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Hesperia Fabricius, 1793
Branded Skippers
Hesperia Fabricius, 1793; Ent. Syst. 3 (1): 258, TS: Papilio comma Linnaeus
Pamphila Fabricius, 1807; Magazin f. Insektenk. (Illiger) 6: 287, TS: Papilio comma Linaneus
Diorthosus Rafinesque, 1815
= ; [NL4A, 68]
Steropes Rafinesque, 1815; (unavail.)
Phidias Rafinesque, 1815; (repl. name)
Symmachia Sodoffsky, 1837; Bull. Soc. imp. Nat. Moscou 1837 (6) : 82 (repl. Hesperia Fabricius, preocc. Hübner, [1819])
Ocytes Scudder, 1872; 4th Ann. Rep. Peabody Acad. Sci. (1871): 76, TS: Hesperia metea Scudder
= ; [NL4A, 68]
Anthomaster Scudder, 1872; 4th Ann. Rep. Peabody Acad. Sci. (1871): 78, TS: Hesperia leonardus Harris
= ; [NL4A, 68]
Urbicola Tutt, 1905; Nat. Hist. Br. Lepid. 8 : 84, TS: Papilio comma Linnaeus
= ; [NL4A, 69]
Pamphilus Ihering, 1908
= ; [NL4A, 69]

Herpena ; Edwards, 1872, (missp.)
Anthosmastes ; Scudder, 1881, (missp.)
Phamphila ; Mabille, 1891, (missp.)
Pmaphila ; Maynard, 1891, (missp.); Mabille, 1891, (missp.)
Pamphala ; Maynard, 1891, (missp.)
Pamphilia ; Moffat, 1894, (missp.)
Hesperis ; Brunner von Wattenwyl, 1897, (missp.)
Pamhpila ; Maynard, 1901, (missp.)
Pamhila ; Maynard, 1901, (missp.)
Herperia ; Woodworth, 1913, (missp.)
Pamphilv ; Rocha, 1936, (missp.)
Hesperria ; Justino, 1949, (missp.)
Hespena ; dos Passos, 1964, (missp.)
Uribicola ; Bridges, 1988, (missp.)
Hesperia ; Godman & Salvin, [1899], Biol. centr.-amer., Lep. Rhop. 2: 449
Hesperia (Hesperiinae) ; [NL4A, 68]


[sp-1.jpg] 640x546 (~33Kb) USA: Table Mtn, (~ 47°19N 120°42W), Kittitas Co., WA, 30.7.1999, Photo © Markku Savela [sp-1x.jpg] 500x401 (~22Kb) USA: Table Mtn, (~ 47°19N 120°42W), Kittitas Co., WA, 30.7.1999, Photo © Markku Savela [sp-2.jpg] 500x630 (~38Kb) USA: Table Mtn, Kittitas Co., WA, 30.7.1999, Photo © Markku Savela
[sp-3.jpg] 600x505 (~39Kb) USA: FR9708, Table Mtn (47°18'28N 120°41'25W), Kittitas Co., WA, 19.7.1999, Photo © Markku Savela [sp-4.jpg] 800x577 (~47Kb) USA: Table Mtn (~ 47°19N 120°42W), Kittitas Co., WA, 30.7.1999, Photo © Markku Savela [sp-5.jpg] 500x642 (~48Kb) USA: Mima Mounds Natural Area Preserve, Thurston Co., WA, 13.8.2001, Photo © Markku Savela

The exact identification of these species is still unknown, but tentatively assumed to belong into this group.



fiHesperia comma (Linnaeus, 1758)
NAF, EU, Temperate Asia, NA. See [About maps] Valkotäpläpaksupää Silver-spotted Skipper Common Branded Skipper Holarctic Grass Skipper Allmän ängssmygare Kommasmygare Kommafalter le Comma Dorada manchas blancas Karłątek klinek
Papilio comma Linnaeus, 1758; Syst. Nat. (Edn 10) 1 : 484, TL: [Sweden]
Hesperia comma, E.H. Strickland Ent. Mus, U of [ALBERTA]
Hesperia comma, Lepidoptera collection [SZMN]
Hesperia comma, Butterflies and Moths of North America [BAMONA]
Hesperia comma, Larvae of North-European Lepidoptera [Kimmo Silvonen]
Hesperia comma, Svenska fjärilar [SVF]
Hesperia comma, Norges Sommerfugler [NORLEP]
Hesperia comma, Moths and Butterflies of Europe [MBE]
Hesperia comma, Captain's European Butterfly Guide [Simon Coombes]
Erynnis comma ; Godman & Salvin, [1900], Biol. centr.-amer., Lep. Rhop. 2: 477; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 49
Hesperia comma ; [BOW: pl. 11, f. 18-19]; [BRU, 131]; [NACL, #4020]; [MRS, 732]; [BOC, 74]; [Opler]; [NL4A, #1629]; [Otakar Kudrna]
[comma-1u.jpg] 500x458 (~84Kb) upperside Martin down, Dorset UK 06/08/95, Photo © Simon Coombes [comma-1v.jpg] 500x350 (~52Kb) underside Martin down, Dorset UK 06/08/95, Photo © Simon Coombes [comma-2v.jpg] 350x500 (~24Kb) underside A patch of a meadow steppe at the village Fadino, 10 km S of the city Omsk, West Siberia, Russia. 15th August 1995, Photo © Oleg Kosterin

Larva on Holcus, Poa [SPRK]


H. c. catena (Staudinger, 1861)
H. c. benuncas (Oberthür, 1912)
Morocco, Algeria Augiades benuncas Oberthür, 1912; Bull. Soc. ent. Fr. 1912 : 349-351, TL: Lambessa (Algeria)
Hesperia comma maroccana Picard, 1950; Bull. Soc. Sci. nat. Maroc 28: 129-130, TL: Afraou des Beni Abdallah, Daya Chicker, Tameghilt, Col du Taghzeft (Morocco)
Hesperia comma benuncas ; [BMAT: 98, pl. 31, f. 50-53]

Larva on (in Spain) Lotus corniculatus, Ornithopus perpusillus, Poaceae [BMAT]

H. c. pallida Staudinger, 1901
Transcaucasia Hesperia comma pallida Staudinger, 1901; in Staudinger & Rebel, Cat. Lepid. palaearct. Faunengeb., 1: 92, TL: Turkey

In [BMAT], a synomym of Hesperia comma benuncas.

H. c. mixta Alpheraky, 1881
comma mixta Alpheraky, 1881; , TL: Tian-Shan
H. c. shandura Evans, 1949²
comma shandura Evans, 1949²; , TL: Chitral
H. c. lena Korshunov & P. Gorbunov, 1995
comma lena Korshunov & P. Gorbunov, 1995; , TL: Yakutia, Chukot Peninsula
Hesperia comma lena, Lepidoptera collection [SZMN]
H. c. planula Korshunov, 1995
comma planula Korshunov, 1995; , TL: W. Siberia
H. c. sushkini Korshunov, 1995
H. c. dimila (Moore, [1875])
Pamphila dimila Moore, [1875]; Proc. zool. Soc. Lond. 1874 (4) : 576, TL: Runang Pass, Busahir
Hesperia comma dimila ; [MRS, 732]
H. c. borealis Lindsey, 1942
Hesperia comma borealis ; [NACL, #4020d]
H. c. dodgei (Bell, 1927)
Hesperia comma dodgei ; [NACL, #4020i]
H. c. hulbirti Lindsey, 1939
W.Washington (Olympics) Hesperia comma hulbirti Lindsey, 1939; , TL: Hurrican Hill, Olympic Mts., Washington
Hesperia comma hulbirti ; [NACL, #4020k]; [BOC, 74]
H. c. laurentina (Lyman, 1892)
Canada, Minnesota comma laurentina (Lyman, 1892); Can. Ent. 24 (3) : 58 (?1893, vol. 25?)
Erynnis comma laurentina ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 50
Hesperia laurentina ; [BOW: pl. 22, f. 2]
Hesperia comma laurentina ; [NACL, #4020c]
H. c. manitoba (Scudder, 1874)
N.Cascades (N.Washington, S.British Columbia) comma manitoba (Scudder, 1874); Mem. Boston Soc. nat. Hist. 2 (Pt 3, No. 4): 351, TL: "Colorado; Canada"
Hesperia colorado Dyar, 1905, not Scudder, 1874
Erynnis comma manitoba ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 50
Hesperia comma manitoba ; [NACL, #4020a]; [BOC, 74]

Larva on Festuca ovina [BOC]

H. c. ochracea Lindsey, 1941
Hesperia comma ochracea ; [NACL, #4020l]
H. c. oroplata Scott, 1981
H. c. susanae Miller, 1962
Arizona Hesperia comma susanae Miller, 1962; , TL: Arizona
Hesperia comma susanae, Butterflies of Southeastern Arizon [Bruce Walsh]
Hesperia comma susanae ; [NACL, #4020n]; [NL4A, #1629b]
H. c. yosemite Leussler, 1933
Hesperia colorado Garth, 1935, not Scudder, 1874
Hesperia harpalus Lindsey, 1940, not Edwards, 1881
Hesperia comma yosemite ; [NACL, #4020f]

Hesperia assiniboia (Lyman, 1892)
assiniboia (Lyman, 1892); Can. Ent. 24 (3) : 57
Hesperia assiniboia, E.H. Strickland Ent. Mus, U of [ALBERTA]
Hesperia assiniboia, Butterflies and Moths of North America [BAMONA]
Erynnis comma assiniboia ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 50
Hesperia comma assiniboia ; [NACL, #4020b]
Hesperia assiniboia ; [Opler]

Hesperia colorado (Scudder, 1874)
California - S.British Columbia, Alberta. See [About maps] Western Branded Skipper Common Branded Skipper
colorado (Scudder, 1874); Mem. Boston Soc. nat. Hist. 2 (Pt 3, No. 4): 349
Hesperia colorado, Butterflies and Moths of North America [BAMONA]
Erynnis comma colorado ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 49
Hesperia comma colorado ; [NACL, #4020m]
Hesperia colorado ; [BOC, 74]; [Opler]; Shapiro & Forister, 2005, J. Lep. Soc. 59 (3): 161-165
H. c. harpalus (Edwards, 1881)
Idaho Branded Skipper
colorado harpalus (Edwards, 1881); Trans. Amer. Ent. Soc. 9 : 3, TL: Carson City, Nevada
= comma idaho (Edwards, 1883); [BOC, 74]
idaho (Edwards, 1883); Can. Ent. 15 (8) : 148, TL: Lake Tahoe, Placer Co., California
?cabelus (Edwards, 1881); Trans. Amer. Ent. Soc. 9 : 4
= comma harpalus (Edwards, 1881); [NACL]
?idaho (Edwards, 1883)
= comma harpalus (Edwards, 1881); [NACL]
?manitoba (W. G. Wright, 1905, not Scudder, 1874)
= comma harpalus (Edwards, 1881); [NACL]
?colorado (W. G. Wright, 1905, not Scudder, 1874)
= comma harpalus (Edwards, 1881); [NACL]
?oregonia (W. G. Wright, 1905, not Edwards, 1883)
= comma harpalus (Edwards, 1881); [NACL]
?viridis (Comstock, 1927, not Edwards, 1883)
= comma harpalus (Edwards, 1881); [NACL]
?leussleri (Lindsey, 1942, not Lindsey, 1940); (part)
= comma harpalus (Edwards, 1881); [NACL]
?Hesperia ruricola dos Passos, not Boisduval, 1852
= comma harpalus (Edwards, 1881); [NACL]

Erynnis harpalus ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 50
Erynnis comma idaho ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 50
Hesperia comma idaho ;
Hesperia colorado idaho ; [BOC, 74]
Erynnis cabelus ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 50
Hesperia harpalus ; [BOW: pl. 22, f. 9 (text)]
Hesperia comma harpalus ; [NACL, #4020e]
Hesperia colorado idaho ; [BOC, 74]

[idaho-1vU.jpg] identification uncertain 700x830 (~68Kb) underside USA: FR 9705 nr. Liberty and Lion Gulch, (47°17'44N 120°39'54W ±0.5km), Kittitas Co., WA, 14.8.2001, Photo © Markku Savela

Larva on Muhlenbergia, Stipa, Andropogon, Lolium [BNA, (for idaho?)]

H. c. oregonia (Edwards, 1883)
Oregon Branded Skipper
Pamphila oregonia Edwards, 1883; Can. Ent. 15 (8) : 150, TL: Trinity Co, California
?ruricola (Wright, 1905, not Boisduval, 1852)
= comma oregonia (Edwards, 1883); [NACL]
?Hesperia juba Comstock, 1927, not Scudder, 1872
= comma oregonia (Edwards, 1883); [NACL]
?Hesperia nevada Comstock, 1927, not Scudder, 1872
= comma oregonia (Edwards, 1883); [NACL]
?Hesperia manitoba Blackmore, 1927, not Scudder, 1874
= comma oregonia (Edwards, 1883); [NACL]
?Hesperia colorado Blackmore, 1927, not Scudder, 1874
= comma oregonia (Edwards, 1883); [NACL]
?Hesperia idaho Leighton, 1946, not Edwards, 1883
= comma oregonia (Edwards, 1883); [NACL]
Erynnis comma oregonia ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 49
Hesperia comma oregonia ; [NACL, #4020j]
Hesperia colorado oregonia ; [BOC, 74]

Larva on Festuca idahoensis, Lolium, Bromus [BOC]

H. c. mattoonorum McGuire, 1998
Hesperia colorado mattoonorum McGuire, 1998; in Emmel, Systematics of W. North Am. Butts: (461-474), TL: Smith R., Del Norte Co., California
Hesperia colorado mattoonorum ; [BOC, 74]
H. c. tildeni Freeman, 1956
Hesperia tildeni Freeman, 1956; Lepid. News. 9 (6): (196-198), TL: Cherry Flat Reservoir, Santa Clara Co., California
Hesperia comma tildeni ; [NACL, #4020h]
Hesperia colorado tildeni ; Opler & Warren, 2002
H. c. leussleri Lindsey, 1940
California Hesperia comma leussleri Lindsey, 1940; , TL: California
Hesperia colorado leussleri, Listing of Mexican Butterflies [ILMB]
Hesperia colorado leussleri, Butterflies of America [BOA]
Hesperia comma leussleri ; [NACL, #4020g]; [NL4A, #1629a]
Hesperia colorado leussleri ;

Hesperia caucasica Riabov, 1926²

Hesperia florinda (Butler, 1878)
Transbaikalia - Korea, Amur, Ussuri, Japan. See [About maps] florinda (Butler, 1878); Cistula Entomologica, 2(19): 285, TL: Japan
repugnans Staudinger, 1892
mikado Leech, 1895
Hesperia florinda ; [BRU, 132]
Hesperia comma florinda ; [MRS, 732]

Larva on Carex [BRU, Fukuda et al., 1984]

H. f. rozhkovi Kurentzov, 1970
Transbaikalia
[rozhkovi-1m.jpg] 800x600 (~85Kb) male A forb meadow on a SE slope 12 km SE of the village Gazimurskii Zavod, Gazimurskozavodskoi District, E Chita Province, E Transbaikalia, Siberia, Russia. 24th July 1997, on Dianthus versicolor Fischer ex Link, Photo © Oleg Kosterin


Hesperia uncas Edwards, 1863
W.Great Plains, Nevada - Mexico. See [About maps] Uncas Skipper White-vein Skipper
Hesperia uncas Edwards, 1863; Proc. ent. Soc. Philad. 2 (1): 19, TL: Denver, Colorado
Hesperia uncas, E.H. Strickland Ent. Mus, U of [ALBERTA]
Hesperia uncas, Listing of Mexican Butterflies [ILMB]
Hesperia uncas uncas, Butterflies of America [BOA]
Hesperia uncas, Butterflies and Moths of North America [BAMONA]
Hesperia unkas[sic, recte uncas] ; Mabille
Hesperia uncus[sic, recte uncas] ; Brown, Eff & Rotger, 1956
Erynnis uncas ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 50
Hesperia uncas ; [NACL, #4018]; [BOW: pl. 22, f. 10]; [Opler]; [NL4A, #1634]

Larva on Stipa nevadensis, Bouteloua gracilis [NAB]

Colorado, ...? Hesperia ridingsii Reakirt, 1866; , TL: USA(Colorado)
= ; [NL4A, #1634a]
unkas ; Mabille, 1904, (missp.)
ridingsi ; Lindsey, 1921, (missp.)
riddingsii ; Draudt, 1923, (missp.)
uncus ; Brown et al., 1956, (missp.)
Hesperia uncas uncas ; [NL4A, #1634a]
H. u. lasus (Edwards, 1884)
Arizona uncas lasus (Edwards, 1884) (Pamphila); Papilio 4 (3) : 54, TL: Arizona
Hesperia uncas lasus, Listing of Mexican Butterflies [ILMB]
Hesperia uncas lasus, Butterflies of America [BOA]
Hesperia uncas lasus, Butterflies of Southeastern Arizon [Bruce Walsh]
Erynnis lasus ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 50
Hesperia uncas lasus ; [NACL, #4018a]; [NL4A, #1634c]
H. u. macswaini MacNeill, 1964
Hesperia uncas macswaini MacNeill, 1964; Blancos Corral, White Mts., Mono Co., California
Hesperia uncas macswaini, Butterflies of America [BOA]
Hesperia uncas macswaini ; [NACL, #4018b]; [BOC, 72]

Larva on Stipa nevadensis, S. pinetorum [BOC]

H. u. gilberti MacNeill, 1964
Mexico Hesperia uncas gilberti MacNeill, 1964; , TL: Mexico(Distrito Federal)
Hesperia uncas gilberti, Listing of Mexican Butterflies [ILMB]
Hesperia uncas gilberti, Butterflies of America [BOA]
Hesperia uncas gilberti ; [NL4A, #1634b]

Hesperia juba (Scudder, 1872)
WNA, S.British Colombia. See [About maps] Juba Skipper Yuba Skipper Jagged-border Skipper
Pamphila juba Scudder, 1872; 4th Ann. Rep. Peabody Acad. Sci. (1871): 77, TL: neighborhood of Salt Lake City, Utah [or California?]
ab. ogdenensis (Holland, 1931) (Erynnis); , TL: Utah
= ; [NL4A, #1630]
Hesperia nevada Garth, 1934, not Scudder, 1874
Hesperia juba, Listing of Mexican Butterflies [ILMB]
Hesperia juba, Butterflies of America [BOA]
Hesperia juba, Butterflies and Moths of North America [BAMONA]
suba ; Möschler, 1878, (missp.)
Erynnis comma juba ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 50
Hesperia juba ; [NACL, #4019]; [Opler]; [NL4A, #1630]
Misidentified/misapplied:
Hesperia viridis Wright, 1905; ( not Edwards, 1883)
Hesperia colorado Wright, 1905; (not Scudder, 1874)
[juba-1vU.jpg] identification uncertain 500x609 (~45Kb) underside USA: Mima Mounds Natural Area Preserve, Thurston Co., WA, 13.8.2001, Photo © Markku Savela

Larva on Stipa, Poa pratensis, Bromus, Deschampsia [BOC]



Hesperia lindseyi (Holland, 1930)
SW.Oregon - California - SW.Arizona. See [About maps] Lindsey's Skipper Lindsey's Branded Skipper Lost-egg Skipper
Erynnis lindseyi Holland, 1930; , TL: Nellie and Ukiah, California
Hesperia lindseyi, Butterflies and Moths of North America [BAMONA]
Hesperia lindseyi ; [NACL, #2032]; [BOC, 78]; [Opler]; [NL4A, #1631]

Larva on Festuca, Danthonia [NAB], Festuca idahoensis, Danthonia californica [BOC]

California Hesperia lindseyi lindseyi ; [NL4A, #1631a]
H. l. macneillii Emmel, Emmel & Mattoon, 1998
California Hesperia lindseyi macneillii Emmel, Emmel & Mattoon, 1998; in Emmel, Systematics W. North Am. Butts.: (475-480)
Hesperia lindseyi macneillii ; Severns & Severns, 2005, J. Lep. Soc. 59 (3): f. 1E-F
H. l. eldorado Emmel, Emmel & Mattoon, 1998
California Hesperia lindseyi eldorado Emmel, Emmel & Mattoon, 1998; in Emmel, Systematics W. North Am. Butts.: (475-480)
H. l. septentrionalis Emmel, Emmel & Mattoon, 1998
S.Oregon, N.California Hesperia lindseyi septentrionalis Emmel, Emmel & Mattoon, 1998; in Emmel, Systematics W. North Am. Butts.: (475-480), TL: Little Shata R., E of Montague, Siskiyou Co., California
Hesperia lindseyi septentrionalis, Butterflies of America [BOA]
Hesperia lindseyi septentrionalis ; [BOC, 78]; Severns & Severns, 2005, J. Lep. Soc. 59 (3): f. 1C-D
H. l. mccorklei Severns & Severns, 2005
SW.Oregon Hesperia lindseyi mccorklei Severns & Severns, 2005; J. Lep. Soc. 59 (3): 166, f. 1A-B, TL: Oregon, Douglas Co., Hillside N of Jackson Creek Road, 3.8mi E of junction with South Umpqua Road

?Hesperia liberia Plötz, 1883
Hesperia liberia Plötz, 1883; Stett. ent. Ztg 44 (4-6): 220

Hesperia woodgatei (Williams, 1914)
Arizona, New Mexico, S.Texas, N.Mexico. See [About maps] Apache Skipper
Pamphila woodgatei Williams, 1914; , TL: New Mexico
Hesperia liberia Evans, 1955, not Plötz, 1883
Hesperia woodgatei, Listing of Mexican Butterflies [ILMB]
Hesperia woodgatei, Butterflies of America [BOA]
Hesperia woodgatei, Butterflies and Moths of North America [BAMONA]
Hesperia woodgatei, Butterflies of Southeastern Arizon [Bruce Walsh]
Hesperia woodgatei ; [NACL, #4021]; [Opler]; [NL4A, #1636]

Larva on Poaceae [NAB]


Hesperia ottoe Edwards, 1866
Nebraska, Arizona, Kansas, ?. See [About maps] Ottoe Skipper
Hesperia ottoe Edwards, 1866; Proc. ent. Soc. Philad. 6 (4) : 207, TL: Kansas
Hesperia ottoe, Butterflies and Moths of North America [BAMONA]
Erynnis ottoe ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 50
Hesperia ottoe ; [NACL, #4022]; [BOW: pl. 22, f. 5 (text)]; [Opler]

Larva on Poaceae, Leptoloma cognatum [NAB]


Hesperia leonardus Harris, 1862
Canada - Florida. See [About maps] Leonardus Skipper
Hesperia leonardus Harris, 1862; Treatise Ins. injur. Veget. (3rd edn): 314, f. 138
Hesperia leonardus f. stallingsi Freeman, 1944; Bull. Brooklyn Ent. Soc. 38 (5): (153-154), TL: Blendon, Franklin Co., Ohio
Hesperia leonardus, Butterflies and Moths of North America [BAMONA]
Anthomaster leonardus ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 51
Hesperia leonardus ; [NACL, #4023]; [BOW: pl. 22, f. 3]; [Opler]

Larva on Agrostis, Panicum, Eragrostis [NAB]


Hesperia pawnee Dodge, 1874
Colorado. See [About maps] Pawnee Skipper
Hesperia pawnee Dodge, 1874; Can. Ent. 6 (3) : 44
montana (Skinner, 1911)
ogallala (Leussler, 1921)
Erynnis pawnee ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 50
Hesperia pawnee ; [NACL, #4024]; [BOW: pl. 22, f. 5]
Hesperia leonardus pawnee ;

Hesperia pahaska Leussler, 1938
-. See [About maps] Pahaska Skipper
Hesperia pahaska Leussler, 1938; , TL: Nebraska
Hesperia pahaska, Butterflies and Moths of North America [BAMONA]
Hesperia pahaska ; [NACL, #4025]; [Opler]; [NL4A, #1633]

Larva on Poaceae, Bouteloua gracilis, Tridens pulchella [NAB]

Nebraska Hesperia pahaska pahaska ; [NL4A, #1633a]
H. p. williamsi Lindsey, 1940
Arizona Hesperia pahaska williamsi Lindsey, 1940; , TL: Arizona
Hesperia pahaska williamsi, Listing of Mexican Butterflies [ILMB]
Hesperia pahaska williamsi, Butterflies of America [BOA]
Hesperia pahaska williamsi, Butterflies of Southeastern Arizon [Bruce Walsh]
Hesperia pahaska williamsi ; [NACL, #4025a]; [NL4A, #1633b]
Misidentified/misapplied:
Hesperia columbia Evans, 1955; (not Scudder, 1872)
H. p. martini MacNeill, 1964
Hesperia pahaska martini ; [NACL, #4025b]

Hesperia columbia (Scudder, 1872)
Baja California - SW.Oregon. See [About maps] Columbian Skipper Columbia Skipper Chaparral
Pampihla columbia Scudder, 1872; 4th Ann. Rep. Peabody Acad. Sci. (1871): 56, TL: "California"
sylvanoides (Scudder, 1874, not Boisduval, 1852)
Pamphila california Wright, 1905; , TL: California
= ; [NL4A, #1628]
Thymelicus erynnioides Dyar, 1907; , TL: California
= ; [NL4A, #1628]
idaho (Seitz, 1924, not Edwards, 1883)
Hesperia columbia, Butterflies of America [BOA]
Hesperia columbia, Butterflies and Moths of North America [BAMONA]
californica ; Skinner, 1920, (missp.)
colnmbia ; Kirby, 1877, (missp.)
erynnoides ; Evans, 1955, (missp.)
Erynnis comma columbia ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 49
Hesperia columbia ; [NACL, #4026]; [BOC, 76]; [Opler]; [NL4A, #1628]
Misidentified/misapplied:
zabulon; Kirby, 1877
colorado; Edwards, 1877

Larva on Koeleria cristata [NAB]


Hesperia metea Scudder, 1863
E.USA. See [About maps] Cobweb Skipper
Hesperia metea Scudder, 1863; Proc. Essex Inst. 3: 177
Hesperia metea metea, Butterflies of America [BOA]
Hesperia metea, Butterflies and Moths of North America [BAMONA]
Erynnis metea ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 50
Hesperia metea ; [NACL, #4027]; [BOW: pl. 22, f. 4]; [Opler]

Larva on Andropogon scoparius [NAB]

H. m. licinus (Edwards, 1871)
Texas Pamphila licinus Edwards, 1871; Trans. amer. ent. Soc. 3 : 275, TL: nr. Waco, Texas
Hesperia horus Edwards, 1871; Trans. amer. ent. Soc. 3 : 277, TL: Dallas, Texas
= ; Freeman, 1948, Ent. News 59 (8): (203-206)
Hesperia metea f. belfragei Freeman, 1944; Field and Lab. 12 (1): (20-22), TL: Cedar Hill, Dallas Co., Texas
= ; Freeman, 1948, Ent. News 59 (8): (203-206)
Erynnis licinus ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 50
Lerema horus ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 53
Hesperia metea licinus ; [NACL, #4027a]

Hesperia viridis (Edwards, 1883)
Mexico, New Mexico. See [About maps] Green Skipper
Pamphila juba var. viridis Edwards, 1883; Can. Ent. 15 (8) : 147, TL: New Mexico
Hesperia viridis, Listing of Mexican Butterflies [ILMB]
Hesperia viridis, Butterflies of America [BOA]
Hesperia viridis, Butterflies and Moths of North America [BAMONA]
Hesperia viridis, Butterflies of Southeastern Arizon [Bruce Walsh]
Erynnis comma viridis ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 50
Hesperia viridis ; [NACL, #2028]; [Opler]; [NL4A, #1635]

Larva on Bouteloua gracilis? [NAB]


Hesperia attalus (Edwards, 1871)
S.USA. See [About maps] Dotted Skipper
Pamphila attalus Edwards, 1871; Trans. amer. ent. Soc. 3 : 276, TL: nr Waco, Texas
seminole (Scudder, 1872)
quaiapen (Scudder, 1889)
Hesperia attalus, Butterflies and Moths of North America [BAMONA]
Erynnis attalus ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 50
Hesperia attalus ; [NACL, #2029]; [BOW: pl. 22, f. 1]; [Opler]
H. a. slossonae (Skinner, 1890)
Hesperia attalus slossonae ; [NACL, #2029a]

Hesperia meskei Edwards, 1877
Texas - Florida. See [About maps] Meske's Skipper Dixie Skipper
Hesperia meskei Edwards, 1877; Can. Ent. 9 (3) : 58
Hesperia meskei, Butterflies and Moths of North America [BAMONA]
Anthomaster meskei ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 51
Hesperia meskei ; [NACL, #2030]; [Opler]
H. m. straton (Edwards, 1881)
Florida Pamphila straton Edwards, 1881; Papilio 1 (5) : 78, TL: Florida
stratton[sic, recte straton] ; Bell, 1938
Hesperia meskei straton ; [NACL, #2030a]

Hesperia dacotae (Skinner, 1911)
Dakota Skipper
Hesperia dacotae, Butterflies and Moths of North America [BAMONA]
Hesperia dacotae ; [NACL, #2031]; [Opler]

Larva on Bouteloua, Andropogon, Aristida, Poaceae [NAB]


Hesperia sassacus Harris, 1862
E.USA. See [About maps] Indian Skipper
Hesperia sassacus Harris, 1862; Treatise Ins. injur. Veget. (3rd edn): 315
Hesperia sassacus sassacus, Butterflies of America [BOA]
Hesperia sassacus, Butterflies and Moths of North America [BAMONA]
Erynnis sassacus ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 50
Hesperia sassacus ; [NACL, #2033]; [BOW: pl. 22, f. 9]; [Opler]

Larva on Panicum, Festuca, Digitaria [NAB]

H. s. manitoboides (Fletcher, 1889)
Pamphila manitoboides Fletcher, 1889; Rept. Ent. Soc. Ont. 19: 85
Pamphila manitoboides Fletcher, 1903; Trans. R. Soc. Canad. (2) 9 (4) : 215
Pamphila manitoboides Fletcher, 1904; Can. Ent. 36 (5) : 128
Hesperia sassacus manitoboides, Butterflies of America [BOA]
Erynnis comma manitoboides ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 50
Hesperia sassacus manitoboides ; [NACL, #2033a]

Hesperia miriamae MacNeill, 1959
Sierra Skipper
Hesperia miriamae ; [NACL, #2034]; [Opler]

Hesperia nevada (Scudder, 1874)
Colorado. See [About maps] Nevada Skipper Montane Skipper
nevada (Scudder, 1874); Mem. Boston Soc. nat. Hist. 2 (Pt 3, No. 4): 347, TL: "Colorado, Oregon"
colorado (Scudder, 1874, not Scudder, 1874); (part)
Hesperia nevada, E.H. Strickland Ent. Mus, U of [ALBERTA]
Hesperia nevada nevada, Butterflies of America [BOA]
Hesperia nevada, Butterflies and Moths of North America [BAMONA]
Erynnis comma nevada ; Dyar, 1903, Bull. U.S. nat. Mus. 52 : 50
Hesperia nevada ; [NACL, #2035]; [BOW: pl. 22, f. 8 (text)]; [Opler]

Larva on Stipa occidentalis, Festuca ovina [NAB]

Hesperia nevada nevada ; [BOC, 80]
H. n. sierra Austin, Emmel, Emmel and Mattoon, 1998
Hesperia nevada sierra Austin, Emmel, Emmel and Mattoon, 1998; , TL: Tahoe Meadows, Carson Range, Washoe Co., California
Hesperia nevada sierra ; [BOC, 89]

Hesperia nabokovi (Bell & Comstock, 1948)
Hispaniola Atalopedes nabokovi Bell & Comstock, 1948; Amer. Mus. Novit. No. 1379: 19, TL: Hispaniola
Atalopedes nabokovi ; Evans, 1955, Cat. amer. Hesp. Brit. Mus. 4: 339; Riley, 1975, Field Guide butts West Indies: 186
Hesperia nabokovi ; Burns, 1987, J. Lep. Soc. 41 (4): 174, f. 1-15; [NL4A, #1632]


Unknown or unplaced taxa

Hesperia eburus Plötz, 1885; Berl. ent. Z. 29 (2) : 226


31.1.2013 (24)

References:
[ALBERTA]
University of Alberta, E.H. Strickland Entomological Museum; Browse Entomology Collection
[About maps]
Warning! The maps are automatically generated from the textual information, and the process does not always produce acceptable result; See about maps for more info.
[BAMONA]
Opler, Pavulaan, Stanford & Pogue; Butterflies and Moths of North America; Mountain Prairie Information Node
[BCA]
electronic; BIOLOGIA CENTRALI-AMERICANA
[BMAT]
Tennent, 1996; The butterflies of Morocco, Algeria and Tunisia; ISBN 0 906802 05 9
[BNA]
Scott, 1986; The Butterflies of North America
[BOA]
Interactive Listing of American Butterflies; Listado Interactivo de la Mariposas Americanas; Butterflies of America
[BOC]
Robert Michael Pyle, 2002; The Butterflies of Cascadia
[BOW]
Lewis, 1974; Butterflies of the World; ISBN 0 245 52097 X
[BRU]
Tuzov, Bogdanov, Devyatkin, Kaabak, Korolev, Murzin, Samodurov, Tarasov, 1997; Guide to the Butterflies of Russia and adjacent territories; Hesperiidae, Papilionidae, Pieridae, Satyridae; Volume 1
[Bruce Walsh]
Bruce Walsh; e-mail: Bruce Walsh; The Moths of Southeastern Arizona; Butterflies of Southeastern Arizona
[ILMB]
Interactive Listing of Mexican Butterflies
[Kimmo Silvonen]
Espoo, Finland; Kimmo's Lep Site
[MBE]
Moths and Butteflies of Europe; Paolo Mazzei, Diego Reggianti and Ilaria Pimpinelli; http://www.leps.it/
[MRS]
Chou Io (Ed.); Monographia Rhopalocerum Sinensium, 1-2
[NAB]
Pyle, 1981; Field Guide to North American Butterflies, National Audubon Society; ISBN 0-394-51914-0
[NACL]
Hodges, 1983; Check List of the Lepidoptera of America North of Mexico
[NL4A]
Gerardo Lamas, 2004; Atlas of Neotropical Lepidoptera; Checklist: Part 4A; Hesperioidea - Papilionoidea
[NORLEP]
Norges Sommerfugler
[Oleg Kosterin]
Institue of Cytology & Genetics; Siberian Division of Russian Academy of Sciences; Novosibirsk; e-mail: Oleg Kosterin; http://pisum.bionet.nsc.ru/kosterin/
[Opler]
Opler & Warren, 2003 (April); Butterflies of North America. 2. Scientific Names List for Butterfly Species of North America, north of Mexico.
[SPRK]
Seppänen, E. J, 1970; Suomen suurperhostoukkien ravintokasvit, Animalia Fennica 14
[SVF]
Svenska fjärilar; Naturhistoriska riksmuseet; (Sweden)
[SZMN]
Lepidoptera; Siberian Zoological Museum
[Simon Coombes]
1 Park Street, Plymouth, PL3 4BL, UK; 01752 607854; e-mail: Simon Coombes; http://www.butterfly-guide.co.uk
[²]
This may require parentheses or not. I don't have the necessary information for this taxon.


Some related literature:
Bell & Comstock, 1948
A new genus and some new species and subspecies of American Hesperiidae Amer. Mus. Novit. No. 1379: 23pp
Burns, 1987
The big shift: nabokovi from Atalopedes to _Hesperia J. Lep. Soc. 41 (4): 173-186
Dodge, 1874
Hesperia pawnee N. sp. Can. Ent. 6 (3) : 44-45
Dyar, 1903
A List of North American Lepidoptera and Key to the Literature of this Order of Insects Bull. U.S. nat. Mus. 52 : 1-723
Edwards, 1863
Descriptions of certain species of Diurnal Lepidoptera, found within the limits of the United States and British America. (No. 1-3) Proc. ent. Soc. Philad. 2 (1): 14-22, pl. 1, 11 (1863), (2): 78-82 (1863), (4): 501-507 (1864)
Edwards, 1866
Descriptions of certain species of Diurnal Lepidoptera found within the limits of the United States and British America. No. 5 Proc. ent. Soc. Philad. 6 (4) : 200-208
Edwards, 1871
Descriptions of new species of North American butterflies Trans. amer. ent. Soc. 3 : 266-277
Edwards, 1877
Description of a new species of Hesperia from Texas Can. Ent. 9 (3) : 58-59
Edwards, 1881
Description of a new hesperian from Florida Papilio 1 (5) : 78-79
Edwards, 1881
Descriptions of new species of diurnal Lepidoptera found within the United States Trans. Amer. Ent. Soc. 9 : 1-8
Edwards, 1883
Notes on Dr. Speyer's paper Can. Ent. 15 (8) : 147-151
Edwards, 1884
Description. of new species of butterflies mostly from Arizona Papilio 4 (3) : 53-58
Emmel, Emmel & Mattoon, 1998
Three new subspecies of Hesperia lindseyi (Lepidoptera: Hesperiidae) in California and northwestern Nevada in Emmel, Systematics W. North Am. Butts.: 475-480
Evans, 1955
A Catalogue of the American Hesperiidae, indicating the classification and nomenclature adopted in the British Museum. Part IV. Hesperiinae and Megathyminae Cat. amer. Hesp. Brit. Mus. 4: 499pp, pls. 54-88
Fabricius, 1807
Systema Glossatorum. Die neueste Gattungs-Eintheilung der Schmetterlinge aus den Linnéischen Gattungen Papilio und Sphinx Magazin f. Insektenk. (Illiger) 6: 277-295
Fletcher, 1904
Descriptions of some new species and varieties of Canadian butterflies Can. Ent. 36 (5) : 121-129
Freeman, 1944
A new form of Hesperia metea Scudder from Texas (Lepidoptera Hesperiidae) Field and Lab. 12 (1): 20-22
Freeman, 1944
A new form of Hesperia leonardus (Harris) from the middle western United States (Lepidoptera, Rhopalocera, Hesperiidae) Bull. Brooklyn Ent. Soc. 38 (5): 153-154
Freeman, 1948
Notes on some North American skippers, with the description of a new species from Kansas (Lepidoptera: Hesperiidae) Ent. News 59 (8) : 203-206
Freeman, 1956
A new species of Hesperia from Califormia (Hesperiidae) Lepid. News. 9 (6): 196-198
Godman & Salvin, [1899]; Godman & Salvin, [1900]
Biologia Centrali-Americana. Rhopalocera. (1887-1901) Biol. centr.-amer., Lep. Rhop. 2: 1-782, 3: pl. 1-112
Harris, 1862
A treatise on some of the insects injurious to vegetation Treatise Ins. injur. Veget. (3rd edn): 1-640, 8pls
Linnaeus, 1758
Systema Naturae per Regna Tria Naturae, Secundum Clases, Ordines, Genera, Species, cum Characteribus, Differentiis, Symonymis, Locis. Tomis I. 10th Edition Syst. Nat. (Edn 10) 1 : 1-338, : 339-824
Lyman, 1892
Pamphila manitoba, Scud., and its varieties Can. Ent. 24 (3) : 57-59
McGuire, 1998
Description of three new subspecies of Hesperia (Lepidoptera: Hesperiidae) from the western United States in Emmel, Systematics of W. North Am. Butts: 461-474
Moore, [1875]
Descriptions of new Asiatic Lepidoptera Proc. zool. Soc. Lond. 1874 (4) : 565-579, pl. 66-67
Oberthür, 1912
Description d'une nouvelle espèce algérienne d'_Hesperiidae Bull. Soc. ent. Fr. 1912 : 349-351
Plötz, 1883
Die Hesperiinen -- Gattung Hesperia Aut. und ihre Arten. (3) & (4) Stett. ent. Ztg 44 (1-3): 26-64, (4-6): 195-233
Plötz, 1885
Neue Hesperiden des indischen Archipels und Ost-Africa's aus der Collection des Herrn H. Ribbe in Blasewitz-Dresden, gesammelt von den Herren: C. Ribbe auf Celebes, Java un den Aru-Inseln, Künstler auf Malacca (Perak); Kühn auf West-Guinea (Jekar); Menger auf Ceylon Berl. ent. Z. 29 (2) : 225-232
Riley, 1975
Field guide to the butterflies of the West Indies Field Guide butts West Indies: 244pp
Scudder, 1872
A systematic Revision of some of the American Butterflies; with brief notes on those known to occur in Essex County, Mass. 4th Ann. Rep. Peabody Acad. Sci. (1871): 24-83
Scudder, 1874
The species of the lepidopterous genus Pamphila Mem. Boston Soc. nat. Hist. 2 (Pt 3, No. 4): 341-353
Severns & Severns, 2005
Hesperia lindseyi mccorklei (Hesperiidae): a new subspecies of skipper from southwestern Oregon, USA J. Lep. Soc. 59 (3): 166-169
Shapiro & Forister, 2005
Phenological "races" of the Hesperia colorado complex (Hesperiidae) on the west slope of the California Sierra Nevada J. Lep. Soc. 59 (3): 161-165
Sodoffsky, 1837
Entomologische untersuchungen ueber die Gattungsnamen der Schmetterlinge Bull. Soc. imp. Nat. Moscou 1837 (6) : 76-97
Tutt, 1905
A natural history of the British Lepidoptera. A text-book for students and collectors Nat. Hist. Br. Lepid. 8 : 1-479, pl. 1-20


If you have corrections, comments or information to add into these pages, just send mail to Markku Savela
Keep in mind that the taxonomic information is copied from various sources, and may include many inaccuracies. Expert help is welcome.