 872x600 (~56Kb) upperside female Highland dwarf birch tundras on of the southerm principal slope of the Yuzhno-Chuiskii mountain at the the Chikty Rivulet headwaters, 2400 m above sea level, the upper Dzhazator River basin, Kosh-Agach District, Altai Republic, West Siberia, Russia. 15th July 1998, Photo © Oleg Kosterin [alpestris-1fux.jpg] 900x592 (~98Kb) upperside female A flowery alpine meadow in a valley of the rivulet descending from the southerm principle slope of the Yuzhno-Chuiskii mountain range between the Chikty and Akbul Rivulets, 2600 m above sea level, the upper Dzhazator River basin, Kosh-Agach District, Altai Republic, West Siberia, Russia. 11th July 1998, Photo © Oleg Kosterin [alpestris-1mv.jpg] 475x500 (~33Kb) underside male An alpine meadow vegetation at a brook flowing in a moss/sedge tundra in a cirque of the sources of a rivulet descending from the southerm principle slope of the Yuzhno-Chuiskii mountain range between the Chikty and Akbul Rivulets, 2650 m above sea level, the upper Dzhazator River basin, Kosh-Agach District, Altai Republic, West Siberia, Russia. 16 July 1998, Photo © Oleg Kosterin

C. f. maritima (Kardakov, 1928)
Amur, Ussuri Clossiana frigga maritima ; [BARU]; [BRU2, 47]
C. f. annae (Sushkin, 1906)
Tarbgatai Mts. Clossiana frigga annae ; [BRU2, 47 (?)]
C. f. saga (Staudinger, 1861)
Canada Argynnis frigga saga Staudinger, 1861; Stettin Ent. Ztg 22 (10-12): 350 (Kaden)
Brenthis frigga saga Kaden, 1861
Brenthis frigga saga ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 16
Clossiana frigga saga ; [NACL, #4466a]
Boloria frigga saga ; [BNA, #244]
C. f. gibsoni (Barnes & Benjamin, 1926)
Alaska - Hudson Bay alaskensis (Lehmann, 1913)
gibsoni (Barnes & Benjamin, 1926)
lehmanni (Holland, 1928)
Boloria frigga gibsoni, Butterflies of America [BOA]
Clossiana frigga alaskensis ; [NACL, #4466b]
Boloria frigga gibsoni ; [BNA, #244]
C. f. sagata (Barnes & Benjamin, 1923)
Wyoming - Colorado Boloria frigga sagata, Butterflies of America [BOA]
Clossiana frigga sagata ; [NACL, #4466c]
Boloria frigga sagata ; [BNA, #244]

Clossiana dia (Linnaeus, 1767)
-. See [About maps] Violet Fritillary Hainveilchen-Scheckling La Petite Violette Perlada Violeta Dostojka dia
Papilio dia Linnaeus, 1767; Syst. Nat. (Edn 12) 1 (2): 785, TL: Austria
Boloria dia, NSG Voucher Specimen [Wahlberg]
Boloria dia, Larvae of North-European Lepidoptera [Kimmo Silvonen]
Clossiana dia, Moths and Butterflies of Europe [MBE]
Clossiana dia, Europaische Schmetterlinge [Mario Maier]
Clossiana dia, Captain's European Butterfly Guide [Simon Coombes]
Clossiana dia ; [BOW: pl. 1, f. 28]; [EBW]; [BARU, #200]; [MRS, 477]; [BRU2: 49, pl. 33, f. 13-21]
Boloria dia ; [Otakar Kudrna]
[dia-1v.jpg] 533x400 (~44Kb) underside A dwarf birch (Betula rotundifolia Spach) montane tundra at 2200 m above sea level in the Argem (Direntai) Stream basin, 15 km W of the village Dzhazatr, on the eastern spurs of the Katunskiy Mt. Range, Kosh-Agach District, Central Altai Mts., West Siberia, Russia. 129 - 15th, 130 - 12th July, 1988, Photo © Oleg Kosterin [dia-2l.jpg] 598x400 (~44Kb) larva Central Europe 1990-2000, Photo © Michael Borsch [dia-2u.jpg] 598x400 (~45Kb) upperside Central Europe 1990-2000, Photo © Michael Borsch
[dia-u.jpg] 465x500 (~43Kb) upperside Axat, Pyrenees orientales, France 24/07/96, Photo © Simon Coombes [dia-v.jpg] 500x503 (~50Kb) underside Axat, Pyrenees orientales, France 24/07/96, Photo © Simon Coombes

Larva on Viola, Rubus idaeus, Prunella vulgaris [BRU2, 49]


W.Europe Clossiana dia dia ; [BRU2, 49]
C. d. disconota (Krulikovsky, 1909)
CEU, W.Siberia Argynnis dia var. disconota Krulikovsky, 1909; Rev. russ. ent. 9 (3) : 298
Clossiana dia disconota ; [BRU2, 49]
C. d. calida (Jachontov, 1911)
Caucasus, Transcaucasus Clossiana dia calida ; [BRU2, 49]
C. d. alpina (Elwes, 1899)
Dzhungarsky Alatau, Altai, Sayan, Transbaikalia dia alpina (Elwes, 1899); , TL: Altai
Clossiana dia alpina ; [BARU]; [BRU2, 49]
C. d. semota Tuzov, 2000
Far East Clossiana dia semota ; [BRU2, 49]
C. d. setania (Fruhstorfer, 1909)
dia setania (Fruhstorfer, 1909) (Argynnis); Int. ent. Zs. 3 (4): 21, TL: 'Marokko' [mislabeling?]
Clossiana dia setania ; [BMAT: 105, pl. 32, f. 19-20 (not in N.Africa)]

Clossiana gong (Oberthür, 1884)
Tibet, W.China, N.China. See [About maps] ?gong (Oberthür, 1914); Etudes d'Ent. 9: 15
Clossiana gong ; [BOW: pl. 194, f. 11]; [MRS, 477]
Clossiana gong splendens ; [MRS, 477]
C. g. xizangensis Huang, 2000

Clossiana jerdoni (Lang, 1868)
W.Himalayas, Chitral - Kashmir. See [About maps] Jerdons Silverspot
Argynnis jerdoni Lang, 1868; Ent. Montrhly Mag. 5: 34
Clossiana jerdoni ; [BOW: pl. 194, f. 13]

fiClossiana improba (Butler, 1877)
. See [About maps] Kääpiöhopeatäplä Dusky-windged Fritillary Dinky Fritillary Dingy Arctic Fritillary Dvärgpärlefjäril
Argynnis improba Butler, 1877; Ent. mon. Mag. 13 : 206, TL: "Winter Cove and Cambridge Bay" [NW Territories, Canada]
Boloria improba, E.H. Strickland Ent. Mus, U of [ALBERTA]
Boloria improba, Butterflies and Moths of North America [BAMONA]
Boloria improba, Larvae of North-European Lepidoptera [Kimmo Silvonen]
Boloria improba, Svenska fjärilar [SVF]
Boloria improba, Norges Sommerfugler [NORLEP]
Clossiana improba, Moths and Butterflies of Europe [MBE]
Brenthis frigga improba ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 16
Clossiana improba ; [NACL, #4467]; [BARU, #197]; [BOW: pl. 13, f. 20]; [BRU2: 47, pl. 33, f. 22-24]
Boloria improba ; [BNA: 321, #245, pl. 5, 26, 60]; [Opler]; [Otakar Kudrna]
[improba-1.jpg] 600x450 (~38Kb) Gohpascurro/Bjorkliden, 1100-1400m, Torne Lapmark, Sweden 9 June 1985, Photo © Lars Andersen [improba-2v.jpg] 800x583 (~75Kb) underside Gohpascurro/Bjorkliden, 1100-1400m, Torne Lapmark, Sweden 9 June 1985, Photo © Lars Andersen [improba-3u.jpg] 800x549 (~56Kb) upperside Gohpascurro/Bjorkliden, 1100-1400m, Torne Lapmark, Sweden 9 June 1985, Photo © Lars Andersen
[improba-4u.jpg] 800x709 (~76Kb) upperside Gohpascurro/Bjorkliden, 1100-1400m, Torne Lapmark, Sweden 9 June 1985, Photo © Lars Andersen

Larva on Bistorta vivipara, Salix herbacea [BRU2, 47], Polygonum viviparum, Salix [BARU], Salix, S. reticulata?, (NW.Terr.) [BNA, 321]


Arctic Europe, Arctic Asia Clossiana improba improba ; [BRU2, 47]
C. i. improbula (Bryk, 1920)
C. i. youngi (Holland, 1900)
Alaska, E.Chukotka? improba youngi (Holland, 1900); Ent. News 11: 283, TL: Alaska
Brenthis youngi ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 16
Clossiana improba youngi ; [NACL, #4467a]
Wyoming (Wind River Mts.) Boloria improba harryi
Boloria improba harryi, Butterflies of America [BOA]
Boloria improba harryi ; [BNA, #245]

Larva on Salix arctica [BNA, 321]

?C. i. acrocnema (Gall & Sperling, 1980)
Colorado (San Juan Mts.) Uncompaghre Fritillary
Boloria acrocnema Gall & Sperling, 1980; J. Lep. Soc. 43: (230-252), TL: near Uncompahgre Peak, San Juan Mts., Hinsdale Co., Colorado
Boloria improba acrocnema, Butterflies of America [BOA]
Boloria improba acrocnema ; [BNA, #245]

Larva on Salix nivalis [BNA]


fiClossiana titania (Esper, 1794)
. See [About maps] Lehtohopeatäplä Titania's Fritillary Purple Bog Fritillary Amathusias pärlemofjäril Lundpärlefjäril Natterwurz-Perlmutterfalter Dostojka titania
Papilio titania Esper, 1794; Die Schmett., Suppl. Th 1 (3-4): 58, (5-6): pl. 103, f. 4, TL: Sardinia
Boloria titania, NSG Voucher Specimen [Wahlberg]
Boloria titania, Larvae of North-European Lepidoptera [Kimmo Silvonen]
Clossiana titania, Moths and Butterflies of Europe [MBE]
Clossiana titania, Europaische Schmetterlinge [Mario Maier]
Clossiana titania, Captain's European Butterfly Guide [Simon Coombes]
Clossiana titania ; [BOW: pl. 2, f. 5]; [EBW]; [BARU, #198]; [BRU2: 49, pl. 33, f. 31-36]
Boloria titania ; [Otakar Kudrna]
Misidentified/misapplied:
Clossiana titania (= Clossiana chariclea); [NACL, #4474]
Boloria titania (= Clossiana chariclea); [BNA: 319, #240]
[titania-1a.jpg] 571x653 (~100Kb) Russia, Moscow area (36°25'E, 56°00'N), 24.7.2009, Photo © D. Smirnov [titania-1v.jpg] 604x640 (~97Kb) underside Russia, Moscow area (36°25'E, 56°00'N), 24.7.2009, Photo © D. Smirnov

Sardinia, ?
C. t. bivina (Fruhstorfer, 1908)
CEU, SEU Argynnis amathusia bivina Fruhstorfer, 1908; Int. ent. Zs. 1 (41) : 310, TL: Saratow
Papilio amathusia Esper, 1783; Die Schmett. Th. I, Bd. 2 (8): 170, pl. 88, f. 1-2 (preocc.)
rossica (Hemming, 1934)
Clossiana titania bivina ; [BARU]; [BRU2, 50]

Larva on Viola, Vaccinium uliginosum [BRU2, 50]

C. t. staudingeri (Wnukowsky, 1929)
S.Siberia, Altai, Sayan, mountains (Tuva), Transbaikalia, Amur, Sakhalin? sibirica (Staudinger, 1892); (preocc.)
altaica (Seitz, [1909]); (preocc.)
julia (Hemming, 1934)
Clossiana titania staudingeri ; [BARU]; [BRU2, 50]

Larva on Bistorta major, Filipendula ulmaria, Trollius asiaticus [BRU2, 50]

C. t. miyakei (Matsumura, 1919)
Sakhalin, Low Amur Clossiana titania miyakei ; [BARU]

fiClossiana euphrosyne (Linnaeus, 1758)
. See [About maps] Pursuhopeatälä Pearl-bordered Fritillary Prydlig pärlemorfjäril Skogspärlefjäril Silberstrich Grand Collier Argenté Perlada rojiza Dostojka eufrozyna
Papilio euphrosyne Linnaeus, 1758; Syst. Nat. (Edn 10) 1 : 481, no. 142
Papilio euphrosyne Linnaeus, 1761; Fauna Suecica (Edn 2): 282, TL: Sweden
Boloria euphrosyne, NSG Voucher Specimen [Wahlberg]
Boloria euphrosyne, Larvae of North-European Lepidoptera [Kimmo Silvonen]
Boloria euphrosyne, Svenska fjärilar [SVF]
Boloria euphrosyne, Norges Sommerfugler [NORLEP]
Clossiana euphrosyne, Moths and Butterflies of Europe [MBE]
Clossiana euphrosyne, Europaische Schmetterlinge [Mario Maier]
Clossiana euphrosyne, Captain's European Butterfly Guide [Simon Coombes]
Clossiana euphrosyne ; [BOW: pl. 1, f. 26]; [EBW]; [BARU, #190]; [BRU2: 44, pl. 31, f. 13-24]
Boloria euphrosyne ; [Otakar Kudrna]
[euphrosyne-1u.jpg] 500x349 (~71Kb) upperside Luckett, Cornwall, UK 20/05/95, Photo © Simon Coombes [euphrosyne-1v.jpg] 417x500 (~57Kb) underside Luckett, Cornwall, UK 20/05/95, Photo © Simon Coombes [euphrosyne-2l.jpg] 533x400 (~50Kb) larva An open pine grove above the bluff of the valley of the Bol'shoi Elbash River, 5 km NW of the village Ust'-Chem, Iskitim dist., Novosibirsk region, West Siberia, 9th May 1995 (mature larva), Photo © Oleg Kosterin
[euphrosyne-3v.jpg] 600x832 (~62Kb) underside Finland, Loppi, Luutasuo 673:35, 2.6.1979, Photo © Pertti Pakkanen [euphrosyne-4fu.jpg] 900x745 (~146Kb) upperside female FINLAND: Li: Inari, Luttojoki, 760:54, 11.7.1998, Photo © Markku Savela [euphrosyne-5u.jpg] 600x722 (~61Kb) upperside FINLAND, Ab: Suomusjärvi, 669:31, 15.6.1997, Photo © Markku Savela
[euphrosyne-6.jpg] 900x1291 (~125Kb) Finland, Ab: Suomusjärvi (669:31), 6.6.2004, Photo © Markku Savela [euphrosyne-v.jpg] 800x631 (~57Kb) underside FINLAND: Ka: Virolahti, 671:53 (Reskopinsuo), 14.6.1997, Photo © Markku Savela

Larva on Viola, V. palustris, V. riviniana, Vaccinium uliginosum, Ledum palustre [SPRK], Viola hirta, Waldsteinia ternata [BRU2, 45]


CEU, Siberia Argynnis euphrosyne euphrosyne ; Fruhstorfer, 1907, Ent. Zs. 21 (27) : 164
Clossiana euphrosyne euphrosyne ; [BRU2, 45]
C. e. fingal (Herbst, 1804)
NEU, Siberia Argynnis euphrosyne fingal ; Fruhstorfer, 1907, Ent. Zs. 21 (27) : 164
Clossiana euphrosyne fingal ; [BRU2, 45]
C. e. rusalka (Fruhstorfer, 1909)
SEU, W.Siberia euphrosyne rusalka (Fruhstorfer, 1909) (Argynnis); Int. ent. Zs. 3 (20): 112
Clossiana euphrosyne rusalka ; [BRU2, 45]
C. e. orphana (Fruhstorfer, 1907)
Transbaikalia, Amur, Ussuri Argynnis euprhosyne orphanus Fruhstorfer, 1907; Ent. Zs. 21 (27) : 164
Clossiana euphrosyne orphanus ; [BARU]
Clossiana euphrosyne orphana ; [BRU2, 45]
C. e. kamtschadalus (Seitz, [1909])
Kamchatka, N.Sakhalin euphrosyne kamtschadalus (Seitz, [1909]); , TL: Kamchatka
Clossiana euphrosyne kamtschadalus ; [BARU]; [BRU2, 45]
C. e. umbra (Seitz, [1909])
Altai, Sayan Clossiana euphrosyne umbra ; [BRU2, 45]
C. e. dagestanica (Sovinsky, 1905)
Caucasus, Transcaucasia Argynnis euphrosyne var. dagestanica Sovinsky, 1905; Revue Russe Ent. 5 (3-4) : 108
Argynnis euphrosyne anka Fruhstorfer, 1907; Ent. Zs. 21 (27) : 164, TL: Caucasus
Clossiana phaenea Jachontov, 1908
?C. e. nephele (Herrich-Schäffer, [1847])
Ural, Siberia euphrosyne nephele (Herrich-Schäffer, [1847]) (Argynnis); Syst. Bearb. Schmett. Europ. 1 (26): (ii) f. 371-372
Argynnis euphrosyne nephele ; Fruhstorfer, 1907, Ent. Zs. 21 (27) : 164
Clossiana euphrosyne nephele ; [BARU]
C. e. orphanoides Huang & Murayama, 1992
Xinjiang Clossiana euphrosyne orphanoides Huang & Murayama, 1992; Tyō to Ga 43 (1) : 7, f. 17, TL: Jiadengyu, Altai, 1380-1600m

fiClossiana chariclea (Schneider, 1794)
. See [About maps] Tundrahopeatäplä Arctic Fritillary Tundrapärlefjäril
chariclea (Schneider, 1794); Neuest. Mag. Lieb. Ent. 1 (5): 588, TL: Lappland
Boloria chariclea, E.H. Strickland Ent. Mus, U of [ALBERTA]
Clossiana chariclea, Lepidoptera collection [SZMN]
Boloria chariclea, NSG Voucher Specimen [Wahlberg]
Boloria chariclea, Butterflies and Moths of North America [BAMONA]
Boloria chariclea, Larvae of North-European Lepidoptera [Kimmo Silvonen]
Boloria chariclea, Svenska fjärilar [SVF]
Boloria chariclea, Norges Sommerfugler [NORLEP]
Clossiana carichlea ; Zetterstedt, 1839, (missp.)
Clossiana churiclea ; Johansen, 1921, (missp.)
Clossiana churchilea ; dos Passos, 1964, (missp.)
Brenthis chariclea ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 16
Clossiana chariclea ; [NACL, #4475]; [BARU, #199]; [BOW: pl. 1, f. 25]; [BRU2: 48, pl. 33, f. 4-9]
Boloria chariclea ; [BOC, 286]; [Opler]; [Otakar Kudrna]
[chariclea-1u.jpg] 550x424 (~48Kb) upperside FINLAND: Li: Utsjoki Puollamoaivi, 775:50, 8.7.1997, Photo © Markku Savela [chariclea-1v.jpg] 500x564 (~35Kb) underside FINLAND: Li: Utsjoki Ailigas, 775:50, 8.7.1997, Photo © Markku Savela [chariclea-2u.jpg] 700x506 (~69Kb) upperside FINLAND: Li: Utsjoki Ailigas, 775:50, 6.7.1997, Photo © Markku Savela

Larva on Dryas integrifolia, Salix arctica, S. reticulata, (N.America) [BRU2, 49]


Arctic Europe Clossiana chariclea chariclea ; [BRU2, 48]

Larva on Cassiope tetragona [BRU2, 49]

C. c. arctica (Zetterstedt, 1839)
Arctic Asia, Wrangel Is., Chukotka chariclea arctica (Zetterstedt, 1839); Insecta Lapponica : 899
Cclossiana chariclea tshuktsha Korshunov, 1998
oenome (Scudder, 1875)
obscurata (McLachlan, 1878); J. Linn. Soc. Zool. Lond. 14 : 110
Brenthis chariclea arctica ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 16
Brenthis chariclea obscurata ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 16
Clossiana chariclea arctica ; [NACL, #4475a]
C. c. butleri (Edwards, 1883)
Arctic America, Chukotka, Kamchatka? Argynnis butleri Edwards, 1883; Can. Ent. 15 (2) : 32, TL: "Cape Thompson, Kotzebue Sound, W.Alaska"
Clossiana chariclea butleri ; [BARU]; [NACL, #4475b]
Clossiana butlerii ; Edwards, 1884, (missp.)
Clossiana butleri ; [BRU2: 49, pl. 33, f. 10-12]
C. c. boisduvalii (Duponchel, 1832)
Alaska, Alberta, Labrador, Newfoundland, Minnesota, British Columbia chariclea boisduvalii (Duponchel, 1832); in Godart, Hist. nat. Lépid. Fr. (Suppl.) 1 : pl. 20, f. 4
labradorensis (Gunder, 1928)
Brenthis chariclea boisduvalii ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 16
Clossiana titania boisduvalii ; [NACL, #4474a]
Boloris titania boisduvalii ; [BNA, #240]
C. c. rainieri (Barnes & McDunnough, 1913)
Washington Brenthis chariclea rainieri Barnes & McDunnough, 1913; Contr. nat. Hist. Lep. N. Am. 2 (3) : 96, pl. 2, f. 1-4, TL: Paradise Valley, Mt. Ra[i]nier, 6-7000ft, [Pierce or Lewis Co.] Wash[ington]
Boloria chariclea rainieri, Butterflies of America [BOA]
Brenthis chariclea ranieri[sic, recte rainieri] ; Barnes & McDunnough, 1913, Contr. nat. Hist. Lep. N. Am. 2 (3) : pl. 2, f. 1-4 (orig. missp.)
ranieri[sic, recte rainieri] ; Guppy & Kondla, 2002, J. Lep. Soc. 56 (3): 180
Clossiana titania rainieri ; [NACL, #4474b]
Boloria chariclea rainieri ; [BOC, 286]
C. c. grandis (Barnes & McDunnough, 1916)
N.British Columbia, Ontario chariclea grandis (Barnes & McDunnough, 1916); , TL: Hymers, Ontario
Boloria chariclea grandis, Butterflies of America [BOA]
Clossiana titania grandis ; [NACL, #4474c]
Boloria chariclea grandis ; [BOC, 286]
C. c. montina (Scudder, 1863)
montinus (Scudder, 1863); Proc. Essex Inst. 3: 166
Brenthis montinus ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 16; dos Passos, 1936, Can. Ent. 68: 239-241
Clossiana titania montina ; [NACL, #4474d]
C. c. ingens (Barnes & McDunnough, 1918)
martini (Gunder, 1934)
Clossiana titania ingens ; [NACL, #4474e]
C. c. helena (Edwards, 1871)
Rocky Mountains Argynnis helena Edwards, 1871; Trans. amer. ent. Soc. 3 : 268, TL: Mosquito Pass, Park/Lake cos, CO
Boloria chariclea helena, Butterflies of America [BOA]
Brenthis helena ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 16
Clossiana titania helena ; [NACL, #4474f]
Boloria titania helena ; [BNA, #240]
Boloria chariclea helena ; [BOC, 286]
[helena-1fu.jpg] 802x949 (~107Kb) upperside female USA: Colorado, Clear Creek Co., Summit Lake, Mt. Evans, 23.7.2012 (39°35'52"N 105°38'23"W), Photo © Markku Savela [helena-2av.jpg] 800x998 (~123Kb) underside USA: Colorado, Clear Creek Co., Summit Lake, Mt. Evans, 23.7.2012 (39°35'52"N 105°38'23"W), Photo © Markku Savela [helena-2bv.jpg] 800x608 (~84Kb) underside USA: Colorado, Clear Creek Co., Summit Lake, Mt. Evans, 23.7.2012 (39°35'52"N 105°38'23"W), Photo © Markku Savela


Clossiana bellona (Fabricius, 1775)
Rocky Mountains, Canada, Washington, Oregon. See [About maps] Meadow Fritillary
Papilio bellona Fabricius, 1775; Syst. Ent.: 517
fasciata (Cockerell, 1889)
kleeni (Watson, 1921)
ammiralis (Hemming, 1933)
pardopsis (Holland, 1928)
Boloria bellona, E.H. Strickland Ent. Mus, U of [ALBERTA]
Boloria bellona bellona, Butterflies of America [BOA]
Boloria bellona, NSG Voucher Specimen [Wahlberg]
Boloria bellona, Butterflies and Moths of North America [BAMONA]
Brenthis bellona ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 16
Clossiana bellona ; [NACL, #4465]; [BOW: pl. 13, f. 19]
Boloria bellona ; [BNA: 320, #243, pl. 26]; [BOC, 282]; [Opler]
Misidentified/misapplied:
myrina; Martyn, 1797

Larva on Viola sororia, V. pallens [BNA, 320]

Boloria bellona bellona
Boloria bellona bellona ; [BNA, #243]
C. b. toddi (Holland, 1928)
Labrador, N.Manitoba, Northwest Territories bellona toddi (Holland, 1928); , TL: St. Margaret's River, Quebec
Clossiana bellona toddi ; [NACL, #4465a]
Boloria belona toddi ; [BNA, #243]
C. b. jenistae (D. Stallings & Turner, 1947)
Clossiana bellona jenistae ; [NACL, #4465b]

Clossiana kriemhild (Strecker, 1879)
Montana, Wyoming. See [About maps] Relict Fritillary Kriemhild Fritillary
kriemhild (Strecker, 1879); in Ruffner, Rept. Surv. Missouri. app.: ?
laurenti (Skinner, 1913)
Clossiana luecki Reuss, 1923
Boloria kriemhild, Butterflies of America [BOA]
Boloria kriemhild, Butterflies and Moths of North America [BAMONA]
Clossiana kriemeld ; McHenry, 1964, (missp.)
Clossiana lucki ; dos Passos, 1964, (missp.)
Brenthis epithore kreimhild[sic] ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 16
Clossiana kriemhild ; [NACL, #4468]
Boloria kriemhild ; [BNA: 320, #241, pl. 26]; [Opler]

Larva on Viola [NAB]


Clossiana epithore (Edwards, 1864)
Yukon - California. See [About maps] Western Meadow Fritillary Pacific Fritillary
epithore (Edwards, 1864); Proc. ent. Soc. Philad. 2 (4): 504, TL: Saratoga, Santa Clara, California
?epithore Boisduval, 1869; Ann. Soc. ent. Belg. 12 : 58
Brenthis epithore Boisduval, 1869
Boloria epithore, E.H. Strickland Ent. Mus, U of [ALBERTA]
Boloria epithore, Butterflies of America [BOA]
Boloria epithore, Butterflies and Moths of North America [BAMONA]
Brenthis epithore ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 16
Clossiana epithore ; [NACL, #4469]
Boloria epithore ; [BNA: 320, #242, pl. 26]; [BOC, 283]; [Opler]
[epithore-1uU.jpg] identification uncertain 600x539 (~44Kb) upperside USA: Near Crystal Mtn, (46°57'10N 121°28'27W), Pierce Co., WA, 27.7.1999, Photo © Markku Savela [epithore-1vU.jpg] identification uncertain 400x369 (~20Kb) underside USA: Near Crystal Mtn, (46°57'10N 121°28'27W), Pierce Co., WA, 27.7.1999, Photo © Markku Savela

Larva on Viola sempervirens, V. glabella, V. ocellata, V. adunca, V. nephrophylla? [BOC, 283]


California Boloria epithore epithore
Boloria epithore epithore ; [BNA, #242]
C. e. chermocki (E. & S. Perkins, 1966)
Oregon epithore chermocki (E. & S. Perkins, 1966); , TL: 2.9 mi E of Dolph, Yamhill Co., Oregon
Boloria epithore chermocki, Butterflies of America [BOA]
Clossiana epithore chermocki ; [NACL, #4469a]
Boloria epithore chermocki ; [BNA, #242]
C. e. uslui (Koēak, 1984)
British Columbia Boloria epithore uslui Koēak, 1984; (preocc.)
epithore borealis (E. Perkins, 1973); (preocc.), TL: Shingle Creek Rd., Keremeos, BC
obscuripennis (Gunder, 1926); (ab.)
Clossiana epithore borealis ; [NACL, #4469b]
C. e. sierra (E. Perkins, 1973)
California epithore sierra (E. Perkins, 1973); , TL: Sentinel Dome, Yosemite Nat. Park, California
eldorado (Strand, 1915); (ab.)
wawnae (Gunder, 1924); (ab.)
Clossiana epithore sierra ; [NACL, #4469c]

Clossiana alberta (Edwards, 1890)
-. See [About maps] Alberta Fritillary Alberta Alpine Fritillary
Argynnis alberta Edwards, 1890; Can. Ent. 22 (6) : 113, TL: Laggan, Alberta, Canada
banffensis (Gunder, 1932)
Boloria alberta, E.H. Strickland Ent. Mus, U of [ALBERTA]
Boloria alberta, Butterflies of America [BOA]
Boloria alberta, Butterflies and Moths of North America [BAMONA]
Brenthis alberta ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 16
Clossiana alberta ; [NACL, #4472]; [BARU, #204]; [BOW: pl. 13, f. 17]; [BRU2: 50, pl. 35, f. 1-3]
Boloria alberta ; [BNA: 322, #246, pl. 26]; [Opler]
[alberta-1fu.jpg] 1600x1200 (~262Kb) upperside female Canada, Mt. Tripoli, 18.7.1989, Photo © Norbert Kondla leg. [alberta-1fv.jpg] 1600x1200 (~248Kb) underside female Canada, Mt. Tripoli, 18.7.1989, Photo © Norbert Kondla leg. [alberta-1mu.jpg] 1600x1200 (~233Kb) upperside male Canada, Mt. Tripoli, 18.7.1989, Photo © Norbert Kondla leg.
[alberta-1mv.jpg] 1600x1200 (~226Kb) underside male Canada, Mt. Tripoli, 18.7.1989, Photo © Norbert Kondla leg.

Larva on Dryas octopetala, Vaccinium [BARU], Dryas octopetala, Dryas integrifolia [BRU2, 50]


Alberta, British Columbia
C. a. kurentzovi (Wyatt, 1961)
Chukotka alberta kurentzovi (Wyatt, 1961); , TL: Chukotka
Clossiana alberta kurentzovi ; [BARU]; [BRU2, 50]

Clossiana astarte (Doubleday, [1848])
Alberta, British Columbia - Alaska, Washington, Montana. See [About maps] Astarte Fritillary Arctic Ridge Fritillary
astarte (Doubleday, [1848]); Gen. diurn. Lep. (2): pl. 53, f. 5, TL: Rock Lake, Alberta
Argynnis victoria Edwards, 1891; Can. Ent. 23 (9): (195-199)
Boloria astarte, E.H. Strickland Ent. Mus, U of [ALBERTA]
Boloria astarte astarte, Butterflies of America [BOA]
Boloria astarte, Butterflies and Moths of North America [BAMONA]
Brenthis astarte ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 16
Clossiana astarte ; [NACL, #4473]; [BOW: pl. 13, f. 18]
Boloria astarte ; [BNA: 322, #247, pl. 26]; [BOC, 285]; [Opler]

Larva on Saxifraga bronchialis [NAB]

NE.British Columbia, Alberta Boloria astarte astarte
Boloria astarte astarte ; [BNA, #247]

The tritonia species-group

Superspecies tritonia. [BARU]


Clossiana tritonia (Böber, 1812)
-. See [About maps] tritonia (Böber, 1812); Mém. Soc. Imp. Nat. Moscou 3: 20, TL: [Lake Baikal, Irkutsk, Russia]
Clossiana tritonia, Lepidoptera collection [SZMN]
Boloria tritonia, NSG Voucher Specimen [Wahlberg]
Clossiana tritonia ; [BARU, #206a]; [BRU2: 51, pl. 34, f. 7-9, 19-21]
Boloria tritonia ; [Otakar Kudrna]

Larva on Saxifraga bronchialis [BARU]

Baikal Clossiana tritonia tritonia ; [BRU2, 51]
C. t. amphilochus (Ménétriés, 1859)
Amur, Ussuri Clossiana tritonia amphilochus ; [BRU2, 51]

Clossiana elatus (Staudinger, 1892)
Transbaikalia. See [About maps] elatus (Staudinger, 1892); , TL: Mongolia, Kentey
Clossiana elatus ; [BARU, #205]
Clossiana tritonia elatus ; [BRU2, 51]

Larva on Saxifraga bronchialis, Vaccinium uligonosum [BARU]


Clossiana distincta (Gibson, 1920)
-. See [About maps] distincta (Gibson, 1920); Rep. Can. arctic exped. 1913-18. 3 (1): 25(i), 54(i), TL: Harrington Creek, Yukon Territory, Canada: lat 65°05'
Clossiana astarte distincta ; [NACL, #4473a]
Clossiana distincta ; [BARU, #206b]; [BRU2: 51, pl. 34, f. 10-18]

Larva on Saxifraga spinula, (Russia) Salix, (N.America) [BRU2, 51]

Yukon - British Columbia
Misidentified/misapplied:
Boloria astarte distincta; [BNA, #247]
C. d. machati Korshunov, 1987
Polar Urals Clossiana distincta machati ; [BARU]; [BRU2, 51]
C. d. dubatolovi Korshunov, 1987
Transbaikalia Clossiana distincta dubatolovi ; [BARU]; [BRU2, 51]
C. d. tschukotkensis (Wyatt, 1961)
NE.Yakutia, Chukotka Clossiana distincta suntara Dubatolov, 1997
Clossiana distincta tschukotkensis ; [BARU]; [BRU2, 51]
Misidentified/misapplied:
?Boloria astarte tschukotkensis; [BNA, #247]
C. d. ershovi Korshunov & Gorbunov, 1995
C.Yakutia Clossiana distincta ershovi Korshunov & Gorbunov, 1995; , TL: Yakutsk
Clossiana distincta ershovi ; [BRU2, 51]

Clossiana matveevi Gorbunov & Korshunov, 1995
Altai. See [About maps] Clossiana matveevi Gorbunov & Korshunov, 1995; [BARU, #206c], TL: Altai, Kurayskiy Mountain rnage, Aktash, 2700m
Clossiana matveevi, Lepidoptera collection [SZMN]
Clossiana matveevi ; [BRU2: 51, pl. 35, f. 10-11]

 

Clossiana erda (Christoph, 1893)
-. See [About maps] erda (Christoph, 1893); Dt. Ent. Zeit. Iris, 6 (1) : 87, TL: "Vilui" [Vilui R., Yakutia]
Clossiana erda ; [BARU, #203]; [BRU2: 50, pl. 35, f. 4-9]

Larva on Vaccinium vitis-idaea [BRU2, 50]

Yakutia Clossiana erda erda ; [BRU2, 50]
C. e. kitoica Belik, 1996
E.Sayan Clossiana erda kitoica ; [BRU2, 50]
C. e. puella Churkin, 1999
Transbaikalia (Barguzinsky Mts.) Clossiana erda puella Churkin, 1999; , TL: Transbaikalia (Barguzinsky Mts.)
Clossiana erda puella ; [BRU2, 50]
C. e. dulkeiti Kurenzov, 1970
Magadan Clossiana dulkeiti Kurenzov, 1970; , TL: Omsukchansky Mts., Magadan
Clossiana erda dulkeiti ; [BARU]; [BRU2, 50 (?)]

Clossiana iphigenia (Graeser, 1888)
Japan. See [About maps] iphigenia (Graeser, 1888); Berl. Ent. Zs. 32 (1): 90, TL: Amur "downstream of Khabarovsk"
Clossiana iphigenia ; [BARU, #191]; [BRU2: 45, pl. 32, f. 10-12]

Larva on Viola selkirkii, (mainland,Japan) Viola grypoceras, (Sakhalin,Japan) [BARU] [BRU2, 45]

E.Amur, Ussuri, NE.China Clossiana iphigenia iphigenia ; [BRU2, 45]
Sakhalin, Kunashir Argynnis sachalinensis Matsumura, 1908; Ent. Zs. 22 (39) : 157-158:Hokkaido
Clossiana iphigenia sachalinensis ; [BARU]
Clossiana iphigenia alpharatoria ; [BRU2, 45]


Unknown or unplaced taxa

Brenthis morrisii Reakirt, 1866; Proc. Acad. nat. Sci. Philad. 18 (3) : 246, TL: California

Brenthis nenoquis Reakirt, 1866; Proc. Acad. nat. Sci. Philad. 18 (3) : 247, TL: California


21.2.2013 (29)

References:
[ALBERTA]
University of Alberta, E.H. Strickland Entomological Museum; Browse Entomology Collection
[About maps]
Warning! The maps are automatically generated from the textual information, and the process does not always produce acceptable result; See about maps for more info.
[BAMONA]
Opler, Pavulaan, Stanford & Pogue; Butterflies and Moths of North America; Mountain Prairie Information Node
[BARU]
Korshunov & Gorbunov; Butterflies of the Asian part of Russia; English translation by Oleg Kosterin; [Dnevnye babochki aziatskoi chasti Rossii. Spravochnik]
[BHL]
The; Biodiversity Heritage Library; and on; Internet Archive; Note: The linking to the content of Internet Archive is somewhat delicate -- notify me when links break!
[BMAT]
Tennent, 1996; The butterflies of Morocco, Algeria and Tunisia; ISBN 0 906802 05 9
[BNA]
Scott, 1986; The Butterflies of North America
[BOA]
Interactive Listing of American Butterflies; Listado Interactivo de la Mariposas Americanas; Butterflies of America
[BOC]
Robert Michael Pyle, 2002; The Butterflies of Cascadia
[BOW]
Lewis, 1974; Butterflies of the World; ISBN 0 245 52097 X
[BRU2]
Tuzov (ed.), Bogdanov, Churkin, Devyatkin, Dantchenko, Murzin, Samodurov, Zhdanko, 2000; Guide to the Butterflies of Russia and adjacent territories; Libytheidae, Danaidae, Nymphalidae, Riodinidae, Lycaenidae; Volume 2
[CiNii]
http://ci.nii.ac.jp/
[D. Smirnov]
Russia, Moscow; e-mail: D. Smirnov
[EBW]
Smart, Paul, 1976; The Illustrated Encyclopedia of the Butterfly World; ISBN 0 600 31381 6
[ESP]
Heppner, 1981; The dates of E. J. C. Esper's Die Schmetterlinge in Abblidungen... 1776-[1830]; Archives of Natural History (1981) 10 (2): 251-254
[HBN]
Francis Hemming, 1937; HÜBNER, A bibliographical and systematic account of the entomological works of Jacob Hübner and of the supplements thereto by Carl Geyer, Gottfried Franz von Frölich and Gottlieb August Wilhelm Herrich-Schäffer, Vol 1 & 2
[Harri Arkkio]
Finland; For the time being these pictures are only for this site. Other usage not allowed.
[Joseph Belicek]
e-mail: Joseph Belicek
[Kimmo Silvonen]
Espoo, Finland; Kimmo's Lep Site
[LAMAS-2005]
Gerardo Lamas; Entomofauna 26 (6): 57-100; A Bibliography of the Zoological Publications of Hans Fruhstorfer (1886* - 1922+)
[LAMAS-NEO]
Gerardo Lamas; Bibliography of butterflies; An Annocated Bibliography of the Neotropical Butterflies and Skipper (Lepididoptera: Papilionoidea and Hesperioidea); Revised Electronic Edition (2007)
[Lars Andersen]
http://www.pedersen-bike.dk/indextre.html
[MBE]
Moths and Butteflies of Europe; Paolo Mazzei, Diego Reggianti and Ilaria Pimpinelli; http://www.leps.it/
[MRS]
Chou Io (Ed.); Monographia Rhopalocerum Sinensium, 1-2
[Mario Maier]
Europäische Schmetterlinge; e-mail: Mario Maier
[Michael Borsch]
Geesthacht, Germany; contact through; e-mail: Francisco Welter-Schultes
[NAB]
Pyle, 1981; Field Guide to North American Butterflies, National Audubon Society; ISBN 0-394-51914-0
[NACL]
Hodges, 1983; Check List of the Lepidoptera of America North of Mexico
[NORLEP]
Norges Sommerfugler
[Norbert Kondla]
Canada, BC; e-mail: Norbert Kondla
[Oleg Kosterin]
Institue of Cytology & Genetics; Siberian Division of Russian Academy of Sciences; Novosibirsk; e-mail: Oleg Kosterin; http://pisum.bionet.nsc.ru/kosterin/
[Opler]
Opler & Warren, 2003 (April); Butterflies of North America. 2. Scientific Names List for Butterfly Species of North America, north of Mexico.
[Pertti Pakkanen]
e-mail: Pertti Pakkanen; Butterfly and moth observations from Finland
[SPRK]
Seppänen, E. J, 1970; Suomen suurperhostoukkien ravintokasvit, Animalia Fennica 14
[SVF]
Svenska fjärilar; Naturhistoriska riksmuseet; (Sweden)
[SZMN]
Lepidoptera; Siberian Zoological Museum
[Simon Coombes]
1 Park Street, Plymouth, PL3 4BL, UK; 01752 607854; e-mail: Simon Coombes; http://www.butterfly-guide.co.uk
[Wahlberg]
Niklas Wahlberg; The NSG's voucher specimen database; Nymphalidae Systematics Group
[²]
This may require parentheses or not. I don't have the necessary information for this taxon.


Some related literature:
Alphéraky, 1881
Lépidoptčres du district de Kouldją et des montagnes environnantes. (1-3) Horae Soc. ent. Ross. 16 (3-4): 334-435, pl. 14-15 (1881), 17 (1-2): 15-103, pl. 1-3 (1882), 17 (3-4): 156-227, pl. 8-9 (1883)
O. Bang-Haas, 1915
Rhopalocera der Chotan-Ausbeite 1914 Dt. ent. Z. Iris 29 (2/3) : 92-100
Barnes & Benjamin, 1925
A new subspecies of Brenthis myrina Cram. (Lepid.: Nymphalidae) Ent. News 36 : 44
Barnes & McDunnough, 1913
New North American Lepidoptera with notes on described species Contr. nat. Hist. Lep. N. Am. 2 (3) : 93-165, pl. 1-9, I-IX
Boisduval, 1869
Lépidoptčres de la Californie Ann. Soc. ent. Belg. 12 : 5-28,37-94
Butler, 1877
Description of a new species of Argynnis from Arctic America Ent. mon. Mag. 13 : 206
Butler, 1882
On Lepidoptera collected in Japan and the Corea by Mr. W. Wykeham Perry Ann. mag. nat. Hist. (5) 9 (49) : 13-20
Böber, 1812
Contribution de la description de quelques nouvelles espčces de papillons découverts en Sibérie ... Mém. Soc. Imp. Nat. Moscou 3: 20-21,?72
Christoph, 1893
Lepidoptera Nova Faunae Palaearcticae Dt. Ent. Zeit. Iris, 6 (1) : 86-96
Cramer, [1777]
Uitlandsche Kapellen (Papillons exotiques) Uitl. Kapellen 2 (9-16): 1-152, pl. 97-192 (1777)
dos Passos, 1936
Some early stages of Brenthis montinus Scudder Can. Ent. 68: 239-241
Doubleday, [1848]
The genera of diurnal Lepidoptera, comprising their generic characters, a notice of their habitats and transformations, and a catalogue of the species of each genus; illustrated with 86 plates by W. C. Hewitson Gen. diurn. Lep. (1): 1-18, pl. A, 1-4 (1-2) (1846), (1): 19-132, pl. 4*, 5-25, 28 (3-14) (1847), (1): 133-200, pl. 1*, 26, 27, 29, 31-44 (15-23) (1848), (1): 201-242, pl. 30, 45-52, 56-58, 60-62, 64 (24-31) (1849), (1): 243-250 [?], (2): 243-326, pl. 53-55, 63, 65, 66 (32-38) (1850), (2): 327-466, pl. 59, 67-77 (39-50) (1851), (2): 467-534, 143, 144, pl. 78-80 (Index) (51-54) (1852)
Duponchel, 1832
Histoire naturelle des Lépidoptčrs ou Papillons de France - Supplement. Diurnes in Godart, Hist. nat. Lépid. Fr. (Suppl.) 1 : 1-466, pl. 1-50
Dyar, 1903
A List of North American Lepidoptera and Key to the Literature of this Order of Insects Bull. U.S. nat. Mus. 52 : 1-723
Edwards, 1864
Descriptions of certain species of Diurnal Lepidoptera, found within the limits of the United States and British America. (No. 1-3) Proc. ent. Soc. Philad. 2 (1): 14-22, pl. 1, 11 (1863), (2): 78-82 (1863), (4): 501-507 (1864)
Edwards, 1871
Descriptions of new species of North American butterflies Trans. amer. ent. Soc. 3 : 266-277
Edwards, 1883
Descriptions of new species of diurnal Lepidoptera found in British America and the United States Can. Ent. 15 (2) : 32-36
Edwards, 1890
Description of a new species of Argynnis from Canada Can. Ent. 22 (6) : 113-114
Edwards, 1891
Description of a new species of Argynnis from Alberta Territory Can. Ent. 23 (9) : 198-199
Esper, 1783
Die Schmetterlinge in Abbildungen nach der Natur mit Beschreibungen. Theil I. Die Tagschmetterlinge. Fortsetzung. Band 2 Die Schmett. Th. I, Bd. 2 (1): 1-36, pl. 51-56 (1780), (2): 37-68, pl. 57-62 (1781), (3): 69-100, pl. 63-68 (1781), (4): 101-124, pl. 69-74 (1781), (5): 125-140, pl. 75-80 (1782), (7): 141-156, pl. 81-82 (1783), (8): 157-172, pl. 83-88 (1783), (9): 173-184, pl. 89-92 (1784), (28): 185-190, pl. 93 (1786)
Esper, 1789; Esper, 1794; Esper, 1798
Die Schmetterlinge in Abbildungen nach der Natur mit Beschreibungen. Theil I. Die Tagschmetterlinge. Supplement Theil 1. Abschnitt 1 Die Schmett., Suppl. Th 1 (1-2): 1-8, pl. 94 (1789), (3-4): 9-60, pl. 95-101 (1789), (5-6): 61-68, pl. 102-106 (1794), ([7]): [69-88], pl. [107-109] (1798), (8-9): 89-104, pl. 110-112 (1800), (10): 105-120, pl. 113-116 ([1803-1804])
Erschoff, 1870
Description d'un Argynnisnouveau de la Sibérie orientale Bull. Soc. Imp. Nat. Moscou 43 (1) : 112-114
Eversmann, 1837
Kurze notizen ueber einige Schmetterlinge Russlands Bull. Soc. imp. Nat. Moscou 1837 (1): 5-32
Eversmann, 1844
De quibsudam Lepidopteria Rossicis Bull. Soc. Imp. Nat. Moscou 17 (3) : 588-604, pl. 14-16
Fabricius, 1775
Systema Entomologiae, sistens Insectorum Classes, Ordines, Genera, Species, Adiectis Synonymis, Locis, Descriptionibus, Observationibus. Syst. Ent.: 832pp
Fruhstorfer, 1907
Neue Argynnis Ent. Zs. 21 (27) : 163-164 (5 October)
Fruhstorfer, 1908
Neue Argynnis und Melitaea Int. ent. Zs. 1 (41) : 310 (11 January)
Fruhstorfer, 1909
Neue palaearktische Rhopaloceren Int. ent. Zs. 3 (4): 20-21 (24 April)
Fruhstorfer, 1909
Neue Palaearkten Int. ent. Zs. 3 (20): 112-113 (14 August), (21): 120-121 (21 August)
Gall & Sperling, 1980
A new high altitude species of Boloria from southwestern Colorado, with a discussion of phenetics and hiearchical decisions J. Lep. Soc. 43: 230-252
Gibson, 1920
The Lepidoptera collected by the Canadian arctic expedition 1913-1918. Part I in Report of the Canadian arctic expedition 1918-18. Vol. III, Insects Rep. Can. arctic exped. 1913-18. 3 (1): 21i-22i
Graeser, 1888
Beiträge zur Kenntnis der Lepidopteren-Fauna des Amurlandes Berl. Ent. Zs. 32 (1): 33-153 (1888), (2): 309-414 (1889)
Gunder, 1926
Several new aberrant Lepidoptera (Rhopalocera) Ent. News 37 (1) : 1-9, pl. 1
Gunder, 1928
Additional transitional forms Can. Ent. 60 : 162-168, 2pls
Guppy & Kondla, 2002
The correct spellings and type localities of butterflies named from Mount Rainier, Washington, USA J. Lep. Soc. 56 (3): 180-131
Herrich-Schäffer, [1847]
Systematische Bearbeitung der Schmetterlinge von Europa, zugleich als Text, Revision und Supplement zu J. Hübner's Sammlung europäischer Schmetterlinge, 1843 (-1855), Die Tagfalter Syst. Bearb. Schmett. Europ. 1 (9): Title ("1843"), (1): [1-3], 4-16, (ii) pl. 1-5, f. 1-29 (1843), (2): 17-40, (ii) pl. 29-34, f. 131-156 (1843), (3): 41-56, (ii) pl. 6-12, f. 30-60, pl. 35-36, f. 157-164 (1844), (4): 57-80, (ii) pl. 13-22, f. 61-107 (1844), (5): 81-90, (ii) pl. 23-28, f. 108-130, pl. 37-41, f. 165-189 (1844), (6): 91-98, (ii) pl. 42-44, f. 190-203 (1844), (7): 99-114, (ii) pl. 45-52, f. 204-248 (1844), (9): 115-122 (1844), (10): 123-146, (ii) pl. 53-57, f. 249-274 (1845), (11): 147-162 (1845), (12): (iii) pl. 1-3, f. 1-17 (1845), (13): 163-164, (ii) pl. 58-65, f. 275-315 (1846), (14): (ii) pl. 66-70, f. 316-338, (iii) pl. 4-5, f. 18-30 (1846), (26): (ii) pl. 71-77, f. 339-376 (1847), (32): (ii) pl. 78-82, f. 377-400 (1848), (36): (ii) pl. 83-89, f. 401-426, (iii) pl. 6, f. 31-36 (1848), (37): (ii) pl. 90-91, f. 427-437 (1849), (43): (ii) pl. 92-94, f. 438-450, 452-453 (1850), (45): (ii) pl. 95-102, f. 453-489 (1850), (47): (ii) pl. 103, f. 490-495 (1851), (49): (ii) pl. 104-109, f. 496-528 (1851), (51): (ii) pl. 110-118, f. 529-570 (1851), (54): (ii) pl. 119-123, f. 571-596 (1852), (56): (ii) pl. 124-127, f. 597-618, pl. 130-132, f. 628-639, pl. 134, f. 646-651 (1852), (59): (ii) pl. 128-129, f. 619-627, pl. 133, f. 640-645 (1853), (65): (iii) pl. 7, f. 37-42, (index) [1], 2-20 (1854), (66): (index) 21-24 (1855)
Huang & Murayama, 1992
Butterflies of Xinjiang province, China Tyō to Ga 43 (1) : 1-22
Hübner, [1799-1800]; Hübner, [1803-1804]
Sammlung europäischer Schmetterlinge. I. Papiliones - Falter ("Erste Band") Samml. eur. Schmett. [1] : 3-74, : pl. 1-88, f. 1-446 (1799-1800), : pl. 89-96, f. 447-490 (1800-1803), : pl. 97-114, f. 491-588 (1803-1804), : pl. 115-119, f. 589-612 (1805), : pl. 120-124, f. 613-636 (1805-1806), : pl. 125-128, f. 637-652 (1806-1808), : pl. 129-144, f. 653-727 (1808-1813), : pl. 145-147, f. 728-741 (1814-1816), : pl. 148-150, f. 742-756 (1816-1817), : pl. 151, f. 757-762 (1818-1819), : pl. 152-154, f. 757-774 (1819-1822), : pl. 155, f. 775-778 (1822-1823), : pl. 156-161, f. 779-806 (1823), : pl. 162-175, f. 807-874 (1823-1824), : pl. 176, f. 869-874 (1824-1825), : pl. 177-181, f. 875-895 (1825-1826), : pl. 182-187, f. 896-981 (1827-1828), : pl. 188-195, f. 922-968 (1828-1832), : pl. 196, f. 969-974 (1832-1833), : pl. 197-201, f. 975-997 (1834-1836), : pl. 202-203, f. 998-1009 (1836-1838), : pl. 204-207, f. 1010-1029 (1837-1838)
Kohler, 1977
Revision of North American Boloria selene (Nymphalidae) with description of a new subspecies J. Lep. Soc. 31 (4): 243-268
Krulikovsky, 1909
[russian] Rev. russ. ent. 9 (3) : 292-323
Linnaeus, 1758
Systema Naturae per Regna Tria Naturae, Secundum Clases, Ordines, Genera, Species, cum Characteribus, Differentiis, Symonymis, Locis. Tomis I. 10th Edition Syst. Nat. (Edn 10) 1 : 1-338, : 339-824
Linnaeus, 1767
Systema Naturae per Regna tria Naturae, secundum Classes, Ordines, .... Editio Duocecima Reformata. Tom. 1. Part II. Syst. Nat. (Edn 12) 1 (2): -,533-1327
Matsumura, 1908
Die Nymphaliden Japans Ent. Zs. 22 (39) : 157-158
Matsumura, 1927
A list of the butterflies of Corea, with description of new species, subspecies and aberrations Insecta Matsumurana 1 (4) : 159-170
Matsumura, 1927
Some new Butterflies Ins. Matsumurana 2 (2) : 114-118
Matsumura, 1929
Some new butterflies from Korea received from Mr. T. Takamuku Insecta Matsumurana 3 (4) : 152-156
McLachlan, 1878
Report on Insecta (including Arachnida) collected by Captain Feilden and Mr. Hart between Parallels of 78° and 83° North Latitude, during the recent Arctic Expedition J. Linn. Soc. Zool. Lond. 14 : 98-122
Reakirt, 1866
Descriptions of some new species of Diurnal Lepidoptera Proc. Acad. nat. Sci. Philad. 18 (3) : 238-249
Reuss, 1920
Die Androconien von Yramea cytheris Drury und nächtststehenden analogen Schuppenbildungen bei Dione Hbn. und Brenthis Hbn. (Lep.) Ent. Mitt. 9 : 192
Sheljuzhko, 1929
Einige neue palaearktische Lepidopteren-Formen Mitt. Münch. Ent. Ges. 19 : 347-362, pl. 27-28
Sovinsky, 1905
Tri novyk kavkazskikh raznovidnosti cheshuekrylykh [Trois nouvelles formes de Lepidopteres du Caucase] Revue Russe Ent. 5 (3-4) : 108-110
Stallings & Turner, 1947
New American butterflies Can. Ent. 78 (7-8): 134-135
Staudinger, 1861
Reise nach Finmarken Stettin Ent. Ztg 22 (10-12): 325-341 (Wocke & Staudinger, 1861), 22 (10-12): 342-404 (Macrolep., Staudinger, 1861), 23 (1-3): 30-78 (Microlep., Wocke, 1862), 23 (4-6): 233-257 (Microlep., Wocke, 1862)
Staudinger, 1881
Beitrag zur Lepidopterenfauna Central-Asiens Stettin ent. Ztg 42 (7-9): 253-300, (10-12): 393-424
Thunberg, 1791
Dissertio Entomologica sistens Insecta Suecica, ... Diss. Ent. sistens Insecta Suecica (1): 1-24 (1 plate) (1784), (2): [25-31] ??], 32-46 (9 unnumbered) (1791), (3): [47],48-52 (2 unnumbered) (1792), (4): 53-62 (1792)
Troubridge & Wood, 1990
Biology and taxonomic status of Boloria natazhati (Gibson) J. Lep. Soc. 44 (3): 180-187
Wagner, 1913
Beitrag zur Lepidopterenfauna des Iligebietes sowie des Sary-Dschas (Asia centr.) Ent. Mitt. 2 (1): 22-30, (2): 50-62, (3): 88-95, (4): 112-126, (5): 153-158, (6): 185-190, (7/8): 244-254, (9): 285-288
Wyatt, 1961
Additions to the Rhopalocera of Afghanistan with descriptions of new species and subspecies J. Lep. Soc. 15 (1): 1-18, 3pls
Zetterstedt, 1839
Insecta Lapponica Insecta Lapponica : 1-1014 ([1839]), : 1015-1140 (1840)


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Keep in mind that the taxonomic information is copied from various sources, and may include many inaccuracies. Expert help is welcome.